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In this paper, we investigate the cross-layer resource optimization for wireless relay networks with dynamic node-selfishness information. The degree of intrinsic selfishness (DeIS) and the degree of extrinsic selfishness (DeES) of the relay node (RN) are defined as the effects of its own residual energy resource and the incentive in an incentive mechanism, respectively. A model of two-timescale node-selfishness dynamics is constructed to describe the time-varying characteristics of the RN's DeIS and DeES. Based on the two-timescale node-selfishness dynamics of the RNs, a two-timescale resource-optimization dynamical algorithm (TRODA) is employed to control the transmit powers of both RNs and end users (EU), as well as the flow rates of the data packets injected by all EUs. Meanwhile, the stability and the tracking error of the TRODA under the two-timescale node-selfishness dynamics are analyzed by Lyapunov theory. Our simulation results demonstrate that the two-timescale optimization algorithm is stable and has trivial tracking error. [ABSTRACT FROM AUTHOR]