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Steppe and grassland ecosystems constitute important biomes that are influenced by multiple factors such as climate, human activity, and fire. Yet how these factors have influenced the plant composition of these biomes through time continues to be understudied. This paper investigates how these mechanisms have transformed the steppe landscape recorded at the mire site of Shenkani, Armenia. This highland site, located in the South Caucasus, has a long human history of permanent settlements near the core site starting at 5500 cal. BP. A variety of biological and geochemical proxies, including pollen, non-pollen palynomorphs, macro-charcoal, 14C age model, X-ray fluorescence, loss-of-ignition, magnetic susceptibility, and a quantitative climate reconstruction inferred from pollen, are utilized in this paper. We find that this area has remained a steppe with small stands of trees throughout the duration of the Holocene. Changes in steppe plant composition primarily occur between semi-desert steppes (Chenopodiaceae), grassland steppes (or Poaceae dominated meadow-steppes), and mixed steppes with forest patches. In this record, two phases of local grassland expansion occurred between 9500 and 7000 cal. BP and 2500–1000 cal. BP, which covaried with local fire events. These grassland steppes were sustained until tree encroachment led to a more mixed steppe landscape around 7000 cal. BP and again at 1000 cal. BP. Climate, primarily precipitation, is the main driver of this persistent steppe landscape and the plant compositional shifts within it. However, fire and human activities contributed to steppe maintenance. • A 10,240 year-old sequence from Shenkani, Armenia documents a persistent Holocene steppe. • Changes in steppe plant composition occur between semi-desert, grassland, and mixed steppes with forest patches. • Poaceae dominated grasslands covary with fires that limit tree and shrub encroachment. • The mid- to late Holocene transition is characterized by drops in Betula and declines in distant arboreal pollen. [ABSTRACT FROM AUTHOR]

• Identify ancient flood events, which may be the reason for decline of the culture. • Pollen and heavy metals can be used to reflect the strength of human activities. • Pollen and heavy metals revealed the process of culture from prosperity to decline. • This paper defined the chronology from prosperity to decline of Shijiahe culture. • Climate change has been revealed by pollen and heavy metals since 4800 cal. yr BP. The Shijiahe Culture (ca. 4800–4100 yr BP), which originated in the middle Yangtze Basin, was a milestone of prehistoric Chinese civilization. To reveal the interactions between climatic changes and human activities during this time interval, the geochronology and multiple proxies (pollen and heavy metals) of samples from a 220-cm-deep outer wall at the Sanfangwan Site (a typical site of the Shijiahe Culture) were analyzed. Radiocarbon dating indicates that the age range of the Sanfangwan Site is ca. 4800 to 4100 cal. yr BP. From ca. 4800 to 4600 cal. yr BP (the early Shijiahe Culture), pollen assemblages were dominated by Gramineae and an anthropogenic plant community (e.g. Ranunculaceae, Chenopodiaceae, Labiatae, and Cruciferae), indicating an expansion in rice agriculture during this interval. High metal concentrations in the same interval indicate a large flux of metals from smelting and farming activities. Between ca. 4600 and 4100 cal. yr BP (the middle and late Shijiahe Culture), decreases in Gramineae and anthropogenic pollen mirrored the recession of farming. Concurrent decreases in heavy metals may be attributable to a decline in the smelting industry. Three episodes of high pollen concentrations and visible increases in hygrophyte pollen, Pinus pollen, and fern spores at 4600–4550, 4450–4350, and 4200–4100 cal. yr BP probably indicate the occurrence of several flooding events during the middle and late Shijiahe Culture, resulting in the migration of local inhabitants. This study highlights the driving effect of flood events on the collapse of the Shijiahe Culture. [ABSTRACT FROM AUTHOR]

The focus of this paper is on the reconstruction of the mid-Holocene climatic changes based on the pollen records contained in a 6.2 m thick section in the southern part of the Tengger Desert, northwestern China. To understand the modern analogues of the mid-Holocene changes, modern pollen spectra across different vegetation zones from the Qilian Mountains to the lowland Tengger Desert were studied first. Four pollen combinations were identified, corresponding to four different modern ecological environments: (1) a Picea-Artemisia combination in a spruce forest and its vicinity; (2) an Artemisia-Chenopodiaceae-Gramineae-Salix combination in a mixture belt of montane grassland and poplar woodland; (3) a Chenopodiaceae-Artemisia-Nitraria combination in a desert steppe belt; and (4) an Elaeagnus-Artemisia-Chenopodiaceae-Nitraria combination in a desert riparian forest. Our study also shows that a relatively low percentage of Picea pollen (30–10%) seems to be associated with water transportation (alluvial deposits) and a very low percentage of Picea pollen (10–5%) appears to be related to long-distance air transportation. With references to the modern relationships between pollen assemblages and associated bioclimates, the pollen sequence covering the period between 8500 and 3000 cal. BP was used to infer the climatic change history. The results show that a warmer and drier climate dominated between 8500 and 7950 cal. BP. Subsequent decreases in Artemisia and warm desert forbs/shrubs taxa and contemporaneous fluctuating increases in montane coniferous and aquatics elements from 7950 to 7400 cal. BP reflect a fluctuating increase in humidity and/or a fluctuating decrease in temperature. The following period from 7400 to 5650 cal. BP was generally warm with the Holocene Climate Optimum occurring between 7400 and 6500 cal. BP. A remarkably cool spell followed between 6500 and 6300 cal. BP that was then succeeded by a warmer and more humid period between 6300 and 5650 cal. BP. The period between 5650 and 4450 cal. BP was characterized by drastic oscillations in climatic conditions with three humid and probably cold spells intervened by three relatively dry and warm spells. The period between 4450 and 3500 cal. BP was cold and humid, followed by a wet spell as indicated by a very high percentage of Typha pollen. [ABSTRACT FROM AUTHOR]