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Understanding the mechanisms of crustal deformation along convergent margins is critical to identifying seismogenic structures and assessing earthquake hazards for nearby urban centers. In the southern central Andes (28-33[Formula: see text]S), differences in the style of middle to upper-crustal deformation and associated seismicity are highlighted by the January 19th, 2021 (Mw 6.5) San Juan earthquake. We integrate waveforms recorded at regional and teleseismic distances with co-seismic displacements calculated from local Global Navigation Satellite System time series, to re-estimate the source parameters of the 2021 San Juan earthquake, confirming a mid-crustal nucleation depth (21 ± 2 km) and right-lateral transpressional mechanism. Considered alongside decades of seismic observations and geological data, this event provides evidence for retroarc deformation partitioning among inherited basement faults and upper-crustal structures in response to oblique convergence of the Nazca and South American plates. As they may transfer shortening to active upper-crustal faults associated with historically devastating shallower earthquakes, a better understanding of seismogenic basement faults such as the mid-crustal structure activated during the 2021 San Juan earthquake earthquake could help future re-assessment of the seismic risk in western Argentina., (© 2022. The Author(s).)