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The deep structure of the continental collision between India and Asia and whether India's lower crust is underplated beneath Tibet or subducted into the mantle remain controversial. It is also unknown whether the active normal faults that facilitate orogen-parallel extension of Tibetan upper crust continue into the lower crust and upper mantle. Our receiver-function images collected parallel to the India-Tibet collision zone show the 20-km-thick Indian lower crust that underplates Tibet at 88.5-92°E beneath the Yarlung-Zangbo suture is essentially absent in the vicinity of the Cona-Sangri and Pumqu-Xainza grabens, demonstrating a clear link between upper-crustal and lower-crustal thinning. Satellite gravity data that covary with the thickness of Indian lower crust are consistent with the lower crust being only ∼30% eclogitized so gravitationally stable. Deep earthquakes coincide with Moho offsets and with lateral thinning of the Indian lower crust near the bottom of the partially eclogitized Indian lower crust, suggesting the Indian lower crust is locally foundering or stoping into the mantle. Loss of Indian lower crust by these means implies gravitational instability that can result from localized rapid eclogitization enabled by dehydration reactions in weakly hydrous mafic granulites or by volatile-rich asthenospheric upwelling directly beneath the two grabens. We propose that two competing processes, plateau formation by underplating and continental loss by foundering or stoping, are simultaneously operating beneath the collision zone., Competing Interests: The authors declare no competing interest.

In classic orogenic models, the mountain range is underlain by a deep crustal root. Here we present the crustal and upper mantle structures along two receiver function profiles across Qilian, an orogen experiencing recent growth at the northern margin of the Tibetan plateau. Opposite to an expected crustal root beneath the orogen, the Moho beneath Qilian is arch-like, shallower beneath the center and deepens by up to 10 km beneath its southern and northern boundaries. Additional velocity interfaces sub-parallel to the Moho are observed in the lower crust of the basins south of Qilian, which we interpret as the top of a mechanically strong lower crust thrusting several tens of kilometers underneath Qilian. In the north, the small lateral offset between the surface and mantle traces of the thrust system reveals a steep boundary, indicating that the North China cratonic crust acts as a strong resistance to the northward growth of the plateau, forcing the development of the left-lateral strike-slip Haiyuan fault south of the northern Qilian suture. The young Qilian orogen thus has been rising and growing progressively from the boundaries to the center, squeezed up by more rigid tectonic blocks in the north and south.