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Wetting and drying cycles (WDCs) have a significant impact on the shear behavior of discontinuities with different joint wall materials (DDJMs). This influence is crucial for the reasonable evaluation of the long-term stability of soft and hard interbedded rock slopes under water level fluctuations. As the surface topographies of natural discontinuities collected from the field vary, conducting comparative experiments on natural discontinuity specimens with identical surface topographies is challenging. To solve this problem, a 3D surface topography reconstruction technique was employed to obtain DDJM specimens with three types of surface topographies collected from a typical sliding-prone stratum in the Three Gorges Reservoir area, China. A series of experiments, including computed tomography scanning, 3D laser scanning, and direct shear tests, were conducted to investigate the influence of WDCs on the micro- and macroproperties of joint walls, surface topographies, and shear behavior of DDJMs. The experimental results showed that repeated WDC treatments caused the degradation of the microstructures and macroscopic physical properties of the studied joint walls, and the more severely weakened joint wall played a predominant role in reducing the shear strength of DDJMs. The influence of WDCs on the surface topographies of DDJMs was negligible in this study; changes in the shear behavior of DDJMs were closely associated with the weakening of joint walls induced by WDCs; and the impact degree of joint wall weakening on the deterioration of the shear behavior of DDJMs was interactively influenced by and positively correlated with both the joint roughness coefficient and normal stress. These results will contribute to a better understanding of the evolution of the stability of soft and hard interbedded rock slopes induced by water level fluctuations in the Three Gorges Reservoir area and other reservoir regions. [ABSTRACT FROM AUTHOR]