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• The Madden-Julian Oscillation (MJO) affects the DC global electric circuit (GEC). • MJO-related patterns are seen in both simulations and electric field measurements. • On average, GEC parameters manifest sine-like variations with the phase of the MJO. • Regional contributions to the GEC reflect deep convection changes during MJO events. • Key GEC-reflected convection changes are over the Maritime Continent and Indian Ocean. The Madden–Julian Oscillation (MJO) is the most dominant component of the climate variability in the tropics on the timescale of tens of days. Occurring irregularly, the MJO consists of an eastward propagation of certain large-scale coupled structures in atmospheric circulation and deep convection. Here we investigate the effect of the MJO on the direct current global electric circuit (GEC), using both numerical simulations and the results of electric field measurements. We have reproduced the atmospheric dynamics for 1980–2020 with the help of the Weather Research and Forecasting model and meteorological reanalysis data; this allowed us to simulate 41 years of the GEC variation by parameterising contributions to the ionospheric potential (IP) in terms of precipitation and convection. Besides, we have analysed the results of surface potential gradient (PG) measurements at the Vostok station in Antarctica during 2006–2020. When averaged over many days, both the simulated IP and measured fair-weather PG show sinewave-like variations with the MJO phase, which manifest a statistically significant correlation with the MJO cycle. A more elaborate investigation involving an application of empirical orthogonal functions shows that the dynamics of contributions to the IP follows the patterns of variability of deep convection typical for the MJO. The variation of the IP on the MJO scale can largely be represented as a superposition of two basic oscillating patterns of convection, one of which is located over the Maritime Continent and its surrounding seas and the other is located over the Indian Ocean and part of South-East Asia. These oscillating patterns together describe the eastward progression of perturbations and can be naturally associated with the two components of the Real-time Multivariate MJO index, widely used to describe the MJO. Together with earlier results about links between the GEC and the El Niño—Southern Oscillation, the results concerning the MJO obtained in this study indicate that the GEC is an important part of the Earth system, which reflects climate variability on different timescales. [ABSTRACT FROM AUTHOR]