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Responding to China's major environmental challenges, researchers are increasingly exploring dynamics between international non-governmental organisations, the nation-state, and local communities. Much less understood are domestic environmental non-governmental organisations (eNGOs) and their interactions with actors across scales. This paper responds by exploring domestic eNGOs and interactions with institutional actors, including international donors, and with levels of Chinese government. The practices of eNGOs in China are influenced by interactions at various scales. Rather than being directly influenced by international partners and simply constrained by state regulations, Chinese eNGOs are embedded in more complex power relations. These interrelationships are exemplified in the operations of Green Watershed, a domestic eNGO working in Yunnan Province. Green Watershed has extensive interactions with international donors and various levels of government as a result of its national campaign and long-term local projects. Relationships between eNGOs’ international donors and the Chinese government are significant in influencing eNGO practice. NGO–government relations in China are inconsistent and constantly changing owing to the diverse interests and priorities of the various levels of government. The impacts of the local practice of Chinese eNGOs are deeply informed by their relationships both at and beyond the local level. Complex interrelationships make it difficult to achieve bottom-up, participatory resource management in China. [ABSTRACT FROM PUBLISHER]