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The United States Department of Agriculture Forest Service International Institute of Tropical Forestry (the Institute) celebrates its 75th Anniversary with the publication of this Special Issue of Forests. This Issue is based on presentations delivered in a symposium held in San Juan, Puerto Rico in 2014. It augments a quarter century of scientific knowledge and capitalizes on a unique set of synergies chartered by a strategy based on shared stewardship, innovative transdisciplinary collaborations, and breakthroughs in science and technology. The manuscripts contained here present advancements in our approach to the development of policies for effective governance and stewardship, long-term focus for the understanding of ecosystem processes and functions, novelties given attention to cross-boundary collaborative approaches to science, and proposed alternative institutional visions in the Anthropocene. As the Institute continues to collaboratively explore new frontiers in science, we recognize advances in forestry, atmospheric sciences, modeling, hydrology, plant physiology, and microbial ecology as core to the understanding of tropical forests in the Anthropocene. [ABSTRACT FROM AUTHOR]