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This paper presents the results of the explorations carried out in the western sector of the Iberian Central System (western Gredos, Béjar, Gata and Estrela mountain ranges). The macroremains found allow detailing the model of the landscape transformation, the composition and extension of their vegetal formations throughout the Holocene. Ninety-nine woody macroremains, 3 bark remains and 8 pinecones were located from eight sites between 875 and 1860 m above sea level, among which 15 radiocarbon dates were made. The taxonomic identification of two trunks found in Estrela allows us to document the local presence of Pinus cf. sylvestris ca. 6600 cal BP in this site. The same taxon also appears at the western end of Gredos between ca. 1700–2800 cal BP at approximately 1300 m. These remains expand the Holocene distribution of P. cf. sylvestris in the Central System known to date, both temporally and geographically. In parallel, the macroremains analysed document the composition of high mountain scrub (Cytisus type, Erica cf. arborea) in Béjar (ca. 1860 m), as well as mixed scrub-woodland formations in Gata (975–1180 m), which included deciduous Quercus , Betulaceae and Ericaceae. The presence of P. pinaster at ca. 6200 cal BP and 1180 m in this mountain range constitutes the earliest palaeobotanical evidence of this species recorded in Gata. The main results support the continuous persistence of plant formations in which P. sylvestris , P. nigra , and P. pinaster would have formed part throughout the Iberian Central System during the Holocene in a wide altitudinal range (1100–1840 m). • Above one hundred Holocene macrofossils from eight different sites were analysed. • Pinus sylvestris recently disappeared from the western Central System. • P. gr. sylvestris remained in Serra da Estrela until ˜6650 years ago. • First evidence of Pinus pinaster has been found in Gata ˜6200 years ago. • Betula , Quercus , Alnus and Ericaceae Holocene macroremains were found in Gata. [ABSTRACT FROM AUTHOR]

The fossil plant Montsechia vidalii is well known from the Barremian of Spain and was identified as an early angiosperm. It is interpreted as an aquatic plant that thrived in freshwater wetlands, with very limited or no connection with the marine realm, and under a seasonal semi-arid tropical climate. This research paper reports the youngest known occurrence of the genus Montsechia , in upper Albian amber-bearing strata from the Maestrazgo Basin (NE Spain), demonstrating the broad adaptation and evolutionary stability of the genus during the Early Cretaceous. The studied upper Albian material is identified as M. aff. vidalii since it differs from the Barremian Montsechia vidalii in having a medium-thick cuticle and paracytic stomata on the abaxial leaf cuticle. In addition, associated seeds, fruit and pollen grains have been found. The fruit is indehiscent and ascidiate in shape with a single seed inside. The seeds are orthotropous and have been identified as Spermatites sp. The pollen grains were found adhering to a sterile twig. They are monocolpate and assigned to Transitoripollis sp., reinforcing the early angiosperm hypothesis. Unlike its Barremian relative, the late Albian M. aff. vidalii developed under predominantly arid conditions in a fore-erg setting characterised by an interaction of aeolian dune and tide-dominated coastal deposits that reflected general eustatic rise conditions of the mid-Cretaceous. In this regard, the studied remains were found as a minor component of a supratidal coastal plant association largely dominated by conifers of the genera Frenelopsis and Dammarites. • Montsechia cf. vidalii from upper Albian strata of the Maestrazgo Basin, Spain. • Mid-Cretaceous supratidal plant communities of the Western Tethys coast. • Early angiosperm expansion into coastal environments during the mid-Cretaceous. [ABSTRACT FROM AUTHOR]

The present paper reports a complete palynological study of the Upper Turolian sediments outcropping at the Venta del Moro site (eastern Iberian Peninsula). The pollen assemblages observed mainly reflect the local and regional vegetation, which must have formed part of a wetland ecosystem within a general setting of open steppe, as indicated by the great abundance and diversity of herbaceous pollen. This reflects the persistence of open vegetation in the Iberian Peninsula after the Middle Miocene when it became extensive. We here report the most recent record of Disanthus for the Iberian Peninsula. This shrub played an important role in an unusual, edaphically-mediated vegetation linked to lacustrine–palustrine or riparian shore conditions. A shift in the water level led to a flooding episode, extending wetland and causing a drastic reduction in Disanthus and Amaranthaceae. The results of quantitative (multivariate and coexistence) and semi-quantitative (integrated plant record) analyses of the pollen dataset support these interpretations of the palaeovegetation and provide insight into the regional palaeoclimate. Stable climatic conditions would appear to have reigned before the onset of the Messinian salinity crisis. The palynoflora of Venta del Moro is compared with counterparts in other areas of the Mediterranean Basin. [ABSTRACT FROM AUTHOR]

Abstract: This paper focuses on the diversity and palaeontological significance of the Anisian floral record in the Triassic of the northeastern Iberian Peninsula. A summary of published information is followed by new records from macrofloral localities at Rodanas (Aragonese Branch of the Iberian Range) and at Maya del Baztán (Basque–Cantabrian Domain, Pyrenees). Both of these new localities also yielded three Anisian microfloral assemblages. These records provide new evidence of Anisian (basal Middle Triassic) floras in Spain. They are found to be somewhat similar to the classical flora of the Grès à Voltzia Formation (Upper Buntsandstein) of the northern Vosges in north-eastern France, of whose age has been shown to be early Anisian. [ABSTRACT FROM AUTHOR]

Abstract: Plant fossils are rarely preserved within the Upper Permian of western Europe as the majority of continental deposits consists of siliciclastic redbeds. Because organic matter such as plant debris and palynomorphs is destroyed by oxidizing conditions, these red sedimentary facies were often considered as “devoid of life”. This paper presents new palaeobotanical data from the Upper Permian strata of Minorca (Balearic Islands, Spain). In addition, a rich and diverse “Thuringian” palynoflora was recovered from the Permian sediments, and several megafloral taxa are recorded. Additionally, a detailed sedimentological analysis of the studied section is provided. The Permian succession shows a general evolution from a ponded floodplain environment to sand-sheet rivers flowing across a semi-arid alluvial plain. Additionally, new palaeobotanical elements, added to the study of palaeosols, allow reconstruction of Late Permian plant growth within the terrestrial environment, as well as a palaeoenvironment reconstruction of the preserved sedimentary interval. At the transition between ponds and alluvial plain deposits, root systems surprisingly similar to those known in extant mangroves are described for the first time within Permian deposits. As there are no other palaeobotanical remains preserved in association, the type of plant that inhabited this environment remains conjectural. [Copyright &y& Elsevier]

Abstract: In this paper we present a review of the available pollen analyses in the north-western Iberian Range with the aim of studying Fagus glacial refugia and Holocene expansion. The work collects the available pollen analyses in the area (24 sites and 58 14C dates). Based on the available palynological data, we try to reconstruct the changes that occurred in the distribution of Fagus sylvatica in the north-western Iberian Range during the late Pleistocene and the Holocene, its spread patterns and the possible causes for such changes. Results from pollen analyses allow establishing at the Middle Holocene the beginnings of Fagus expansion mostly in relation to anthropic activities. Also, the pollen data demonstrate the existence of Fagus glacial refuge during the Late Pleistocene and the Early Holocene. Therefore, we hypothesize that present-day beech forests, in the area under study, are the direct result of the relic populations that survived the last glaciations and that have been favoured by both environmental conditions and human activities. [Copyright &y& Elsevier]

Abstract: This paper presents a mid- to late-Holocene vegetation sequence of a Juniperus thurifera-dominated area in the eastern, continental plateaux of the Segura Mountains of southeastern Spain. A mid-Holocene maximum of mesic tress is recorded in the pollen diagram between c. 6640 and 4790 cal years BP with the vegetation dominated by deciduous oaks. A dramatic decline in deciduous oaks occurred from approximately 4700 cal years BP to be replaced initially by evergreen oak, and then junipers and other xerophytes from c. 4500 cal years BP. This trend of xericness in the vegetation is coherent with regional and extra-regional palaeoclimatic records for increased mid-Holocene aridity. Significant anthropogenic modification of the vegetation occurred in this region from c. 1350 cal years BP represented by a large reduction in all tree taxa (except Juniperus) and increases of thorny scrub and nitrophilous assemblages. Increased fire incidence, pastoralism, and arboriculture were associated with this anthropogenic activity. We conclude that present-day J. thurifera-dominated communities in this region have become established through a combination of two predominant processes; increased aridification from c.4500 cal years BP and anthropogenic activity from c. 1200 cal years BP. [Copyright &y& Elsevier]

This paper presents a palynological study of the archeological layers from the Neanderthal site Abrigo 3 del Complejo del Humo, in southern Spain (Málaga), with the aim of reconstructing the environmental conditions in the vicinity of this hominin site. The Upper Pleistocene vegetation and its variability are described, revealing a high diversity of thermophilous plant taxa throughout the cold dry phases, together with a long-term persistence of woody taxa, including Mediterranean, mesophytes, xerothermics and conifers. With the pollen records of Maytenus senegalensis as an outstanding finding, this study demonstrates the co-existence of temperate, Mediterranean and Ibero-Maghrebian angiosperms on the southern coastal plains of the Iberian Pleistocene where Neanderthals survived for a long time. It is therefore clear that Neanderthals and early Upper Paleolithic modern humans lived in a litoral refugium, which was a propitious environment for maintaining a high biodiversity, including potentially edible plant species. Besides, this coastal refugium offers broad possibilities for hunting, and interpopulational relationships through coastal platforms. • A palynological study of the archeological layers from the Neanderthal site of Abrigo 3 del Complejo del Humo. • A high diversity of thermophilous plant taxa throughout the cold dry phases. • The co-existence of temperate, Mediterranean and Ibero-Maghrebian angiosperms in the southern of the Iberian Pleistocene. • Neanderthals and early Upper Paleolithic people lived in a litoral refugium. [ABSTRACT FROM AUTHOR]

This paper reviews the past distribution of Aleppo pine woodlands in the Ebro Basin, Northeastern Iberia, from the Mesolithic to Modern times based on wood charcoal data. The aim is to detail the chronological timing and the drivers explaining the long-term presence of Aleppo pine woodlands and associated thermophilous flora. The available charcoal data support the early spread of Pinus halepensis during the Mesolithic (ca. 9000 cal BP) accompanied by Mediterranean trees and shrubs like Quercus sp. evergreen, Juniperus sp., Arbutus unedo, Pistacia lentiscus, Rhamnus/Phillyrea , Cistaceae, and Rosmarinus officinalis , as a local response to global climate change in the Early Holocene. During the arrival and the propagation of the Neolithic culture (ca. 7500–5500 cal BP), anthracological records, as well as regional palynological sequences, demonstrate the progressive replacement of an conifer-dominated open parkland by both Quercus sp. deciduous and evergreen woodlands in response to the Middle Holocene rise in temperature and humidity. This evidence, however, converges with the general idea that the presence and the spread of Pinus halepensis and associated scrubland have usually been attributed to the onset of landscape anthropization. The frequency of xero-thermophilous open scrubland and the use of Aleppo pine for fuel and woodcrafting progressively increased during the Bronze and Iron Ages, and especially in Ibero-Roman and Medieval/Islamic times, when the vegetation landscape in the Middle Ebro Basin was largely deforested as a consequence of increasing demographic pressure, grazing and the establishment of proto-urban centers. • We review the past distribution of Pinus halepensis in the Ebro Basin (NE Iberia) based on anthracological records. • Wood charcoal data support the early spread of Pinus halepensis during the Mesolithic. • The xero-thermophilous open scrubland and the use of Pinus halepensis as woodcrafting is attested during Metal Ages and historic times. [ABSTRACT FROM AUTHOR]