A city in Umbria (central Italy), situated upon a hill 1620 feet above sea level overlooking the fertile valley of the upper Tiber. The region had at one time been inhabited, according to Servius, by the Sarsinates, one of the tribes of the Umbrians (speaking an Indo-European Italic dialect), from whom Pliny the Elder records that the Etruscans seized three hundred towns. Perusia was among them, founded, apparently, by Clusium (Chiusi), which had only become a city in the sixth or fifth century BC. The place was largely responsible for the spread of the cultural influence of the Etruscans across the Tiber into Umbria, importing for example, their alphabet in the fourth century BC. The inhabitants of Perusia retained pretentious foundation legends that conceal the comparatively late date of its foundation, but testify to its importance in the subsequent epoch, when the stories were devised. In particular, these traditions tell of legendary northern expansion by the Perusians—asserting that their city was founded by Aulestes, the father or brother of Ocnus and mythical founder of Felsina (Bononia) and Mantua: the purpose of such tales was to eclipse the more authentic achievements of Clusium (Chiusi), at a time when the Perusians had thrown off its suzerainty. The artistic influence of Clusium is still to be seen in their sarcophagi of c 475–450 when they also produced important bronze work, including bizarre elongated statuettes that have appealed to modern sculptors.