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The ad 79 eruption of the Vesuvius severely affected the floodplain surrounding the ancient city of Pompeii, i.e. the Sarno River floodplain. The landscape was covered with volcaniclastic materials that destroyed the ecosystem but, at the same time, preserved the traces of former environmental conditions. This study provides—for the first time—a pollen sequence reconstructing the environmental evolution and the plant landscape of the Sarno floodplain between 900 and 750 cal bc and ad 79, i.e. before and during the foundation of the city, and during its life phases. Previous geomorphological studies revealed that the portion of the Sarno floodplain under the “Pompeii hill” was a freshwater backswamp with patchy inundated and dry areas. Palynology depicts a thin forest cover since the Early Iron Age, suggesting an open environment with a mosaic of vegetation types. The local presence of Mediterranean coastal shrubland, hygrophilous riverine forest and mesophilous plain forest is combined with the regional contribution of mountain vegetation through the sequence. Oscillations between inundated and wet ground characterized the studied area until the ad 79 eruption. Such a natural environment shows anthropogenic traits since pre-Roman times: pasturelands, cultivated fields and olive groves, which probably occupied drier soils. The most important change in the land use system was the introduction of cabbage cultivation in the fourth century bc and its intensification from the second century bc, when Roman influence grew. The presence of tree crops and of ornamental trees reveals the opulence of the Imperial age until the catastrophic eruption.

The present study concerns the Phoenician-Punic site of Motya, a small island set in Western Sicily (Italy), in the Marsala Lagoon (Stagnone di Marsala), between Trapani and Marsala. A big disposal pit, datable to between the first half of the 8th and the mid-6th century bc, was identified in Area D. This context was sampled for plant macro-remains through bucket flotation. Palynological treatment and analysis were also performed on soil samples collected from each of the identified filling layers. The combination of the study of macro- and micro-remains has shown to be effective in answering questions concerning introduced food plants and agricultural practices, and native plants, including timber use. Here we investigate if a waste context can provide information about Phoenicians at Motya and their impact on the local plant communities. We found that human diet included cereals (mostly naked wheat), pulses and fruits. A focus was placed on weeds (including Lolium temulentum and Phalaris spp.) referable to different stages of crop processing. This aspect was enriched by the finding of cereal pollen, which suggests that threshing (if not even cultivation) was carried out on site. Palynology also indicates an open environment, with little to no forest cover, characterized by complex anthropogenic activities. Anthracology suggests the presence of typical Mediterranean plant taxa, including not only the shrubs Pistacia lentiscus and Erica multiflora, but also evergreen oaks. The presence of a stone pine nut and of Pinus pinea/pinaster in the pollen rain is noteworthy, suggesting the local occurrence of these Mediterranean pines outside their native distribution range. This represents the first such find in the central Mediterranean. Finally, the present study allows us to compare Motya’s past environment with the present one. The disappearance of Juniperus sp. and Erica arborea from the present-day surroundings of the Marsala lagoon appears to be related to land-overexploitation, aridification or a combination of both processes.

European land-use indicators in North America typically include pollen from Poaceae (grasses), Ambrosia (ragweed), Rumex (dock), and/or Plantago (plantain). A potential complement to this list is Fabaceae (legumes), members of which were introduced by colonists to eastern North America. Historical records show that Trifolium (clover) was widespread from the earliest days of colonization, and several other Eurasian legumes were also introduced for livestock forage by 1840 ce. As an insect-pollinated family with taxa native to the region, Fabaceae is not usually emphasized in palynological reconstructions. However, Plantago is commonly used as an indicator despite being represented by native and insect-pollinated varieties. Likewise, it can be difficult to distinguish between native and introduced Poaceae, yet an increase in grass pollen is generally taken as a disturbance indicator. In order to assess the value of Fabaceae as an indicator of the settlement horizon, the R package neotoma was used to retrieve Fabaceae pollen data from the Neotoma Paleoecology Database to assess regional trends in abundance and for comparison to the usual suite of taxa. Results confirm the possible utility of Fabaceae as an anthropogenic marker in the study area. Two case studies with good chronological control illustrate how all five taxa can be found at the settlement horizon across the study area and over time: a 14C and 210Pb-dated pond in Connecticut anthropogenically impacted by 1700 ce, and a varved lake in Ontario impacted by 1840 ce.

Multi-proxy analysis of the coprolites which were found during excavations at two Late Neolithic (fourth millennium bc) pile-dwelling sites (Crnelnik and Stare gmajne) in Slovenia yielded some new insights into human–dog relations and behaviour. The digested content is presented in a multidisciplinary approach, in which palynological, palaeoparasitological, archaeobotanical and archaeozoological features are studied and genetic signs are tested. Beside the origin of the coprolites, the size of an animal and the diet, the faeces provided some additional information, such as health, status, nutrition habits, environment and season.

The Spanish conquest of the Cuchumatanes Highlands (1524–1541 ce) led to dramatic land use changes adhering to colonial practices and values, which included the rearing of livestock, agriculture, timber extraction, mining, and the relocation of indigenous populations to new settlements. These changes are often recorded in historical accounts and official records; however, these are sparse, incomplete, and have been lost over the passage of time. Here, we present a high-resolution palaeoenvironmental reconstruction for Cenote Kail (Guatemala) since the Spanish Conquest, to provide additional evidence of land use changes from an integrated multi-proxy perspective. We analysed: (i) fossil pollen; (ii) macroscopic and microscopic charcoal; and (iii) dung fungal spores (Sporormiella) from a lake sediment core extracted from Cenote Kail in the Cuchumatanes highlands, combining these analyses with remotely sensed satellite data. We reconstructed: (i) forest composition and dynamics; (ii) burning, (iii) fauna abundance, and (iv) agricultural activities. High resolution age-depth modelling was conducted using a combination of 210Pb and 14C dates. The high temporal resolution enabled a novel integrated validation of the charcoal data sets with remotely sensed satellite data and the historical record. Three stages of floral compositional change were discerned from the palynological assemblage data encompassing: (i) the decline of mixed hard wood forests (MHWF), associated with the building of new settlements, agriculture and timber extraction for fuel (1550–1675 ce); (ii) pastoral expansions involving the rearing of livestock (1700–1800 ce); and (iii) the expansions of urban settlements and increasing management of the land (1821–2015 ce). Seed predation is suggested as the dominant factor preventing MHWF from re-establishing in the Cuchumatanes Highlands over the past 500 years. Burning is limited locally and regionally and in line with the modern regime, which suggests that fire has been managed and controlled since European contact.

Waldgirmes in Hesse (Germany) is one of the oldest Roman towns east of the Rhine River. It was founded in 3 bc and abandoned after ad 9, probably in ad 16, when the departing soldiers set fire to the town. In one of the two excavated wells, partially laminated, organic-rich sediments have been preserved which were deposited during the colonisation of the town. The pollen, plant macro-remains and micromorphological sampling strategy of the well sediment allows us to draw a detailed picture of the vegetation and its development in and around the settlement area. Through thin section analyses, the stratigraphy and composition of sediments could be differentiated in more detail. Deriving from different source areas, the pollen and plant macro-remains assemblages gave insights into the tree species composition of the surroundings as well as the vegetation in the settlement. The town was founded within a cultural landscape that had existed for centuries, and not in the dense dark Germanic forests described by Tacitus around ad 100. The contemporaneous woodlands were dominated by oaks. In the well within the town, not only characteristic components of urban waste (bones, charcoal, ash), but also plant remains from the immediate surroundings (pollen, phytoliths, seeds, fruits, leaves) were preserved. Typical urban vegetation from trodden paths, nutrient-enriched sites, and grazing areas could be reconstructed. Thick leaf layers prove the presence of trees, especially oaks, close to the well. Horticulture is evident through cultural indicators, and there are also signs of cereal processing within the settlement.

Located in the Romanian Banat region, the Late Bronze Age (LBA) fortification Cornesti-Iarcuri is the largest known prehistoric settlement in Europe. Archaeobotanical and geoarchaeological investigations have targeted the reconstruction of vegetation, land use practices and subsistence strategies at the site, together with related human impact and environmental changes in the wider study area. Since colluvia constitute valuable archives in terms of landscape history and anthropogenic disturbance, one major focus was put on floodplain profiles. In the valleys, two generations of colluvium prevailed which were separated by fossil topsoils. Based on several radiocarbon datings, a chronology of events, including distinct phases of geomorphological activity and stability, has been established. Some of the buried palaeosurfaces contained pollen in sufficient concentrations to confirm off-site data from the Vinga area, where the regional vegetation during the Middle Copper Age consisted of Tilia-dominated woodlands with steppe elements. Following a major Late Copper Age deforestation phase that also led to considerable soil erosion, the gradual formation of a cultural landscape is documented by a progressive decline in tree cover in which Quercus gained relative importance, and a continuous presence of land use indicators. Plant macro-remains from archaeological excavations underpin both the openness of the semi-natural woodlands during the pre-fortification era and the increase of animal husbandry and farming in the LBA. Despite evident settlement pressure, it proved to be a geomorphologically stable phase. Towards the Early Iron Age, the values of anthropogenic markers in on-site pollen spectra rose to values comparable to those in surface samples.

In a paper from 1942 Johannes Iversen described a warm phase prior to the Allerod warming from lake Bolling in western Denmark and thereby extended the common classification of Lateglacial biostratigraphy. Iversen’s assumptions concerning the Bolling-Oscillation were based on sedimentological features and high Betula (birch) pollen values in two pollen samples prior to the onset of the Allerod warming. Questions regarding the nature of the Bolling warming later became a discussion due to the expansion of terms into different fields and especially with advances in palynology. To further refine the knowledge on the Bolling-Oscillation, Hartmut Usinger investigated the locus classicus in 1982. The method of pollen-size-frequency distribution was applied on birch pollen in order to distinguish between B. nana- and B. pubescens-type pollen, however, the results were never published and got lost over time. Therefore, these data are presented here and the method as described by Usinger (Pollenanalytische und stratigraphische Untersuchungen an zwei Spatglazial-Vorkommen in Schleswig-Holstein, 1975, Danm Geol Unders Arbog, 1977:5–29, 1978a, Pollen Spores 23:389–432, 1981b) is performed on the reassembled dataset again. In order to solve existing ambiguities in Lateglacial terminology, especially concerning the terms Bolling and Meiendorf, we used the data originally compiled by Usinger to discuss it against the background of the previous work by Iversen. Furthermore, the dataset enabled a comparison of different ways to determine between different Betula pollen types. The general focus is set on the distinction of Betula pollen types in order to shed light on the question of the migration of tree birches into northern Europe, especially before the Allerod warming. The results show that the dataset can immensely refine Iversen’s pollen zones. A clear dominance of dwarf birch pollen during the Bolling-Oscillation is demonstrated. Furthermore, the record reveals that Iversen’s assumption concerning the existence of a climatic oscillation prior to the Allerod warming can be confirmed at the locus classicus.

In the Orinoco Llanos, archaeological studies indicate a continuous late Holocene human occupation, including the development of ranked societies, from about 1,500 year bp (ad 500) However, until now palaeoecological studies dealing with the impact of anthropogenic activities in the region have been scarce. Here we present a palynological analysis carried out in the El Cedral archaeological region, Western Venezuelan Llanos. The 145 cm sedimentary record obtained comprises the last ca. 4,200 cal bp (2245 cal bc). Between 4,200–3,800 cal bp (2250–1850 cal bc), wet savannas interspersed with gallery forests dominated in the study area. A wet climate, similar to the present, is suggested by pollen of aquatics and by non-pollen palynomorphs (NPP). Frequent/intense local fires were inferred from charcoal concentrations. Between 3,800–2,100 cal bp (1850–150 cal bc), rapid alternation between forest and savanna-dominated phases occurred. The low diversity of NPP and the low values of pollen from aquatics suggest drier climate conditions than the present. Low charcoal concentrations point to low fire frequencies. Two events of forest expansion at 3,400–3,200 and 2,800–2,100 cal bp were recorded, alternating with savanna-dominated phases (at 3,800–3,600 and 3,200–3,000 cal bp). From 2,100 cal bp (150 cal bc) to the present the expansion of savannas is recorded. Zea mays occurred at 1,880 cal bp (ca. ad 70), accompanied by the increase in charcoal concentration, supporting the hypothesis of intensification of human activities and anthropogenic forest clearance. Different climatic phases recorded in El Cedral match with those described from the Cariaco Basin, indicating that El Nino Southern Oscillation variability was the main climatic forcing. However, the intensification of anthropogenic activities limited the forest expansion expected during the last two millennia, yielding evidence of landscape domestication in this region.

Today, the East European forest-steppe is an agricultural landscape with very few remains of its former natural vegetation. The history of the transformation from natural vegetation to a human-made landscape in the area of Sudzha (Kursk region, Russia) is studied here. We compare the off-site pollen record Sudzha with three on-site pollen records obtained from the archaeological site Kurilovka-2. The sediment core Sudzha covering the last 2,500 years was taken from an oxbow lake in an area with archaeological sites of the early Slavonic period (3rd–8th centuries ce). The Sudzha pollen record indicates dominance of broadleaf forests and meadow steppes in the area from 2,500 to 200 cal year bp with two major settlement phases one between ~ 2,000 and 1,600 cal year bp (~ 50 bce to 350 ce) and the other between 1,100 and 600 cal year bp (850 and 1350 ce) followed by a total deforestation and transformation to an agricultural landscape over the last 200–300 years. Similar changes in the last 300–400 years are indicated by the three on-site pollen records. It is noteworthy, however, that the record Sudzha does not provide an intensive signal of human impact during the 5th–8th centuries ce. This points to a quite restricted spatial influence of the Early Slavonic settlements on the vegetation, leading to a relatively low contribution of palynological anthropogenic indicators to the regional pollen rain signal.

Conventional wisdom states Cannabis sativa originated in Asia and its dispersal to Europe depended upon human transport. Various Neolithic or Bronze age groups have been named as pioneer cultivators. These theses were tested by examining fossil pollen studies (FPSs), obtained from the European Pollen Database. Many FPSs report Cannabis or Humulus (C/H) with collective names (e.g. Cannabis/Humulus or Cannabaceae). To dissect these aggregate data, we used ecological proxies to differentiate C/H pollen, as follows: unknown C/H pollen that appeared in a pollen assemblage suggestive of steppe (Poaceae, Artemisia, Chenopodiaceae) we interpreted as wild-type Cannabis. C/H pollen in a mesophytic forest assemblage (Alnus, Salix, Populus) we interpreted as Humulus. C/H pollen curves that upsurged and appeared de novo alongside crop pollen grains we interpreted as cultivated hemp. FPSs were mapped and compared to the territories of archaeological cultures. We analysed 479 FPSs from the Holocene/Late Glacial, plus 36 FPSs from older strata. The results showed C/H pollen consistent with wild-type C. sativa in steppe and dry tundra landscapes throughout Europe during the early Holocene, Late Glacial, and previous glaciations. During the warm and wet Holocene Climactic Optimum, forests replaced steppe, and Humulus dominated. Cannabis retreated to steppe refugia. C/H pollen consistent with cultivated hemp first appeared in the Pontic-Caspian steppe refugium. GIS mapping linked cultivation with the Copper age Varna/Gumelniţa culture, and the Bronze age Yamnaya and Terramara cultures. An Iron age steppe culture, the Scythians, likely introduced hemp cultivation to Celtic and Proto-Slavic cultures.

Utilising a multi-profile palynological approach and a rapid scanning technique, this paper examines whether or not cereal cultivation is representative of a medieval Icelandic farmstead’s social status; first as a correlate by confirming that cereals were grown in association with the archaeological features characteristic of high status and second, as an indicator in its own right through comparison with other datasets from inferred lower status farms in the same valley. The results suggest that medieval cereal cultivation in Mosfellsdalur was confined to the landholding of the Mosfell Estate. This is probably a direct consequence of the locale being settled early during Iceland’s colonisation and thereby allowing settlers there to secure the prime location in the valley for agriculture. The later abandonment of cereal cultivation on the Estate also appears to be linked to social circumstances in Mosfellsdalur c. ad 1200. An evaluation of other pollen studies and historical sources intimates a transition in cereal cultivation from inland toward coastal (and perhaps geothermal) sites in the mid-13th century, probably as a consequence of the onset of the Little Ice Age. These sites may also be linked with high status institutions. Despite this effort to adapt to altered climate conditions, cereal cultivation in Iceland is believed to have been completely abandoned by ad 1500. Overall, the temporal and spatial dynamics of cereal cultivation are shown to be complex, subject to both societal and environmental changes.

Lake Kinneret, also known as the Sea of Galilee and Lake Tiberias, is located in the northeast of Israel. At a lake level of 211 m b.s.l. (below mean sea level), the central basin is 43 m deep. The maximum length of the lake is 21 km (N–S) and its maximum width is 12 km (W–E). Lake Kinneret’s surface area is 166 km2. A new 17.8 m long sedimentary core was drilled in 2010. Here, we present the entire palynological record from it, which covers the last ~ 9,000 years. Special emphasis is given to the natural and human-influenced vegetation history of Galilee in comparison to that of the more southerly Dead Sea region. Significant signs of human impact are the first Olea (olive) increase during the beginning of the Chalcolithic period between 7,000 and 6,500 years ago, as well as the prominent Olea phase during the Hellenistic/Roman/Byzantine period between 2,300 and 1,500 years ago. Mediterranean macchia and bathas scrub vegetation, as known in the area today, has developed in the southern Levant under human impact since the last ca. 1,500 years.

Archaeological evidence of Cannabis sativa is comprised of textiles, cordage, fibre and seeds, or pottery impressions of those materials, as well as pseudoliths and phytoliths (pollen is not addressed here). Previous summaries of this evidence connect hemp with Bronze and Iron Age cultures in Europe. This study improves upon earlier summaries by: (1) accessing a larger database; (2) relying on original studies instead of secondary sources; (3) stratifying evidence by its relative robustness or validity. We coupled digital text-searching engines with internet archives of machine-readable texts, augmented by citation tracking of retrieved articles. The database was large, so we limited retrieval to studies that predated 27 bce for west-central Europe, and pre-ce 400 for eastern Europe. Validity of evidence was scaled, from less robust (e.g., pottery impressions of fibre) to more robust (e.g. microscopic analysis of seeds). Archaeological sites were mapped using ArcGIS 10.3. The search retrieved 136 studies, a yield four-fold greater than previous summaries when parsed to our geographic/time constraints. Only 12.5% of studies came from secondary literature. No robust evidence supports claims of Neolithic hemp usage. One Copper Age site in southeastern Europe shows robust evidence (from the Gumelniţa-Varna culture). More robust evidence appears during the Bronze Age in southeastern Europe (Yamnaya and Catacomb cultures). An Iron Age steppe culture, the Scythians, likely introduced hemp cultivation to Celtic, Slavic and Finno-Ugric cultures. The results correlate with a recent palynology study of fossil pollen in Europe. We discuss possible autochthonous domestication of Cannabis in Europe.

Tracking changes in biodiversity through time requires an understanding of the relationship between modern diversity and how this diversity is preserved in the fossil record. Fossil pollen is one way in which past vegetation diversity can be reconstructed. However, there is limited understanding of modern pollen-vegetation diversity relationships from biodiverse tropical ecosystems. Here, pollen (palynological) richness and diversity (Hill N (1)) are compared with vegetation richness and diversity from forest and savannah ecosystems in the New World and Old World tropics (Neotropics and Palaeotropics). Modern pollen data were obtained from artificial pollen traps deployed in 1-ha vegetation study plots from which vegetation inventories had been completed in Bolivia and Ghana. Pollen counts were obtained from 15 to 22 traps per plot, and aggregated pollen sums for each plot were > 2,500. The palynological richness/diversity values from the Neotropics were moist evergreen forest = 86/6.8, semi-deciduous dry forest = 111/21.9, wooded savannah = 138/31.5, and from the Palaeotropics wet evergreen forest = 144/28.3, semi-deciduous moist forest = 104/4.4, forest-savannah transition = 121/14.1; the corresponding vegetation richness/diversity was 100/36.7, 80/38.7 and 71/39.4 (Neotropics), and 101/54.8, 87/45.5 and 71/34.5 (Palaeotropics). No consistent relationship was found between palynological richness/diversity, and plot vegetation richness/diversity, due to the differential influence of other factors such as landscape diversity, pollination strategy, and pollen source area. Palynological richness exceeded vegetation richness, while pollen diversity was lower than vegetation diversity. The relatively high global diversity of tropical vegetation was found to be reflected in the pollen rain.

Little is known about the timing and the vegetation dynamics shortly after the Last Glacial Maximum (LGM) on the Swiss Plateau 19,000–15,000 cal BP. Subsequent Late Glacial and Holocene vegetation changes are better known; however, it is unclear if the few available palynological and macrofossil records are able to capture the entire vegetation variability of the region. A new palaeoecological multi-proxy study using pollen, spores, charcoal and X-ray fluorescence (XRF) from Burgäschisee (Swiss Plateau, 465 m a.s.l.) is applied to reconstruct vegetation, fire and land use for the past 19,000 cal years. Steppe tundra vegetation established at c. 18,700 cal BP only c. 300 years after the end of the LGM and deglaciation. A shift from steppe tundra (Artemisia, Helianthemum) to shrub tundra (Betula nana, Salix, Juniperus) with sporadic tree Betula stands occurred around 16,000 cal BP, most likely in response to climate warming after the end of Heinrich event 1. Abundant spores of coprophilous fungi (Sporormiella, Cercophora) may reflect the presence of Pleistocene large herbivores (e.g. Mammuthus primigenius, Bison bonasus, Rangifer tarandus). Afforestation started more than 2,000 years later with Juniperus and tree Betula around 14,500 cal BP. Mixed Betula and Pinus sylvestris forests persisted until 10,800 cal BP, when mixed elm forests expanded into the region in response to climate warming. Around 8,200 cal BP, mesophilous Fagus sylvatica and Abies alba partly replaced more heliophilous species in the forests, when climate became less continental and more moist. Pollen of Cerealia, Plantago lanceolata and other crops and weeds suggest that agricultural activities became significant during the Neolithic around 6,500 cal BP (4550 cal BC). Archaeological findings from Neolithic pile dwellings around 5,950 cal BP (4000 cal BC) indicate local settlements around the lake. The lake sediments are laminated for most of the last c. 6,800 years. With two independent proxies (XRF and pollen), we can demonstrate that these laminations are annual, suggesting short-term mixing of the lake water due to a more open landscape in response to land use. Our study shows that the annually laminated (varved) sediments from Burgäschisee have a great potential for high-resolution multi-proxy analyses covering the past c. 6,800 years. They can provide accurate ages of cultural phases that might be compared with dendrochronologically dated evidence from lake dwellings.

The development of palynology since its recognised launch in 1916 by Lennart von Post is examined in terms of its historiography, the biographies of pollen analysts and the role of those who have influenced the discipline. Emphasis is placed upon research beginnings in Scandinavia (especially Sweden), Great Britain and Ireland. Within an analytical narrative which includes archival and published sources, special consideration is given to a ‘proclamation’ on methodology which appeared in 1909, on a supposed geology/botany dichotomy stemming from von Post’s background, on the forgotten early practitioners in Britain and Ireland and their connections, on the role of women up to the end of the Second World War and on issues related to wartime hostilities. Present day palynology can trace a continuity from von Post and palynologists are part of an extended disciplinary genealogy. Ignorance of these can be seen as a loss of heritage and to represent an intellectual impoverishment.

In 2016 the 100th birthday of palynology was celebrated: Lennart von Post introduced the scientific discipline in 1916 at lectures in Oslo and Stockholm. However, von Post was not the first to study pollen, and research dates back to the works of Nehemiah Grew and Marcello Malpighi in the late 17th century. Afterwards studies focussed mainly on the role of pollen for plant reproduction, and on pollen morphology. The first scientist who observed fossil pollen was Heinrich Goppert in 1836. However, it was not until the 1890s and 1900s that the study of pollen became a regular component in the reconstruction of past vegetation. Lennart von Post subsequently developed theoretical and practical concepts and methods of pollen analysis, and raised palynology from its infancy into the mature scientific discipline that flourished greatly during the century that followed.

The limited availability of historical and archaeological evidence means that much is still unknown about the development of Sami reindeer herding in Fennoscandia in both the recent and more distant past. To address this problem, high-resolution palynological analyses, 14C and 210Pb dating were undertaken on two adjacent (

The infill of abandoned river channels is the main information source on postglacial vegetation and landscape evolution in inland northern Belgium. Palynological studies of these deposits also contribute to a better understanding of the environmental context of the numerous archaeological sites in this area. The results are presented of palynological analyses, including pollen, spores, non-pollen palynomorphs and microcharcoal from the palaeochannel infill at seven sites in the Scheldt and Durme valleys in northern Belgium. Based on the regional vegetation evolution and radiocarbon dating, a biozonation for the Scheldt Basin is constructed encompassing the final Late-glacial and Holocene period. The local taxa show a hydroseral succession from open-water environment, through reed or sedge fen to carr forest. Salix and Viburnum opulus were important elements in the Early Holocene riparian and carr vegetation, but are often underrepresented in pollen diagrams from larger valleys. There are indications for human disturbance and agriculture between 7,000 and 6,400 cal bp. The simultaneous decline of elm and lime in the study area around 4,000 cal bp can be linked to human impact.

The paper presents a critical review of the zooarchaeological, macrobotanical, palynological and archaeological data from Lithuania and their previous interpretations, which formerly served as the basis for the concept of development of pre-Neolithic or Subneolithic low intensity farming and/or livestock breeding in the eastern Baltic region. Moreover, it presents the first direct AMS dates from the crop remains and domestic animal bones discovered in Lithuanian Subneolithic and Neolithic settlements. An investigation proved that most of, or possibly all, the early farming “evidence” came from the wrong identification of the plant and animal species and incorrect dating of crop remains and domestic animal bones. The errors of dating were caused by the fresh water reservoir effect being ignored when dating the bulk lacustrine sediment samples, by the failure to evaluate the impact of the palimpsest and bioturbation phenomena on the formation of an archaeological layer, and by insufficient attention to stratigraphy and spatial documentation of the finds during very extensive archaeological excavations in the second half of the 20th century. To date, no credible evidence is available in Lithuania that domestic animals had been kept and crops grown before the Neolithic Globular Amphora and Corded Ware cultures in 3200/2700 cal bc. However, this does not mean such evidence may not appear in the future, provided direct AMS dating of animal and crop residues from Subneolithic contexts continues, and systematic macrobotanical studies finally start not only in the lake settlement and fishing sites, but also in higher altitude areas.

This multi proxy-study attempts to illuminate land use changes resulting from ore exploitation. The rationale is to use different types of proxy data to evaluate interactions between human activities and the environment in order to gain deeper insight into the environmental impact of prehistoric mining activities. This is approached through palynological and supplementary geochemical analyses of a peat core from a mire, situated in the immediate vicinity of a former ore exploitation area in the Kitzbuhel (Tyrol, Austria) mining district, together with radiocarbon dating, stratigraphic analysis and loss on ignition. The results indicate the first clear human interference in the natural vegetation at approximately 2000 cal bc. During the Late Bronze Age (c. 1400–950 cal bc), palynologically proven changes of the vegetation were found, including two time periods with forest clearing where values of anthropogenic pollen indicators increased synchronously with rising heavy metal values. Through geochemical analysis a connection to mining activities could be made. Subsequently this period is synchronous with dated mining activities. During the Late Bronze Age and the Iron Age (c. 950–50 cal bc), humans are still present in the region but to a lesser degree. An intense anthropogenic phase follows during the Late Iron Age and Roman Times (c. 50 cal bc–cal ad 200) and is indicated by pasture and tillage. In the following period anthropogenic impact decreases, then enhanced human activities are visible through rising pasture, settlement and cultural indicators from the second half of the Early Middle Ages (since c. cal ad 900). From this point on, eight more clearing episodes up to the 19th century could be detected palynologically. Geochemical analyses and historical sources document a second mining phase during Early Modern Times (15th–16th centuries). This study shows that the combination of palynological, geochemical and archaeological data enables a better understanding of the impact of mining on the environment in the Kitzbuhel mining district and is thus applicable to other comparable mining districts.

At the turn of the second to the third millennium bc, fortified and systematically organized settlements along with a developed metallurgy emerged in the Trans-Ural steppe. In order to reconstruct the related vegetation and climatic changes in the area and to detect effects of human impact during the respective Bronze-Age Sintashta-Petrovka and Srubnaya-Alakul cultures (2100–1650 cal bc), palynological and sedimentological investigations accompanying archaeological excavations were carried out. Statistical analyses of pollen spectra from two sediment cores in the immediate vicinity of the settlement Kamennyi Ambar demonstrate substantial similarities in Bronze Age and present vegetation covers. Higher percentages of arboreal pollen, especially in respect of deciduous trees, and consistently high values of Artemisia within the regional spectra suggest that Bronze Age cultures developed in a relatively humid environment. Simultaneously, higher values of Chenopodiaceae, Plantago and Cichorioideae point to steppe degradation at local scales, whereas regional vegetation appears to have been relatively unaffected by human activities. Although more than 30 localities in the micro-region of Karagaily Ayat were analyzed, none of these sediment cores accounts for a continuous sedimentation during the last 9,000 years. Our results reveal that climatic fluctuations are not always directly reflected by sediment formation, with many non-climatic factors, such as post-sedimentary processes, local morphology and erosion believed to be crucial. Only the use of high resolution AMS radiocarbon dating is suitable to detect hiatuses, hence providing a valuable clue to the interpretation of palaeoenvironmental conditions.

The relationship between farming communities in south-eastern Europe and wild plant resources, fruit and nut trees in particular, is explored in this paper, based on charred plant remains from House 1 at late Neolithic Dikili Tash in eastern Macedonia, northern Greece, retrieved between 2010 and 2012. Within the rubble of a burnt destruction level dated to the second half of the 5th millennium cal BC, a wide range of cultivated crops like cereals, pulses and flax were stored together with a variety of fruit and nuts, such as acorns, wild pears, grapes, including grape pips and grape pressings and possibly figs, too. These finds provide a rare opportunity to investigate the use of fruit as well as the origins and context of wine making and consumption in the Neolithic of south-eastern Europe. Human interference with natural vegetation in relation to use of wild trees is discussed in light of the archaeobotanical, palynological and charcoal evidence from the wider area of the site. It is suggested that the remains from Dikili Tash may be pointing towards some early form of arboriculture in the region. The interplay of wild and domesticated plant resources encountered at the site is discussed within the framework of established oppositions between ‘wild’ and ‘domesticated’ in archaeological discourse. It is suggested that fruit and nut use at Dikili Tash might correspond to old traditions dating back to the hunter-gatherers of south-eastern Europe while wine, for which there is evidence at the site, might have acted as a mediator between human communities, cultivated landscapes and wild vegetation, inducing altered states of consciousness and cultivated/wild boundary transitions.

Pollen diagrams from Lake Yenicaga in northern Turkey published by Beug (Quaternary Paleoecology, Yale University Press, New Haven, 1967) and Bottema et al. (Palaeohistoria 35/36(1993/1994):13–72, 1995) are connected, reflecting the development of the vegetation over the time from the Older Dryas period to about ad 800. The sequence can be divided into four pollen zones with several subzones. Six periods of continuous human impact are described.

Palynology provides the opportunity to make inferences on changes in diversity of terrestrial vegetation over long time scales. The often coarse taxonomic level achievable in pollen analysis, differences in pollen production and dispersal, and the lack of pollen source boundaries hamper the application of diversity indices to palynology. Palynological richness, the number of pollen types at a constant pollen count, is the most robust and widely used diversity indicator for pollen data. However, this index is also influenced by the abundance distribution of pollen types in sediments. In particular, where the index is calculated by rarefaction analysis, information on taxonomic richness at low abundance may be lost. Here we explore information that can be extracted from the accumulation of taxa over consecutive samples. The log-transformed taxa accumulation curve can be broken up into linear sections with different slope and intersect parameters, describing the accumulation of new taxa within the section. The breaking points may indicate changes in the species pool or in the abundance of high versus low pollen producers. Testing this concept on three pollen diagrams from different landscapes, we find that the break points in the taxa accumulation curves provide convenient zones for identifying changes in richness and evenness. The linear regressions over consecutive samples can be used to inter- and extrapolate to low or extremely high pollen counts, indicating evenness and richness in taxonomic composition within these zones. An evenness indicator, based on the rank-order-abundance is used to assist in the evaluation of the results and the interpretation of the fossil records. Two central European pollen diagrams show major changes in the taxa accumulation curves for the Lateglacial period and the time of human induced land-use changes, while they do not indicate strong changes in the species pool with the onset of the Holocene. In contrast, a central Swedish pollen diagram shows comparatively little change, but high richness during the early Holocene forest establishment. Evenness and palynological richness are related for most periods in the three diagrams, however, sections before forest establishment and after forest clearance show high evenness, which is not necessarily accompanied by high palynological richness, encouraging efforts to separate the two.

The age of the introduction of cereal cultivation in northern Europe has long been debated by researchers from many disciplines, in particular archaeology and palaeoecology. Over the past 40 years extensive palynological data have been collected concerning pre-industrial land use in northern Fennoscandia. This paper reviews palynological studies that include records of fossil cereal pollen from northernmost Sweden, Finland and Norway at latitudes north of 63°N. The geographical extent of known early cultivation sites is constantly expanding, with more than 100 records of cereal pollen pre-dating ad 1700. The oldest records of scattered cereal pollen derive from Neolithic times. Periods of continuous cultivation, indicated by cereal pollen recorded recurrently in the sediment profiles, derive from the Late Neolithic and Early Bronze Age. Collectively, the reviewed pollen records indicate that cereal cultivation was first introduced into areas close to the coast and later to the interior, and that it may have been practiced locally long before sedentary settlements based on intensive cultivation were established during medieval times. The data do not indicate a latitudinal spread of cultivation from south to north. However, methodological problems relating to pollen morphology of cereals, site characteristics and lack of connections to archaeologically excavated sites imply that the value of many early cereal pollen finds remains unclear. To increase our understanding of the context in which cereal cultivation was introduced in northernmost Fennoscandia, multidisciplinary studies integrating palaeoecology, archaeology and history are needed.

A palynological and sedimentological record from the Mahwaqa Mountain in KwaZulu-Natal, South Africa, provides evidence of the vegetation dynamics in this part of the Grassland Biome during the last c. 18,000 years. The wetland is located at 1,850 m on an isolated outlier of the Ukhahlamba–Drakensberg Mountain range on an ecotone along a climatic gradient. The vegetation responded to humidity and temperature changes during the late Pleistocene and Holocene. The period c. 18,000–13,500 cal. bp is characterized by high Ericaceae and Restionaceae percentages and decreasing values of charred particles, indicating cool conditions. Around 13,500–8,500 cal. bp, Ericaceae were gradually replaced by Poaceae, signaling climate warming. Growing environmental wetness during the same time period is inferred from Phragmites-type and Cliffortia pollen percentages. Since c. 8,500 cal. bp, Cliffortia, Restionaceae, and Phragmites-type percentages have maintained low levels. Ericaceae were almost completely replaced by grasses and Asteraceae by c. 7,500 cal. bp. All indications are that warm and fluctuating moisture conditions followed until 4,600 cal. bp but they became driest between c. 4,600 and 3,500 cal. bp, when high Asteraceae, Pentzia-type and Scabiosa percentages were prominent. From c. 3,500–800 cal. bp, the increase of sedges, Aponogeton and grass pollen (including Phragmites-type) at the expense of Asteraceae pollen suggests the return of slightly more humid conditions. Since c. 1,000 cal. bp an increase of water demanding Podocarpus and Cliffortia occurred. Pine pollen indicates the recent introduction of alien plants in the 19th and 20th centuries.

Castellum Fectio was one of the largest fortifications along the Limes, the northern border of the Roman Empire. The castellum, situated 5 km southeast of Utrecht, the Netherlands, was occupied from around the start of our Era to ca. a.d. 260. It was situated along a river bend of the Rhine that was cut off from the main stream during the occupation of the Roman fort. A 6 m long sediment sequence was recovered from the infill of the residual channel and pieces of Roman wall plaster, glume bases of Triticum spelta and radiocarbon dates indicate that the sediments were deposited during the period of Roman occupation. The combined palaeoecological analyses—palynological, macrobotanical, entomological and geochemical—allow a detailed reconstruction of changing environmental conditions as a consequence of the Roman occupation. The pollen record reveals a dramatic decrease in arboreal pollen, suggesting that the Romans were involved in large-scale deforestation, transforming semi-open parkland to a landscape of meadows and agricultural fields. Non-pollen palynomorphs, botanical macrofossils and insect remains support this conclusion. The recorded mycoflora shows a shift from assemblages characterised by the tree pathogen Kretzschmaria deusta to assemblages dominated by spores of fungi associated with herbaceous plants, concurrent with the decrease in arboreal pollen. The presence of masticated bran fragments of cereals, clover remains, eggs of intestinal parasites and entomological and geochemical data in the upper part of the sequence indicates that these sediments largely consist of faeces that were dumped into the former channel. Surprisingly, seeds of salt tolerant species are encountered in the sediments of this inland site, which was situated outside the influence of the sea. Horses may have brought these seeds to Fectio in their intestinal tracts after grazing in coastal meadows.

Human coprolites from Birka, Sweden and Durrnberg, Austria, have been found, dated and palynologically analysed as a part of interdisciplinary studies. All their pollen spectra are dominated by insect-pollinated taxa well-known as nectar producing flowers, suggesting some consumption of honey. Among those spectra, some show significantly high values of Filipendula ulmaria (meadowsweet) pollen, which was historically used as flavouring in mead production, and which together with other indicators for honey, suggest that mead was part of the historic and prehistoric diet both in Birka and Durrnberg. An evaluation of the background pollen suggests for the Birka specimen that honey was imported to the site from southern Baltic areas. The use of mead based on written sources is known at least from the Roman period. Archaeological studies demonstrate mead as an old crust residue on the inside of pots and other earthenware used as important funeral gifts from at least the 27th–25th centuries b.c. in Georgia. A comparison of the pollen records of European honey/mead samples strongly suggests that Filipendula is indicative of mead.

Palynological investigations on two well-dated peat profiles provide insights into Neolithic vegetation and settlement history from Hummling in north-western Germany. The site selections allow comparisons between local and regional vegetation changes and are used to estimate the extent of Neolithic influence on the vegetation. The interpretation of the fossil spectra relied on radiocarbon dating, evaluation of pollen indicator taxa, non-pollen palynomorphs and multivariate techniques. During the late Mesolithic the vegetation was dominated by mixed oak forests while openings in forest cover were detected, with a decline in elm reflected in the regional pollen record around 4250 cal. b.c. The presence of humans is shown by settlement indicators that are first recorded at ca. 3800 cal. b.c. Vegetation changes were small between 4300 and 3600 cal. b.c. This suggests that regional vegetation was relatively resilient to small-scale disturbances. Possible indications of grazing were recorded in the spectra of the local pollen profile but there is no clear-cut evidence for Neolithic activity. Between 3520 and 2260 cal. b.c. decreases in forest cover were inferred from both profiles and increases in settlement indicators reflect farming activity. These changes coincide with the emergence in the area of the Funnel Beaker Culture and the subsequent Single Grave Culture. Both profiles suggest that settlement probably ceased between ca. 3230 and 3050 cal. b.c. This lull or cessation in activity was probably regional in character. After 2260 cal. b.c. human impact on the vegetation decreases and woodlands regenerate. The longevity of the regeneration phase—ca. 690 years—was probably connected with the low resilient capability of the vegetation on the poor soils.

The geomorphological analysis of the river Tocantins alluvial area was carried out by remote sensing techniques, jointly with studies of sedimentary facies, mineralogy and geochemistry, pollen and spores, C and N isotopes data and radiocarbon dating of a sediment core from a floodplain lake, near the river Tocantins in the southeastern Amazon region. The aim was to identify and discuss the relationship between the main geomorphological processes and climatic factors, as well as their influence on vegetation patterns. The data indicate three phases of sediment and organic matter accumulation associated with vegetation changes during the Holocene: in Phase 1 (>7760 ± 80 cal yrs b.p.), the channel was gradually being disconnected from its main course, accumulating organic matter from terrestrial woody plants; in Phase 2 (>7760 ± 80 to 5145 ± 175 cal yrs b.p.), the very low energy flows allowed the deposition of autochthonous organic mud from suspension. Herbaceous vegetation and a Mauritia palm-swamp dominated the study area, followed by an increase in the contribution of freshwater dissolved organic carbon (DOC) to the lake; Phase 3 (5145 ± 175 cal yrs b.p. until present) records the expansion of ferns and arboreal plants, and greater freshwater organic matter input into the lake with anoxic water conditions. The development of the studied lake occurred by geomorphological changes and the filling process of an abandoned channel according to hydrodynamics and vegetation changes which were probably influenced by climatic changes during the Holocene.

The lowland heathlands of southern England comprise ca. 14 % of the total area of this habitat in Europe yet their history is poorly understood. This paper presents the first detailed palaeoecological evidence (combining palynological, microscopic charcoal and radiocarbon data) relating to the origin and long-term dynamics of heathland vegetation in southern England. Valley peat sites, situated on the Lower Greensand Group (coarse-grained sandstones) at Conford (Hampshire) and Hurston Warren (West Sussex) have been investigated. The sequence from Conford indicates the unusually late survival of Pinus sylvestris (to as late as ca. 6050 cal. b.p.) in southern England. This is attributed to edaphic factors and, after ca. 7050 cal. b.p., to frequent fires. After intervening phases of dominance by deciduous woodland, heathland vegetation became established in the proximity of both sites in the Late Bronze Age (ca. 3000 cal. b.p.) with increases in indicators of grazing and burning demonstrating an association between the development of heathland and human activity. Thereafter, the pollen and charcoal records show that the vegetation remained in a dynamic state as the scale and nature of human activity varied through time. Major expansions in the extent of heathland occurred relatively recently; after ca. 1450 cal. b.p. at Hurston Warren and after ca. 850 cal. b.p. at Conford. A review of the palaeoecological evidence suggests that the most intense use and greatest coverage of heathland in southern England probably occurred during the medieval to post-medieval periods.

Palaeoecological and archaeobotanical studies of past environments are essential for understanding the kinds of natural settings in which agriculture and prehistoric societies evolved. They increase our knowledge of how human activities have changed environment and vice versa. Human impact on vegetation cover was recognised from the beginning of modern palynological research (Firbas 1937; Iversen 1941). Based on plant communities described from traditional agriculture and the evidence from archaeobotanical records, Behre (1981) identified palynological anthropogenic indicators. They comprise pollen of cultivated plants together with their associated arable weeds and ruderals. These indicators have been widely used and adapted in different European regions (e.g. Rosch 1996; Brun 2011) and the Near East (e.g. Behre 1990). Apart from reconstructing the vegetation dynamics caused by humans, current palynological research attempts to identify the spatial scale and variation of human activities, and their precise duration and location in given landscapes. Recent advances in quantification and numerical analyses of pollen data including climate reconstruction and vegetation/land-cover modelling, which allow for more precise estimates of the opening of woodlands in past times as well as simulating possible scenarios of human impact on the vegetation and landscapes (e.g. Gaillard et al. 2008). With these methodologies, past and future anthropogenic land cover change can be explored (Gaillard et al. 2010). However, until now most applications of these approaches have mainly concentrated on northwestern Europe. Extending them to further regions would be of great interest. Numerical analyses are of increasing importance for estimating human impact on vegetation and to allow disentangling the human and climatically induced vegetation changes in some cases (Feurdean et al. 2010; Bakker et al. 2012). Climate models can be used for the interpretation of human–environment interactions (e.g. Riehl et al. 2009). However, climatic changes of the last millennia are usually too small in amplitude to be detected by most of the quantitative climate reconstructions using pollen data (Kuhl et al. 2010). Also, microand macro-charcoal records in palaeoecological and archaeobotanical studies serving as climate and human indicators are becoming increasingly relevant (Carcaillet et al. 2002). Investigations of charcoal in soil have focussed on the tropics, the boreal zone and around the Mediterranean, however recently the temperate zone has been integrated as well. In addition, on-site wood charcoal analyses can contribute to reconstructing the human impact on vegetation, especially in areas where no suitable pollen archives are available near human occupation sites (Riehl and Marinova 2008). E. Marinova (&) Centre for Archaeological Sciences, University of Leuven, Celestijnenlaan 200E, bus 2408, 3001 Leuven, Belgium e-mail: elena.marinova@bio.kuleuven.be

Past vegetation change and the influence of climate change and anthropogenic pressure during the Holocene is constructed from a series of palynological records sampled from three locations within the territory of the antique site of Sagalassos. The disappearance of the original deciduous oak woodlands and increases in anthropogenic indicator species around 5300 and 4300 b.c. correspond with an increase in settlements in the region. A period of drought following the deforestation may have hampered the recovery of deciduous oak. The timing of the onset of the Beysehir Occupation Phase (BO-Phase) in the territory differs between locations, estimates ranging from ca. 1000–800 b.c. to the start of the Hellenistic period (334 b.c.). The most intense period of arboriculture coincides with the Roman and late-Roman periods. Increases in human pressure on the landscape as reflected in the pollen record correspond with an increased rate of sedimentation and fire activity. The timing of the end of the BO-Phase again differs between locations. Estimates range from the 4th century a.d. to the mid 7th century a.d., when a region-wide shift to dry environmental conditions is observed. Numerical analyses show that post BO-Phase vegetation change is largely driven by climate and displays a succession of dry and wet periods that coincided with well-defined European climate shifts, including the Medieval Climate Anomaly and the Little Ice Age. Current agricultural activities in the region are of a very recent (20th century) origin.

This paper focuses on the impact of Norse settlement on vegetation and landscape around the head of Tunulliarfik (Eriksfjord) in southern Greenland. Pollen, radiocarbon, microscopic charcoal and fungal spore data are presented from a peat monolith which was collected close to the ruins of a large Norse farm complex (group O39 at Qinngua in the former Eastern Settlement). Landnam is identified at ca. cal. a.d. 1020 by a small decrease in pollen from Betula, a slight increase in Poaceae, and the appearance of pollen from Norse apophytes (native plants favoured and spread by human activity) and anthropochores (not native and unintentionally introduced by people). Increases in microscopic charcoal and palynological richness are also apparent. This pattern is broadly consistent with that seen in other pollen diagrams from this region. The sequence is unusual for Greenland, however, in that relatively high Betula pollen percentages (average 20% TLP) are recorded throughout the period of settlement, up to the end of the 14th century a.d. before the profile becomes truncated. If these data are primarily representative of the dry land vegetation, then they suggest that birch woodland and scrub may well have persisted close to the farm throughout the Norse period. Given the potential resource value of woodland to the settlers, this may imply that birch was being managed sustainably, as was the case in Iceland during the medieval period. Coprophilous fungal spores typically found on animal dung are abundant during the early phase of settlement, yet subsequently decline in abundance. This may indicate a likely decrease in grazing intensity or livestock numbers over time, possibly in response to climatic deterioration and/or soil erosion that is expected to have placed increased stress on the pastoral farming system.

As part of a regional research project on wetlands, a analysis was carried out on non pollen palynomorphs (NPP) from the upper sediments of a mountain peat bog in the Ligurian Apennines, northwest Italy. The aim of this project was to see if NPPs could yield useful results on local ecology and human activity in connection with the use of resources. More than 60 NPP types were found, among which a new type is described. A good agreement was observed between pollen assemblages and different NPP types, which are still an under-exploited source of information. Coprophilous fungi suggest the presence of livestock at the site after cal. a.d. 770–1160, while various algae and other water-demanding organisms show an increasing eutrophication of the environment in the most recent phase after cal. a.d. 1500. This approach, which is new for this region, confirms the hypothesis that local farming practices, such as management of pasture and woodland, were not recently established here. The site features and both NPP variety and significance show that further improvement of this additional tool could contribute to answering some questions about past cultural landscapes.

Pollen, micro-charcoal and non-pollen palynomorph (NPP) data from the mid Holocene Ulmus decline and the preceding millennium have provided evidence of repeated fire disturbance of the upland woodland at Bluewath Beck Head, on the North York Moors in northeast England. Woodland disturbance coincides with the Ulmus decline, which at several similar upland sites in northern England is dated to ca. 4800 uncal b.p. (ca. 5550 cal b.p.), and so to the early Neolithic period. Two fire events occur within a cycle of disturbance and regeneration between about 6100 (ca. 6950 cal b.p.) and 5700 b.p. (ca. 6475 cal b.p.), placing them in the later stages of the Late Mesolithic hunter-gatherer occupation of the upland and near the start of the transition to early Neolithic agricultural economies. Increased Melampyrum and Corylus pollen percentages characterise the post-fire vegetation response. These disturbances probably resulted from human activity, suggesting that fire was an integral part of the Late Mesolithic ecology. The local origin of some NPPs greatly enhances the palaeoecological interpretation, showing variations in the hydrological responses to disturbance that are much less visible in the pollen record, and helping to distinguish between local and regional vegetation changes. Other NPPs indicate burning near to the site. A substantial peak in spores of the wood-rot fungus Kretzschmaria deusta across the Ulmus decline may indicate girdling and other woodland management techniques as part of Neolithic woodland farming.

Information from pollen and non-pollen palynomorphs was used to reconstruct ritual burial traditions in the first half of the third millennium b.c. The floor of the burial chamber of the Paravani barrow and a ceremonial wagon were covered with flax textiles (Linum), as evidenced by the abundance of flax microfibres. The wagon was made of pine (Pinus), oak (Quercus) and juniper (Juniperus) wood. This is demonstrated by parenchyma cells of wood which were linked to the destroyed remains of the wagon. The presence of an animal skin could be established. The presence of the skin is evidenced by large amounts of pollen, ascospores of coprophilous fungi, hairs of dermestid beetle larvae and the bristle of a moth grub. According to the taxonomic composition of the flowers used for decoration of the ceremonial wagon and numerous mite remains, the burial occurred during the first half of the summer.

International audience; This paper presents results from a modern dataset of non-pollen palynomorphs and its application to aca. 2,000 year peat record from the same area in the western Pyrenees (Basque Country, France). The modern dataset is composed of 35 surface samples (moss polsters) from a mountainous pasture-woodland landscape. Airborne fungal spores (ascospores and conidia), found dominant in the dataset, are linked to the degree of landscape openness and grazing pressure. The complete spectrum of 13 selected spore-types of dung-related Ascomycetes is positively linked with grazing pressure. However, different dung affinities between the spore-types have been identified. These are types clearly related to high grazing pressure and types with no or unclear dung indicative value. The modern dataset is used to aid interpretation of the local fossil pollen record as an independent 'proxy' to assess past pastoral dynamics. This study confirms the utility of modern nonpollen palynomorphs from terrestrial ecosystems in the reconstruction of historical local pastoral activities but also shows their limitation. It may be necessary to extend such study to wetland ecosystems and to investigate the spatial dimension of some fungal spores.

Agrarian history and local cultural landscape dynamics have been documented through pollen analysis of a peat core within the hamlet of Yttra Berg, which is situated in an upland area of southwest Sweden. The sequence covers the last 5,000 years, from Neolithic to modern time. Wood pasturing started at 2000 b.c., followed by grazing and small-scale cultivation with 500 year cycles from 650 b.c., and permanent fields of agriculture from a.d. 1150. The area was abandoned during the period a.d. 1350–1550. Three cycles of succession related to land-use have been identified for the period 650 b.c. to a.d. 1550. Correlation with frequent clearance cairns in the area is discussed. Recessions of agriculture/settlement during the late Middle Ages and late modern time are documented. Pollen data indicate increased landscape and plant diversity since the Neolithic, closely linked to openness of the agrarian landscape. These results are important for landscape restoration.

The cultural landscape development of a farming community in western Norway was investigated through pollen analyses from a lake and a peat/soil profile. The pollen record from the lake indicates that there was a decrease in arboreal pollen (AP) by the end of the Mesolithic period (ca. 4200 cal b.c.), and that a substantial forest clearance occurred during the Bronze Age (ca. 1500 cal b.c.). The latter, together with grazing indicators and cereals, suggests a widespread establishment of farming. At the beginning of the Roman Iron Age there is an increase in heath communities. The pollen diagram from the peat/soil profile shows the forest clearance in the Bronze Age more clearly than the lake profile. This local pollen diagram is compared with modern pollen samples from mown and grazed localities in western Norway. Both analogue matching and ordination (PCA) indicate that the site was characterised by pastures and cereal fields from the Late Bronze Age to the Late Iron Age. An expansion of cereal cultivation took place during the Pre-Roman Iron Age, and an arable field was established at the site after ca. a.d. 800. This investigation illustrates the potential of selecting pollen sites reflecting different spatial scales, and complements the cultural history of the area as inferred from archaeological and historical records.

The layers studied, parts of banquettes, consist of limnic clayey-silty fine sediments with some intercalated horizons of shaly coal and lacustrine mud flow rubble. In the lower part of the record the Bornette interglacial has been established. This is characterised by high values of Corylus and a significant presence of Quercus and Hedera in addition to a wide-spread presence of Buxus,Ulmus and Carpinus. A linkage of this interglacial to the Eemian (MIS 5e) can be considered an applicable hypothesis. Quantitatively, however, Alnus and Picea exceed all these thermophytics. However the lower part of Lathuile Gare is also characterised by palynological duplications untypical of the Eemian, for example two distinct peaks of Corylus linked to a significant decline in Picea in PZ 3 and 5 and higher values of Abies and Picea in PZ 4 and 6. Possibly these duplications can be explained by a rapid sub-aquatic flow causing a doubling of the beds. In the upper part of the Lathuile Gare record two mean climatic recession periods in the lowest and the second uppermost section (PZ 9 and 15) were detected with increased values of NAP and Pinus in PZ 9 and decreased proportions of Picea in PZ 15. They represent the Chaperon b interstadial and the Lathuile stadial correlated hypothetically with the St. Germain Ib (MIS 5c2) and the Melisey II (MIS 5b) stadial. The enclosed middle part, the Chaparon c interstadial, revealing similarities with the St. Germain Ic (MIS 5c1), is dominated by Picea and Pinus. Only a few thermophilous taxa occur in the middle of this interstadial in PZ 11.

Data from 59 sequences studied through pollen analysis were used to examine the decline in Alnus in Estonia during the Iron Age. Between a.d. 300 and 1300, the Alnus pollen frequency declined markedly in 30 records distributed evenly across the investigated area. The beginning of the decline was time transgressive, coincidental with the start of extensive cultivation, and was frequently connected with the commencement of rye cultivation and the availability of land suitable for cultivation. The greatest reduction in Alnus abundance occurred during the Late Iron Age between a.d. 900 and 1000. This spatially random asynchrony suggests that one or more factors affected Alnus populations across the whole northern region. Human impact is discussed as a plausible cause of the decline. To determine the initiation of extensive crop farming in the eastern Baltic area, pollen diagrams from Latvia, Lithuania and the Novgorod region were also examined.

A pollen diagram from a site in the Esbjerg area, western Denmark, is used for reconstruction of the Holocene vegetational and environmental history there. During the Atlantic there was a parallel development of the landscape to that of other areas in Jylland (Jutland). From the late Neolithic onwards the development took its own course related to the approaching North Sea, which periodically inundated parts of the Esbjerg area. The record reflects landscape development in a formerly marine valley where sediments seem to be missing from parts of the Bronze Age and the early Iron Age. Consequently the landscape development during these times is only reflected in glimpses in the vegetation record, which shows gradually more open woodland and increasing human impact. During the late part of the Iron Age, Viking period and Middle Ages, the woodland was diverse in taxa but became increasingly open, finally reaching a stage during which there may have been too little wood even for daily use. At the same time the use of the land intensified. During the Sub-Atlantic, the Esbjerg area offered good natural resources with extensive grazing areas in the marine marshes in addition to good possibilities for farming and use of the woodland on higher ground, but devastating floods occurred.

A palaeoecological study was conducted close to the forest limit in the northern Scandinavian mountain range. The aim was to elucidate the degree to which human impact has affected the vegetation at Hiednikvalta, a Stallo settlement site. Stallo settlements consist of round hut foundations that have a hearth in the middle and are surrounded by a low turf wall. They were probably established by Sami people using the mountain areas for hunting and/or reindeer herding. In order to separate the effects of humans and climate on the vegetation, a reference area approach was adopted, i.e. the vegetation development at the Stallo settlement site Hiednikvalta was compared with the vegetation development in a forested reference area AvvuhatjAyenhkkAyen, at the same altitude as Hiednikvalta but with no archaeological remains of settlements. Peat stratigraphies were retrieved at the two sites and pollen analysis, loss-on-ignition (LOI), pollen accumulation rates (PAR), macrofossil analysis and Betula pollen size statistics were all examined. The results indicate that Hiednikvalta was forested with Betula trees prior to the Stallo settlement period, which occurred between the eighth and thirteenth centuries. Human activities resulted in a decrease in tree cover at the site, as found in a previous study at Adamvalta, another Stallo settlement site in the region. However, the magnitude of vegetation change, and the post-Stallo vegetation development differed between the two areas, suggesting that site-specific factors are important. The use of reference areas in palynological studies is also discussed.

Vegetation changes during the late Holocene are interpreted from four fossil pollen sequences from two caves at the Los Toldos archaeological locality, Santa Cruz province, Argentina. Taphonomic processes are particularly taken into account in order to analyze the effects on the fossil pollen records of biotic factors such as human occupation and animals, and abiotic ones such as volcanic ash fall. Fossil pollen assemblages are interpreted using local modern pollen data. The main vegetation change occurred at ca. 3750 uncal b.p., when a shrub steppe of Asteraceae subf. Asteroideae with Schinus, Ephedra frustillata and a high proportion of grasses was replaced by a shrub steppe of Colliguaja integerrima and Asteraceae subf. Asteroideae. This change is synchronous with an archaeological record change and could be related either to moderate climatic variations or the effects of ash fall on the environment. Plant communities similar to the present-day ones were established in the Los Toldos area from ca. 3750 uncal b.p.

Early and mid Holocene local vegetational history, with special reference to woodland communities, was revealed by pollen analysis of a radiocarbon dated lake sediment profile from Lake Milkowskie (Jezioro Milkowskie) in northeastern Poland. The main factor controlling the dynamics of woodland composition changes until ca. 1950 b.c. was climate. After that, the role of human activity became increasingly important. The results of high-resolution pollen analyses provide evidence for early woodland disturbances caused by Mesolithic people at ca. 6950 b.c. Several episodes of human impact, differing in scale, and separated by subsequent episodes of woodland regeneration/stabilization were noted. The first traces of local crop farming, shown by the presence of Cerealia pollen, were recorded at ca. 3800 b.c. in the Paraneolithic/Neolithic period. Animal husbandry as well as cereal cultivation played only a marginal role in the economy, which was traditionally based on hunting, fishing and gathering through the Neolithic and the early Bronze Age. The change in economic strategies from foraging towards farming, starting around 3750 b.c., was a long-lasting process. An increase of productive economy took place in the middle Bronze Age at ca. 1400 b.c.

We use a data set of 35 surface pollen samples from lake sediments, moss polsters and top soils on the north- eastern Tibetan Plateau to explore the relationship between modern pollen assemblages and contemporary vegetation patterns. The surface pollen transect spanned four vegetation zones--alpine meadow, steppe, steppe desert and desert-- under different climatic/elevational conditions. Relative representation (R (rel)) values and Principal Components Analysis (PCA) were used to determine the relationships between modern pollen and vegetation and regional climate gradients. The results show that the main vegetation zones along the regional and elevational transects can be distinguished by their modern pollen spectra. Relative to Poaceae, a high representation of Artemisia, Nitraria and Chenopodiaceae was found, while Cyperaceae and Gentiana showed values in the middle range, and Ranunculaceae, Asteraceae, Ephedra and Fabaceae had low relative representation values. PCA results indicate a high correlation between the biogeoclimatic zones and annual precipitation and annual temperature and July temperature. The Artemisia/ Chenopodiaceae ratio and the Artemisia/Cyperaceae ratio are useful tools for qualitative and semi-quantitative palaeoenvironmental reconstruction on the north-eastern Tibetan Plateau. Surface lake sediments are found to have different palynomorph spectra from moss cushion and soil samples, reflecting the larger pollen source area in the contemporary vegetation for lakes.