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The restriction of the Mediterranean–Atlantic marine connection over the Messinian stage, which led to a salinity crisis and the deposition of the youngest salt giant on Earth, impacted the composition and structure of marine biota in unprecedented ways, but its effects on the biological functions of marine organisms remains unchartered territory. By analyzing the stable oxygen and carbon isotopic composition of the otoliths of two carefully selected fish species, a pelagic (surface) and a benthic (bottom-dwelling) one, we were able to infer not only the sea surface and bottom salinity and oxygenation conditions, but also the fishes’ metabolic response to the paleoceanographic changes in the eastern Mediterranean from 7.2 to 6.5 Ma. The high salinity and stratification of the Mediterranean water column during this interval hampered the ability of the marine fishes to grow, particularly those dwelling in the sea bottom. A first event (6.82–6.81 Ma) of high temperature and salinity induced an increase in the metabolism of fishes across the water column, meaning that they consumed and respired more. Subsequent warming, increased salinity and stratification at 6.70–6.69 Ma further intensified the environmental stress for bottom-water fishes. To cope with these extreme conditions at the sea floor, benthic fishes show increased metabolic carbon despite low food availability, which implies that fishes metabolized (i.e. consumed) their own tissue to survive. The disappearance of benthic fishes in the study area after 6.8 Ma further reinforces this conclusion. Our study promotes the importance of otoliths stable isotopic analyses as tools for reconstructing complex paleoenvironmental histories.

Molluscs, and among them bivalves, are organisms known for their ability to precisely record paleoenvironmental changes, both in shallow and deep marine settings. When looking into the recent geological past, bivalve assemblages offer information on the climatic changes that have impacted their taxonomic compositions. In the eastern Mediterranean, assemblages of bathyal bivalves are scarce. In order to investigate the impact of climatic changes on deep-water bivalve communities during the Early-Middle Pleistocene Transition, we focus here on two well-dated sections on Rhodes Island (Greece) corresponding to the Lindos Bay Formation. The sections of Lindos and Lardos present a continuous sedimentation of fine, marly sediments, and cover the Marine Isotopic Stages (MIS) 32 to 18. A total of 15 samples were analysed, resulting in the recovery of 31 species of bathyal bivalves. The depositional depths of these samples are estimated to be between 150 and 500 m. All samples are dominated by Protobranch bivalves, with the larger diversity found in families Nuculanidae and Yoldiidae. Three species, found only in cool intervals, are now extinct: Ledella nicotrae, Katadesmia confusa, and Pseudoneilonella pusio. Differences in sample composition are thought to be due mainly to climatic rather than bathymetric conditions. Although the associations in most MIS are similar to those found in the Italian Pleistocene deposits, those of the MIS 21 interglacial (Nucula nucleus – Saccella commutata – Cyclopecten hoskynsi – Limea crassa) and the MIS 20 glacial (Saccella commutata – Bathyspinula excisa –Yoldiella curta – Bathyarca spp.) are new for the Mediterranean region. These results imply that there were significant changes in bathyal bivalve associations during the climatic transitions of the Early-Middle Pleistocene and that modern bathyal associations of bivalves have been stabilized after the Middle Pleistocene.

Evaluation of the impact of climatic changes on the composition of fish assemblages requires quantitative measures that can be compared across space and time. In this respect, the mean temperature of the catch (MTC) approach has been proven to be a very useful tool for monitoring the effect of climate change on fisheries catch. Lack of baseline data and deep-time analogues, however, prevent a more comprehensive evaluation. In this study, we explore the applicability of the mean temperature approach to fossil fish faunas by using otolith assemblage data from the eastern Mediterranean and the northern Adriatic coastal environments corresponding to the last 8000 years (Holocene) and the interval 2.58–1.80 Ma B. P. (Early Pleistocene). The calculated mean temperatures of the otolith assemblage (MTO) range from 13.5 to 17.3 °C. This case study shows that the MTO can successfully capture compositional shifts in marine fish faunas based on variations in their climatic affinity driven by regional climate differences. However, the index is sensitive to methodological choices and thus requires standardized sampling. Even though theoretical and methodological issues prevent direct comparisons between MTO and MTC values, the MTO offers a useful quantitative proxy for reconstructing spatial and temporal trends in the biogeographic affinity of fossil otolith assemblages.

Stable oxygen and carbon isotopic composition of the otoliths of a pelagic and a benthic fish species are used as indicators of sea surface and bottom salinity-oxygenation conditions, as well as proxies of the fishes’ metabolic response to the paleoceanographic changes in the eastern Mediterranean from 7.2 to 6.5 Ma. The step-wise restriction of the Mediterranean–Atlantic gateway impacted the ability of the Mediterranean fishes to grow, particularly those dwelling in the sea bottom. The most important changes identified are: 1) a negative water budget (evaporative) event at 6.82–6.81 Ma that resulted in an increase in fish metabolism and respiration, and 2) a warming event at 6.70–6.69 Ma that further intensified the environmental stress for bottom-water fish and increased their metabolic activity despite low food availability. The results provide proof that the resilience of the benthic fish population was hampered, particularly after the 6.70–6.69 Ma event, due to the increased salinity, temperature and stratification of the water column in the eastern Mediterranean as a result of the marine gateway restriction.

9 pages, 4 figures.-- Data accessibility: The raw dataset produced in this study is available at https://doi.org/10.5061/dryad.gxd2547pn [84]; the R script at https://doi.org/10.5281/zenodo.6643635 [85]; and electronic supplementary material can be accessed at https://doi.org/10.5281/zenodo.6643637 [60], Mesopelagic fishes are an important element of marine food webs, a huge, still mostly untapped food resource and great contributors to the biological carbon pump, whose future under climate change scenarios is unknown. The shrinking of commercial fishes within decades has been an alarming observation, but its causes remain contended. Here, we investigate the effect of warming climate on mesopelagic fish size in the eastern Mediterranean Sea during a glacial–interglacial–glacial transition of the Middle Pleistocene (marine isotope stages 20–18; 814–712 kyr B.P.), which included a 4°C increase in global seawater temperature. Our results based on fossil otoliths show that the median size of lanternfishes, one of the most abundant groups of mesopelagic fishes in fossil and modern assemblages, declined by approximately 35% with climate warming at the community level. However, individual mesopelagic species showed different and often opposing trends in size across the studied time interval, suggesting that climate warming in the interglacial resulted in an ecological shift toward increased relative abundance of smaller sized mesopelagic fishes due to geographical and/or bathymetric distribution range shifts, and the size-dependent effects of warming, This work was funded by the Austrian Science Fund (FWF) project M2894-N ‘Deep-time climate change impact on marine food webs’ (P.I.: K.A.). It was also supported by the National programmes Tellus-INTERRVIE and Tellus-SYSTER of CNRS-INSU (F.Q.). M.C. acknowledges the ‘Severo Ochoa Centre of Excellence’ accreditation (grant no. CEX2019-000928-S) to the Institute of Marine Science (ICM-CSIC) and partial funding from the European Union's Horizon 2020 research and innovation programme under grant agreement no. 869300 (FutureMARES project)

Fossil records of nearshore, shallow marine fish communities are rare. Here, we present the rich and diverse fish fauna of a coastal setting in the eastern part of the Mediterranean during the early Pliocene, which comprises 54 taxa, 77% of which are extant and currently occupy the same shores. We analyse these assemblages to estimate the palaeodepth, the substratum and the climatic and oceanographic conditions prevailing in the region at the time. Furthermore, we review the stratigraphic and geographic distribution of the identified taxa from the Tortonian until today, to establish patterns and trends in the evolution of the Mediterranean coastal fish fauna. Contrary to expectations, the Pliocene coastal fauna is very similar to the Miocene and to the Pleistocene in terms of functional traits as well as taxonomically. Replacements of species seem to have been gradual, through multiple extirpations and reintroductions that led to the final extinction of mostly tropical species from the basin, while subtropical–temperate taxa invaded to take their place.

Aspects of paleoclimatic and paleoceanographic evolution of the north Aegean Sea through the Holocene are revealed by the study of quantitative variations in planktonic foraminiferal, pteropodal, and palynomorph assemblages; the isotopic composition of planktonic foraminifera; and hydrographic-related indices, extracted from two high-sedimentation rate cores from the North Aegean Trough. Focusing on the last ~10 cal ka BP, the current Holocene subdivision (Greenlandian, Northgrippian, and Meghalayan) confirms the traditional understanding of an evolution from wetter (Greenlandian) to gradually drier (Northgrippian and Meghalayan) climatic conditions and further highlights the role of changing seasonality during this time. The most warm and humid phase corresponds to the time of the sapropel S1 deposition (9.6–6.1 cal ka BP). The Holocene climatic instability of the study area is further supported by six episodes of brief cooling (North Aegean cooling; NAEGC6–NAEGC1) centered at 9.30, 8.05, 7.05, 4.55, 3.55, and 2.05 cal ka BP, reflected by significant faunal changes and oxygen isotope enrichments. These cold/arid events are coeval with equivalent cooling events that have been described in different basins of the Mediterranean Sea, while signal similarities with equivalent changes in the intensity of the Siberian high suggest a climatic link between the studied area and the high-latitude areas.

Tindaria kretensis n. sp., a minute deep-water protobranch, is described from Early Pliocene marls cropping out in the Voutes section, northern Crete Island. This is one of the few fossil records of Tindaria from the Mediterranean area, the first well documented for the Neogene. Tindaria is an almost cosmopolitan deep-water genus, absent from the modern Mediterranean. Its occurrence in the Early Pliocene is in agreement with what is known about the past psychrospheric conditions in the Mediterranean.

Conidae is a very diverse family of carnivorous gastropods with over 900 extant species, that present beautiful colour patterns. These patterns can be useful to biologists for the systematic approach, but lack usually in fossil samples, when they are only observed in natural light. Therefore conid species were difficult to distinguish.To resolve that problem, we use ultraviolet light to reveal the colour patterns of these fossil shells. Here, we present the second part of a PhD project that attempts to clarify the diversity of the Tortonian Conidae from Crete, Greece. The first part of this project already revealed eleven species of Conilithes Swainson, 1840 and Conus (Kalloconus) da Motta, 1991 five of them being new. In the second part, we found seventeen species, of those seven are considered as new. Among them, ten are attributed to Conus (Lautoconus) Monterosato, 1923 and three in Conus (Plagioconus) Tucker & Tenorio, 2009. Finally, a group of four species with special characteristics is discussed and is proposed as a new subgenus.This project will greatly enrich the knowledge of the family in the Tortonian of Eastern Proto-Mediterranean. The results will be compared to similar faunas from the Middle-Miocene to Pliocene of Europe, in order to assess the biogeography of this family.

Reconstructing paleoceanographic conditions for the entire water column remains challenging, due to the lack of proxies for seawater parameters below surface waters, which could be used to validate models. Fish otoliths and ostracods are used here to obtain biogeochemical proxy data of sea surface/bottom temperature and productivity, as well as the biological response of marine fishes to paleoenvironmental change. Our study area is located in the eastern Mediterranean Sea (Heraklion basin, Crete Island). During the Messinian age (specifically between 7.2 and 6.5 Ma), the Mediterranean–Atlantic connection was restricted leading to a strongly stratified water column. We study the sea surface and bottom conditions under these conditions.Stable oxygen isotopes on ostracod valves (Bairdoppilata sp.) reflect the combined effect of bottom-water temperature and salinity changes. For fishes however, this depends on each species lifestyle. We analyzed two very common species: 1) Bregmaceros albyi, a surface-water pelagic species, and 2) Lesueurigobius friesii, a demersal fish dwelling on the sea bottom. Our hypothesis was that the stable oxygen isotopic ratios on B. albyi otoliths would reflect surface-water conditions, whereas those on L. friesii would correspond to bottom-water conditions. Furthermore, we obtained δ13C values for the same ostracod and otolith specimens. Stable carbon isotopic ratios in invertebrate shells indicate biological productivity, since carbon fractionation takes place as a single-step process during biomineralization. However, fish otolith aragonite mineralization is more complicated, involving more than one fractionation steps, and carbon is obtained from seawater, but also from diet. Therefore, otolith δ13C is considered a proxy of the fish’s field metabolic rate, reflecting its ability to continue to grow and reproduce despite environmental change.Our results show that B. albyi δ18O values correlate well with those derived from planktonic foraminifera shells, whereas L. friesii δ18O is in agreement with ostracod values, thereby confirming our hypothesis. Moreover, ostracod and foraminifera δ13C follow the same decreasing pattern. However, otolith δ13C remains stable, even after 6.8 Ma, when high-amplitude salinity variations prevail. This suggests that fish maintained their capacity to grow and reproduce despite significant changes in seawater conditions at least until 6.5 Ma. However, whether this reflects their resilience to these environmental changes or an adaptation mechanism such as reducing their growth rate or shifting their trophic preferences remains a mystery.

During the Late Miocene the Mediterranean Sea experienced severe disruption of its connectivity to the Atlantic Ocean, highlighted by a rapid sea-level drop, culminating to the Messinian Salinity Crisis (MSC; 5.97-5.33 Ma). Such a paleoceanographic change, triggered by the cumulative effect of climate and tectonics, caused high-amplitude fluctuations in the hydrology of the entire basin, and further influenced the geological history of the Mediterranean Sea. Although a consistent pattern of the paleoclimate has started to emerge, we currently lack a continuous sea surface salinity (SSS) record linking the timing of sea surface temperature (SST) variations, sea-level fluctuations, and the overall environmental change, particularly for the pre-evaporitic period. Initial viewpoints of a linear and gradual salinity increase prior to the onset of the MSC have been recently revised and replaced by highly variable salinity-related patterns representative of the stepwise restriction of the Mediterranean Sea. Here we use the combined Tetra Ether (TEX86-) and/or alkenone unsaturation ratio (UK′37) based SSTs and oxygen isotopes (δ18O) from the cyclic marl-sapropel sedimentary succession of Agios Myron section (north-central Crete, Greece) to assess hydroclimate changes during that time, and we finally present the first record of SSS in the eastern Mediterranean Sea for the earliest Messinian (7.2–6.5 Ma). The relatively stable marine conditions after the Tortonian/Messinian boundary, expressed through a cool and fresh upper water column, significantly changed at ∼6.9 Ma, when an important reversal in the heart of the Messinian cooling trend supplemented by a coherent hypersaline water column took place. The observed SST and SSS patterns provide context for a two-fold evolution of this event (centered at 6.9–6.8 and 6.72 Ma), which finally led to the onset of a brine pool into the eastern Mediterranean basin. The transitional character of the following time interval (6.7–6.5 Ma) marks another important step in the basin restriction with a wider range of salinity fluctuations from highly saline to diluted conditions and enhanced water column stratification prior to the deposition of evaporites. Overall, this evolution supports the concept of a stepwise restriction of the Mediterranean Sea associated with substantial hydroclimate variability and increasing environmental (thermal and salinity) stress, and further confirms its position as a preferred laboratory for developing new conceptual models in paleoceanography, allowing the investigation and scale assessment of a phenomenon with high chances of representing a future analogue scenario.

Marine biodiversity is controlled by depth, climate, and ocean connectivity. The early Miocene marine fauna of the proto-Mediterranean is particularly important because this is the time when the connection with the Indo-Pacific realm was disrupted , and the marine biodiversity hotspot shifted toward the southeast, marking the onset of a subtropical gradually oligotrophic regime in the proto-Mediterranean Sea. The Mesohellenic basin in particular, at the northern part of the proto-Mediterranean, was located at the intersection with the Paratethys epicontinental sea. Therefore, the fauna in this area was influenced by the conditions in both basins. In this study, we present the mollusc and fish fauna of the Aquitanian Mesohellenic basin and discuss its paleobiogeographic and evolutionary implications. Comparisons and assessment of faunal similarities of the proto-Mediterranean and adjacent provinces is carried out using ordination methods to treat available data from relevant publications and the Paleobiology Database (PBDB).This research has been co-financed by Greece and the European Union (European Social Fund- ESF) through the Operational Programme “Human Resources Development, Education and Lifelong Learning 2014–2020” in the context of the project “Mollusc and fish migrations in the dynamic environments of the early to middle Miocene in the Mediterranean” (MIS 5047960).

Degradation of coastal environments is an issue that many areas in Europe are facing. In the present work, an ancient coastal lake wetland is investigated, the so-called Lake Lerna in NE Peloponnese, Greece. The area hosted early agricultural populations of modern Greece that started modifying their environment as early as the early–middle Neolithic. Two drill cores in the area of the ancient lake were analysed to establish the sedimentological succession and the depositional environments using sub-fossil assemblages (molluscs and ostracods). Three lithological and faunal units were recovered, the latter being confirmed by the statistical ordination method (non-metric multidimensional scaling). The usage of sub-fossil mollusc species for the first time in the region enriched the dataset and contributed significantly to the delimitation of the faunas. These consist of environments characterised by various levels of humidity (from stagnant waters to freshwater lake) and salinity, with ephemeral intrusions of salt water to the lake, documented by mollusc and ostracod populations. We conclude that the lake and its included fauna and flora were mostly affected by climatic fluctuations rather than human intervention in the area.

The Messinian Salinity Crisis (MSC) was an environmental perturbation with dramatic environmental consequences that greatly affected marine organisms. Messinian deposits are found in several locations around the Mediterranean, but few offer marine faunas rich in molluscs. A section near Heraklion, central Crete, has provided new material that contains a well preserved and rich molluscan fauna that includes many micromorphic species. The section is of early Messinian age, belongs to Agios Miron Formation, and bears several layers of fossiliferous marly sands.Molluscs from a fossiliferous bed of the section are presented here for the first time. Gastropods and bivalves are most common, but scaphopods and chitons are not infrequent. The assemblage seems to be composed of transported elements from nearby environments and the most frequent species are present in comparable abundances for gastropods and bivalves. The gastropod fauna is represented by Bittium sp. and Gibbula sp., accompanied by Diodora cf. graeca, Turritella sp., Jujubinus sp., species of Pyramidellidae and rarer Homalopoma sp. and Haliotis sp. The presence of Bittium sp. together with Jujubinus sp. suggests vegetated environments. Bivalves are represented by species dwelling mostly in sandy environments such as Glycymeris cf. inflata (also occurring in larger specimens), Spisula sp., Timoclea sp. and various cardiids. Exceptionally well-preserved chitons indicate the presence of hard substrates such as rocks, pebbles or roots of seagrass beds. This is confirmed by the presence of the gastropods Diodora cf. graeca and Haliotis sp.The assemblage points towards normal salinity shallow marine conditions of sandy bottoms with patches of seagrass-type vegetation before the onset of the MSC.

Marine environmental status can be assessed through the study of bio-indicator species. Here, we monitor natural environmental stress by the occurrence of morphologically abnormal planktonic foraminiferal specimens from a suite of surface sediments in the eastern Mediterranean Sea. We also compare Scanning Electron Microscopy (SEM) abnormality observations from sapropel S1-derived sediments in the Aegean, Libyan and Levantine basins, since they provide a direct record of a natural stress experiment that took place over past time scales. At initial sapropel deposition levels, we observe increased growth asymmetry in Globigerinoides ruber twinned and twisted individuals, possibly associated with eutrophication and anoxia. In modern material, a range of malformations and aberrant morphologies from slight deformity with smaller or overdeveloped chambers to more severe deformity with abnormally protruding or misplaced chambers, distorted spirals, and double tests is also observed, as a result of the hypersaline, oligotrophic and oxygen-depleted nature of the Mediterranean Sea water column. Overall, we highlight the current use of the relative abundance of abnormal tests as a bio-indicator for monitoring natural stress, especially the occurrence of twin specimens as indicative of high-salinity stress conditions, and further illustrate the necessity to map both their spatial and temporal distribution for accurate paleoenvironmental reconstructions. Such an approach presents the advantage to rapidly provide information over wide spatial and temporal scales, extending our ability to monitor a wide variety of environments (from coastal to the open-sea). However, further investigations should extend this approach to test the robustness of our findings in a number of similar oceanic settings.

Diversified deep-water corals and associated communities occur in early Pleistocene (early Calabrian; ∼1.7–1.6 Ma) calcareous crusts discovered on the south-western coast of Karpathos Island (eastern Greece). Apart from abundant and conspicuous solitary and colonial corals (12 species), the accompanying fauna comprises mostly bivalves (10 species), serpulid worms (4 species), and bryozoans (40 species). The growth and deposition of the studied organisms occurred in an upper bathyal environment at water depths around 400–600 m. The calculated mean uplift rate after deposition of the crusts indicates that Karpathos was drowned during the earliest Calabrian and experienced severe uplifts by the late Calabrian. A comparison between these communities and those of the Pleistocene to Recent Mediterranean and north-east Atlantic shows that a common stock of taxa exist(ed); among them two azooxanthellate colonial scleractinians (Lophelia pertusa and Madrepora oculata) are predominant.

We here report on the systematics, paleoecology and biostratigraphy of a rare, but diversified gastropod fauna found in the early Pliocene sediments of the Agia Triada section in the Falanthi Basin (Peloponnese, Greece). Sixty-one species belonging to forty-eight genera are identified, allowing biostratigraphic assignment to the Mediterranean Plio-Pleistocene Molluscan Unit 1 (MPPMU1). Planktonic foraminiferal biostratigraphic analyses indicate that the Agia Triada section correlates with Zones MPL2 and MPL3, corresponding to the latest Zanclean. A comparison is further carried out between the studied assemblages and those previously documented from the Atlantic coasts of Europe and the Mediterranean region. New information on the palaeobiogeography of early Pliocene gastropod fauna is given, as seven species from the Mediterranean, thirty-nine species from the eastern Mediterranean and forty-four species from Greece, are documented here for the first time.

Alonissos (Iliodroma) Island is really famous to palaeobotanists because of the type locality of Glyptostrobus europaeus (Brongniart) Unger only. Since the first report of the latter holotype no palaeobotanical work has taken place in Alonissos. This is the first research of the fossiliferous locality, along with new palaeobotanical, palynological and gastropod findings, revealed after 200 years. Among the collected material, a xylitic specimen was studied and identified as Pinuxylon alonissianum Mantzouka & Sakala, sp. nov. The presence of this species points to botanical affinities with sections of Diploxylon Pines (Pinus leiophylla var. chihuahuana (Engelm.) Shaw, P. arizonica Engelm., P. arizonica var. cooperi (C.E.Blanco) Farjon, P. engelmannii Carriere and P. jeffreyi Balf.) especially from Central America (Sierra Madre Occidental and Oriental of Mexico, Arizona, California, Oregon and Sierra Nevada). This is the first research of this kind revealing as the Nearest Relative bioprovince of the studied Greek fossil pine wood Central America and not Himalayas or Asia. Moreover, a palynological study and an analysis of fossil molluscs were carried out for the first time, giving more data regarding the palaeovegetation and palaeoclimate of the area. Finally, the holotype of Glyptostrobus europaeus (Brongniart) Unger, housed in the collections of the Museum national d'Histoire naturelle, Paris (MNHN), is found, photographed and presented for the first time.

The Pliocene Eastern Mediterranean fish record is revealed through the study of a 60-m thick stratigraphic sequence near the village Voutes (Heraklion, Crete). Forty-two species belonging to twenty families are identified. Calcareous nannoplankton biostratigraphy places the studied sequence within the biozone MNN16a (latest Zanclean). The stratigraphic distribution of 31 species is modified. Among these, 12 species are reported for the first time in the Eastern Mediterranean Zanclean, while 19 species are first reported outside the Ionian Sea. The Voutes fish fauna presents a diversified benthic and benthopelagic assemblage filling a significant gap in the fossil record.

International audience; The marine Messinian deposits of Tunisia cover a narrow littoral strip some 300 km long between the northern Bizerte and Cap Bon areas and the central-eastern Sahel region. Litho- and biofacies analysis of six stratigraphic sections reveals the distinctive features of these deposits.;The lower Messinian deposits are characterized by ubiquitous siliciclastics and abundant oolitic/bioclastic limestones organized in an eastward facing ramp. Westward (landward), this ramp changes into coastal lagoons, sometimes containing evaporites. Eastward, the ramp passes to the reefal Pelagian Platform extending as far as Lampedusa.;Two main sedimentary cycles are distinguished: 1) an early Messinian siliciclastic retrogradational then oolitic/bioclastic progradational cycle (Beni Khiar Formation and lower Oued bel Khedim Formation): 2) a late Messinian brackish to continental cycle that probably accumulated in rapidly subsiding lagoons (Cued el Bir Formation and upper Oued bel Khedim Formation). The Tunisian early Messinian cycle is partly eustatically controlled, but the late Messinian cycle cannot be confidently correlated to other well-known Messinian series because of tectonic movements.;The lower Messinian deposits of Tunisia are also characterized by abundant suspension-feeding organisms (molluscs and bryozoans) and rare corals, calcareous algae, echinoids, and larger benthic foraminifers. The proposed palaeoenvironmental model shows that the lower Messinian ramp of Tunisia was located on a current-protected margin and subjected to continent-derived sediment and nutrient supply. Eastward, nutrient influx diminished and a shallow-water isolated carbonate platform with coralgal facies developed between the western and the eastern Mediterranean basins. The main hydrological connection between these two basins occurred through a narrow seaway situated to the northeast of the Pelagian Platform, south of Sicily and Malta.

Changes in benthic foraminiferal and mollusk assemblages from the lower part of the Upper Pliocene of the Atsipades Section (Iraklion Basin, central Crete) were studied. The Atsipades Section represents a shallowing-upward sequence from outer shelf blue–gray clays at the bottom of the sequence, deposited below the storm wave base, to shallow inner shelf deposits affected by storm waves at the top. The foraminiferal assemblage at the bottom of the sequence is dominated by Bolivina spathulata , Bolivina dilatata and Uvigerinidae, a microfossil assemblage corresponding to the deepest deposits formed under dysoxic sea-floor conditions. Foraminiferal assemblages of the middle part of the section are highly diversified, predominantly Haynesina depressula , Cassidulina carinata and Reusella spinulosa . The top of the section is mainly characterised by Asterigerinata planorbis , Bolivina pseudoplicata , Cibicides lobatulus and Elphidium sp., a typically epiphytic foraminiferal assemblage which can be correlated with the presence of an algal covered sea-bottom. Within this general environmental trend, a minor shallowing cycle can be differentiated. The boundaries of this cycle can be inferred, based on a substantial microfossil assemblage change and on the coincidence of species diversity maximum and a planktonic/benthic (P/P + B) ratio peak. Nonetheless, upwelling currents and/or over-abundance of nutrients due to continental outflow could also contribute to increased diversity and P/P + B ratio. The character of the mollusk assemblages is in accordance with these trends. Moreover, the increase in diversity and in sculpture constitutes a clear indication of an increase in hydrodynamic energy related to a shallowing-upward trend.

Στην εργασία αυτή μελετήθηκε η πλούσια και καλά διατηρημένη πανίδα μαλακίων που περιέχεται σε αμμώδες στρώμα ρηχών θαλάσσιων αποθέσεων του Κατωτέρου Πλειστοκαίνου της νήσου Ρόδου. Πιστοποιήθηκαν 4 διαφορετικές συγκεντρώσεις μαλακίων γειτονικών περιβαλλόντων καθώς και οι οικολογικές τους φωλεές οι οποίες καθορίζονται από την πλευρική μεταβολή του τύπου του υποστρώματος. Αναγνωρίστηκαν αντιπρόσωποι μαλακίων σκληρού υποστρώματος έως και αμμώδους βυθού της παράκτιας ζώνης. Προσδιορίστηκαν 55 είδη μαλακίων εκ των οποίων 29 είδη γαστερόποδων. Πιστοποιήθηκε η παρουσία της Ervilia castanea και της αντιπροσωπευτικής φάσης της στο Κατώτερο Πλειστόκαινο της Ελλάδας για πρώτη φορά., Highly diverse and well preserved marine mollusc associations from a sandy horizon of the Lower Pleistocene shallow water deposits of Rhodes island are here presented. Four significantly different molluscan assemblages characterize neighboring environments and document a variety of niches provided by the lateral change of bottom type. Molluscs of hard substrate to sandy seabed of the Infralittoral Zone were documented. A total of 55 molluscan species were identified. Gastropods dominate with 29 species. For the first time, the Ervilia castanea assemblage is described for the Lower Pleistocene of Greece.

Στην παρούσα εργασία παρουσιάζεται η παλαιοβαθυμετρική ανάλυση των συναθροίσεων ωτολίθων Ιχθύων, που προσδιορίστηκαν στα ιζήματα ηλικίας Ζαγκλίου στην τομή Βούτες (Ηράκλειο, Κρήτη). Η περιοχή Βούτες αποτελούσε μία ακτή με νηριτική Ιχθυοπανίδα, με υψηλή συμμετοχή μεσοπελαγικών ειδών, κυρίων Myctophidae. Τα βενθικά και βενθοπελαγικά ψάρια εμφανίζουν επίσης μεγάλη ποικιλότητα. Κατά το ανώτερο Ζάγκλιο, η περιοχή μελέτης αντιστοιχούσε σε βαθύ νηριτικό περιβάλλον. Σταδιακά η περιοχή ανυψώθηκε σε πολύ μικρά βάθη, όπως φαίνεται από την αυξημένη συμμετοχή στην πανίδα των Gobidae, αλλά και την παρουσία Bothidae. Στη συνέχεια, η περιοχή βάθυνε ξανά, καθώς παρατηρείται σημαντική μείωση της συμμετοχής των ρηχών βενθικών-βενθοπελαγικών ειδών. Το συμπέρασμα αυτό ενισχύεται από την παρουσία στο επίπεδο αυτό των Πτεροπόδων, αλλά και την απόθεση διατομιτών. Σε ανώτερο τμήμα της τομής, το βάθος της θάλασσας μειώνεται και πάλι, πριν τον ανώτερο διατομιτικό ορίζοντα, ο οποίος ωστόσο εμφανίζει ιδιαίτερα φτωχή συνάθροιση ωτολίθων., The fish otolith assemblages identified in the Zanclean sediments of Voutes section (Heraklion, Crete) are analyzed in order to estimate the depth of deposition. The assemblages indicate that Voutes area was in fact a coast inhabited by a neritic fish fauna with significant mesopelagic component, mostly Myctophids. The benthic and benthopelagic group exhibits great diversity. In the late Zanclean, the study area corresponds to a deep neritic environment, which gradually uplifts. At the end of the Zanclean, Voutes area becomes a very shallow coast, as shown by the increased contribution to the assemblage by members of the Gobiid family, as well as the notable presence of Bothidae. Thereafter, the area deepens again, as the shallow benthic-benthopelagic component significantly decreases. This is consistent with the appearance of Pteropods and reaches a peak during the deposition of diatomites at maximum depth. At the upper part of the section, the sea bottom depth decreases again to allow for the deposition of marls with various molluscs remains, where Gadids and Gobiids again reappear. Finally, the upper diatomitic horizon is characterized by a rather poor fish fauna. These observations allow reconstructing the evolution of Voutes coastal area during the latest Zanclean until the Piacenzian.

To μeγάλο ποσοστό eξαφάνισης των Μαλακίων του Πλeιόκαινου, το οποίο ξeπeρνά το 30%, δίνeι τη δυνατότητα της χρήσης τους για την αναγνώριση κλιματικών-ωκeανογραφικών και χρονολογικών στοιχeίων. Στην παρούσα eργασία, η δυνατότητα χρήσης του μοντέλου των Raffi & Monegatti (1993) στην πeριοχή της ανατολικής Μeσογeίου, αποδeικνύeται από τη μeλέτη δύο παραδeιγμάτων από τον Eλλαδικό χώρο και την Κύπρο. Το μοντέλο αυτό αφορά στα βιοοτρωματογραφικά γeγονότα eξαφάνισης των Μαλακίων του Πλeιόκαινου, διακρίνοντας 4 eνότητeς Μαλακίων διαφορeτικής κλιματικής-ωκeανογραφικής σημασίας. Δύο πeριοχές που έχουν καλά μeλeτηθeί από αυτή την άποψη, η τομή Ατσιπάδeς Ηρακλeίου Κρήτης και η τομή Αθηένου Κύπρου, δίνουν στοιχeία για το συσχeτισμό των αποθέσeων τους μe το μοντέλο αυτό. Η τομή Ατσιπάδeς συσχeτίζeται μe το πρώτο βιοστρωματογραφικό γeγονός eξαφάνισης των taxa των βeνθονικών Μαλακίων των eνοτήτων των Raffi & Monegatti το οποίο μπορeί να θeωρηθeί ως αποτέλeσμα του πρώτου γeγονότος ψύχρανσης κατά το Πλeιόκαινο. Η τομή Αθηένου μπορeί να συσχeτιστeί μe την MPMU2 eνότητα των Raffi & Monegatti, μeτά το γeγονός της πρώτης eξαφάνισης και πριν το γeγονός της δeύτeρης eξαφάνισης των Μαλακίων του Πλeιόκαινου.