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Augyles (Augyles) hispidulus (Kiesenwetter, 1843) Material examined. 250 exs. RUSSIA. Astrakhan Oblast: Enotaevsky Distr., Zamyany vill., at light, 16.09.2007, leg. O.G. Brekhov, 30 exs. (IBIW). Ivanovo Oblast: Pestryakovsky Distr., Purezhka River, 56°28'30.3"N 42°38'4.2"E, at light, 24– 25.06.2011, leg. D.V. Potanin, 2 exs. (IBIW). Kaliningrad Oblast: Nemansky Distr., 3 km E of Bolshoe Selo vill., Neman River, 55°03'07.3"N 22°09'02.1"E, 12.06.2019, leg. A.A. Przhiboro, 12 exs. (ZISP). Kirov Oblast: Kilzemsky Distr., Tautovo vill., Kilmez River, 7.06.2005, leg. E.A. Korobeynikova, 1 ex. (IBIW). Leningrad Oblast: Kingiseppsky Distr., Kingisepp (Yamburg), 6.07.1904, leg. Barovsky, 2 exs. (ZISP); Gatchinsky Distr., env. Malkovoe vill., Suyda River, 16.07.2018, leg. A.G. Koval, 3 exs. (VIZR). Lipetsk Oblast: Eletsky Distr., 30 km E from Elets, Don River left bank, on sand, 26.07.2004 leg. A.O. Bieńkowski, 1 ex. (PCB). Republic of Mari El: Tsaryovokokshaysky Uyezd, 12.07.1913, unknown collector, 1 ex. (ZISP). Moscow Oblast: Serpukhovsky Distr., 3 km W of Luzhi vill., 18.07.2011, leg. A.V. Matveev, 1 ex. (ZISP). Nizhniy Novgorod Oblast: Dzerzhinsky Distr., Gavrilovka vill. 6.07.2013, leg. D.V. Potanin, 8 exs. (IBIW). Perm Krai: Perm, Balmoshnaya River, soil traps, 17.09.– 5.10.2017, leg. V.O. Kozminykh, 1 ex. (IBIW). Rostov Oblast: Sholokhovsky Distr., Kruzhilinsky vill., at light, 26– 30.06.2003, leg. O.V. Brekhov, 2 exs. (IBIW); Sholokhovsky Distr., Vyoshinskaya vill., at light, 3– 4.07.2003, leg. O.V. Brekhov, 2 exs. (IBIW). Ryazan Oblast: Spassky Distr., Brykin Bor vill., bank of Pra River, 54°42'48.1"N 40°51'10.8"E, on sand, 28.06.2017, leg. A.S. Sazhnev, 2 exs. (IBIW). Saratov Oblast: Lysogorsky Distr., 3 km NE Semyonovka vill., bank of Medveditsa River, on sand, 25– 28.06.2012, leg. A.S. Sazhnev, 13 exs. (IBIW); Voskresensky Distr., Sinodskoe vill., Tereshka River, 51°58'06.4"N 46°39'44.5"E, 15.07.2012, leg. N.V. Rodnev, 3 exs. (IBIW); Balashovsky Distr., Repnoe vill., pools, 29.05.2017, leg. A.N. Volodchenko, 1 ex. (IBIW). Republic of Tatarstan: Kazan, Gavrilova str., Kazanka River, on sand, 13.07.2010, leg. D.A. Klyomin, 1 ex. (IBIW). Vladimir Oblast: Vozhegodsky Distr., Grudino (Zarechie) vill., sandy bank of Vozhega River, 28.07.2016, leg. D. Vtorushin, 81 exs. (IBIW), same place, 31.07.2016, leg. D. Vtorushin, 55 exs. (IBIW). Volgograd Oblast: Uryupinsky Distr., 5 km W pf Mikhaylovskaya vill., bank of Medveditsa River, 21.06.2011, leg. A.V. Matveev, 1 ex (ZISP). Ilovlinsky Distr., Baybaev vill., at light, 23.05.2002, leg. O.G. Brekhov, 1 ex. (PCBr). Voronezh Oblast: Novousmansky Distr., 2 km W Maklok vill., bank of Usmanka River (env. Chistoe Lake), sand, 12.07.2016, leg. A.S. Sazhnev, 10 exs. (IBIW). Yaroslavl Oblast: Yaroslavsky Distr., Krasnye Tkachi settlement, bank of Kotorosl River, 5.06.1993, leg. D.V. Vlasov, 3 exs. (YMR); Nekouzsky Distr., Borok settlement, Radovsky Island, sandy bank of Rybinskoe Reservoir, 14.06.2016, leg. A.S. Sazhnev, 13 exs. (IBIW). Addition material. 10 exs. KAZAKHSTAN. Aktyubinsk Oblast: [Uralsk Oblast: Temirsky Ueazd], Chatarly River, 6.06.1908, leg. B. Uvarov & D. Borodin, 2 ex. (ZISP); same place, necropolis Daumshar, 9.06.1908, leg. B. Uvarov & D. Borodin, 3 exs. (ZISP); same place, Kokdzhida sands, 10.06.1908, leg. B. Uvarov & D. Borodin, 1 ex. (ZISP); same place, 17.06.1908, leg. B. Uvarov & D. Borodin, 4 ex. (ZISP). Distribution in European Russia. NT: ARH SPB VLG CT: IVA KIR KOS KÖN MOS NNO* RZN TUL VLD YAR VL: CHV MAR MRD SAM TAT ST: AST LIP ROS SAR VOL* VRN UR: ORB PER (Lindemann 1871; Zaitzev 1907; Pomerantsev 1908; Jacobson 1913; Lebedev 1925; Gussakovsky 1926; Semyonov, 2009; Tsurikov 2009; Alekseev & Shapoval 2019; Litovkin 2012; Litovkin et al., 2013; Dedyukhin & Kholmogorova, 2014; Sazhnev 2014, 2016c, 2018c; Egorov 2019; Sazhnev & Kozminykh 2019; Sazhnev et al., 2018; Egorov et al., 2020) Distribution. The species was described from Austria, distributed in Europe, Near East (Israel, Iraq), Asian Turkey, and Middle Asia (Turkmenistan, Uzbekistan) (Mascagni 2016). Records from the south Siberia and the Russian Far East (Kirejtshuk 2001) should be attributed to other species of the Augyles cribratellus species group sensu Charpentier (1965) (Sazhnev 2018c). The record of this species from Issyk-Kul hollow in Kyrgyzstan (Ovchinnikov 1996) require verification (Litovkin & Sazhnev 2018). Remarks. The species recorded for two subjects of Russia (Nizhniy Novgorod and Volgograd Oblasts) for the first time., Published as part of Sazhnev, Alexey S., 2022, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region, pp. 581-593 in Zootaxa 5209 (5) on pages 584-585, DOI: 10.11646/zootaxa.5209.5.5, http://zenodo.org/record/7340995, {"references":["Kiesenwetter, E. von. (1843) Beitrage zur Monographie der Gattung Heterocerus. Zeitschrift fur die Entomologie, 4, 194 - 224.","Lindemann, K. (1871) A review of geographic distribution of beetles in Russian Empire. Part 1. Introduction, Fore word. Northern, Moscowian and Turanian Oblasts. Proceedings of Russian Entomological Society, 6, 41 - 366. [in Russian]","Pomerantsev, D. (1908) List of the beetles around Vel'sk City and other places of the Vologda Oblast. Trudy Russkogo Entomoloshicheskogo Obschestva, 38 (4), 421 - 506. [in Russian]","Jacobson, G. G. (1913) Semeystvo Heteroceridae. Pilousy. In: Zhuki Rossii i zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov. Vol. 10. A. F. Davrien, Sankt-Peterburg, pp. 867 - 869. [in Russian, English translation in: Family Heteroceridae. Mud-loving beetles. Beetles of Russia, Western Europe and neighbouring countries. Guide to the determination of beetles. A. F. Devrien, St. - Petersburg, 10, pp. 867 - 869.]","Lebedev, A. G. (1925) Materials for the fauna of beetles of the Republic Tatarstan. III. Entomological Review, 19, 133 - 138. [in Russian]","Gussakovsky, V. V. (1926) List of beetles (Coleoptera) vicinity of Kostroma city. Biological collection (Proceedings of the Kostroma Scientific Society for the Study of the Local Territory, 37, 21 - 75. [in Russian]","Semyonov, V. B. (2009) An annotated list of coleopteran insects (Insecta, Coleoptera) of Central Meschera. KMK, Moscow, 167 pp. [in Russian]","Tsurikov, M. N. (2009) Beetles of Lipetsk Oblast. Voronezh State University, Voronezh, 332.","Alekseev, V. I. & Shapoval, A. P. (2019) Beetles (Insecta, Coleoptera), sampled with use of light trap on the Curonian Spit: the materials of the sixth season. Acta Biologica Sibirica, 5 (2), 68 - 82. [in Russian] https: // doi. org / 10.14258 / abs. v 5. i 2.5933","Litovkin, S. V. (2012) Preliminary review of aquatic beetles (Insecta, Coleoptera) of the Buzuluksky Bor National Park. Proceedings of the Orenburg Branch of the Russian Entomological Society, 2. 47 - 53. [in Russian]","Litovkin, S. V., Sazhnev, A. S. & Klyomin, D. A. (2013) Variegated mud-loving beetles (Coleoptera, Heteroceridae) of Samarskaya, Saratovskaya and Ulyanovskaya Oblasts and the Republic of Tatarstan, Russia. Euroasian Entomological Journal, 12 (6), 561 - 569. [in Russian]","Sazhnev, A. S. (2014) On the fauna of variegated mud-loving beetles (Coleoptera: Heteroceridae) of the Middle and Lower Volga Region. Gidroentomologija v Rossii i sopredelnykh stranakh. Materialy V Vserossijskogo simpoziuma po amfibioticheskim i vodnym nasekomym, 2014, 161 - 165. [in Russian]","Sazhnev, A. S. (2016 c) The new materials to the fauna of variegated mud-loving beetles (Coleoptera: Heteroceridae) from the south of European part of Russia. Caucasian entomological bulletin, 12 (2), 247 - 252. [in Russian] https: // doi. org / 10.23885 / 1814 - 3326 - 2016 - 12 - 2 - 247 - 252","Egorov, L. V. (2019) Some data concerning the coleopterofauna of the Nature Reserve \" Prisursky \". Information 8. Scientific proceedings of the State Nature Reserve \" Prisursky \", 34, 26 - 167. [In Russian]","Sazhnev, A. S. & Kozminykh, V. O. (2019) Beetles of the family Heteroceridae (Insecta: Coleoptera) in the fauna of Ural. Materialy po flore i faune Respubliki Bashkortostan, 22, 88 - 105. [in Russian]","Sazhnev, A. S., Lychkovskaya, I. Yu. & Prokin, A. A. (2018) New data to the fauna of aquatic and semi-aquatic beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae, Hydrochidae, Hydrophilidae, Heteroceridae) of Ryazan Province. Eversmannia, 55 - 56. [in Russian]","Egorov, L. V., Ruchin, A. B., Semenov, V. B., Semionenkov, O. I., Semishin, G. B. (2020) Checklist of the Coleoptera of Mordovia State Nature Reserve, Russia. ZooKeys, 962, 13 - 122. https: // doi. org / 10.3897 / zookeys. 962.54477","Mascagni, A. (2016) Family Heteroceridae MacLeay, 1825. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden, Boston, pp. 610 - 616.","Kirejtshuk, A. G. (2001) Family Heteroceridae (Pilousy). In: Tsalolikhin, S. J. (Ed.), Key to freshwater invertebrates of Russia and adjacent lands. Vol. 5. Nauka, St. - Petersburg, pp. 341 - 348 + 742 - 749. [in Russian]","Charpentier, R. (1965) A monograph of the family Heteroceridae (Coleoptera) of the Ethiopian region. South African Animal Life, Lund, 11, 215 - 343.","Ovchinnikov, S. V. (1996) Family Heteroceridae - pilousy. In: Genetical fund cadastre of Kyrghyzstan. Vol. III. Superclassis Hexapoda (Entognatha and Insecta). Institute for Biology and Pedology National Academy of Sciences Kyrghyz republic, \" ALEINE \" oecological movement of Kyrghyzstan, Entomological Society of Kyrghyzstan, Bishkek, pp. 129 - 130. [in Russian]","Litovkin, S. V. & Sazhnev, A. S. (2018) The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist, 372, 25 - 32. https: // doi. org / 10.25221 / fee. 372.2"]}

Augyles (Augyles) obliteratus (Kiesenwetter, 1843) Material examined. 8 exs. CRIMEA. Donuzlav bay, Krasnoyarskoe vill., 24.07.2016, leg. V.V. Shaporinsky, 1 ex. (IBIW). RUSSIA. Astrakhan Oblast: Krasnoyarsky Distr., env, Dosang vill., Akhtuba River, 46°54'30.50"N 47°54'25.50"E, at light, 1– 8.05.2011, leg. I.A. Zabaluev, 1 ex. (IBIW); same place, in sediments, 28.04.2014, leg. A. Zaitsev & M. Pilipenko, 1 ex. (ZMMU); Akhtubinsky Distr., Bolkhuny vill., bank of Akhtuba River, 17.07.2014, leg. E.A. Negrobova, 1 ex. (IBIW). Karachay-Cherkess Republic: Urup Distr., Urup River near Pregradnoye vill., 43°58'25.2"N 41°12'04.7"E, 15.08.2016, leg. A.A. Prokin, 1 ex. (IBIW); Dagestan Republic: Magaramkent Distr., Gazardkam-Kazmalar vill., 19.07.2014, leg. E.V. Ilyina, 2 exs. (IBIW). Orenburg Oblast: Gaysiy Distr., Khmelevka vill., at light, 7.05.2012 leg. A.S. Ukrainsky, 1 ex. (IBIW); Orenburg, bank of Ural River, Petropavovskaya Grove, at light, 15.07.2012, leg. A.S. Ukrainsky, 1 ex. (IBIW). Distribution in European Russia and Caucasus. ST: AST CA: DAG KCH UR: ORB (Sazhnev 2017; Sazhnev & Ilyina 2017; Sazhnev & Kozminykh 2019) Georgia (Mascagni 2016). Distribution. The species was described from Iraq, is distributed in South Europe, Caucasus, Near East (Iraq, Iran, Syria, Yemen), Asian Turkey, Middle Asia (Afghanistan, Kazakhstan, Turkmenistan, Uzbekistan) and Mongolia (Mascagni 2016; Sazhnev 2020)., Published as part of Sazhnev, Alexey S., 2022, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region, pp. 581-593 in Zootaxa 5209 (5) on page 587, DOI: 10.11646/zootaxa.5209.5.5, http://zenodo.org/record/7340995, {"references":["Kiesenwetter, E. von. (1843) Beitrage zur Monographie der Gattung Heterocerus. Zeitschrift fur die Entomologie, 4, 194 - 224.","Sazhnev, A. S. & Ilyina, E. V. (2017) The variegated mud-loving beetles (Coleoptera: Heteroceridae) of the Republic of Dagestan, Russia. Caucasian Entomological Bulletin, 13 (2), 151 - 154. [in Russian] https: // doi. org / 10.23885 / 1814 - 3326 - 2016 - 12 - 2 - 247 - 252","Sazhnev, A. S. & Kozminykh, V. O. (2019) Beetles of the family Heteroceridae (Insecta: Coleoptera) in the fauna of Ural. Materialy po flore i faune Respubliki Bashkortostan, 22, 88 - 105. [in Russian]","Mascagni, A. (2016) Family Heteroceridae MacLeay, 1825. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden, Boston, pp. 610 - 616.","Sazhnev, A. S. (2020) Variegated mud-loving beetles (Heteroceridae) of the Russia and abject countries: Additions and corrections to the Catalogue of Palaearctic Coleoptera, Volume 3 (2016). Zootaxa, 4810 (2), 368 - 374. https: // doi. org / 10.11646 / zootaxa. 4810.2.10"]}

Augyles (Augyles) turanicus (Reitter, 1887) Material examined. 37 exs. AZERBAIJAN. Baku: Sangachal, shore of Caspian Sea, 40°09'39.53"N 49°28'20.53"E, 6.06.2013, leg. D.G. Kasatkin, 5 exs. (IBIW); Gobustal, 6.06.2013, leg. I.V. Shokhin & D.G. Kasatkin, 8 exs. (IBIW). RUSSIA. Dagestan Republic: Magaramkent Distr., State Nature Zakaznik “Samursky”, 15– 17.05.2015, leg. E.V. Ilyina, 7 exs. (IBIW); Magaramkent Distr., estuary of Samur River, Caspian Sea coast, at light, 5.06.2013, leg. E.V. Ilyina, 7 exs. (IBIW); Magaramkent Distr., Gazardkam-Kazmalar vill., 19.07.2014, leg. E.V. Ilyina, 9 exs. (IBIW); Magaramkent Distr., Tagirkent-Kazmalar vill., 20.07.2014, leg. E.V. Ilyina, 1 ex. (IBIW). Distribution in Caucasus. CA: DAG (Sazhnev 2016b, 2017; Sazhnev & Ilyina 2017). Azerbaijan: Baku (Sazhnev 2018b); Georgia (Mascagni 2016). Distribution. The species was from Turkmenistan, distributed in Caucasus, Near East (Arab Emirates, Iran, Iraq, Israel, Syria, Saudi Arabia), Asian Turkey, Middle Asia (Kazakhstan, Kirgizstan, Turkmenistan and Uzbekistan) and North Africa (Algeria) (Mascagni 2016; Sazhnev 2020)., Published as part of Sazhnev, Alexey S., 2022, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region, pp. 581-593 in Zootaxa 5209 (5) on page 588, DOI: 10.11646/zootaxa.5209.5.5, http://zenodo.org/record/7340995, {"references":["Reitter, E. (1887) Beschreibung der neuen Arten. Horae Societatis Entomologicae Rossicae, 21, 209 - 210. https: // doi. org / 10.1002 / mmnd. 48018870114","Sazhnev, A. S. (2016 b) New to the territory of Russia species of variegated mud-loving beetles (Coleoptera: Heteroceridae). Eversmannia, 47 - 48, 104. [in Russian]","Sazhnev, A. S. & Ilyina, E. V. (2017) The variegated mud-loving beetles (Coleoptera: Heteroceridae) of the Republic of Dagestan, Russia. Caucasian Entomological Bulletin, 13 (2), 151 - 154. [in Russian] https: // doi. org / 10.23885 / 1814 - 3326 - 2016 - 12 - 2 - 247 - 252","Mascagni, A. (2016) Family Heteroceridae MacLeay, 1825. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden, Boston, pp. 610 - 616.","Sazhnev, A. S. (2020) Variegated mud-loving beetles (Heteroceridae) of the Russia and abject countries: Additions and corrections to the Catalogue of Palaearctic Coleoptera, Volume 3 (2016). Zootaxa, 4810 (2), 368 - 374. https: // doi. org / 10.11646 / zootaxa. 4810.2.10"]}

[Augyles (Augyles) dilutissimus (Reitter, 1887)] Distribution in Caucasus. Azerbaijan (Skalický 1998; Sazhnev 2018b). Distribution. The species was described from China (Xinjiang) (Reitter 1887), distributed in Middle Asia (Kazakhstan, Tajikistan, Turkmenistan), Near East (Israel, Iran, Iraq) (Derwesh 1965; Mascagni 2016; Litovkin & Sazhnev 2018) and Caucasus (Azerbaijan) (Skalický 1998)., Published as part of Sazhnev, Alexey S., 2022, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region, pp. 581-593 in Zootaxa 5209 (5) on page 584, DOI: 10.11646/zootaxa.5209.5.5, http://zenodo.org/record/7340995, {"references":["Reitter, E. (1887) Beschreibung der neuen Arten. Horae Societatis Entomologicae Rossicae, 21, 209 - 210. https: // doi. org / 10.1002 / mmnd. 48018870114","Skalicky, S. (1998) Two new Heteroceridae species, collected by the National Natural History Museum of Prague expeditions to Iran (Coleoptera). Opuscula Zoologica Fluminensia, 160, 1 - 11.","Derwesh, A. I. (1965) A preliminary list of identified insects and some arachnids of Iraq. Directorate of Agricultural Research and Projects, 112, 123.","Mascagni, A. (2016) Family Heteroceridae MacLeay, 1825. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden, Boston, pp. 610 - 616.","Litovkin, S. V. & Sazhnev, A. S. (2018) The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Kyrgyzstan. Far Eastern Entomologist, 372, 25 - 32. https: // doi. org / 10.25221 / fee. 372.2"]}

Augyles (Augyles) marmota (Kiesenwetter, 1850) Material examined. 4 exs. RUSSIA: Orenburg Oblast: Orenburg, bank of Ural River, Petropavovskaya Grove, at light, 15.07.2012, leg. A.S. Ukrainsky, 4 exs. (IBIW). Distribution in European Russia and Caucasus: VL: SAM UR: ORB (Litovkin et al. 2013: as Augyles sp.; Sazhnev & Kozminykh 2019). Armenia (Mascagni 2016). Distribution. Species distributed in Europe (including southern Ural) Asian Turkey, Mongolia and North Africa (Mascagni 2016; Sazhnev & Kozminykh 2019). Remarks. Comparison of examined specimens (as well as Mongolian specimens) with European material and types is required. The diagnosis of the taxon Augyles marmota and the list of its synonyms needs revision. It is possible that more than one species is hidden under this name. [Augyles (Augyles) nebulosus (Kuwert, 1890)] Distribution in Caucasus. Azerbaijan: Agdash (Aresh, Geok Tepe), Mingachevir (Zaitsev 1946; Sazhnev 2018b). Distribution. The species was described from Turkmenistan (Kuwert, 1890), is distributed in Caucasus (Azerbaijan) and Middle Asia (Turkmenistan, Uzbekistan) (Mascagni 2016). Remarks. There is no indication for Azerbaijan in the Palaearctic Catalogue (Mascagni 2016), it is necessary to add E: AB., Published as part of Sazhnev, Alexey S., 2022, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region, pp. 581-593 in Zootaxa 5209 (5) on page 587, DOI: 10.11646/zootaxa.5209.5.5, http://zenodo.org/record/7340995, {"references":["Litovkin, S. V., Sazhnev, A. S. & Klyomin, D. A. (2013) Variegated mud-loving beetles (Coleoptera, Heteroceridae) of Samarskaya, Saratovskaya and Ulyanovskaya Oblasts and the Republic of Tatarstan, Russia. Euroasian Entomological Journal, 12 (6), 561 - 569. [in Russian]","Sazhnev, A. S. & Kozminykh, V. O. (2019) Beetles of the family Heteroceridae (Insecta: Coleoptera) in the fauna of Ural. Materialy po flore i faune Respubliki Bashkortostan, 22, 88 - 105. [in Russian]","Mascagni, A. (2016) Family Heteroceridae MacLeay, 1825. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden, Boston, pp. 610 - 616.","Kuwert, A. (1890) Bestimmungstabelle der Heteroceren Europas und der angrenzenden Gebiete, soweit dieselben bisher bekannt wurden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 40, 517 - 548.","Zaitsev, F. A. (1946) Rasprostranenie v Zakavkaz'e vidov sem. Pilousov (Coleoptera, Heteroceridae). Trudy Zoologicheskogo instituta AN GruzSSR, 6, 213 - 220. [in Russian, English translation in: Distribution in Transcaucasia species of the variegated mud-loving beetles (Coleoptera, Heteroceridae). Proceedings of Zoological Institute of the Georgian Academy of Sciences, 6, 213 - 220.]"]}

Augyles (Augyles) maritimus (Guérin-Méneville, 1844) Material examined. 22 exs. ABKHAZIA. Sukhumi, 30.03.1960 leg. Chernyshev, 1 ex. (IBIW); Bzyb’ River near Pitsunda, 43°11'34.4"N 40°17'14.6"E, leg. A.A. Prokin, 7 exs. (IBIW). AZERBAIJAN. Lankaran District: Avrora vill., Hirkan National Park, at light, 4.05.1980, leg. S.G. Korolev, 1 ex. (IBIW). RUSSIA. Astrakhan Oblast: Krasnoyarsky Distr., env, Dosang vill., at light, 8.05.2009, leg. A.V. Kovalev 4 exs. (IBIW, ZISP); same place, Akhtuba River, 46°54'30.50"N 47°54'25.50"E, at light, 1– 8.05.2011, leg. I.A. Zabaluev, 2 exs. (IBIW). KarachayCherkess Republic: Urup Distr., Urup River near Pregradnoye vill., 43°58'25.2"N 41°12'04.7"E, 15.08.2016, leg. A.A. Prokin, 4 exs. (IBIW). Krasnodar Krai: 3 km N Aderbievka vill., W slope of Kotsekhur mountain range, shore of Aderba River, h ~ 160 m, 44˚37’52"N 38˚06’06"E, 3.05.2016, 1 ex, K.V. Makarov & O.L. Makarova leg. (IBIW). Republic of North Ossetia-Alania: Alagirsky Distr., Barzikau vill., shore of Fiagdon River, h ≈ 1200 m, 42°50'37"N 44°18'44"E, 11.05.2016, leg. A.S. Sazhnev, 1 ex. (IBIW). Saratov Oblast: [Saratov Uezd], on the clay bank, 2.06.1900, leg. A.I. Jakovlev, 1 ex. (ZISP). Ulyanovsk Oblast: Novospassky Distr., 3 km E of Marievka vill., 53°08'N 48°09'E, 114 m, 7.07.2009, leg. A.V. Kovalev, 1 ex. (IBIW). CRIMEA. Tauria, 14.06.1911, leg. V. Pliginskiy, 1 ex (ZISP). Distribution in European Russia and Caucasus. ST: AST SAR* VL: SAM ULY CA: KCH KRD NOS (Isaev et al. 2001; Sazhnev 2014, 2017; Sazhnev & Prokin 2022). Crimea (state with limited recognition); Abkhazia (state with limited recognition); Azerbaijan (Sazhnev 2018b). Distribution. West-Palearctic species was described from France, distributed in Europe, Caucasus, Near East (Syria), Asian Turkey and North Africa (Mascagni 2016; Sazhnev 2020). Remarks. The species recorded for Saratov Oblast for the first time. This species was recorded from Crimea [Tauria] as Augyles obliteratus (Plyginskiy 1928)., Published as part of Sazhnev, Alexey S., 2022, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region, pp. 581-593 in Zootaxa 5209 (5) on pages 586-587, DOI: 10.11646/zootaxa.5209.5.5, http://zenodo.org/record/7340995, {"references":["Guerin-Meneville, F. E. (1844) Iconographie du regne animal de G. Cuvier, vol. 7, Insects. J. B. Bailliere, Paris, 576 pp., 104 pls.","Isaev, A. Yu., Anikin, V. V. & Zolotukhin, V. V. (2001) To the beetle fauna (Coleoptera) of Krasnoyarsky district Astrakhan Oblast. Entomological and parasitological research in the Volga Region, 1, 41 - 44. [In Russian]","Sazhnev, A. S. (2014) On the fauna of variegated mud-loving beetles (Coleoptera: Heteroceridae) of the Middle and Lower Volga Region. Gidroentomologija v Rossii i sopredelnykh stranakh. Materialy V Vserossijskogo simpoziuma po amfibioticheskim i vodnym nasekomym, 2014, 161 - 165. [in Russian]","Sazhnev, A. S. & Prokin, A. A. (2022) New data on Dytiscidae, Hydrophilidae, Hydraenidae, Dryopidae and Heteroceridae (Coleoptera) of North Ossetia. T ransactions of Papanin Institute for Biology of Inland Waters RAS, 98 (101), 38 - 45. https: // doi. org / 10.47021 / 0320 - 3557 - 2022 - 38 - 45.","Mascagni, A. (2016) Family Heteroceridae MacLeay, 1825. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden, Boston, pp. 610 - 616.","Sazhnev, A. S. (2020) Variegated mud-loving beetles (Heteroceridae) of the Russia and abject countries: Additions and corrections to the Catalogue of Palaearctic Coleoptera, Volume 3 (2016). Zootaxa, 4810 (2), 368 - 374. https: // doi. org / 10.11646 / zootaxa. 4810.2.10"]}

[Augyles (Augyles) pruinosus (Kiesenwetter, 1851)] Material examined. 2 exs. AZERBAIJAN. Lankaran District: Lenkoran, leg. Leder (Reitter), 1 ex., (ZISP); Masally Distr.: Goryachie vody, Vilyash-chay River, 20.08.1932, leg. Znoyko, 1 ex. (ZISP). Distribution in Caucasus. Azerbaijan (this paper). Distribution. The species was described from France, is distributed in Europe, Near East (Iran, Iraq), Caucasus (Azerbaijan), Asian Turkey and Middle Asia (Afghanistan, Turkmenistan, Uzbekistan) (Mascagni 2016). Remarks. The species recorded for Azerbaijan (add E: AB) for the first time., Published as part of Sazhnev, Alexey S., 2022, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region, pp. 581-593 in Zootaxa 5209 (5) on page 587, DOI: 10.11646/zootaxa.5209.5.5, http://zenodo.org/record/7340995, {"references":["Kiesenwetter, E. von. (1851) Revision der Kafergattung Heterocerus. Linnaea Entomologica, 5, 281 - 300.","Mascagni, A. (2016) Family Heteroceridae MacLeay, 1825. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden, Boston, pp. 610 - 616."]}

Augyles (Augyles) sericans (Kiesenwetter, 1843) = Augyles scutellatus (Motschulsky, 1854) = Augyles vitticollis (Reitter, 1884) comb. nov. Material examined. 18 exs. AZERBAIJAN. Qabala District: Nidzh vill, shore of Turianchay River, 38°34'58.6"N 48°47'38.2"E, 6.06.2017, leg. A.A. Prokin, 1 ex. (IBIW); Lankaran District: Lenkoran, leg. Leder (Reitter), 2 exs., as Augyles scutellatus (ZISP); Alekseevka vill., Hirkan National Park, at light, 11.07.1978, leg. V.V. Belov, 1 ex. (IBIW), same place, 13.07.1978, leg. V.V. Belov, 1 ex. (IBIW), same place, 15.07.1978, leg. V.V. Belov, 1 ex. (IBIW). GEORGIA. Mtskheta-Mtianeti Krai: Mtskheta Distr., Kura River opposite Dzegvi vill., 41°51.199'N 44°35.737'E, 10.08.2016, leg. D. Palatov, 3 exs. (IBIW). Samtskhe – Javakheti Krai: Borjomi, 3.05.1911, unknown collector, 1 ex. (ZISP). RUSSIA. Dagestan Republic: Magaramkent Distr., State Nature Zakaznik “Samursky”, 15– 17.05.2015, leg. E.V. Ilyina, 1 ex. (IBIW), 29– 30.07.2015, leg. E.V. Ilyina, 1 ex. (IBIW); Magaramkent Distr., Gazardkam-Kazmalar vill., 19.07.2014, leg. E.V. Ilyina, 2 exs. (IBIW); Magaramkent Distr., Tagirkent-Kazmalar vill., 20.07.2014, leg. E.V. Ilyina, 1 ex. (IBIW). Krasnodar Krai: Cauc. occ., Adler, 08, leg. Dr. Lgocki, 1 ex., as Augyles scutellatus (ZISP); Tuapse Distr., env. Tuapse, Agoy River, 44.15°N, 39.06°E, 20.08.2016, leg. A.A. Prokin, 1 ex. (IBIW); Sochi, Adlersky Distr., Adler, Mzymta River, 2– 3.09.2004, leg. S.A. Svobodny, 2 exs. (IBIW). Orenburg Oblast: Gaysiy Distr., Khmelevka vill., at light, 8.05.2012 leg. A.S. Ukrainsky, 1 ex. (IBIW). Distribution in European Russia and Caucasus. CT: RZN VL: MAR/TAT ST: AST CA: DAG KRD UR: ORB (Jacobson 1913; Zaitsev 1916; Lebedev 1925; Litovkin et al. 2013; Sazhnev 2014; Sazhnev & Shapovalov 2016; Sazhnev 2017; Sazhnev & Ilyina 2017; Sazhnev & Kozminykh 2019). Azerbaijan: Agdash (Geok Tepe) (Zaitsev 1946, 1953; Sazhnev 2018b), Elisabethpol Governorate (including as Augyles scutellatus) (Jacobson 1913), Ganja (Kirovabad, as Augyles vitticollis) (Zaitsev 1946), Lankaran (as Augyles vitticollis) (Zaitsev 1946), Talysh (as Augyles scutellatus) (Jacobson 1913); Georgia: Borjomi (Zaitsev 1946, 1953), Khashuri (Mikhailovo) (Schneider & Leder 1877; Zaitsev 1946), Kvemo Kartli Krai (Bolnisi—former Ekaterinenpheld, including as Augyles vitticollis) (Zaitsev 1946), Matani (Zaitsev 1946, 1953), Tbilisi (Tiflis, including as Augyles scutellatus) (Jacobson 1913). Distribution. European species (described from Switzerland), distributed in Caucasus, Near East (Israel, Syria) (Mascagni, 2016; Sazhnev 2020). Remarks. By F.A. Zaitsev (1946) typical material of Augyles scutellatus (Motschulsky, 1854) was studied and this taxon to Augyles sericans was synonymized. Since Augyles vitticollis (Reitter 1884) is junior synonym of Augyles scutellatus (Mascagni 2016), both taxons should be referred to synonyms of Augyles sericans., Published as part of Sazhnev, Alexey S., 2022, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region, pp. 581-593 in Zootaxa 5209 (5) on page 588, DOI: 10.11646/zootaxa.5209.5.5, http://zenodo.org/record/7340995, {"references":["Kiesenwetter, E. von. (1843) Beitrage zur Monographie der Gattung Heterocerus. Zeitschrift fur die Entomologie, 4, 194 - 224.","Motschulsky, V. I. (1854) Sur les Heteroceres de Russie. Etudes Entomologiques, Imprimerie de la Societe de Literature Finnoise, Helsingfors, 2, 14 - 21.","Jacobson, G. G. (1913) Semeystvo Heteroceridae. Pilousy. In: Zhuki Rossii i zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov. Vol. 10. A. F. Davrien, Sankt-Peterburg, pp. 867 - 869. [in Russian, English translation in: Family Heteroceridae. Mud-loving beetles. Beetles of Russia, Western Europe and neighbouring countries. Guide to the determination of beetles. A. F. Devrien, St. - Petersburg, 10, pp. 867 - 869.]","Zaitsev, F. A. (1916) Zametki o nekotorykh predstavitelyakh semeistva Heteroceridae (Coleoptera). Russkoe entomologicheskoe obozrenie, 15 (4), 569 - 571. [in Russian, English translation in: Notes on some of the representatives of the family Heteroceridae (Coleoptera)]. Russian Entomological Review, 15 (4), 569 - 571.","Lebedev, A. G. (1925) Materials for the fauna of beetles of the Republic Tatarstan. III. Entomological Review, 19, 133 - 138. [in Russian]","Litovkin, S. V., Sazhnev, A. S. & Klyomin, D. A. (2013) Variegated mud-loving beetles (Coleoptera, Heteroceridae) of Samarskaya, Saratovskaya and Ulyanovskaya Oblasts and the Republic of Tatarstan, Russia. Euroasian Entomological Journal, 12 (6), 561 - 569. [in Russian]","Sazhnev, A. S. (2014) On the fauna of variegated mud-loving beetles (Coleoptera: Heteroceridae) of the Middle and Lower Volga Region. Gidroentomologija v Rossii i sopredelnykh stranakh. Materialy V Vserossijskogo simpoziuma po amfibioticheskim i vodnym nasekomym, 2014, 161 - 165. [in Russian]","Sazhnev, A. S. & Ilyina, E. V. (2017) The variegated mud-loving beetles (Coleoptera: Heteroceridae) of the Republic of Dagestan, Russia. Caucasian Entomological Bulletin, 13 (2), 151 - 154. [in Russian] https: // doi. org / 10.23885 / 1814 - 3326 - 2016 - 12 - 2 - 247 - 252","Sazhnev, A. S. & Kozminykh, V. O. (2019) Beetles of the family Heteroceridae (Insecta: Coleoptera) in the fauna of Ural. Materialy po flore i faune Respubliki Bashkortostan, 22, 88 - 105. [in Russian]","Zaitsev, F. A. (1946) Rasprostranenie v Zakavkaz'e vidov sem. Pilousov (Coleoptera, Heteroceridae). Trudy Zoologicheskogo instituta AN GruzSSR, 6, 213 - 220. [in Russian, English translation in: Distribution in Transcaucasia species of the variegated mud-loving beetles (Coleoptera, Heteroceridae). Proceedings of Zoological Institute of the Georgian Academy of Sciences, 6, 213 - 220.]","Zaitsev, F. A. (1953) Vodnye zhestkokrylye v faune Gruzii. Trudy Zoologicheskogo instituta AN GruzSSR, 11, 87 - 126. [in Russian, English translation in: Water beetles in the fauna of Georgia. Proceedings of Zoological Institute of the Georgian Academy of Sciences, 11, 87 - 126.]","Schneider, O. & Leder, H. (1877) Beitrage zur Kenntniss der kaukasischen Kaferfaun. Verhandlungen des Naturforschenden Vereins in Brunn, 16, 3 - 258.","Mascagni, A. (2016) Family Heteroceridae MacLeay, 1825. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden, Boston, pp. 610 - 616.","Sazhnev, A. S. (2020) Variegated mud-loving beetles (Heteroceridae) of the Russia and abject countries: Additions and corrections to the Catalogue of Palaearctic Coleoptera, Volume 3 (2016). Zootaxa, 4810 (2), 368 - 374. https: // doi. org / 10.11646 / zootaxa. 4810.2.10"]}

Augyles (Augyles) flavidus (Rossi, 1794) Material examined. 52 exs. GEORGIA. Mtskheta-Mtianeti Krai: Mtskheta Distr., Kura River opposite Dzegvi vill., 41°51.199'N 44°35.737'E, 10.08.2016, leg. D.M. Palatov, 2 ex. (IBIW). RUSSIA. Astrakan Oblast: Krasnoyarsky Distr., Dosang vill., 8.05.2009, leg. A.V. Kovalev, 4 exs. (ZISP); same place, bank Akhtuba River, 46°54'30.50"N 47°54'25.50"E, at light, 1– 8.05.2011, leg. I.A. Zabaluev, 7 exs, (IBIW). Dagestan Republic: Kumtorkalinsky Distr., Kumtorkala vill., Sarykum barkhan, at light, 9.06.2013, leg. E.V. Ilyina, 1 ex. (IBIW); same place, Kumtorkala station, 31.07.2016, E.V. Ilyina, 1 ex. (IBIW), 19.05.2018, leg. E.V. Ilyina, 5 exs. (IBIW); same place, right bank Shura-Ozen River, near the railway bridge, night at light, 23.05.2019, leg. I.A. Zabaluev, 21 exs, (IBIW); same place, Kumtorkala station, 2– 3.06.2019, leg. I.A. Zabaluev, 4 exs. (IBIW); Magaramkent Distr., Gazardkam-Kazmalar vill., 19.07.2014, leg. E.V. Ilyina, 2 exs. (IBIW); Magaramkent Distr., Tagirkent-Kazmalar vill., 20.07.2014, leg. E.V. Ilyina, 2 exs. (IBIW); Magaramkent Distr., State Nature Zakaznik “Samursky”, 29– 30.07.2015, leg. E.V. Ilyina, 1 ex. (IBIW); Nogaysky Distr., sands Karagaily-Kum, 44°02'15.4"N 45°38'34.2"E, 16.08.2016, leg M.V. Nabozhenko, S.V. Nabozhenko & G.M. Abdurakhmanov, 1 ex. (IBIW). Volgograd Oblast: Palassovsky Distr., Elton vill., at light, 21– 28.05.2006, leg A.V. Matalin, 1 ex. (IBIW). Distribution in European Russia and Caucasus. ST: AST VOL (Sazhnev 2016c) CA: ADG DAG KAB (Tersky Distr.) KCH/STA (Podkumok River) (Motschulsky 1854; Jakobson 1913; Zaitsev 1946; Nikitsky 2010; Sazhnev 2017; Sazhnev & Ilyina 2017). Armenia: Elisabethpol, Governorate (Jacobson 1913; Mascagni 2016); Azerbaijan: Agdash (Geok Tepe) (Schneider & Leder 1877; Jacobson 1913; Zaitsev 1946; Sazhnev 2018b); Georgia: Karayaz, Tbilisi (Zaitsev 1946, 1953; Mascagni 2016). Distribution. The species was described from Italy (Etruria), distributed in South Europe, Caucasus (Russian North Caucasus, Armenia, Azerbaijan, Georgia), Near East (Arab Emirates, Iran, Iraq, Lebanon, Saudi Arabia, Egypt (Sinai)), Middle Asia (Kazakhstan, Kirgizstan, Tajikistan, Turkmenistan and Uzbekistan), Pakistan and North Africa (Mascagni 2016; Sazhnev 2020). In Russia, this species occurs only in the extreme south of European part and North Caucasus (Sazhnev & Shapovalov 2016; Sazhnev 2016b). The records for most part of Siberia (species is possible in the southwest West Siberia) and the Russian Far East are erroneous and require confirmation (Sazhnev 2020). Remarks. The species recorded for Mtskheta-Mtianeti Krai of Georgia for the first time. The record from Khashuri (Mikhailovo) (Schneider & Leder 1877) on the opinion by F.A. Zaitsev (1953) is other species Augyles sericans (Kiesenwetter, 1843)., Published as part of Sazhnev, Alexey S., 2022, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region, pp. 581-593 in Zootaxa 5209 (5) on page 584, DOI: 10.11646/zootaxa.5209.5.5, http://zenodo.org/record/7340995, {"references":["Rossi, P. (1794) Mantissa Insectorum, exhibiens species nuper in Etruria collectas a Petro Rossio adiectis faunae Etruscae illustrationibus, ac emendationibus. Tom. II. Pisis: Typographia Prosperi, 154 pp., 8 pls.","Sazhnev, A. S. (2016 c) The new materials to the fauna of variegated mud-loving beetles (Coleoptera: Heteroceridae) from the south of European part of Russia. Caucasian entomological bulletin, 12 (2), 247 - 252. [in Russian] https: // doi. org / 10.23885 / 1814 - 3326 - 2016 - 12 - 2 - 247 - 252","Motschulsky, V. I. (1854) Sur les Heteroceres de Russie. Etudes Entomologiques, Imprimerie de la Societe de Literature Finnoise, Helsingfors, 2, 14 - 21.","Zaitsev, F. A. (1946) Rasprostranenie v Zakavkaz'e vidov sem. Pilousov (Coleoptera, Heteroceridae). Trudy Zoologicheskogo instituta AN GruzSSR, 6, 213 - 220. [in Russian, English translation in: Distribution in Transcaucasia species of the variegated mud-loving beetles (Coleoptera, Heteroceridae). Proceedings of Zoological Institute of the Georgian Academy of Sciences, 6, 213 - 220.]","Nikitsky, N. B. (2010) Heteroceridae - Pilousy. In: Zhestkokrylye nasekomye (Insecta, Coleoptera) Respubliki Adygeya. Izdatel'stvo Adygeyskogo universiteta, Maykop, pp. 129 - 130. [in Russian, English translation in: Beetles (Insecta, Coleoptera) of Republic of Adygea (annotated catalogue of species) (Abstracts Fauna of Adygea. No. 1). In: Zamotaylov, A. S. & Nikitsky, N. B. (Eds.), Adyghei State University Publishers, Maykop, pp. 129 - 130.]","Sazhnev, A. S. & Ilyina, E. V. (2017) The variegated mud-loving beetles (Coleoptera: Heteroceridae) of the Republic of Dagestan, Russia. Caucasian Entomological Bulletin, 13 (2), 151 - 154. [in Russian] https: // doi. org / 10.23885 / 1814 - 3326 - 2016 - 12 - 2 - 247 - 252","Jacobson, G. G. (1913) Semeystvo Heteroceridae. Pilousy. In: Zhuki Rossii i zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov. Vol. 10. A. F. Davrien, Sankt-Peterburg, pp. 867 - 869. [in Russian, English translation in: Family Heteroceridae. Mud-loving beetles. Beetles of Russia, Western Europe and neighbouring countries. Guide to the determination of beetles. A. F. Devrien, St. - Petersburg, 10, pp. 867 - 869.]","Mascagni, A. (2016) Family Heteroceridae MacLeay, 1825. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden, Boston, pp. 610 - 616.","Schneider, O. & Leder, H. (1877) Beitrage zur Kenntniss der kaukasischen Kaferfaun. Verhandlungen des Naturforschenden Vereins in Brunn, 16, 3 - 258.","Zaitsev, F. A. (1953) Vodnye zhestkokrylye v faune Gruzii. Trudy Zoologicheskogo instituta AN GruzSSR, 11, 87 - 126. [in Russian, English translation in: Water beetles in the fauna of Georgia. Proceedings of Zoological Institute of the Georgian Academy of Sciences, 11, 87 - 126.]","Sazhnev, A. S. (2020) Variegated mud-loving beetles (Heteroceridae) of the Russia and abject countries: Additions and corrections to the Catalogue of Palaearctic Coleoptera, Volume 3 (2016). Zootaxa, 4810 (2), 368 - 374. https: // doi. org / 10.11646 / zootaxa. 4810.2.10","Sazhnev, A. S. (2016 b) New to the territory of Russia species of variegated mud-loving beetles (Coleoptera: Heteroceridae). Eversmannia, 47 - 48, 104. [in Russian]","Kiesenwetter, E. von. (1843) Beitrage zur Monographie der Gattung Heterocerus. Zeitschrift fur die Entomologie, 4, 194 - 224."]}

Augyles (Augyles) intermedius (Kiesenwetter, 1843) Material examined. 36 exs. RUSSIA. Kaluga Oblast: Kirovsky Distr., Berezhki vill., 54°07'16.0"N 34°26'29.0"E, at light, 27– 28.06.2016, leg. V.V. Perov, 3 exs. (IBIW). Komi Republic: Ust-Tsilemsky Distr., Ust-Tsilma vill., 29.08.1904, leg. Zhuravsky, 1 ex. (ZISP); Syktyvkar, Vezhdino vill., 20.07.1991, leg. A. Zheleznyakov, 3 exs. (MPSU). Krasnodar Krai: Novorossiysk, 18.08.1878 (var. basimaculatus Reitter, 1908), unknown collector, 1 ex. (ZISP). Leningrad Oblast: Kingiseppsky Distr., Romanovka vill., env. Kingisepp (Yamburg), 30– 31.05.1904, leg. Barovsky, 4 exs. (ZISP). Moscow Oblast: [Klinsky uezd, Babkovo], 3.06.1905, D.S., 1 ex. (ZISP); 23 km W Zvenigorod, env. Glubokoe Lake, at light, 14.06.1998, leg. A.O. Bieńkowski, 2 exs. (PCB); 5 km N Zelenograd, Nikolskoe vill., at light, 23.06.2006, leg. A.O. Bieńkowski, 1 ex. (PCB). Nenets Autonomous Okrug: Gulf of Khaydypur, More-Yu River, 7– 15.08.2015, leg. O. Makarova & N. Bizin, 2 exs. (MSPU). Nizhniy Novgorod Oblast: Arzamas Distr., Volchikha vill., [Gorbatovsky uezd], 4– 9.05.1894, leg. Jacobson, 1 ex. (ZISP). Saint Petersburg: Lesnoy, 20.05.1862, leg. А. Moravits, 1 ex. (ZISP); env. Pavlovsk, 5.06.1906, leg. V. Barovsky, 2 exs. (ZISP). Saratov Oblast: Tatishchevo Distr., env. Storozhovka vill., Kurdyum River, 4.09.2014, leg. I.A. Zabaluev, 1 ex. (IBIW). Tambov Oblast: 20 km NNE Tambov, Zarechie vill., at light, 16.08.2008, leg. R.N. Ishin, 1 ex. (PCL). Tver Oblast: Bologoe, 26.06.1903, unknown collector, 1 ex. (ZISP). Ulyanovsk Oblast: Radishchevsky Distr., 6 km N Vyazovka vill., saline steppe, 3.05.2005, leg. A.Yu. Isaev, 1 ex. (ZISP); Radishchevsky Distr., 8 km S Vyazovka vill., 29.06.2008, leg. A.V. Kovalev, 1 ex. (ZISP). Vladmir Oblast: Murmansky Distr., Berdishchevo, unknown collector, 1 ex. (ZISP). Vologda Oblast: Vozhegodsky Distr., env. Gridino vill., sand bank Vozhega River, 31.07.2016, leg. D. Vtorushin, 1 ex. (IBIW). Yaroslavl Oblast: Yaroslavl, 10.05.1923, unknown collector, 1 ex. (ZMMU), same place, 14.05.1923, unknown collector, 1 ex. (ZMMU); Rostov Distr., Melenki station, 4.06.2006, leg. V.D. Titov, 1 ex. (YMR); Uglich Distr., env. Uglich, Uglich Reservoir, 23.05.1999, leg. Maslov, 1 ex. (YMR); Uglich Distr., Meteevo vill., Uleyma River, 14.06.1997, leg. D.V. Vlasov, 1 ex. (YMR); Yaroslavl Distr., 9.08.1926, leg. Jakovlev, 2 exs. (ZISP); Yaroslavl Distr., Berditsyno vill., 3.05.1989, leg. A. Jakovlev, 1 ex. (ZISP); Yaroslavl Distr., Belkino vill., Tallitsa River, 2.07.1999, leg. D.V. Vlasov, 1 ex. (YMR). Distribution in European Russia and Caucasus. NT: KAR NEN SPB VLG CT: KÖN KLG MOS NNO* NOV TAM* TVR* VLD YAR VL: TAT ULY ST: LIP SAR CA: KRD (Novorossiysk) UR: EKB KOM (Motschulsky 1854; Lindemann 1871; Yakovlev 1902; Zaitsev 1907, 1916, 1946; Jacobson 1913; Zaitsev 1915, 1915a, 1930; Lebedev 1925; Bercio & Folwaczny 1979; Semyonov 2009; Tsurikov 2009; Litovkin et al. 2013; Sazhnev 2016a, 2017, 2018; Sazhnev & Shapovalov 2016; Sazhnev et al. 2019; Sazhnev & Kozminykh 2019). Distribution. Holarctic species (Charpentier 1979; Mascagni 2016) with a disjunction of the range on the Black Sea coast of the North Caucasus (Zaitsev 1946). Species was described from Germany, distributed in North and Central Europe, Central Asia (northern Kazakhstan and Mongolia), West and East Siberia, Russian Far East, North America (Mascagni 2016; Sazhnev 2020). Remarks. The first record this species for Caucasus was make as Augyles seriepilosus Motschulsky, 1860 (Zaitsev 1916), which F.A. Zaitsev considered as probably synonym of Augyles intermedius species (Zaitsev 1946). But Augyles seriepilosus (species dubius) was described from East Siberia (Transbaikal Region) (Motschulsky 1860), specimen of « Augyles seriepilosus » with Motschulsky labels is in ZISP collection, and it possible refers to Asian species Augyles interspidulus (Charpentier, 1979). The species recorded for Nizhniy Novgorod, Tambov and Tver Oblasts for the first time., Published as part of Sazhnev, Alexey S., 2022, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region, pp. 581-593 in Zootaxa 5209 (5) on pages 585-586, DOI: 10.11646/zootaxa.5209.5.5, http://zenodo.org/record/7340995, {"references":["Kiesenwetter, E. von. (1843) Beitrage zur Monographie der Gattung Heterocerus. Zeitschrift fur die Entomologie, 4, 194 - 224.","Motschulsky, V. I. (1854) Sur les Heteroceres de Russie. Etudes Entomologiques, Imprimerie de la Societe de Literature Finnoise, Helsingfors, 2, 14 - 21.","Lindemann, K. (1871) A review of geographic distribution of beetles in Russian Empire. Part 1. Introduction, Fore word. Northern, Moscowian and Turanian Oblasts. Proceedings of Russian Entomological Society, 6, 41 - 366. [in Russian]","Zaitsev, F. A. (1907) Water scavenger beetles of St. - Petersburg Governorate. Ezhegodnik zoologicheskago muzeya Imperatorskoy akademii nauk, 12 (2), 144 - 176. [in Russian]","Zaitsev, F. A. (1916) Zametki o nekotorykh predstavitelyakh semeistva Heteroceridae (Coleoptera). Russkoe entomologicheskoe obozrenie, 15 (4), 569 - 571. [in Russian, English translation in: Notes on some of the representatives of the family Heteroceridae (Coleoptera)]. Russian Entomological Review, 15 (4), 569 - 571.","Zaitsev, F. A. (1946) Rasprostranenie v Zakavkaz'e vidov sem. Pilousov (Coleoptera, Heteroceridae). Trudy Zoologicheskogo instituta AN GruzSSR, 6, 213 - 220. [in Russian, English translation in: Distribution in Transcaucasia species of the variegated mud-loving beetles (Coleoptera, Heteroceridae). Proceedings of Zoological Institute of the Georgian Academy of Sciences, 6, 213 - 220.]","Jacobson, G. G. (1913) Semeystvo Heteroceridae. Pilousy. In: Zhuki Rossii i zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov. Vol. 10. A. F. Davrien, Sankt-Peterburg, pp. 867 - 869. [in Russian, English translation in: Family Heteroceridae. Mud-loving beetles. Beetles of Russia, Western Europe and neighbouring countries. Guide to the determination of beetles. A. F. Devrien, St. - Petersburg, 10, pp. 867 - 869.]","Lebedev, A. G. (1925) Materials for the fauna of beetles of the Republic Tatarstan. III. Entomological Review, 19, 133 - 138. [in Russian]","Bercio, H. & Folwaczny, B. (1979) Verzeichnis der Kafer Preussens. Parzeller & Co, Fulda, 194.","Semyonov, V. B. (2009) An annotated list of coleopteran insects (Insecta, Coleoptera) of Central Meschera. KMK, Moscow, 167 pp. [in Russian]","Tsurikov, M. N. (2009) Beetles of Lipetsk Oblast. Voronezh State University, Voronezh, 332.","Litovkin, S. V., Sazhnev, A. S. & Klyomin, D. A. (2013) Variegated mud-loving beetles (Coleoptera, Heteroceridae) of Samarskaya, Saratovskaya and Ulyanovskaya Oblasts and the Republic of Tatarstan, Russia. Euroasian Entomological Journal, 12 (6), 561 - 569. [in Russian]","Sazhnev, A. S. (2016 a) Composition and structure of the Heteroceridae (Coleoptera) population in the shore zone of water objects in the Saratov region. Povolzhskiy Journal of Ecology, 1, 85 - 93. [in Russian] https: // doi. org / 10.18500 / 1684 - 7318 - 2016 - 1 - 85 - 93","Sazhnev, A. S. & Kozminykh, V. O. (2019) Beetles of the family Heteroceridae (Insecta: Coleoptera) in the fauna of Ural. Materialy po flore i faune Respubliki Bashkortostan, 22, 88 - 105. [in Russian]","Charpentier, R. (1979) Heteroceridae (Coleoptera) from Mongolia with description of Heterocerus kaszabi n. sp. and Heterocerus interspidulus n. sp. Entomologica scandinavica, 10 (3), 229 - 237. https: // doi. org / 10.1163 / 187631279794814913","Mascagni, A. (2016) Family Heteroceridae MacLeay, 1825. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Brill, Leiden, Boston, pp. 610 - 616.","Sazhnev, A. S. (2020) Variegated mud-loving beetles (Heteroceridae) of the Russia and abject countries: Additions and corrections to the Catalogue of Palaearctic Coleoptera, Volume 3 (2016). Zootaxa, 4810 (2), 368 - 374. https: // doi. org / 10.11646 / zootaxa. 4810.2.10","Motschulsky, V. I. (1860) Coleopteres de la Siberie orientale et en particulier des rives de l'Amour. Schrenk's Reisen und Forschungen im Amurlande, 2, 77 - 257."]}

Platypsyllus castoris Ritsema, 1869 (Fig 1) Material examined. Only adults: 1♂, 2♀♀, Russia: Western Siberia, Altai Krai, Kytmanovsky Distr., env. Sosnovy Log vill., Kamenka River, on beaver, 23 Aug 2019, coll. F. Budaev, IBIW RAS; 2♂♂, 5♀♀, same place, env. Staraya Taraba vill., Balakhcha River, on beaver, 27 July 2020, coll. F. Budaev, BPC. Distribution. The species have Holarctic distribution. In North America species know from Canada, United States (Peck, 2006), in Palaearctic Platypsyllus castoris recorded for Europe (Belgium, Belarus, The Czech Republic, France, Germany, Latvia, The Netherlands, Norway, Poland, Slovakia, Sweden and Switzerland) (Perreau 2015) and the European part of Russia, where the species is known from Voronezh (Prokin & Kirejtshuk 2007), Rostov (Arzanov et al. 2013) and Saratov (Sazhnev et al. 2019) Oblasts., Published as part of SAZHNEV, Alexey S. & BUDAEV, Fyodor A., 2020, First record of Platypsyllus castoris Ritsema, 1869 (Coleoptera: Leiodidae) from Siberia, pp. 125-127 in Fragmenta Faunistica 63 (2) on pages 125-126, DOI: 10.3161/00159301FF2020.63.2.125, http://zenodo.org/record/6252527, {"references":["PECK ST. B. 2006. Distribution and biology of the ectoparasitic beaver beetle Platypsyllus castoris Ritsema in North America (Coleoptera: Leiodidae: Platypsyllinae). Insecta Mundi 20 (1 - 2): 85 - 94.","PROKIN A. A. & KIREJTSHUK A. G. 2007. On Platypsyllus castoris Ristema, 1869 (Coleoptera, Leiodidae, Platypsyllinae) in Voronezh Province and notes on other leiodid beetles connected with mammals. Problems and perspectives of general entomology. Abstracts of the 13 th Congress of Russian Entomological Society: 296 - 297.","ARZANOV YU. G., VALOV G. V. & BAKHTADZE G. B. 2013. Platypsyllus castoris Ritsema, 1869. (Coleoptera: Leiodidae) - new species from Rostov Region (Russia). Caucasian Entomological Bulletin 9 (1): 47 - 49. DOI: 10.23885 / 1814 - 3326 - 2013 - 9 - 1 - 47 - 49","SAZHNEV A. S., VOLODCHENKO A. N. & ZABALUEV I. A. 2019. New data to the fauna of beetles (Coleoptera) of the Saratov Province. Report 2. Eversmannia 57: 9 - 1."]}

The species Berosus geminus Reiche & Saulcy, 1856 (Hydrophilidae) is recorded for the first time for European part of Russia. A map with the general distribution of this species is provided.

Augyles (Augyles) letovi sp. nov. (Figs 1, 2) Type locality. N Vietnam, H��a B��nh Province. Type material. Holotype: ��� 15.01.1976 Bьeтнaм, Xao бинь, Л. H. Meдвeдeв, УФ cвeт, 36��� [15.01.1976 Vietnam, H��a B��nh, L.N. Medvedev, UV light, 36���in Russian], ♂ (ZISP). Paratype: ��� 15.01.1976 Bьeтнaм, Xao бинь, Л. H. Meдвeдeв, УФ cвeт, 33��� [18.01.1976 Vietnam, H��a B��nh, L.N. Medvedev, UV light, 33���in Russian], ♀ (IBIW). Description. Holotype, ♂. Total length 3.2 mm (from apex of labrum to apex of elytra). Body (Fig. 1) moderately elongated, dorsoventrally compressed. Head light brown, with dense whitish pubescence and longer scattered setae; mandibles average, not hypertrophied, with acute apex; eyes well developed, antennae with 10 antennomeres, apical 7 antennomeres forming club; clypeus without pair of anterior horns, with dense and short setae, labrum dark brown. Pronotum light brown, with disc darker, evenly wide or slightly narrower than elytra at base, rectangular, densely punctured, bordered posteriorly, laterally with dense whitish pubescence and longer setae. Elytra with pale brown spots (Fig. 1), densely and uniformely punctate, with dense whitish pubescence; elytral longitudinal ridges only slightly indicated. Setae of elytra short, sparse, semierect. Legs light brown, densely pubescent; tibiae with numerous spines and long setae. Abdomen light brown, stridulatory arch marked; postmetacoxal line present. Aedeagus (0.7 mm long) and spiculum gastrale (V-shaped) as in Fig. 2. Paratype, ♀. Total length 3.3 mm (from apex of labrum to apex of elytra). Externally similar to male, but mandible, head and pronotum smaller. Head and pronotum brown. Variability. Body size 3.2���3.3 mm. Males and females are very difficult to separate without examination of the genitalia. Elytral markings slightly differ in two studied specimens (spots of holotype specimen is more contrasting). Differential diagnosis. Based on the shape of the male genitalia, and shape of the elytral pattern, Augyles letovi sp. nov. is probably most related to the south-east Asian species Augyles rejseki Skalick��, 1999 (described from Thailand) and Augyles hiekei (Mascagni, 1995) (described from Vietnam). A. letovi differs from A. rejseki by the tegmen, having lateral margin evenly curved (not dentate) and possessing paired sclerotized elements in basal part (Fig. 2). A. letovi differs from A. hiekei also by the structure of tegmen presence possessing paired sclerotized elements in basal part. Ecology. All specimens were collected at light. Distribution. So far known only from the type locality. Etymology. Dedicated to Igor Fedorovich ���Yegor��� Letov, Russian poet, musician, singer, songwriter, audio engineer and conceptual art painter, best known as the founder and leader of the post-punk/psychedelic rock band Grazhdanskaya Oborona (Civil Defense)., Published as part of Sazhnev, Alexey S., 2018, A new species of the genus Augyles Schi��dte, 1866 (Coleoptera: Heteroceridae) from Vietnam, pp. 597-599 in Zootaxa 4521 (4) on pages 597-599, DOI: 10.11646/zootaxa.4521.4.9, http://zenodo.org/record/2610192, {"references":["Skalicky, S. (1999) New species of Heteroceridae from Thailand and Namibia (Coleoptera: Heteroceridae). Koleopterologische Rundschau, 69, 119 - 123.","Mascagni, A. (1995) Heteroceridae: Check list of the Heteroceridae of China and neighbouring countries, and description of two new species (Coleoptera). In: Jach M. A. & Ji L. (Eds.). Water Beetles of China. Uol. 1. Zoologisch-Botanische Gesellschaft and Wiener Coleopterologenverein, Wien, pp. 341 - 348"]}

Heterocerus fausti Reitter, 1879 (Figs. 1, 2B, 3) Heterocerus fausti Reitter, 1879: 546 stat. nov. Type locality. Azerbaijan: Baku. Type material examined. Syntypes: ��� Heterocerus Fausti Rtt ��� [Reitter's handwriting], ���Caucasus��� [red label], ���к. А. Яковлева��� [cоllеctiоn оf А. Jаcоvlеv���in Russiаn], 2♂♂ (ZISP); ���Bаku Fаust.���, gоldеn squаrе plаtе, ��� Heterocerus Fausti Reitt. ��� [Reitter's handwriting], 2♀♀ [one specimen without head] (ZISP). Additional material examined. 9 exs. TURKMENISTAN. Balkan Region (Velayat): Uzun-Ada (now the territory of Hazar Nature Reserve), 15.04.1895, leg. Varentsov, 7 exs. (ZISP); Turkmenbashi (Krasnovodsk), 0 6.1900, leg. P. Maksimovitsh, 1 ex. (ZISP); Esenguly, 3.07.1932, unknown collector, 1 ex. (IBIW). Diagnosis. Body size 3.8���5.0 mm. Body moderately elongated, dorsal-ventrally compressed (Fig. 1). Males and females do not differ. Aedeagus as in Fig. 3. Heterocerus fausti is similar in body form and drawing of elytra on Heterocerus parallelus but can be easily distinguished by the shape of its aedeagus (Fig. 4A���B) and smaller size. Redescription. Color. Ground color is yellow or red-yellow; a combination of black-brown blotches on the pronotum, and brown blotches of elytra and abdomen form a characteristic pattern. Head and labrum are black-brown. Antennae completely yellow. Pronotum is black-brown with a more or less wide yellow edging on the sides and narrow yellow stripe along the front edge. Scutellum is black-brown. Drawing of elytra characteristic of flexuosus species-group sensu Charpentier (1965). Yellow pattern of each elytron normally forms edging, S-shaped basal mark, M-shaped medial mark, apical spot and suture stripe. Dark pattern of the elytra brownish almost blending with the yellow background. Prosternum black-brown with yellow episternae and yellowish front margin. Mesepisternum and metepisternum blackbrown. Abdomen black-brown with yellow pattern in form of edging. Legs, except dark metacoxa, yellow. Head. Clypeus transversal, widely incised on the front edge. Antennae 11-segmented with 7-segment club. Hairs whitish. Puncturing uniform. Thorax. Pronotum not bordered basally; slightly narrower or comparable in width with base of elytra in females, and slightly wider than elytra base in males. Hairs on the disc are double: comprise dense, short, pressed, and long and standing hairs; whitish. Puncturing very dense, fine and uniform. Elytra covered with erect hairs; hair length is comparable with the length of few first spines of mesotibia; hairs slightly longer along sides and near the elytral apex; hairs whitish. Legs of structure typical for family, covered with whitish hairs. Claws of tarsus noticeably longer or equal to the last tarsomere (Fig. 2). Abdomen. Abdomen covered with whitish hairs. Postmetacoxal lines absent. Remarks. The records of Heterocerus fausti for Uzbekistan (Jacobson, 1913; Zaitsev, 1946) are not confirmed. Specimen from Uzbekistan (Qamashi) signed of F.A. Zaitsev as ��� Heterocerus fausti ��� is Heterocerus heydeni Kuwert, 1890. Ecology. The long claws are probably an adaptation to the psammophilous way of life. Distribution. Pre-Caspian Region including Caucasus: Azerbaijan, West Turkmenistan (Fig. 5)., Published as part of Sazhnev, Alexey S., 2018, Redescription of Heterocerus fausti Reitter, 1879, bona species (Coleoptera, Heteroceridae), pp. 597-600 in Zootaxa 4441 (3) on pages 597-600, DOI: 10.11646/zootaxa.4441.3.12, http://zenodo.org/record/1301560, {"references":["Reitter, E. (1879) Neue Coleopteren aus dem sudostlichen Russland aufgefunden von Herrn J. Faust aus Helsingfors. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 29, 543 - 546. [in German]","Charpentier, R. (1965) A monograph of the family Heteroceridae of the Ethiopian Region. In: South African Animal Life. II. Result of the Lund University expedition in 1950 - 1951. Vol. XI. Swedish Natural Science Research Council, Stockholm, VA, pp. 215 - 343.","Jacobson, G. G. (1913) Semeystvo Heteroceridae. Pilousy. In: Zhuki Rossii i zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov. Vol. 10. A. F. Davrien, Sankt-Peterburg, pp. 867 - 869. [in Russian, English translation in: Family Heteroceridae. Mud-loving beetles. Beetles of Russia, Western Europe and neighbouring countries. Guide to the determination of beetles. A. F. Devrien, St. - Petersburg, 10, pp. 867 - 869]","Zaitsev, F. A. (1946) Rasprostranenie v Zakavkaz'e vidov sem. Pilousov (Coleoptera, Heteroceridae). Trudy Zoologicheskogo instituta AN GruzSSR, 6, 213 - 220. [in Russian, English translation in: Distribution in Transcaucasia species of the variegated mud-loving beetles (Coleoptera, Heteroceridae). Works of Zoological Institute of the Georgian Academy of Sciences, 6, 213 - 220]"]}

Genus Allopygmephorus Cross, 1965 Type species: Pygmephorus matthesi Krczal, 1959, by original designation., Published as part of Khaustov, Alexander A. & Sazhnev, Alexey S., 2016, Mites of the families Neopygmephoridae and Scutacaridae associated with variegated mud-loving beetles (Coleoptera: Heteroceridae) from Russia and Kazakhstan, pp. 261-273 in Zootaxa 4175 (3) on page 264, DOI: 10.11646/zootaxa.4175.3.5, http://zenodo.org/record/265766, {"references":["Cross, E. A. (1965) The generic relationships of the family Pyemotidae (Acarina, Trombidiformes). The University of Kansas Science Bulletin, 45, 29 - 215.","Krczal, H. (1959) Systematik und Okologie der Pyemotiden. Beitrage zur Systematik und Okologie mitteleuropaischer Acarina, 1, 385 - 625."]}

Genus Scutacarus Gros, 1845 Type species: Scutacarus femoris Gros, 1845, by monotypy.

Scutacarus sphaeroideus Karafiat, 1959 Scutacarus longitarsus sphaeroideus Karafiat, 1959: 679, fig. 23. This cosmopolitan species inhabits wet soil and is phoretic on the staphylinid beetles Stenus ruralis Erichson, S. bipunctatus Erichson and Carpelimus fuliginosus (Gravenhorst) (Karafiat 1959; Khaustov 2008). During this study we recorded phoresy of S. sphaeroideus on Heteroceridae for the first time. Material examined. One female, KAZAKHSTAN: Aktyubinsk Province, Bezhetsk region, vicinity of settlement Emba, N 48��38'38.7���, E 057��54'58���, on Heterocerus fenestratus, 25 April 2012, coll. A.V. Ivanov; four females, same locality and date, on H. flexuosus; 1 female RUSSIA: Astrakhan Province, Yenotayevsk region, settlement Zamyany, in light trap, on H. fenestratus, 16 September 2007, coll. О.G. Brekhov; one female, same locality and date, on Augyles hispidulus., Published as part of Khaustov, Alexander A. & Sazhnev, Alexey S., 2016, Mites of the families Neopygmephoridae and Scutacaridae associated with variegated mud-loving beetles (Coleoptera: Heteroceridae) from Russia and Kazakhstan, pp. 261-273 in Zootaxa 4175 (3) on page 272, DOI: 10.11646/zootaxa.4175.3.5, http://zenodo.org/record/265766, {"references":["Karafiat, H. (1959) Systematik und Okologie der Scutacariden. Beitrage zur Systematik und Okologie mitteleuropaischen Acarina, 1, 627 - 712.","Khaustov, A. A. (2008) Mites of the family Scutacaridae of Eastern Palaearctic. Akademperiodyka, Kiev, 291 pp."]}

Allopygmephorus punctatus sp. nov. (Figs 7–9) Description. FEMALE (Figs 7–9). Body well sclerotized. Length of idiosoma 270 (270–280), width 170 (170– 190). Gnathosoma similar to that of P. heteroceri sp. nov. Idiosomal dorsum (Fig. 7 A). Only posterior part of prodorsum covered by anterior margin of tergite C. All dorsal plates with numerous small dimples (punctate). Stigmata round, with transverse slit-like opening. Trichobothria clavate and weakly barbed. Setae sc 2 smooth, needle-like; other dorsal setae sparsely barbed; setae e and h 2 blunt-ended, other dorsal setae pointed. Posterior margin of tergite H rounded. Cupules ia on tergite D and ih on tergite H small, round. Lengths of dorsal setae: sc 2 19 (18–22), c 1 42 (42–45), c 2 51 (50–54), d 42 (40–44), e 26 (26–27), f 49 (48–52), h 1 43 (42–49), h 2 31 (30–33). Distances between setae: sc 2 –sc 2 54 (53–56), c 1– c 1 52 (52–54), c 1– c 2 39 (38–44), d–d 37 (36–39), e–f 17 (16–17), f– f 74 (72–76), h 1– h 1 33 (32–35), h 1– h 2 23 (23–25). Idiosomal venter (Fig. 7 B). All ventral plates with numerous small dimples. Setae 1 b thichened, lanceolate and barbed. Setae ps 2 very small, vestigial; other ventral setae sparsely barbed, pointed. Setae ps 1 and ps 3 weakly thickened in basal part. Apodemes as in A. spinisetus sp. nov. Posterior margin of posterior sternal plate weakly concave in middle part. Posterior margin of aggenital plate rounded. Ags small, triangular, pgs large, triangular, with indistinct anterior margin. Posterior sternal plate with distinct scale-like structure posteriorly and laterally to bases of legs IV. Lengths of ventral setae: 1 a 34 (29–35), 1 b 26 (25–27), 2 a 30 (30–33), 2 b 30 (28–31), 3 a 35 (33–36), 3 b 32 (30–33), 3 c 26 (26–29), 4 a 43(40–45), 4 b 49 (47– 50), 4 c 40 (39–42), ps 1 26 (25–27), ps 3 24 (24–27). Legs (Figs 8, 9). Leg setation as in P. heteroceri sp. nov. Leg I (Fig. 8 A). Tip of tarsal claw thin, pointed. Seta k smooth, pointed. Seta l’ of femur smooth, blunt-ended; v” on femur I very short, smooth, spiniform; v’ on trochanter short, smooth, blunt-ended; other leg setae (except eupathidia) pointed and sparsely barbed. Lengths of solenidia ω 1 9 (9–10)> ω 2 4 (4–5) < φ 1 7 (6–7) = φ 2 7 (6–7); ω 1 finger-shaped, ω 2 and φ 2 baculiform, φ 1 weakly clavate. Leg II (Fig. 8 B). Solenidion ω 13 (13–15), finger-shaped, solenidion φ 5 (5–6) weakly clavate. Seta tc” of tarsus smooth, pointed; u’ weakly blunt-ended, barbed; other leg setae pointed and sparsely barbed. Leg III (Fig. 9 A) Solenidion φ 3 (3–4) weakly clavate. Seta tc” of tarsus smooth, pointed; u’ weakly blunt-ended, barbed; other leg setae pointed and sparsely barbed. Leg IV (Fig. 9 B). Solenidion φ 2 (2) very small. All leg setae barbed; seta v” of tibia blunt-ended, other leg setae pointed. MALE and LARVA unknown. Type material. Female holotype, slide No. 270715, RUSSIA: Dagestan, Biryuzyak station, on Heterocerus fenestratus, 27 July 2015, coll. V.V. Ilyina; paratypes: 2 females, same data; 2 females, KAZAKHSTAN, Aktyubinsk Province, Bezhetsk region, vicinity of settlement Emba, N 48°38'38.7”, E 057°54'58”, on Heterocerus flexuosus, 25 April 2012, coll. A.V. Ivanov. Etymology. The name of the new species is derived from Latin word punct meaning dot referring to punctate dorsal and ventral plates. Differential diagnosis. The new species is most similar to A. persicus Khaustov and Hajiqanbar, 2006, in having pointed setae h 1. It differs from A. persicus by its blunt-ended setae h 2 (pointed in A. persicus); setae 1 b thickened and lanceolate (not thickened in A. persicus), and distinctly punctate dorsal and ventral plates (smooth in A. persicus).

Khaustov, Alexander A., Sazhnev, Alexey S. (2016): Mites of the families Neopygmephoridae and Scutacaridae associated with variegated mud-loving beetles (Coleoptera: Heteroceridae) from Russia and Kazakhstan. Zootaxa 4175 (3): 261-273, DOI: http://doi.org/10.11646/zootaxa.4175.3.5

Allopygmephorus spinisetus sp. nov. (Figs 4���6) Description. FEMALE (Figs 4���6). Body well sclerotized. Length of idiosoma 240 (220���270), width 170 (165��� 180). Gnathosoma similar to that of P. heteroceri sp. nov. Idiosomal dorsum (Fig. 4 A). Only posterior part of prodorsum covered by anterior margin of tergite C. All dorsal plates with numerous small dimples (punctate). Stigmata round, with transverse slit-like opening. Trichobothria clavate and weakly barbed. Setae sc 2 smooth, needle-like; setae c 1 and c 2 smooth or with 1-2 small barbs, pointed; setae d and f sparsely barbed, pointed; other dorsal setae blunt-ended and sparsely barbed. Posterior margin of tergite H rounded. Cupules ia on tergite D and ih on tergite H small, round. Tergite C with one pair of pore-like structures situated anteromedially to c 1. Lengths of dorsal setae: sc 2 15 (15���16), c 1 37 (36���43), c 2 44 (43���46), d 34 (34���39), e 22 (21���23), f 42 (41���44), h 1 28 (27���31), h 2 28 (27���29). Distances between setae: sc 2 ���sc 2 53 (53���54), c 1��� c 1 61 (59���63), c 1��� c 2 37 (36���38), d���d 26 (26���36), e��� f 17 (14���17), f���f 68 (66���73), h 1��� h 1 28 (27���32), h 1��� h 2 18 (17���19). Idiosomal venter (Fig. 4 B). All ventral plates with numerous small dimples. Setae 1 b thichened, lanceolate and barbed. Setae ps 2 very small, vestigial; other ventral setae sparsely barbed, pointed. Setae ps 1 and ps 3 weakly thickened in basal part. Ap1 well-developed and joined with appr; ap2 well-developed, joined with appr; appr and apsej well-developed; ap3 well-developed, straight. Ap4 well sclerotized, long and joined with appo; apodemes 5 absent. Posterior margin of posterior sternal plate weakly concave in middle part. Posterior margin of aggenital plate rounded. Ags small, triangular, pgs large, triangular. Posterior sternal plate with distinct scale-like structure posteriorly and laterally to bases of legs IV. Lengths of ventral setae: 1 a 29 (28���30), 1 b 27 (25���28), 2 a 32 (31���34), 2 b 29 (28���32), 3 a 33 (32���35), 3 b 37 (35���38), 3 c 32 (30���33), 4 a 47 (45���49), 4 b 56 (53���58), 4 c 40 (38���42), ps 1 27 (25���28), ps 3 26 (24���28). Legs (Figs 5, 6). Leg setation as in P. heteroceri sp. nov. Leg I (Fig. 5 A). Tip of tarsal claw thin, pointed. Seta k smooth, pointed. Seta l��� of femur smooth, blunt-ended; v��� of femur I smooth, spiniform; other leg setae (except eupathidia) pointed and sparsely barbed. Lengths of solenidia �� 1 10 (10���11)> �� 2 4 (4) < �� 1 7 (7���8)> �� 2 6 (6���7); �� 1 finger-shaped, �� 2 and �� 2 baculiform, �� 1 weakly clavate. Leg II (Fig. 5 B). Solenidion �� 13 (13���15), finger-shaped, solenidion �� 6 (5���6) weakly clavate. Seta v��� of femur spiniform, smooth, tc��� of tarsus smooth, pointed; u��� weakly blunt-ended, barbed; other leg setae pointed and sparsely barbed. Leg III (Fig. 6 A) Solenidion �� 5 (4���5) weakly clavate. Setae pv���, tc��� and tc��� of tarsus smooth, pointed; u��� weakly blunt-ended, barbed; other leg setae pointed and sparsely barbed. Leg IV (Fig. 6 B). Solenidion �� 2 (2) very small. Setae u��� smooth; other leg setae barbed; setae v��� and v��� of tibia bluntended, other leg setae pointed. MALE and LARVA unknown. Type material. Female holotype, slide No. 250703 /1, RUSSIA: Moscow Province, Zverinogolovsk region, vicinity of settlement Kubinka, on Heterocerus fenestratus, 25 July 2003, coll. I.M. Vyshinsky; paratypes: nine females, same data; one female, Tver Province, Bezhetsk region, vicinity of settlement Drutskovo, on Heterocerus fenestratus, 2 September 2004, coll. I.M. Vyshinsky. Etymology. The name of the new species is combined from Latin words spina meaning spine and seta referring to presence of spiniform seta v��� on femur II. Differential diagnosis. The new species is most similar to A. matthesi (Krczal, 1959), by having blunt-ended setae e, h 1, h 2, and pointed f. It differs from A. matthesi by its spiniform seta v��� on femur II (not modified in A. matthesi), and setae 1 b thickened and lanceolate (not thickened in A. matthesi)., Published as part of Khaustov, Alexander A. & Sazhnev, Alexey S., 2016, Mites of the families Neopygmephoridae and Scutacaridae associated with variegated mud-loving beetles (Coleoptera: Heteroceridae) from Russia and Kazakhstan, pp. 261-273 in Zootaxa 4175 (3) on pages 264-267, DOI: 10.11646/zootaxa.4175.3.5, http://zenodo.org/record/265766, {"references":["Krczal, H. (1959) Systematik und Okologie der Pyemotiden. Beitrage zur Systematik und Okologie mitteleuropaischer Acarina, 1, 385 - 625."]}

Genus Protoallopygmephorus gen. nov. Type species: Protoallopygmephorus heteroceri sp. nov. Diagnosis. FEMALE. Body oval. Only the posterior part of the prodorsum covered by tergite C. Gnathosomal capsule oval, its length about equal to its width, dorsally with two pairs of cheliceral setae (cha, chb) and postpalpal setae situated anteriorly to setae cha; palps prominent, with two pairs of dorsolateral setae (dFe, dGe) and distinct tibial claw distally. Subcapitulum with one pair of setae m and one pair of oval pits in the posterior half. Palps ventrally with mushroom-like accessory setigenous structure and small solenidion. Prodorsum with two pair of simple setae (v 2, sc 2), one pair of clavate trichobothria (sc 1), and circular stigmata. Dorsal hysterosomal setae not modified. Tergite C with two pairs of setae (c 1, c 2); tergite D with one pair of setae (d) and one pair of round cupules ia; tergite EF with two pairs of setae (e, f); tergite H with two pairs of setae (h 1, h 2) and one pair of round cupules ih. Coxal fields I with two pairs of setae (1 a, 1 b); coxal fields II with two pairs of setae (2 a, 2 b); coxal fields III with three pairs of setae (3 a, 3 b, 3 c); coxal fields IV with three pairs of setae (4 a, 4 b, 4 c). Pseudanal segment with three pairs of setae (ps 1-3). Apodemes 1 (ap1) well-developed, apodemes 2 (ap2) well-developed and fused with welldeveloped prosternal apodeme (appr); appr joined with sejugal apodeme (apsej); apodemes 3 (ap3) straight, diffuse, apodemes 4 (ap4) long, reaching to bases of legs III. Apodemes 5 absent. Secondary transverse apodeme absent. Posterior margin of posterior sternal plate entire. Anterior genital sclerite (ags) small, triangular, posterior genital sclerite (pgs) large, triangular, median genital sclerite (mgs) present, oval. Ventral idiosomal setae long, sparsely barbed. Legs I subequal to leg II. Tibiotarsus I slightly enlarged, with well-developed claw; unguinal setae (u) spiniform, separated, solenidion �� 1 fused with tibiotarsus. Seta d of femur hook-like. Legs II���III with one pair of basally thickened, slightly asymmetrical claws and pointed distally empodium; claws on tarsus IV simple, as large as those on tarsi II and III, empodium pointed, but smaller than on tarsi II-III. Tarsus and pretarsus IV short. Femora III���IV divided into basi- and telofemur. Leg setation: leg I; tr 1 (v���), fe 3 (d, l���, v���), ge 4 (l���, l���, v���, v���), tita 18(4) (d, l���, l���, v���, v���, k, tc���, tc���, p���, ft���, ft���, pl���, pl���, pv���, pv���, s, u���, u���, �� 1, �� 2, �� 1, �� 2); leg II: tr 1 (v���), fe 3 (d, l���, v���), ge 3 (l���, l���, v ���), ti 4(1) (d, l���, v���, v���, ��), ta 6(1) (tc���, tc���, pl���, pv���, pv���, u���, ��); leg III: tr 1 (v���), fe 2 (d, v���), ge 2 (l���, v���), ti 4(1) (d, l���, v���, v���, ��), ta 6 (tc���, tc���, pl���, pv���, pv���, u���); leg IV: tr 1 (v���), fe 2 (d, v���), ge 1 (v���), ti 4(1) (d, l���, v���, v���, ��), ta 6 (pl���, tc���, tc���, u���, pv���, pv���). Differential diagnosis. The new genus is most similar to Allopygmephorus Cross, 1965. Both genera have characteristic pointed empodia on tarsi II-IV, separated spiniform unguinal setae on tibiotarsus I, the same leg setation, and similarly shaped claws on the legs. The new genus differs from Allopygmephorus by the presence of setae v 2 (absent in Allopygmephorus), the presence of the median genital sclerite (absent in Allopygmephorus), and by solenidion �� 1 fused with tibiotarsus I (prominent in Allopygmephorus). Among neopygmephorid genera with median genital sclerite, the new genus is most similar to Pseudopygmephorus Cross, 1965, by having solenidion �� 1fused with tibiotarsus I. The new genus differs from Pseudopygmephorus by the characteristic distally-pointed empodia on tarsi II���IV (flipper-like in Pseudopygmephorus), and non-bifurcate setae 1 b (bifurcate in Pseudopygmephorus). Species included. The genus Protoallopygmephorus gen. nov. includes only type species, P. heteroceri sp. nov. Distribution and habitat. Protoallopygmephorus heteroceri sp. nov. is phoretic on Heterocerus fenestratus (Thunberg) (Coleoptera: Heteroceridae) in the European part of Russia. Etymology. The name of the new genus is combined from Greek ���������� meaning first, and the name of closely related genus Allopygmephorus. It refers to its similarity to the genus Allopygmephorus., Published as part of Khaustov, Alexander A. & Sazhnev, Alexey S., 2016, Mites of the families Neopygmephoridae and Scutacaridae associated with variegated mud-loving beetles (Coleoptera: Heteroceridae) from Russia and Kazakhstan, pp. 261-273 in Zootaxa 4175 (3) on page 262, DOI: 10.11646/zootaxa.4175.3.5, http://zenodo.org/record/265766, {"references":["Cross, E. A. (1965) The generic relationships of the family Pyemotidae (Acarina, Trombidiformes). The University of Kansas Science Bulletin, 45, 29 - 215."]}

Allopygmephorus punctatus sp. nov. (Figs 7���9) Description. FEMALE (Figs 7���9). Body well sclerotized. Length of idiosoma 270 (270���280), width 170 (170��� 190). Gnathosoma similar to that of P. heteroceri sp. nov. Idiosomal dorsum (Fig. 7 A). Only posterior part of prodorsum covered by anterior margin of tergite C. All dorsal plates with numerous small dimples (punctate). Stigmata round, with transverse slit-like opening. Trichobothria clavate and weakly barbed. Setae sc 2 smooth, needle-like; other dorsal setae sparsely barbed; setae e and h 2 blunt-ended, other dorsal setae pointed. Posterior margin of tergite H rounded. Cupules ia on tergite D and ih on tergite H small, round. Lengths of dorsal setae: sc 2 19 (18���22), c 1 42 (42���45), c 2 51 (50���54), d 42 (40���44), e 26 (26���27), f 49 (48���52), h 1 43 (42���49), h 2 31 (30���33). Distances between setae: sc 2 ���sc 2 54 (53���56), c 1��� c 1 52 (52���54), c 1��� c 2 39 (38���44), d���d 37 (36���39), e���f 17 (16���17), f��� f 74 (72���76), h 1��� h 1 33 (32���35), h 1��� h 2 23 (23���25). Idiosomal venter (Fig. 7 B). All ventral plates with numerous small dimples. Setae 1 b thichened, lanceolate and barbed. Setae ps 2 very small, vestigial; other ventral setae sparsely barbed, pointed. Setae ps 1 and ps 3 weakly thickened in basal part. Apodemes as in A. spinisetus sp. nov. Posterior margin of posterior sternal plate weakly concave in middle part. Posterior margin of aggenital plate rounded. Ags small, triangular, pgs large, triangular, with indistinct anterior margin. Posterior sternal plate with distinct scale-like structure posteriorly and laterally to bases of legs IV. Lengths of ventral setae: 1 a 34 (29���35), 1 b 26 (25���27), 2 a 30 (30���33), 2 b 30 (28���31), 3 a 35 (33���36), 3 b 32 (30���33), 3 c 26 (26���29), 4 a 43(40���45), 4 b 49 (47��� 50), 4 c 40 (39���42), ps 1 26 (25���27), ps 3 24 (24���27). Legs (Figs 8, 9). Leg setation as in P. heteroceri sp. nov. Leg I (Fig. 8 A). Tip of tarsal claw thin, pointed. Seta k smooth, pointed. Seta l��� of femur smooth, blunt-ended; v��� on femur I very short, smooth, spiniform; v��� on trochanter short, smooth, blunt-ended; other leg setae (except eupathidia) pointed and sparsely barbed. Lengths of solenidia �� 1 9 (9���10)> �� 2 4 (4���5) < �� 1 7 (6���7) = �� 2 7 (6���7); �� 1 finger-shaped, �� 2 and �� 2 baculiform, �� 1 weakly clavate. Leg II (Fig. 8 B). Solenidion �� 13 (13���15), finger-shaped, solenidion �� 5 (5���6) weakly clavate. Seta tc��� of tarsus smooth, pointed; u��� weakly blunt-ended, barbed; other leg setae pointed and sparsely barbed. Leg III (Fig. 9 A) Solenidion �� 3 (3���4) weakly clavate. Seta tc��� of tarsus smooth, pointed; u��� weakly blunt-ended, barbed; other leg setae pointed and sparsely barbed. Leg IV (Fig. 9 B). Solenidion �� 2 (2) very small. All leg setae barbed; seta v��� of tibia blunt-ended, other leg setae pointed. MALE and LARVA unknown. Type material. Female holotype, slide No. 270715, RUSSIA: Dagestan, Biryuzyak station, on Heterocerus fenestratus, 27 July 2015, coll. V.V. Ilyina; paratypes: 2 females, same data; 2 females, KAZAKHSTAN, Aktyubinsk Province, Bezhetsk region, vicinity of settlement Emba, N 48��38'38.7���, E 057��54'58���, on Heterocerus flexuosus, 25 April 2012, coll. A.V. Ivanov. Etymology. The name of the new species is derived from Latin word punct meaning dot referring to punctate dorsal and ventral plates. Differential diagnosis. The new species is most similar to A. persicus Khaustov and Hajiqanbar, 2006, in having pointed setae h 1. It differs from A. persicus by its blunt-ended setae h 2 (pointed in A. persicus); setae 1 b thickened and lanceolate (not thickened in A. persicus), and distinctly punctate dorsal and ventral plates (smooth in A. persicus)., Published as part of Khaustov, Alexander A. & Sazhnev, Alexey S., 2016, Mites of the families Neopygmephoridae and Scutacaridae associated with variegated mud-loving beetles (Coleoptera: Heteroceridae) from Russia and Kazakhstan, pp. 261-273 in Zootaxa 4175 (3) on pages 269-270, DOI: 10.11646/zootaxa.4175.3.5, http://zenodo.org/record/265766, {"references":["Khaustov, A. A. & Hajiqanbar, H. R. (2006) Redefinition of the genus Allopygmephorus (Acari: Heterostigmata: Neopygmephoridae), with description of a new species from Iran. Acarina, 14, 191 - 194."]}

Genus Scutacarus Gros, 1845 Type species: Scutacarus femoris Gros, 1845, by monotypy., Published as part of Khaustov, Alexander A. & Sazhnev, Alexey S., 2016, Mites of the families Neopygmephoridae and Scutacaridae associated with variegated mud-loving beetles (Coleoptera: Heteroceridae) from Russia and Kazakhstan, pp. 261-273 in Zootaxa 4175 (3) on page 272, DOI: 10.11646/zootaxa.4175.3.5, http://zenodo.org/record/265766, {"references":["Gros, D. (1845) Des Acares. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 1, 397 - 416."]}

Protoallopygmephorus heteroceri sp. nov. (Figs. 1���3) Description. FEMALE (holotype, Figs. 1���3). Body well sclerotized. Length of idiosoma 225 (200���230), width 160 (140���160). Gnathosoma. Gnathosomal capsule semicircular, subequal in length and width, dorsally with two pairs of sparsely barbed, subequal pointed cheliceral setae (cha, chb). Dorsal median apodeme well-developed. Postpalpal setae needle-like. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae m. Palps freely articulated to gnathosomal capsule, with smooth, subequal pointed setae dFe and dGe dorsolaterally. Palps ventrally with large accessory setigenous structure (ass) and small solenidion. Palps with a small terminal claw. Pharyngeal pumps II and III grouped together. Pharyngeal pump I small; butterfly-like, situated inside gnathosomal capsule; pharyngeal pump II large, about double length of pharyngeal pump III. Lengths of gnathosomal setae: cha 20 (19���20), chb 22 (21���22), m 14 (13���14). Idiosomal dorsum (Fig. 1 A). Trichobothria (sc 1) clavate, with minute barbs. All dorsal plates with numerous small dimples (punctate). Setae v 2 smooth, other dorsal setae sparsely barbed; setae sc 2, c 1 and c 2 pointed; v 2 needle-like; other dorsal setae blunt-ended. Posterior margin of tergite H weakly undulate. Cupules ia on tergite D and ih on tergite H small, round. Lengths of dorsal setae: v 2 6 (6���7), sc 2 31 (26���32), c 1 47 (42���49), c 2 58 (54���60), d 33 (32���34), e 30 (26���30), f 46 (42���47), h 1 54 (46���55), h 2 34 (30���35). Distances between setae: v 2 ���v 2 43 (41���44), sc 2 ���sc 2 50 (47���52), c 1��� c 1 70 (66���71), c 1��� c 2 29 (25���30), d���d 75 (72���77), e���f 13 (12���13), f���f 67 (57���69), h 1��� h 1 28 (20���28), h 1��� h 2 16 (15���18). Idiosomal venter (Fig. 1 B). All ventral plates with numerous small dimples. Setae 1 b widened, lanceolate, sparsely barbed, ps 2 smooth, other ventral setae barbed, pointed. Posterior margin of posterior sternal plate slightly convex in middle part. Posterior margin of aggenital plate rounded. Lengths of ventral setae: 1 a 36 (34���37), 1 b 27 (26���28), 2 a 36 (34���37), 2 b 32 (30���33), 3 a 33 (31���33), 3 b 36 (33���36), 3 c 32 (26���33), 4 a 38 (31���38), 4 b 45 (39���46), 4 c 38 (32���39), ps 1 29 (28���30), ps 2 12 (9��� 13), ps 3 20 (17���21). Legs (Figs 2, 3). Leg I. Tip of tarsal claw thin, pointed. Seta k smooth, pointed. Seta l��� of femur smooth, blunt-ended; v��� of femur I smooth, spiniform; other leg setae (except eupathidia) pointed and sparsely barbed. Lengths of solenidia �� 1 9 (9���10) < �� 2 12 (11���12)> �� 1 6 (6���7) = �� 2 7 (6���7); �� 2 and �� 2 baculiform, �� 1 weakly clavate. Leg II (Fig. 2 B). Solenidion �� 14 (14���15), finger-shaped, solenidion �� 5 (4���5) weakly clavate. Seta v��� of femur and tc��� of tarsus smooth, pointed; u��� weakly blunt-ended, barbed; other leg setae pointed and sparsely barbed. Leg III (Fig. 3 A) Solenidion �� 4 (4���5) weakly clavate. Setae pv��� and tc��� of tarsus smooth, pointed; u��� weakly blunt-ended, barbed; other leg setae pointed and sparsely barbed. Leg IV (Fig. 3 B). Solenidion �� 2 (2) very small. All leg setae barbed; setae d of femur and v��� of tibia blunt-ended, other leg setae pointed. MALE and LARVA unknown. Type material. Female holotype, slide No. 250703, RUSSIA: Moscow Province, Zverinogolovskiy vicinity of settlement Kubinka, on Heterocerus fenestratus, 25 July 2003, coll. I.M. Vyshinsky; paratypes: seven females, same data; five females, Tver Province, Bezhetsk region, vicinity of settlement Drutskovo, on Heterocerus fenestratus, 2 September 2004, coll. I.M. Vyshinsky. Etymology. The specific name is derived from Latin name of the genus Heterocerus, the phoretic host of the new species., Published as part of Khaustov, Alexander A. & Sazhnev, Alexey S., 2016, Mites of the families Neopygmephoridae and Scutacaridae associated with variegated mud-loving beetles (Coleoptera: Heteroceridae) from Russia and Kazakhstan, pp. 261-273 in Zootaxa 4175 (3) on pages 262-264, DOI: 10.11646/zootaxa.4175.3.5, http://zenodo.org/record/265766