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This paper presents unique comprehensive quantitative measures of the changes in Spanish rural landscapes, specifically in relation to the Rural-Urban Interface (RUI). Our objective was to investigate the urbanisation process in rural areas of Spain, thus the expansion of the RUI in an environmentally complex study area (it includes arid regions, continental areas or high mountains). The data were produced by the Spanish Rural Landscapes Monitoring System project (SISPARES), which has carried out five separate surveys for 1956, 1984, 1998, 2008 and 2018. SISPARES data provide RUI quantification by identifying urban patches, using aerial images, in a permanent network of 206 stratified random landscape sampling units (4 × 4 km each) deliberately located in rural areas. The cost-effective and spatio-temporal SISPARES approach enables the production of landscape geographical models at each survey date and the modelling of its evolution over each time period. We hypothesised that the RUI expansion did not follow a homogeneous spatio-temporal pattern, and therefore neither did the drivers responsible for it. Building on SISPARES data, we evaluated through CART analyses the relationship of RUI expansion with relevant environmental and anthropogenic driving factors, such as climate, bedrock, landscape composition and human population density. The historical narrative perspective was also used to aid discussion of the results. Historically, Spain contained few areas of RUI, but major changes were determined in the initial study period (1956-1984), which conformed a seminal geographic pattern for subsequent RUI increment. In contrast, the RUI expansion did not increase greatly during the following periods, between 1984 and 2018. The RUI expansion has been primarily driven by anthropogenic factors, constrained by the environmental characteristics of Spain. The expansion pattern is likely to has shifted from a national scale in the 1950s, to a continental scale at later dates as the result of the changes in socio-political scenarios., Competing Interests: Declaration of competing interest The authors declare that they have no known competing financial interests or personal relationships that could have appeared to influence the work reported in this paper., (Copyright © 2020 Elsevier B.V. All rights reserved.)

Semi-natural grasslands harbour high biodiversity and play a key role in the supply of ecosystem services (ES). However, abandonment, changes in traditional management practices and agricultural intensification constitute a major threat to these grasslands worldwide and these practices have led to declines in species diversity. In this paper the multi-functionality of semi natural-grasslands is assessed from the ES perspective, within a range of common semi-natural grassland types throughout Estonia. The analysis follows a stepwise approach based on the ES cascade model. Firstly, analyses of the relationships between plant species distribution patterns and environmental factors are described. Secondly, the effect of grassland abandonment on plant species diversity, as well as on the presence of rare and protected plant species is tested. In order to overcome the lack of data on ES at the national scale, plant species diversity and soil organic carbon are tested as surrogate indicators for five ESS: pollination, herbs for traditional medicinal use, nutrient cycling, nutrient retention and biomass production. In the final step, the spatial distribution of ES is assessed, based on an ES hotspots map obtained by detecting areas where high levels of plant species diversity and soil organic carbon overlap. The results show that the majority of ES hotspots are present in wooded meadows and pastures. However, there is an important threat to these hotspots because 45% are not eligible for agri-environmental support., (Copyright © 2019 Elsevier B.V. All rights reserved.)