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Urban greening is attracting considerable research and practical attention for its contributions to conserving biodiversity, mitigating urban heat and enhancing the liveability and sustainability of cities and urban areas. Much of the urban greening research is concentrated on major cities, with little focus on the needs and experiences with urban greening in regional cities and towns. This paper addresses this by presenting a case study of an urban greening project, 'Naturally Cooler Towns', focused on regional towns in north-east Victoria, Australia. The project was undertaken by the Goulburn Murray Climate Alliance, a regional alliance of local governments, catchment management authorities and a state government department. As the region faces increasing temperatures and impacts of more frequent heatwaves, the project aimed to review tree management practices and identify opportunities for increasing canopy cover with appropriate species for 'climate adapted street trees'. We examine how urban greening is planned and implemented in these towns, and the roles of the regional alliance in providing a forum for collaborative adaptive governance. The regional alliance plays a key role as an intermediary facilitating approaches that demonstrate situatedness: credibility, salience and legitimacy. The paper contributes towards increased understandings of urban greening approaches beyond the major cities. [ABSTRACT FROM AUTHOR]