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The fusion of a low-spatial-resolution hyperspectral image (HSI) (LR-HSI) with its corresponding high-spatial-resolution multispectral image (MSI) (HR-MSI) to reconstruct a high-spatial-resolution HSI (HR-HSI) has been a significant subject in recent years. Nevertheless, it is still difficult to achieve the cross-mode information fusion of spatial mode and spectral mode when reconstructing HR-HSI for the existing methods. In this article, based on a convolutional neural network (CNN), an interpretable spatial–spectral reconstruction network (SSR-NET) is proposed for more efficient HSI and MSI fusion. More specifically, the proposed SSR-NET is a physical straightforward model that consists of three components: 1) cross-mode message inserting (CMMI); this operation can produce the preliminary fused HR-HSI, preserving the most valuable information of LR-HSI and HR-MSI; 2) spatial reconstruction network (SpatRN); the SpatRN concentrates on reconstructing the lost spatial information of LR-HSI with the guidance of spatial edge loss (Lspat); and 3) spectral reconstruction network (SpecRN); the SpecRN pays attention to reconstruct the lost spectral information of HR-MSI under the constraint of spatial edge loss ({Lspec). Comparative experiments are conducted on six HSI data sets of Urban, Pavia University (PU), Pavia Center (PC), Botswana, Indian Pines (IP), and Washington DC Mall (WDCM), and the proposed SSR-NET achieves the superior or competitive results in comparison with seven state-of-the-art methods. The code of SSR-NET is available at https://github.com/hw2hwei/SSRNET. [ABSTRACT FROM AUTHOR]