Many and various methods have been employed at different times for the measurement of light in ecological habitats, and these methods have already been critically reviewed by the writer ('25). In the same paper a new method and instrument were set forth, and later ('27) a land model of this instrument was described, with illustrations. These instruments, which have hitherto had no name, except that of ecological photometer, and which I now refer to, for the sake of brevity, as the ecophotomteter, aquatic model, and ecophotometer, land model, have been employed in making many measurements, and have continued to perform satisfactorily. The land model has been used, not only in various habitats, but in measuring the light in a single habitat at sunset, in the gloaming, in moonlight, and in starlight. This instrument has also been calibrated in terms of total light energy, as measured by a Moll-Richard-Gorczynski pyrheliometer, and the accompanying graph (Fig. I) gives the results of this comparison. It will be seen that a practically " straight-line " relationship exists for full sunlight, for faint sunlight, and for the light from a " cloudy-bright " sky, but that the slope of the lines is different for the different light conditions. This is exactly what was to be expected, because in bright sunlight the red and yellow rays, which have the greatest heating effect, predominate, while they are less in the case of faint sunlight (that is, sunlight which just casts a faint shadow), and least in the light from an overcast, but bright, sky. The comparison for the light from a dull sky could not be made, because the pyrheliometer did not respond at all to such low intensities. It is thus clear that the " photic effect," as distinct from the " thermic effect," is measured far more efficiently by the panchromatic plate than by the pyrheliometer, as the shorter wave-lengths (which include the near ultra-violet, violet and blue) which have a decided influence on many important processes in plants and animals, are adequately recorded. In connection with this graph it should be stated that: (I) full noon sunlight, on July 12th, latitude 450 north, sea level, was taken as standard (ioo per cent) ; (2) that the photometer was in all cases inclined so as to be at rioht angles to the incident radiation, so as to be in the same position as the receiving disc of the pyrheliometer; (3) that two batches of panchromatic plates were used; and (4) that each plate was developed separately. The last two facts are mentioned because they show that no