IEMIZONIA IS A genus of the tribe Madiinae of the family Compositae. Commonly known as the tarweeds, its members are confined in their distribution largely to California including the coastal islands, and partly to adjacent Oregon, Nevada and Baja California. As reconstituted by Keck (unpublished) the genus consists of four sections: Zonamra, Deinandra, Centromadia and Hemizonia. In the earlier taxonomic treatment of Jepson (1925), the spikeweeds, Centromadia, were accorded generic status. Keck (1935) recognized four species under Centromadia: Hemnizonia pungens (Hk. et Arn.) Torr. et Gray, H. parryi Greene, H. fitchii Gray and H. perennis (Greene) Keck. Characters of the pappus, anthers, bracts, inflorescence and herbage are used to distinguish the taxa. Hemizonia pungens has no pappus on its disk florets, H. parryi has three pappus bristles per disk floret and H. fitchii has many. The chromosome ntumbers previousl y recorded are n 9 for H. pungens, n 11 and 12 for H. parryi and n 13 for H. fitchii (Clausen, Keck and Heusi, 1934). The chromosome number of H. perennis is unknown; unlike the other species, it is a perennial and is known from only two localities in Baja California. Figure 1 shows the relative distribution of the species and subspecies. Hemizonia pungens, by far the commonest member of the section, is widely distributed. At least in California, there are three distinct forms of this species which occur in more or less distinct ecological niches and are taxonomically treated as subspecies. The maritime form, H. pungens ssp. mnaritima Keck occurs chiefly in salt marshes around San Francisco bay and along the coast, but is rare elsewhere. Hemizonia pungens ssp. interior Keck, an inland form occurring in the dry inner valleys, is very abundant throughout the San Joaquin and Salinas valleys and probably has been introduced along the western borders of the Mojave desert. The populations occurring in the passes of the Inner Coast Range which separates the two subspecies, contain many intermediates and recombinations. Hemizonia pungens ssp. laevis is