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Learning gain in higher education, specifically the development of subject-adjacent skills, abilities and competencies, is of key scholarly and policy relevance. However, little research focuses on students’ broad understandings of learning gain. This paper takes a phenomenographic approach to explore students’ conceptions of learning gain, and how these may differ by subject and stage of study. Drawing on thirty-three qualitative interviews with a diverse group of undergraduate and postgraduate students across four subjects in three UK universities, this paper derives three distinct categories of understanding in relation to learning gain, increasing in complexity. The first category, naïve understandings, is characterised by short-term learning goals, the prioritisation of surface approaches to learning including memorisation, with subject knowledge more important than skills. Motivations to learn are broadly extrinsic and related to assessments. The second category, emergent understandings, features greater awareness of approaches to learning, including structured planning of learning activities in response to a range of goals, and pragmatic responses to desirable learning outcomes. There is growing recognition of links between subject-specific and generic skills, and adaptation features strongly. The most sophisticated category, comprehensive understandings, is characterised by flexible learning goals, the acknowledgment of the independent nature of higher education learning and strong self-regulation, with deep approaches to learning consciously prioritised. Dynamic views of knowledge prevail, which distinguish this category from the other two. These categories of understanding grow in complexity with study stage, and also vary by subject of study, with interactions present in relation to prioritised aspects of learning gain, views of change and how explicit aspects of learning gain are made within the subject of study. Any attempt to capture learning gain for its improvement would therefore need to encompass a variety of aspects of learning gain, contextualised for subject and stage and include a student perspective. [ABSTRACT FROM AUTHOR]