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• Technological and behavioural developments in Europe at 400,000 years ago. • Model of landscape and resources shaping development of localised material culture. • Model of population movement triggered by shifts in climate or environment. • Estimates of social group and territory size for Middle Pleistocene Europe. Identification of cultural groups is rare in the early Palaeolithic due to site formation processes including taphonomy and the effect of raw material and site function. This paper reviews a critical period in Europe at about 400 ka (MIS 11) when we may be able to identify such groups. This period, sees more sustained occupation and evidence of new technologies, including bone and wooden tools, hunting and fire-use. Importantly, brain size had begun to approach modern capacity. The fine-tuned record from Britain enables correlation of sites and new models of human behaviour to be developed. Millennial-scale changes in material culture can now be recognised, which can be interpreted as brief incursions by different cultural groups into Britain from mainland Europe. We suggest that population movement was primarily driven by changes in climate and environment. We further propose that variation in material culture is a reflection of local resources and landscape and that during stable environment localised expressions of culture emerge. This can be applied to Europe, where it is suggested that a complex mosaic of small-scale cultural groupings can be identified, some with and some without handaxes, but underpinned by a common set of technologies and behaviours. [ABSTRACT FROM AUTHOR]