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We consider the problem of quantum and stochastic confinement for drift-diffusion equations on domains Ω ⊂ R d . We obtain various sufficient conditions on the behavior of the coefficients near the boundary of Ω which ensure the essential self-adjointness or stochastic completeness of the symmetric form of the drift-diffusion operator, − 1 ρ ∞ ∇ ⋅ ρ ∞ D ∇ . The proofs are based on the method developed in [31] for quantum confinement on bounded domains in R d . In particular for stochastic confinement we combine the Liouville property with Agmon type exponential estimates for weak solutions. [ABSTRACT FROM AUTHOR]

The wave equation □ g M , a ψ = 0 on subextremal Kerr spacetimes ( M M , a , g M , a ) , 0 < | a | < M , does not admit real mode solutions, as was established by Shlapentokh-Rothman. In this paper, we show that the absence of real modes does not persist under the addition of an arbitrary short-range non-negative potential V to the wave equation or under changes of the metric g M , a in the far away region of M M , a (retaining the causality of the stationary Killing field T there). In particular, we first establish, for any 0 < | a | < M , the existence of real mode solutions ψ to equation □ g M , a ψ − V ψ = 0 , for a suitably chosen time-independent real potential V with compact support in space, satisfying V ≥ 0 . Exponentially growing modes are also obtained after perturbing the potential V . Then, as an application of the above result, we construct a family of spacetimes ( M M , a , g M , a ( d e f ) ) which are compact in space perturbations of ( M M , a , g M , a ) , have the same symmetries as ( M M , a , g M , a ) and moreover admit real and exponentially growing modes. These spacetimes contain stably trapped null geodesics, but we also construct a more complicated family of spacetimes with normally hyperbolic trapped set, admitting real and exponentially growing modes, at the expense of having conic asymptotics. The aforementioned results are in contrast with the case of stationary asymptotically flat (or conic) spacetimes ( M , g ) with a globally timelike Killing field T , where real modes for equation □ g ψ − V ψ = 0 are always absent, giving a useful zero-frequency continuity criterion for showing stability for a smooth family of equations □ g ψ − V λ ψ = 0 , with λ ∈ [ 0 , 1 ] and V 0 = 0 . We show explicitly that this criterion fails on Kerr spacetime. [ABSTRACT FROM AUTHOR]

Concentrations of cosmogenic 10Be, measured in quartz from chert and river sediment around the Cradle of Humankind (CoH), are used to determine basin-averaged erosion rates and estimate incision rates for local river valleys. This study focusses on the catchment area that hosts Malapa cave with Australopithecus sediba, in order to compare regional versus localized erosion rates, and better constrain the timing of cave formation and fossil entrapment. Basin-averaged erosion rates for six sub-catchments draining the CoH show a narrow range (3.00 ± 0.28 to 4.15 ± 0.37 m/Mega-annum [Ma]; ±1σ) regardless of catchment size or underlying geology; e.g. the sub-catchment with Malapa Cave (3 km²) underlain by dolomite erodes at the same rate (3.30 ± 0.30 m/Ma) as the upper Skeerpoort River catchment (87 km²) underlain by shale, chert and conglomerate (3.23 ± 0.30 m/Ma). Likewise, the Skeerpoort River catchment (147 km²) draining the northern CoH erodes at a rate (3.00 ± 0.28 m/Ma) similar to the Bloubank-Crocodile River catchment (627 km²) that drains the southern CoH (at 3.62 ± 0.33 to 4.15 ± 0.37 m/Ma). Dolomite- and siliciclastic-dominated catchments erode at similar rates, consistent with physical weathering as the rate controlling process, and a relatively dry climate in more recent times. Erosion resistant chert dykes along the Grootvleispruit River below Malapa yield an incision rate of ∼8 m/Ma at steady-state erosion rates for chert of 0.86 ± 0.54 m/Ma. Results provide better palaeo-depth estimates for Malapa Cave of 7-16 m at the time of deposition of A. sediba. Low basin-averaged erosion rates and concave river profiles indicate that the landscape across the CoH is old, and eroding slowly; i.e. the physical character of the landscape changed little in the last 3-4 Ma, and dolomite was exposed on surface probably well into the Miocene. The apparent absence of early Pliocene- or Miocene-aged cave deposits and fossils in the CoH suggests that caves only started forming from 4 Ma onwards. Therefore, whilst the landscape in the CoH is old, cavities are a relatively young phenomenon, thus controlling the maximum age of fossils that can potentially be preserved in caves in the CoH. [ABSTRACT FROM AUTHOR]

Abstract: Weeds with broad distributions and large morphological variation are challenging for systematists and evolutionarily intriguing because their intensive dispersal would likely prevent local morphological differentiation. Study on weeds will help to understand divergence in plants unlikely to be affected by geographical and ecological barriers. We studied Youngia japonica based on nrDNA and cpDNA sequences. This is a widespread native in Asia and invasive worldwide; nevertheless, three subspecies (japonica, longiflora, and formosana) and an undescribed variant occur in Taiwan. Bayesian and the most parsimonious phylogenies revealed that subspecies longiflora is a different linage and independently arrived in Taiwan during the Pleistocene via land connection to the Asian Continent. Bayesian time estimation suggested that Youngia in Taiwan diverged in the lower Pleistocene or more recently. Extreme habitats that emerged in the Pleistocene, i.e., cold mountain ranges for subspecies formosana and xeric, raised coral reefs for the undescribed Youngia variant probably had triggered the divergence. Components of Youngia in Taiwan are not monophyletic; a coalescent-based test suggested incomplete lineage sorting. Nevertheless, the samples within each taxon share unique morphological features suggesting a common gene pool and each taxon has different dominant ITS and/or cpDNA types; these conditions suggest ongoing process toward monophyly via coalescent processes and support the delimitation of intraspecific taxa. [Copyright &y& Elsevier]

Abstract: Paleoindians of North America entered a continent undergoing rapid climatic and environmental changes. This paper is a preliminary contribution toward obtaining a better picture regarding how climate and environmental change might have impacted the first settlers of North America. The Paleoindian sites we analyzed are, from the oldest to the youngest, Ingleside (Texas), Blackwater Draw Locality No. 1 (New Mexico), Hiscock Site (New York), and Plainview Quarry (Texas). Paleoenvironmental reconstruction involves identifying the dietary traits of ungulate species that might reflect the environmental conditions where they were living, and also where they might have been hunted by Paleoindians. Such an approach is realized through tooth microwear and mesowear analyses. Results indicate that a variety of food resources were available for the ungulates at these sites as well as the likely presence of mosaic environments around these sites which allowed Paleoindians to exploit resources in a large variety of habitats. The application of a method that allows for the estimation of the duration of occupation at archaeological sites reveals that results for Blackwater Draw and Hiscock Site indicate a long-term occupation of probably several months. However, during short events, Paleoindians were most likely hunting herds of horse and bison when these prey were available near the site. Results indicate that Plainview Quarry was likely used only for short-term occupations, with large game hunting focusing on bison. These patterns identified at the archeological sites studied are related to the fact that Paleoindians follow a high-technology forager model and frequently shifted their territory depending on the composition and distribution of the large mammal fauna. [Copyright &y& Elsevier]

Abstract: Optically stimulated luminescence (OSL) measurements were made on individual, sand-sized grains of quartz from Middle Palaeolithic deposits at two cave sites (El Harhoura 2 and El Mnasra) on the Atlantic coast of Morocco. We were able to calculate OSL ages for 32 of the 33 samples collected from the Middle Palaeolithic deposits, including the earliest and latest Aterian levels at both sites. These ages reveal periods of occupation between about 110 and 95 ka (thousands of years ago), and at ∼75 ka. A late Middle Palaeolithic occupation of El Harhoura 2 is also recorded at ∼55 ka. Our single-grain OSL chronologies largely support previous age estimates from El Mnasra and other sites along the Atlantic coast of Morocco, but are generally more precise, reproducible and stratigraphically more coherent (i.e., fewer age reversals). We compare the single-grain ages for El Harhoura 2 and El Mnasra with those obtained from single- and multi-grain OSL dating of Middle Palaeolithic deposits in the nearby sites of Contrebandiers and Dar es-Soltan 1 and 2, and with records of past regional environments preserved in sediment cores collected from off the coast of northwest Africa. A conspicuous feature of the new chronologies is the close correspondence between the three identified episodes of human occupation and periods of wetter climate and expanded grassland habitat. Owing to the precision of the single-grain OSL ages, we are able to discern gaps in occupation during Marine Isotope Stages 5 and 4, which may represent drier periods with reduced vegetation cover. We propose that these climatic conditions can be correlated with events in the North Atlantic Ocean that exert a major control on abrupt, millennial-scale fluctuations between wet and dry periods in northwest and central North Africa. [Copyright &y& Elsevier]

Abstract: The Hadar paleoanthropological site in Ethiopia preserves a record of hominin evolution spanning from approximately 3.45Ma to 0.8Ma. An angular unconformity just above the ca. 2.95Ma BKT-2 complex divides the sediments into the Hadar Formation (ca. 3.8–2.9Ma) and the Busidima Formation (ca. 2.7–0.15Ma). The unconformity is likely a response to a major tectonic reorganization in the Afar Depression, and activation of the As Duma fault near the Ethiopian Escarpment (west of Hadar) created a half-graben in which the Busidima Formation was deposited. The pattern and character of sedimentation in the region changed dramatically above the unconformity, as cut-and-fill channel conglomerates and silt-dominated paleosols that comprise the Busidima Formation stand in sharp contrast to the underlying deposits of the Hadar Formation. Conglomerate deposition has been related to both the perennial, axial paleo-Awash and ephemeral, escarpment-draining tributaries. Overbank silts have yielded fossils attributed to early Homo and Oldowan stone tools. Numerous tuffaceous deposits exist within the Busidima Formation, but they are often spatially limited, fine-grained, and reworked. Recent work on the tephrostratigraphic framework of the Busidima Formation at Hadar has identified at least 12 distinct vitric tephras and established the first geochemical-based correlations between Hadar and the neighboring project areas of Gona and Dikika. Compared to Gona and Dikika, where Busidima Formation sediments are exposed over large areas, the highly discontinuous sediments at Hadar comprise less than 40m in composite section and are exposed over an area of <20km2, providing only snapshots into the 2.7–0.15Ma window. The stratigraphic record at Hadar confirms the complex depositional history of the Busidima Formation, and also provides important details on regional stratigraphic correlations and the pattern of deposition and erosion in the lower Awash Valley reflective of its tectonic history. [Copyright &y& Elsevier]

Abstract: Excavations in the late Pleistocene deposits at Liang Bua cave, Flores, have uncovered the skeletal remains of several small-bodied and small-brained hominins in association with stone artefacts and the bones of Stegodon. Due to their combination of plesiomorphic, unique and derived traits, they were ascribed to a new species, Homo floresiensis, which, along with Stegodon, appears to have become extinct ∼17 ka (thousand years ago). However, recently it has been argued that several characteristics of H. floresiensis were consistent with dwarfism and evidence of delayed development in modern human (Homo sapiens) myxoedematous endemic (ME) cretins. This research compares the skeletal and dental morphology in H. floresiensis with the clinical and osteological indicators of cretinism, and the traits that have been argued to be associated with ME cretinism in LB1 and LB6. Contrary to published claims, morphological and statistical comparisons did not identify the distinctive skeletal and dental indicators of cretinism in LB1 or LB6 H. floresiensis. Brain mass, skeletal proportions, epiphyseal union, orofacial morphology, dental development, size of the pituitary fossa and development of the paranasal sinuses, vault bone thickness and dimensions of the hands and feet all distinguish H. floresiensis from modern humans with ME cretinism. The research team responsible for the diagnosis of ME cretinism had not examined the original H. floresiensis skeletal materials, and perhaps, as a result, their research confused taphonomic damage with evidence of disease, and thus contained critical errors of fact and interpretation. Behavioural scenarios attempting to explain the presence of cretinous H. sapiens in the Liang Bua Pleistocene deposits, but not unaffected H. sapiens, are both unnecessary and not supported by the available archaeological and geochronological evidence from Flores. [Copyright &y& Elsevier]

Abstract: The southern Rocky Mountains and adjacent Intermontane Plateau Highlands region of western North America is a geographically diverse area with an active geologic history. Given the topological complexity and extensive geologic activity, organisms inhabiting this region are expected to show some degree of morphological and genetic divergence, especially populations found on the southern montane ‘sky islands’ of this region. Here we examine the phylogeographic history and diversification of a montane forest inhabiting harvestmen, Sclerobunus robustus, using a combination of genetic and morphological data. Divergence time estimates indicate that much of the diversification within and between major groups S. robustus predate the Pleistocene glacial cycles. The most widespread subspecies, Sclerobunus robustus robustus, is recovered as six genetically distinct, geographically cohesive mitochondrial phylogroups. Gene tree data for a single nuclear gene reveals congruent, albeit slightly more conservative, patterns of genetic divergence. Despite high levels of genetic divergence throughout their distribution, phylogroups show extreme conservation in somatic and reproductive morphology. This uncoupling of morphological and genetic differentiation may be due to morphological conservatism associated with a conserved microhabitat preference. Based on these data, it is obvious that S. robustus has undergone some level of cryptic diversification. [Copyright &y& Elsevier]

Abstract: In response to climate changes that have occurred during Pleistocene glacial cycles, taxa associated to steppe vegetation might have followed a pattern of historical evolution in which isolation and fragmentation of populations occurred during the short interglacials and expansion events occurred during the long glacial periods, in contrast to the pattern described for temperate species. Here, we use molecular genetic data to evaluate this idea in a steppe bird with Palaearctic distribution, the little bustard (Tetrax tetrax). Overall, extremely low genetic diversity and differentiation was observed among eight little bustard populations distributed in Spain and France. Mismatch distribution analyses showed that most little bustard populations expanded during cooling periods previous to, and just after, the last interglacial period (127,000–111,000years before present), when steppe habitats were widespread across Europe. Coalescent-based methods suggested that glacial expansions have resulted in substantial admixture in Western Europe due to the existence of different interglacial refugia. Our results are consistent with a model of evolution and genetic consequences of Pleistocene cycles with low between-population genetic differentiation as a result of short-term isolation periods during interglacials and long-term exchange during glacial periods. [Copyright &y& Elsevier]

Abstract: Attirampakkam, an open-air stratified Palaeolithic site in southeastern India is the focus of ongoing studies to investigate the nature of Lower and Middle Palaeolithic occupation in relation to changing Pleistocene environments. The paucity of faunal or palaeobotanical remains at the site required the use of rock magnetic properties (χ lf, χ fd, χ fd %, χ ARM, SIRM, χ ARM/SIRM, SIRM/χ lf, χ ARM/χ lf, χ ARM/χ fd, S-ratio, HIRM and HIRM/IRM300mT) as proxies for deciphering the palaeoenvironments and palaeoclimates of the region. Rock magnetic data from stratified deposits within excavated trenches show that the bulk of the sedimentation was intermittent with no subaerial exposure of sediments for long periods, and points to fluctuating wetter and drier climates associated with the Acheulian to Late Middle Palaeolithic levels. There is a general trend towards a wetter climate from Layer 5 upwards. Layer 7 offers evidence of bacterial magnetite (χ ARM/χ lf >40 and χ ARM/χ fd >1000), suggesting the onset of micro-anaerobic environments as a result of floods. Greigite is indicated in Layer 5 (SIRM/χ lf >30×103 Am−1), which indicates a reducing environment. Layer 2 represents an arid climate (higher proportion of haematite and a coarser magnetic grain size). Layer 1a again is indicative of an arid climate (presence of calcrete, magnetically “hard” minerals and coarser magnetic grain size). Results indicate that the southeast coast of India experienced a mostly dry tropical climate during the Pleistocene. The detected occurrence of a few notably more arid intervals did not disrupt occupation of the site by successive groups of hominins. [Copyright &y& Elsevier]

Abstract: A detailed taphonomic analysis of the large mammal assemblage from 1998 to 2006 excavations at San Teodoro Cave is presented, taking into account the stratigraphic context of the deposits. Three not strictly contemporary fossiliferous levels having different lithological features have been detected, here named B-I, B-II, and B-III. Fossil remains are prevalently accumulated in B-I and B-II. The three levels are characterized by evidence of Crocuta crocuta spelaea occupation, represented by their skeletal remains, coprolites, and distinctive damages on the bones, similar to fossil and modern spotted hyena dens from Europe and Africa. A differential distribution of coprolites and small digested bones, probably due to different humidity conditions, has been recognized in B-I and B-II, and can be related to different topographic locations within the cave or to different climate conditions during the sedimentation phases. The very low density of fossil remains in B-III, which is the oldest level, could indicate an area that was less inhabited by hyenas, probably due to geomorphological conditions. Taphonomic comparison of the three fossiliferous levels of the San Teodoro Cave deposits points to a long-term, perhaps cyclic, occupation of the cave by hyenas and confirms the cave as one of the most important Pleistocene hyena dens in Europe. [Copyright &y& Elsevier]

Abstract: China is one of the few centers in the world where plant domestication evolved independently, but its developmental trajectory is poorly understood. This is because there is considerably less data from the terminal Pleistocene and early Holocene documented in China than in other regions, such as the Near East and Mesoamerica, and previous studies on Paleolithic subsistence in China have largely focused on animal hunting rather than plant gathering. To resolve these problems the current research investigated the range of plants used by late Paleolithic hunter–gatherers in the middle Yellow River region where some of the earliest farmers emerged. We employed usewear and starch analyses on grinding stones to recover evidence for plant use in a hunting–gathering population at a late Paleolithic site, Shizitan Locality 9 in Jixian, Shanxi (ca. 13,800–8500 cal. BP). The usewear analysis shows that all artifacts preserved a range of usewear patterns best matching multiple tasks and indicating multi-functional use. Starch remains recovered from these tools indicate that the Shizitan people collected and processed many types of grass seeds (Panicoideae and Pooideae subfamilies), acorns (Quercus sp.), beans (Phaseoleae tribe) and yams (Dioscorea sp.). The Shizitan people represented some of the last hunter–gatherers in the middle Yellow River region. Their broad spectrum subsistence strategy was apparently carried on by the first Neolithic farmers in the same region, who collected similar wild plants and eventually domesticated millets. The trajectory from intensified collection of a wide range of wild plants to domestication of a small number of species was a very long process in north China. This parallels the transition from the “broad spectrum revolution” to agriculture in the Near East. [Copyright &y& Elsevier]

Abstract: The small mammal accumulations in the Pleistocene deposits of Karain Cave are investigated to identify the predators and possible biases in the fossil assemblages. Seven small mammal assemblages are studied in chronological order from two chambers of the cave, the main chamber E and the side chamber B. The lowermost level within the whole sequence is Proto-Charentien, which corresponds to an early stage of the Middle Palaeolithic. The main part of the material belongs to Middle Palaeolithic layers. The most important aspect of the fossil record in the cave is the human occupation without any interruption through the Pleistocene to Holocene. The small mammal fossil evidence in the cave denotes the presence of opportunistic predators throughout the sequence with one exception, and these produce balanced samples of small mammal faunas in the habitat. The lack of bias in the small mammal faunas allow the interpretation of local environments, showing that partial steppe and arid conditions existed during deposition of the lowermost levels of the Middle Palaeolithic in Karain Cave and that these shifted into more temperate and wooded habitat in the upper levels. Evidence from the side chamber indicates some differences, with a more open grassy environment. In the Mediterranean part of Anatolia the temperate and moist conditions in the Middle Palaeolithic were superseded by more arid conditions in the Upper Palaeolithic, followed by a decrease in steppe conditions during the Epi-Palaeolithic period. [Copyright &y& Elsevier]

Abstract: The first lithic refit studies were carried out at the end of 19th century, but the method was not considered an area of real interest to archaeology until quite recently. Today, lithic refitting is applied in a multitude of areas of enquiry including lithic technology, intra-site and inter-site spatial distribution, archaeostratigraphy and formation processes. In this paper, we present a refit study of the lithic materials recovered in the base of the NW sector of level TD10 at the site of Gran Dolina (Sierra de Atapuerca, Spain). We aim to provide information about spatial distribution through identifying activity areas and the internal connections between those areas. Our work resulted in various refits whose connections reveal the movements of the pieces and/or knappers that once occupied the site. Our results also show the importance of studying the set of materials as a whole and the bias involved in analyzing only a small sample. The refits, connections and directions of movement allow us to infer areas in which an activity took place, but these results must be confirmed in future works covering the entire lithic assemblage of TD10. [Copyright &y& Elsevier]

Abstract: The 11,000-year old lake edge archaeological site of Star Carr in the Vale of Pickering of North Yorkshire is one of the most famous Mesolithic sites in Europe, and one of the earliest, dated to the period of climatic warming that immediately followed the final termination of the last ice age. One of the main reasons for this international importance is the richness of its organic artefacts, faunal assemblage and environmental data. However, recent investigations have demonstrated that these organic remains have severely deteriorated over the last 60 years due to the decay and acidification of the surrounding peat. This paper presents research into the effects on the bone (histological analysis using light and polarising microscopy, and Transmission Electron Microscopy, bulk collagen analysis, and amino acid analysis), antler (visual and metrical analysis, loss on ignition and Scanning Electron Microscopy) and wood (visual analysis, decay assessment tests and Scanning Electron Microscopy). [Copyright &y& Elsevier]

Abstract: Archaeological evidence regarding the presence of obsidian in levels that antedate the food production stage could have been the result of usage or intrusion of small obsidian artifacts from overlying Neolithic layers. The new obsidian hydration dates presented below employing the novel SIMS-SS method, offers new results of absolute dating concordant with the excavation data. Our contribution sheds new light on the Late Pleistocene/Early Holocene exploitation of obsidian sources on the island of Melos in the Cyclades reporting dates c. 13th millennium - end of 10th millennium B.P. [Copyright &y& Elsevier]

Abstract: The presence of very large lithic artifacts at the Acheulian site of Gesher Benot Ya‘aqov is one of the site’s most distinctive and enlightening features. Basalt giant cores and their products, modified by a variety of core methods and found in association with different hominin activities, occur throughout the stratigraphic sequence of the site. In this paper we describe the giant artifacts of Gesher Benot Ya‘aqov together with their reduction sequence, from the nature and acquisition of the raw material, through the shaping of the cores, to the discarded cores and their typical waste products. We then discuss the significance of these finds and their implications for understanding the site and its varied activities, as well as for Acheulian cognitive abilities and behavior during the early Middle Pleistocene on the margins of the paleo-Lake Hula. [Copyright &y& Elsevier]

Abstract: The Chauvet-Pont d''Arc and Deux-Ouvertures caves, located along the Ardèche River (France), contain abundant remains of the extinct cave bear (Ursus spelaeus). Because they also display a variety of Palaeolithic anthropogenic evidences, such as the earliest charcoal drawings recorded to date (Chauvet-Pont d''Arc), and delicate engravings (Deux-Ouvertures), they offer the opportunity of studying the interaction between animals and human beings during a key period for Pleistocene species extinctions. We characterized cave bear specimens from these two sites by radiocarbon dating, stable isotopes, and mitochondrial DNA analysis. In Chauvet-Pont d''Arc, we obtained radiocarbon ages that ranged between 29,000 and 37,300 years before present (BP). The Deux-Ouvertures cave bear specimens clustered to the bottom of this time frame, returning radiocarbon ages of 27,440–30,220 years BP. Cave bear nitrogen isotope values were all compatible with a vegetarian diet. Mitochondrial DNA analysis, carried out on a highly variable domain of the control region, evidenced only two cave bear haplotypes, including a new haplotype, and a common one which largely predominated. We detected both haplotypes in Chauvet-Pont d''Arc, but only recorded the predominant one in the Deux-Ouvertures Cave. Our data put forward the surprising observation that cave bears inhabited Ardèche over a short period of time, from about 37,000 to 27,400 years BP. They were notably present during the first (Aurignacian) phase of human intrusions in Chauvet-Pont d''Arc, 30,000–32,000 years BP. This points to the possible competition for cave sites, presumably on a seasonal scale considering the cave bear habit for hibernation. During this time period, the small number of haplotypes is at variance with the extensive genetic diversity reported elsewhere for much more ancient specimens. [Copyright &y& Elsevier]

Abstract: Neandertals and their immediate predecessors are commonly considered to be the only humans inhabiting Europe in the Middle and early Late Pleistocene. Most Middle Pleistocene western European specimens show evidence of a developing Neandertal morphology, supporting the notion that these traits evolved at the extreme West of the continent due, at least partially, to the isolation produced by glacial events. The recent discovery of a mandible, BH-1, from Mala Balanica (Serbia), with primitive character states comparable with Early Pleistocene mandibular specimens, is associated with a minimum radiometric date of 113 + 72 − 43 ka. Given the fragmented nature of the hemi-mandible and the fact that primitive character states preclude assignment to a species, the taxonomic status of the specimen is best described as an archaic Homo sp. The combination of primitive traits and a possible Late Pleistocene date suggests that a more primitive morphology, one that does not show Neandertal traits, could have persisted in the region. Different hominin morphologies could have survived and coexisted in the Balkans, the “hotspot of biodiversity.” This first hominin specimen to come from a secure stratigraphic context in the Central Balkans indicates a potentially important role for the region in understanding human evolution in Europe that will only be resolved with more concentrated research efforts in the area. [Copyright &y& Elsevier]

Abstract: Our knowledge of early Australasian societies has significantly expanded in recent decades with more than 220 Pleistocene sites reported from a range of environmental zones and depositional contexts. The uniqueness of this dataset has played an increasingly important role in global debates about the origins and expression of complex behaviour among early modern human populations. Nevertheless, discussions of Pleistocene behaviour and cultural innovation are yet to adequately consider the effects of taphonomy and archaeological sampling on the nature and representativeness of the record. Here, we investigate the effects of preservation and sampling on the archaeological record of Sahul, and explore the implications for understanding early cultural diversity and complexity. We find no evidence to support the view that Pleistocene populations of Sahul lacked cognitive modernity or cultural complexity. Instead, we argue that differences in the nature of early modern human populations across the globe were more likely the consequence of differences in population size and density, interaction and historical contingency. [Copyright &y& Elsevier]

Abstract: In order to understand the behaviours and subsistence choices of Palaeolithic hunter–gatherers, it is essential to understand the behavioural ecology of their prey. Here, we present strontium isotope data from sequentially-sampled enamel from three reindeer (Rangifer tarandus ssp.) and a single bison (Bison cf. priscus) from the late Middle Palaeolithic site of Jonzac (Chez-Pinaud), France. The results are used to investigate the ranging and migratory behaviours of these important prey species. We found that the bison had isotope values most consistent with a local range, while the three reindeer had values indicating a seasonal migration pattern. Due to the similarity of the patterning of two of the three reindeer and in conjunction with zooarchaeological results, we suggest that they may have been from the same herd, were likely killed around the same point during their seasonal round and may therefore be the product of a single hunting event or a small number of successive hunting events. The isotope analyses complement the zooarchaeological data and have allowed greater insight into the palaeoecology of these species, the palaeoenvironment, and Neanderthal site use and hunting strategies. [Copyright &y& Elsevier]

Abstract: This paper considers the transition from Howiesons Poort to post-Howiesons Poort technologies at the rock shelter site of Klein Kliphuis, South Africa. The transition at this site is shown to be gradual, with incremental changes in material selection and in the size and shapes of flakes and cores. Implements which appear to blend characteristics of those distinctive of the earlier and later industries appear briefly at the mid-point of the transition. The results suggest that there is unlikely to have been an occupational hiatus between the Howiesons Poort and post-Howiesons Poort. Explanations for the Howiesons Poort phrased in terms of population expansion and contraction are also difficult to support. Technological changes at this time may relate to environmental variation, though the limited nature of terrestrial archives documenting changes in resource productivity necessitate that any such suggestion be made with caution. [Copyright &y& Elsevier]

Abstract: Here we present a detailed palaeopathological study of the hominin mandible ATE9-1 found at the Sima del Elefante site (TE), Sierra de Atapuerca, Spain. This fossil represents the earliest hominin remains from Western Europe with an age of ca. 1.3 Ma. The specimen displays several dento-gnathic lesions; the antiquity and geographic location of this fossil justifies a detailed palaeopathological study to determine if the pathologies have significantly altered taxonomically relevant features. Our study reveals severe dental attrition combined with generalized hypercementosis, alveolar root exposure, mild periodontal disease, tooth dislocation, and an anomalous occlusal plane. We have also observed calculus deposits, two cystic lesions and an anomalous wear facet compatible with tooth picking. The majority of these pathological signs can be explained by compensatory eruption. We propose that these lesions are associated as causes, consequences, and amplifiers of one another within the framework of heavy and even traumatic occlusion, masticatory habits, or both traumatic occlusion and masticatory habits. Despite the severity of these lesions, occlusion was at least partially functional so it was unlikely to influence the survival of this individual. In addition, the lesions do not prohibit the taxonomic assessment of the mandible. [Copyright &y& Elsevier]

Abstract: We present a detailed morphological comparative study of the hominin mandible ATE9-1 recovered in 2007 from the Sima del Elefante cave site in Sierra de Atapuerca, Burgos, northern Spain. Paleomagnetic analyses, biostratigraphical studies, and quantitative data obtained through nuclide cosmogenic methods, place this specimen in the Early Pleistocene (1.2–1.3 Ma). This finding, together with archaeological evidence from different European sites, suggests that Western Europe was colonised shortly after the first hominin expansion out of Africa around the Olduvai subchron. Our analysis of the ATE9-1 mandible includes a geometric morphometric analysis of the lower second premolar (LP4), a combined and detailed external and internal assessment of ATE9-1 roots through CT and microCT techniques, as well as a comparative study of mandibular and other dental features. This analysis reveals some primitive Homo traits on the external aspect of the symphysis and the dentition shared with early African Homo and the Dmanisi hominins. In contrast, other mandibular traits on the internal aspect of the symphysis are derived with regard to African early Homo, indicating unexpectedly large departures from patterns observed in Africa. Reaching the most occidental part of the Eurasian continent implies that the first African emigrants had to cross narrow corridors and to overcome geographic barriers favouring genetic drift, long isolation periods, and adaptation to new climatic and seasonal conditions. Given these conditions and that we are dealing with a long time period, it is possible that one or more speciation events could have occurred in this extreme part of Eurasia during the Early Pleistocene, originating in the lineages represented by the Sima del Elefante-TE9 hominins and possibly by the Gran Dolina-TD6 hominins. In the absence of any additional evidence, we prefer not include the specimen ATE9-1 in any named taxon and refer to it as Homo sp. [Copyright &y& Elsevier]

Abstract: In order to enhance our understanding of the evolutionary history of squirrel monkeys (Saimiri spp.), we newly sequenced and analyzed data from seven complete mitochondrial genomes representing six squirrel monkey taxa. While previous studies have lent insights into the taxonomy and phylogeny of the genus, phylogenetic relationships and divergence date estimates among major squirrel monkey clades remain unclear. Using maximum likelihood and Bayesian procedures, we inferred a highly resolved phylogenetic tree with strong support for a sister relationship between Saimiri boliviensis and all other Saimiri, for monophyly of Saimiri oerstedii and Saimiri sciureus sciureus, and for Saimiri sciureus macrodon as the sister lineage to the S. oerstedii/S. s. sciureus clade. We inferred that crown lineages for extant squirrel monkeys diverged around 1.5million years ago (MYA) in the Pleistocene Epoch, with other major clades diverging between 0.9 and 1.1MYA. Our results suggest a relatively recent timeline of squirrel monkey evolution and challenge previous conceptions about the diversification of the genus and its expansion into Central America. [Copyright &y& Elsevier]

Abstract: The schizothoracine Gymnocypris chilianensis is restricted to the Shiyang, Ruoshui and Shule Rivers, listed from east to west, along the northeast edge of the Tibetan Plateau. This distribution provides a valuable system to test hypotheses about postglacial colonization. We used mitochondrial DNA sequence data (a control region and the cytochrome b gene; 1894bp) to assess the phylogeographic structure of this species based on 278 specimens sampled from throughout the species’ entire geographical range. We found three lineages corresponding geographically to the three rivers, suggesting three independent glacial differentiation centers within the northeast edge of the Tibetan Plateau. The phylogenetic analysis suggested that the Shiyang River population forms a lineage that separated from the other populations of G. chilianensis at the basal phylogenetic split within this species. The molecular data further demonstrated a clear pattern of decreasing genetic diversity from the eastern Shiyang River towards the central Ruoshui River and western Shule River lineages, a pattern consistent with sequential western colonization. We therefore propose a phylogeographic scenario for G. chilianensis of a gradual westerly expansion from the Shiyang River population along the northeast edge of the Tibetan Plateau, with subsequent allopatric evolution at approximately 0.37 and 0.05 million years ago (Ma), through at least two glacial maxima. Together with the genetic evidence reported in other species, our findings suggest that this common biogeographic pattern emphasizes the importance of the northeastern edge region of the Tibetan Plateau as a hotspot of genetic diversity for some taxa. [Copyright &y& Elsevier]

Abstract: The lentic Basommatophora molluscs and hygrophilous land snails of the Early-Middle Pleistocene site of Gesher Benot Ya‘aqov (GBY), 0.78 Ma, crossed by the Matuyama–Brunhes chron boundary [MBB] were studied in order to reconstruct their specific habitats and possible reactions to climate change along the site sequence. Samples of equal sizes from 27 of the 46 layers along the 100 k.yr. time-span of the site were examined. About 2000 specimens of 21 lentic and hygrophilous species belonging to five families: Planorbidae (11), Lymnaeidae (6), Acroloxidae (2), Carychiidae (1) and Succineidae (1) were identified. The family with the largest biodiversity is the Planorbidae and of these, the most abundant species include Gyraulus piscinarum (937), Planorbarius corneus (210) and Radix labiata (160). The recent known zoogeographic origin of 81% of the species is Palaearctic and Holarctic. The MBB coincides with remarkable environmental changes reflected in molluscs, other faunal and floral elements and stable isotopes. The Planorbidae and Lymnaeidae reach greater abundance (90% and 80% of their assemblages, respectively) pre-MBB, while Acroloxidae, Succineidae and Carychiidae are more abundant (74%, 64% and 90%) post-MBB. Our data indicate that GBY molluscs show a two-phase pattern (shallow and deep lake) in each of the five defined cycles. Their numbers increase during the shallow water phases, thus the site climate changes from cold and humid in the oldest layers, to dry and cold up to the MBB and few succeeding layers. Between cycles 2 and 3, post-MBB, we see a short period of warm and humid climate that enables the influx of African and Asian elements. At the same time, cold climate species of Euro-Siberian and Palaearctic origin disappear. The succeeding layers indicate a cooler and humid climate. Finally, the sequence indicates deep water in the oldest layers and desiccation towards MBB and deeper water post-MBB. [Copyright &y& Elsevier]

Abstract: We report the study of the collection of fossil bovid specimens from the Early–Middle Pleistocene Acheulian site of Gesher Benot Ya‘aqov. This locality, situated in the Levantine Corridor (the bottleneck that connects Africa and Eurasia) is a key site to explain the faunal and human dispersals out of Africa during the Matuyama/Brunhes boundary around 0.8Ma. Two species of bovine (Bos sp., and Bovini gen. et sp. indet. cf. Bison sp.), one antelope (Gazella sp. cf. G. Gazella), and another indeterminate Bovidae gen. et sp. indet., have been recorded. The largest species, Bos sp., is an African immigrant related to the species from the Eritrean site of Buia, Bos buiaensis, which evolved from the buffalo of Olduvai Pelorovis oldowayensis, and colonized the Eurasian continent in parallel with the dispersal of the Acheulian culture into the northern continent. Numerous important species first recorded in several localities of Early–Middle Pleistocene transition from Eurasia are included in this dispersal out of Africa, including the megaherbivore, Palaeoloxodon antiquus, and the carnivores Crocuta crocuta, and later, Panthera leo and Panthera pardus. This faunal turnover is coincident with the change to colder climates that dominated the Middle Pleistocene. [Copyright &y& Elsevier]

Abstract: Aquifers dominated by Pleistocene basalts and Jurassic to Cretaceous calcareous rocks feed the Hula basin which is drained by the Jordan River into Lake Kinneret. The sedimentary sequence of Lower–Middle Pleistocene Benot Ya‘akov Formation (BYF) exposed by excavations of the 0.78 Ma lake-side site of Gesher Benot Ya‘aqov (GBY) consists of six cycles representing ca. 100 ka history of the Hula basin. This study characterizes the types of water sources in the catchment, tests the use of the Strontium (Sr) isotopes in the common extant snail Melanopsis sp. as a tracer for water in its habitat, and uses this tracer in the fossil specimens from GBY to investigate the palaeohydrology of the Hula paleolake during the corresponding period. The Sr isotope composition (87Sr/86Sr) of extant Melanopsis shells in the Hula catchment range widely (0.7046–0.7079). These analyses define distinct groups of water sources and aquifers, while the Jordan River at the GBY site has values around 0.70685. The values for fossil Melanopsis from GBY vary along stratigraphy; they are highest around 0.70710 in Cycles 1 and 2, decrease to around 0.70685 in Cycle 3, and exhibit upward trending fluctuations in the subsequent cycles to 0.70703 in Cycle 6. This trend reveals the dominance of the Hermon Jurassic aquifer during the earlier, colder periods before the Matuyama-Brunhes Boundary (MBB) and enhanced influence of the Golan basaltic aquifers, in subsequent warmer periods, indicating that the MBB coincides with climate warming as supported by other indicators. Hence, this global geochronological indicator of 0.78 Ma is also potentially a global palaeoclimatic marker. The similarity between the Sr isotope composition of the Jordan River waters and Melanopsis and those from Cycle 3 suggests that the current climate corresponds to that of the warmest period within the record of GBY, clarifying the comparative interpretation of this 100 k.yr. climate record. [Copyright &y& Elsevier]

Abstract: Preliminary results of the investigation of the microfauna at the Acheulo-Yabrudian Middle Pleistocene site of Qesem Cave, Israel, are presented. Thus far the assemblage includes ca. 10,000 bone and tooth fragments, of which 50% could be identified to the generic and some hundreds to the species level. Based on the current material, the fauna includes the following squamate reptiles: Laudakia sp., Chamaeleo sp., Gekkonidae indet., Lacertidae indet., Scincidae indet., Pseudopus sp., Varanus sp., Colubroidea indet. (at least three species) and micromammals: Suncus etruscus, Crocidura cf. leucodon, Crocidurinae indet. (large form), Chiroptera indet., Sciurus cf. anomalus, Cricetulus cf. migratorius, Microtus guentheri, Nannospalax ehrenbergi, Dipodillus cf. dasyurus, Meriones cf. tristrami, Gerbillidae indet., Mus cf. musculus, Apodemus cf. flavicollis. These results suggest that the fauna includes only taxa that occur recently in the territory of Israel. The ecological preferences of the nearest living relatives of the recorded taxa allow us to infer a paleoenvironment with a mosaic of open and woodland habitats. However, comparing the lower with the upper levels of the microfauna-bearing profile, a slight shift towards more wooded conditions might be detectable. Biostratigraphical inferences from the recorded micromammal taxa cover a rather wide age range, whereas the radiometric (U-series and preliminary TL) dating enable a provisionally estimated date for the microfauna-bearing levels at 360-300 ka. Detailed morphometric comparisons with material from other sites in the region are necessary and may yet provide further insights. [Copyright &y& Elsevier]

Abstract: Climate change is frequently considered an important driver of hominin evolution and dispersal patterns. The role of climate change in the last phase (900–700 ka) of the Middle Pleistocene Transition (MPT) in the Levant and northeast Africa was examined, using marine and non-marine records. During the MPT the global climate system shifted from a linear 41 k.yr. into a highly non-linear 100 k.yr. system, considerably changing its global modulation. Northeast Africa aridity further intensified around 950ka, as indicated by a sharp increase in dust flux, and a jump to overall higher levels thereafter, coinciding with a lack of sapropels in the deep eastern Mediterranean (930–690ka). The increased dust flux centering at ∼800ka corresponds to the minima in 400 k.yr. eccentricity, a minima in 65 °N solar forcing and in the weakest African monsoon precession periodicity. This resulted in expansion of hyper-arid conditions across North Africa, the lowest lake levels in eastern Africa and the lowest rainfall in the Nile River headwaters. In the eastern Mediterranean an increasing continental signature is seen in glacial stages 22 (∼880ka) and 20 (∼800ka). Lower arboreal pollen values also indicate arid conditions during these glacial stages. The southern and eastern parts of the Negev Desert, unlike its northern part, were hyper-arid during the MPT, making them highly unsustainable. The fluctuations in the stands of Lake Amora follow global climate variability but were more moderate than those of its last glacial Lake Lisan successor. In the northern Jordan-Valley Hula Lake, frequent fluctuations in lake level coincide with both global climate changes and minor changes in water salinity varying from fresh to oligohaline. It appears therefore that the most pronounced and widespread deterioration in climate occurred in northeast Africa from 900 to 700ka, whereas in the Levant the corresponding climatic changes were more moderate. [Copyright &y& Elsevier]

Abstract: The dispersal of hominins may have been favored by the opening of the landscape during the Early–Middle Pleistocene transition (EMP) in Western Europe. The structure of the small-vertebrate assemblages of the archaeo-paleontological karstic site of Gran Dolina in Atapuerca (Burgos, Spain) shows important environmental and climatic changes in the faunal succession, across the Matuyama–Brunhes boundary at 780 ka. These changes are interpreted to indicate impoverishment of the forests, along with an increase in dry meadows, and open lands in general that entailed a tendency towards the loss of diversity in small-vertebrate communities above the EMP. We evaluate variation in diversity of the faunal succession of Gran Dolina using Shannon’s Second Theorem as an index of ecosystem structure. The long cultural-stratigraphic sequence of Gran Dolina during the EMP is somewhat similar in its completeness and continuity to that in the locality of Gesher Benot Ya''aqov in the Upper Jordan Valley. We also evaluate related data including faunal and floral (pollen) succession. Both localities present cold, dry and humid, warm fluctuations at the transition between the Early and the Middle Pleistocene. Comparisons between these sites present opportunities to understand large-scale climatic changes. [Copyright &y& Elsevier]

Abstract: The Acheulian Technocomplex exhibits two phenomena: variability and conservatism. Variability is expressed in the composition and frequencies of tool types, particularly in the varying frequencies of bifaces (handaxes and cleavers). Conservatism is expressed in the continuous presence of bifaces along an immense time trajectory. The site of Gesher Benot Ya‘aqov (GBY) offers a unique opportunity to study aspects of variability and conservatism as a result of its long cultural-stratigraphic sequence containing superimposed lithic assemblages. This study explores aspects of variability and conservatism within the Acheulian lithic assemblages of GBY, with emphasis placed on the bifacial tools. While variability has been studied through a comparison of typological frequencies in a series of assemblages from the site, evidence for conservatism was examined in the production modes expressed by the reduction sequence of the bifaces. We demonstrate that while pronounced typological variability is observed among the GBY assemblages, they were all manufactured by the same technology. The technology, size, and morphology of the bifaces throughout the entire stratigraphic sequence of GBY reflect the strong conservatism of their makers. We conclude that the biface frequency cannot be considered as a chrono/cultural marker that might otherwise allow us to distinguish between different phases within the Acheulian. The variability observed within the assemblages is explained as a result of different activities, tasks, and functions, which were carried out at specific localities along the shores of the paleo-Hula Lake in the early Middle Pleistocene. [Copyright &y& Elsevier]

Abstract: This study investigates faunal remains from the site of Gesher Benot Ya‘aqov, analyzing how hominins may have utilized vertebrates and exploring paleoenvironments during the Early and Middle Pleistocene. Based on analyses of a range of vertebrates, results show that the species distribution of terrestrial vertebrates (microvertebrates and mammals) at the site of Gesher Benot Ya‘aqov shows relative stasis above the Matuyama–Brunhes Boundary (MBB) (0.78 Ma). However, consistency in faunal remains at the site reflecting stasis does not seem to reflect accurately the paleoenvironment. Marked changes are demonstrable in the lake-margin sedimentary background, archaeological remains, and agents of accumulation and damage, as well as in the density of medium–large mammals. This study emphasizes the significance of studying assemblages as a whole rather than the species representation on its own. [Copyright &y& Elsevier]

Abstract: The Golan Heights borders the Upper Jordan Valley on its eastern side and likely served as a prime foraging area for hominin groups that inhabited the Upper Jordan Valley during the Mid-Pleistocene. This study tests the hypothesis that Mid-Pleistocene climate in the Golan region was similar to that of the present day. Carbon isotope composition of present day plant communities and soil organic carbon from the Golan were compared to those of paleosols from Nahal Orvim to reconstruct Mid-Pleistocene paleoclimatic conditions. After correcting the paleosol values for recent changes in atmospheric carbon isotope values and potential biodegradation, the isotopic results show a strong similarity to those of present day local plants and soils. These results indicate that during the Mid-Pleistocene, the Golan was dominated by C3 vegetation, shared similar climatic conditions with the present day, and displayed long-term environmental stability. The span of time of paleosol formation is unknown and might cover multiple climatic episodes; thus, although short climatic fluctuations may have occurred, their impact was not substantial enough to be detected in the Nahal Orvim paleosols. This study concludes that the Golan slopes provided hominins and large grazers with a reliable and highly nutritious foraging area that complemented the Jordan Valley riparian ecosystem. [Copyright &y& Elsevier]

Abstract: The molecular phylogeny and biogeography of the East Asian freshwater crabs of the genus Nanhaipotamon (Decapoda: Brachyura: Potamidae) were studied, using two mitochondrial (16S rRNA and cytochrome oxidase I) and one nuclear (28S rRNA) markers, and correlated with various vicariant and dispersal events which have occurred in this region. The results showed Nanhaipotamon to be a monophyletic taxon with four clades which correspond to the topography of the coastal region of southeastern China and Taiwan Island. Mountains appear to play an important role in the distribution. The genus occurs only from east of the Wuyishan Range (Zhejiang and Fujian) and south of the Nanling Range (Guangdong) in southern China, and is also present west of the Central Range in Taiwan. The molecular and geological data suggest that Nanhaipotamon originated in an area between the Wuyishan and Nanling Ranges. In this area, the main and earliest cladogenesis occurred at ∼4.8 million years ago (mya), with speciation probably taking place at around 4mya. The molecular evidence strongly supports the recent invasion of the genus into Taiwan Island from northeastern Fujian, via the paleo-Minjiang River on the landbridge of Taiwan Strait. The presence of the genus in Dongyin Island, however, is through invasion from southeastern Zhejiang, during the Pleistocene glaciation period. Nanhaipotamon reached Taiwan and Dongyin Island at ∼1.0 and 0.4mya, respectively. A small population of Nanhaipotamon formosanum from Penghu Islands (Pescadores) in the central Taiwan Strait has a slightly different genetic constitution and suggests it is a relict of past Pleistocene glaciations. [Copyright &y& Elsevier]

Abstract: Causes previously suggested for the sudden extinction of Neanderthals (Homo neanderthalensis) in Europe, starting around 35,000years ago, comprise food shortage, climatic effects and violence from Modern Humans. The aim here is to formulate a demographic model with reconstructed fertility and death rates, capable of modelling the population development under conditions of changing climate and prey availability, from the early appearance of Neanderthals in Europe about 260,000years ago to their demise. Parameter variation studies are made for the parameters considered to have the highest uncertainty. Finally, the option of regional migration between northern, middle and southern Europe is added, in order to capture population movements away from a region in response to deteriorating or improving climate. This model accounts for population developments, including the re-population of the Middle and Northern regions of Europe during and after the warm Eem period. However, parameter choices that give plausible results during the initial 210,000years also predict that the Neanderthals should have survived the latter part of the Weichselian ice age, despite competing for food with Modern Human newcomers during the last part of the period. The conclusion is that other reasons for extinction than climate or starvation must be sought. [Copyright &y& Elsevier]

Abstract: European Russia has been at the fringe of the hominin dispersal for most of the late Pleistocene. However, by about 20,000 BP this area was settled by modern humans who had highly sophisticated and sometimes unique technologies. Not many Upper Paleolithic human fossils have been described from this area and consequently the morphology of these people remains largely unknown. Here, we present descriptions and a comparative morphological analysis of four possibly late Pleistocene fossils from European Russia: Skhodnya, Khvalynsk, Satanay, and Podkumok. The frontal bone is chosen for study because it is preserved in all of these fossils and is known to provide good discrimination among groups of Pleistocene hominins. All four fossils have been previously claimed to possess ‘archaic’ features of frontal morphology, such as developed supraorbital relief and a flat frontal squama. The results of a 3D geometric morphometric analysis of frontal bone landmarks and semilandmarks indicate that these fossils indisputably belong to modern humans. However, there are good reasons to associate Khvalynsk, Skhodnya, and Podkumok with Upper Paleolithic fossils from central and western Europe, whereas Satanay is more similar to a pooled sample of recent modern humans. [ABSTRACT FROM AUTHOR]

Abstract: We investigated the historical demography of Anopheles albimanus using mosquitoes from five countries and three different DNA regions, the mitochondrial cytochrome oxidase subunit I gene (COI), the single copy nuclear white gene and the ribosomal internal transcribed spacer two (ITS2). All the molecular markers supported the taxonomic status of a single species of An. albimanus. Furthermore, agreement between the COI and the white genes suggested a scenario of Pleistocene geographic fragmentation (i.e., population contraction) and subsequent range expansion across southern Central America. [ABSTRACT FROM AUTHOR]

Abstract: Australia and New Guinea share a common biogeographical history and unique vertebrate fauna. Investigation of genetic relationships among the wet forest-restricted pademelons (Macropodidae: Thylogale) provides insight into the historical connections between the two regions and the evolution of the Australasian marsupial fauna. Molecular phylogenetic relationships among Thylogale species were analysed using mitochondrial (12S rRNA and cytochrome b) and nuclear (omega-globin intron) sequence data with Bayesian and maximum likelihood methods. Australian species were resolved as well-supported, monophyletic clades, whereas endemic New Guinean species did not form clades consistent with current morphological taxonomy. Estimates of divergence using a Bayesian relaxed molecular clock model with standard mammalian nucleotide substitution rates indicated radiation of the genus in Australia in the mid to late Miocene. Persistence of Australian species of Thylogale in both southern temperate and northern tropical forests throughout the drying of the Australian continent can be attributed to their having a greater dietary flexibility than other browsing forest macropods. Divergence of the endemic New Guinean lineage occurred in the late Miocene to early Pliocene, indicating the presence of a partially forested landbridge connecting Australia and New Guinea during the Miocene. Mid-Pleistocene divergence between subspecies of the trans-Torresian T. stigmatica implies gene flow during glacial maxima between forest populations in the southern lowlands of New Guinea and the northern Cape York region of Australia. Complex structuring and relatively limited differentiation among populations of the endemic New Guinean species appears to have been influenced by the uplift of land and climate-induced redistribution of forest habitats during the late Pliocene and Pleistocene period. This is in strong contrast to the long evolutionary history and comparatively deep genetic divergence of Thylogale species in Australia. Further evaluation of the species status of the New Guinean Thylogale using more informative nuclear markers and extensive sampling is required. [Copyright &y& Elsevier]

Abstract: The current paper reports an experimental case study to test the heterogeneity of faunal assemblages from the Early-Middle Pleistocene Layers V-5 and V-6 of the Gesher Benot Ya’aqov Acheulian site (Israel). Tumbling and trampling experiments were initiated to gain qualitative insight into processes of bone modification and to assess the timing of the biostratonomic chronology, as it was assumed that both mechanisms were responsible for the formation of striations documented on the bone surfaces from the site. The tumbling experiments mimicked sediment movement in a calm lacustrine shoreline environment whereas the trampling experiments investigate the role of animal/hominin activities in dry, muddy and wet environments. Models for the internal operational sequence of an abrasional process due to uni- and multidirectional water movement and of a trampling scenario are presented. These models are used for the interpretation of the fauna from Gesher Benot Ya’aqov. [ABSTRACT FROM AUTHOR]

Abstract: Neanderthals have a distinctive suite of dental features, including large anterior crown and root dimensions and molars with enlarged pulp cavities. Yet, there is little known about variation in molar root morphology in Neanderthals and other recent and fossil members of Homo. Here, we provide the first comprehensive metric analysis of permanent mandibular molar root morphology in Middle and Late Pleistocene Homo neanderthalensis, and Late Pleistocene (Aterian) and recent Homo sapiens. We specifically address the question of whether root form can be used to distinguish between these groups and assess whether any variation in root form can be related to differences in tooth function. We apply a microtomographic imaging approach to visualise and quantify the external and internal dental morphologies of both isolated molars and molars embedded in the mandible (n =127). Univariate and multivariate analyses reveal both similarities (root length and pulp volume) and differences (occurrence of pyramidal roots and dental tissue volume proportion) in molar root morphology among penecontemporaneous Neanderthals and Aterian H. sapiens. In contrast, the molars of recent H. sapiens are markedly smaller than both Pleistocene H. sapiens and Neanderthals, but share with the former the dentine volume reduction and a smaller root-to-crown volume compared with Neanderthals. Furthermore, we found the first molar to have the largest average root surface area in recent H. sapiens and Neanderthals, although in the latter the difference between M1 and M2 is small. In contrast, Aterian H. sapiens root surface areas peak at M2. Since root surface area is linked to masticatory function, this suggests a distinct occlusal loading regime in Neanderthals compared with both recent and Pleistocene H. sapiens. [ABSTRACT FROM AUTHOR]

Abstract: Palaeopathological studies of the middle Pleistocene cranium from Florisbad (Free State, South Africa) document the presence of extensive cortical lesions and areas of thinning, a widened medullary cavity with destruction of the diploë, orbital roof lesions, a benign ectocranial neoplasm, and evidence for alveolar destruction, resorption, and antemortem tooth loss. Differential diagnosis suggests one or more possible aetiologies, including a haematological disorder, metabolic condition(s), Paget’s disease of bone, or non-specific infection perhaps following trauma. Moreover, if not directly associated with those on the external vault, orbital lesions alone could have been caused by infection or an indeterminable factor such as pressure from an enlarged organ. Multiple parasagittal lesions on the internal vault cortex probably represent expansile lesions left by enlarged arachnoid granulations. A multifactorial model of pathogenesis may be most appropriate to account for dentoalveolar lesions and antemortem tooth loss. Additionally, there are clear indications of diagenetic alteration deep within the vault, as well as multiple signs of degeneration on the cranium. These complicate the assessment of pathological alterations and identification of their possible aetiology. The Florisbad cranium is the latest specimen to join the growing sample of Pleistocene hominin remains with non-fatal and non-trivial pathological disorders adding to understanding of early human ecology and lifestyle. [ABSTRACT FROM AUTHOR]

Abstract: In 1995–1996 two isolated hominin lower incisors were found at the middle Pleistocene site of Boxgrove in England, with Lower Palaeolithic archaeology. Boxgrove 2 is a permanent lower right central incisor and Boxgrove 3 a permanent lower left lateral incisor. They were found separately, but close to one another and appear to belong to the same individual. The Boxgrove 1 tibia discovered in 1993 came from a different stratigraphic context and is thus believed to represent a different individual. This paper describes the morphology of the incisors, which is similar to other middle Pleistocene hominin specimens and, as with the tibia, suggests that they could be assigned to Homo heidelbergensis (recognising that the taxonomic status of this species is still a matter of debate). The incisors show substantial attrition associated with secondary dentine deposition in the pulp chamber and clearly represent an adult. They also show extensive patterns of non-masticatory scratches on the labial surfaces of both crown and root, including some marks which may have been made postmortem. The roots were exposed in life on their labial sides by a large dehiscence, extending almost to the root apex. This is demonstrated by deposits of calculus, polishing, and scratching on the exposed surfaces. The dehiscence may have been caused by repeated trauma to the gingivae or remodelling of the tooth-supporting tissues in response to large forces applied to the front of the dentition. [ABSTRACT FROM AUTHOR]

Abstract: Miocene to Pleistocene fossiliferous sediments in the Tugen Hills span the time period from at least 15.5 Ma to 0.25 Ma, including time periods unknown or little known elsewhere in Africa. Consequently, the Tugen Hills deposits hold the potential to inform us about crucial phylogenetic events in African faunal evolution and about long-term environmental change. Among the specimens collected from this region are a number of discoveries already important to the understanding of primate evolution. Here, we describe additional cercopithecoid material from the Miocene deposits in the Tugen Hills sequence, including those from securely dated sites in the Muruyur Beds (16–13.4 Ma), the Mpesida Beds (7–6.2 Ma) and the Lukeino Formation (∼6.2–5.7 Ma). We also evaluate previously described material from the Ngorora Formation (13–8.8 Ma). Identified taxa include Victoriapithecidae gen. et sp. indet., cf. Parapapio lothagamensis, and at least two colobines. Specimens attributed to cf. Pp. lothagamensis would extend the species’ geographic range beyond its type locality. In addition, we describe specimens sharing derived characters with modern African colobines (Tribe: Colobina), a finding that is congruent with previous molecular estimates of colobine divergence dates. These colobine specimens represent some of the earliest known members of the modern African colobine radiation and, in contrast to previous hypotheses, suggest that early African colobines were mainly arboreal and that semi-terrestrial Late Miocene and Plio-Pleistocene colobine taxa were secondarily derived in their locomotor adaptations. [ABSTRACT FROM AUTHOR]

Abstract: The classification and phylogenetic relationships of the middle Pleistocene human fossil record remains one of the most intractable problems in paleoanthropology. Several authors have noted broad resemblances between European and African fossils from this period, suggesting a single taxon ancestral to both modern humans and Neanderthals. Others point out ‘incipient’ Neanderthal features in the morphology of the European sample and have argued for their inclusion in the Neanderthal lineage exclusively, following a model of accretionary evolution of Neanderthals. We approach these questions using geometric morphometric methods which allow the intuitive visualization and quantification of features previously described qualitatively. We apply these techniques to evaluate proposed cranio-facial ‘incipient’ facial, vault, and basicranial traits in a middle-late Pleistocene European hominin sample when compared to a sample of the same time depth from Africa. Some of the features examined followed the predictions of the accretion model and relate the middle Pleistocene European material to the later Neanderthals. However, although our analysis showed a clear separation between Neanderthals and early/recent modern humans and morphological proximity between European specimens from OIS 7 to 3, it also shows that the European hominins from the first half of the middle Pleistocene still shared most of their cranio-facial architecture with their African contemporaries. [ABSTRACT FROM AUTHOR]

Abstract: Excavations at a complex of caves and open air sites at Pinnacle Point, Mossel Bay, Southern Africa have uncovered rich stratified assemblages of Middle Stone Age materials, including those from Pinnacle Point Cave 13B (PP13B) that comprises the first modernly excavated assemblage in southern Africa to be securely dated to the Middle Pleistocene. We report here on the complete excavated lithic artifact assemblage from PP13B. Both technological and typological analyses of the complete assemblage were performed. The assemblage-scale analysis allows for intrasite comparison as well as comparison of the PP13B assemblage with other sites from the region. No size-related pattern of change over time was observed within the PP13B assemblage, although there is significant evidence for varying strategies of lithic reduction between excavation areas within the cave. Comparison with other material from the Southern African MSA suggests that there is significant inter- and intra-site variability in the Southern African Middle Stone Age, even between portions of assemblages that are roughly contemporaneous. [ABSTRACT FROM AUTHOR]

Abstract: The Western Cape region of South Africa is home to a unique type of mediterranean vegetation called fynbos, as well as some of the earliest sites of modern human occupation in southern Africa. Reconstructing the paleohabitats during occupations of these early anatomically modern Homo sapiens is important for understanding the availability of resources to the humans during the development of behaviors that are often considered advanced. These reconstructions are critical to understanding the nature of the changes in the environment and resources over time. Here we analyze the craniodental fossils of the larger mammals recovered from two Pleistocene assemblages in the Pinnacle Point complex, Mossel Bay, Western Cape Region, South Africa. We reconstruct the paleohabitats as revealed by multivariate analyses of the mammalian community structures. Pinnacle Point 30 is a carnivore assemblage and Pinnacle Point 13B includes early evidence of a suite of modern human behavior; together they present an opportunity to identify environmental change over time at a localized geographic scale. Further, this is the first such study to include dated Western Cape localities from Marine Isotope Stage 6, a time of environmental pressure that may have marginalized human populations. Results indicate that environmental change in the Western Cape was more complex than generalized C4 grassland expansions replacing fynbos habitats during glacial lowered sea levels, and thus, resources available to early modern humans in the region may not have been entirely predictable. [ABSTRACT FROM AUTHOR]

Abstract: There has been a protracted debate over the evidence for intentional cranial modification in the terminal Pleistocene Australian crania from Kow Swamp and Coobool Creek. Resolution of this debate is crucial to interpretations of the significance of morphological variation within terminal Pleistocene–early Holocene Australian skeletal materials and claims of a regional evolutionary sequence linking Javan Homo erectus and Australian Homo sapiens. However, morphological comparisons of terminal Pleistocene and recent Australian crania are complicated by the significantly greater average body mass in the former. Raw and size-adjusted metric comparisons of the terminal Pleistocene skeleton from Nacurrie, south-eastern Australia, with modified and unmodified H. sapiens and H. erectus, identified a suite of traits in the frontal, parietal, and occipital bones associated with intentional modification of a neonate’s skull. These traits are also present in some of the crania from Kow Swamp and Coobool Creek, which are in close geographic proximity to Nacurrie, but not in unmodified H. sapiens or Javan H. erectus. Frontal bone morphology in H. erectus was distinct from all of the Australian H. sapiens samples. During the first six months of life, Nacurrie’s vault may have been shaped by his mother’s hands, rather than though the application of fixed bandages. Whether this behaviour persisted only for several generations, or hundreds of years, remains unknown. The reasons behind the shaping of Nacurrie’s head, aesthetics or otherwise, and why this cultural practice was adopted and subsequently discontinued, will always remain a matter of speculation. [ABSTRACT FROM AUTHOR]