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Weed prevention is recognized as one of the most cost-effective management strategies for invasive plants. In the field of invasive plant management increasing emphasis is being directed toward proactive management. However, land managers are still somewhat reluctant to aggressively employ prevention programs. Part of this reluctance could be due to lack of understanding of what a comprehensive prevention program entails. The purpose of this paper is to improve strategic decision-making for site-specific prevention programs, such as those on ranches or in watersheds. Our interest is in advancing prevention planning for land managers—the people who are faced with the constant pressure of potential invasive species infestations on a day-to-day basis. To facilitate more widespread use of prevention programs we are proposing definitions for key terminology to standardize and facilitate communication about prevention programs. Additionally, we present a flow model with the steps necessary to successfully implement such programs. The model has three categories from which specific prevention planning occurs: (1) education, (2) early detection and eradication, and (3) interruption of movement. The flow model directs users through a series of interlinked steps. Finally, we provide a case study in which a ranch manager implemented a prevention program using this framework. By using this model, managers are poised to conduct more strategic planning. This model also has applications in outreach and education programs to assist land managers in prevention planning.

Prevention programs are often assumed to be the most cost-effective method for managing invasive plants. However, there is very little information available about economic and biological factors that determine the forage benefits resulting from prevention programs. We developed an easy to use economic model to assess potential savings in livestock forage that might result from implementing prevention programs. The model can be used to determine potential loss in forage production caused by invasive plants and to estimate potential income savings by preventing invasive plant infestations. The model compares a prediction of populations with and without a prevention program using a logistic growth function. Animal unit month (AUM) price and interest rates are the primary economic input variables. The primary biological input variables are amount of invasive plant utilization, size of the initial infestation, and the spread rate with and without prevention. Our model suggests that as the AUM price increases and/or the interest rate decreases, the total savings increases for each AUM that was protected through a prevention program. The model also shows savings per AUM increases as the size of the initial infestation decreases, suggesting that prevention should focus on eliminating seed sources and seed production early in the program. Using our model inputs, the savings per AUM was about $9.20 for each percent reduction in spread rate over 100 yr.

Our objective was to evaluate the cost/benefit of a single herbicide application or targeted grazing of invasive annual grasses during restoration of partially invaded sagebrush steppe ecosystems used for livestock production. The cost/benefit model used is based on estimating the production of vegetation in response to implementing management and modeling cost/benefit economics associated with that prediction. The after-tax present value of added animal unit months (AUMs) obtained was lower than the present value of after-tax treatment costs after 20 yr for a single herbicide treatment, but higher than the present value of after-tax treatment costs for the grazing management scenario. Even at the highest weed utilization level, the value of added AUMs did not offset the cost of the treatment after 20 yr. However, the grazing treatment resulted in a value of added AUMs higher than the costs after 20 yr. Depending on the invasive weed utilization level, break-even points with targeted grazing occurred at anywhere from the first year to 7 yr. This assessment clearly shows that grazing management can be economically viable for managing annual grass-infested rangeland. In the future, models like the one used here can be improved by incorporating the rangeland management and restoration benefits on the wide variety of goods and services gained from rangeland.

The need for practical revegetation strategies for managing medusahead-infested rangeland is substantial and unmet. Our objective was to test the potential for using the single-entry approach (simultaneous application of herbicide and seed with one entry) developed for other invasive-weed infested systems for revegetating medusahead-infested rangeland. Since areas infested with medusahead are frequently burned and burning may help control medusahead and provide a more suitable seedbed for desired species, we also tested the single-entry approach in conjunction with burning. Our primary hypotheses were that (1) the combination of imazapic and seeding would provide best establishment of desired species and (2) burning would enhance the efficacy of imazapic on medusahead and enhance seedling establishment. Treatments included three seeding rates (none, 13.2, or 25.0 kg ha−1 of an even mixture of all species), two herbicides (0 and 52 g a.i. ha−1 imazapic; with and without), and two burning regimes (burned, not burned) applied mid-October 2006 on two sites. In late July, 2008 through 2010, plant density and biomass were sampled. We found that the simultaneous application of imazapic and seeding resulted in establishment of desired species where they were absent. Imazapic application without seeding increased perennial bunchgrass density where enough residual plants existed before treatment. Burning tended to improve the control of medusahead with imazapic and promoted desired species. Even though follow-up treatments may be necessary, applying imazapic and seeding in a single-entry approach may provide a more practical initial revegetation strategy than multi-entry approaches on rangeland devoid of desired residual species. This procedure should reduce the costs of treatment application and make revegetating annual grass-infested rangeland more affordable.

Producers facing infestations of invasive annual grasses regularly voice the need for practical revegetation strategies that can be applied across broad landscapes. Our objective was to determine the potential for scaling up the single-entry approach for revegetating medusahead-infested rangeland to broader, more heterogeneous landscape-scale revegetation of winter annual grass–infested rangeland. We hypothesized, when applied on a highly variable landscape scale, the combination of imazapic and seeding would provide highest abundance of perennial grasses and lowest amount of annual grasses. Treatments included a control, seeding of crested wheatgrass (‘Hycrest’) and Sandberg's bluegrass, spraying (60 g ai ha−1 imazapic), and a simultaneously applied combination of spraying and seeding. The HyCrest and Sandberg's bluegrass seeding rates were 19 and 3.4 kg ha−1, respectively. The treatments were applied to large plots (1.4 to 8 ha) and replicated five times, with each replication located in different watersheds throughout southeastern Oregon. This study shows that the single-entry approach can be scaled up to larger landscapes, but variation within establishment areas will likely be high. This procedure should reduce the costs over multientry treatment applications and make revegetating annual grass–infested rangeland across landscapes more affordable.

Rangeland health assessment provides qualitative information on ecosystem attributes. Successional management is a conceptual framework that allows managers to link information gathered in rangeland health assessment to ecological processes that need to be repaired to allow vegetation to change in a favorable direction. The objective of this paper is to detail how these two endeavors can be integrated to form a holistic vegetation management framework. The Rangeland Health Assessment procedures described by Pyke et al. (2002) and Pellant et al. (2005) currently are being adopted by land managers across the western United States. Seventeen standard indicators were selected to represent various ecological aspects of ecosystem health. Each of the indicators is rated from extreme to no (slight) departure from the Ecological Site Description and/or the Reference Area(s). Successional management identifies three general drivers of plant community change: site availability, species availability, and species performance, as well as specific ecological processes influencing these drivers. In this paper, we propose and provide examples of a method to link the information collected in rangeland health assessment to the successional management framework. Thus, this method not only allows managers to quantify a point-in-time indication of rangeland health but also allows managers to use this information to decide how various management options might influence vegetation trajectories. We argue that integrating the Rangeland Health Assessment with Successional Management enhances the usefulness of both systems and provides synergistic value to the decision-making process.

Land managers have long identified a critical need for a practical and effective framework for designing restoration strategies, especially where invasive plants dominate. A holistic, ecologically based, invasive plant management (EBIPM) framework that integrates ecosystem health assessment, knowledge of ecological processes, and adaptive management into a successional management model has recently been proposed. However, well-defined principles that link ecological processes that need to be repaired to tools and strategies available to managers have been slow to emerge, thus greatly limiting the ability of managers to easily apply EBIPM across a range of restoration scenarios. The broad objective of this article is to synthesize current knowledge of the mechanisms and processes that drive plant community succession into ecological principles for EBIPM. Using the core concepts of successional management that identify site availability, species availability, and species performance as three general drivers of plant community change, we detail key principles that link management tools used in EBIPM to the ecological processes predicted to influence the three general causes of succession. Although we acknowledge that identification of principles in ecology has greatly lagged behind other fields and recognize that identification of ecological principles and the conditions in which they hold are still being developed, we demonstrate how current knowledge and future advances can be used to structure a holistic EBIPM framework that can be applied across a range of restoration scenarios.

Understanding the relative importance of various functional groups in minimizing invasion by medusahead is central to increasing the resistance of native plant communities. The objective of this study was to determine the relative importance of key functional groups within an intact Wyoming big sagebrush–bluebunch wheatgrass community type on minimizing medusahead invasion. Treatments consisted of removal of seven functional groups at each of two sites, one with shrubs and one without shrubs. Removal treatments included (1) everything, (2) shrubs, (3) perennial grasses, (4) taprooted forbs, (5) rhizomatous forbs, (6) annual forbs, and (7) mosses. A control where nothing was removed was also established. Plots were arranged in a randomized complete block with 4 replications (blocks) at each site. Functional groups were removed beginning in the spring of 2004 and maintained monthly throughout each growing season through 2009. Medusahead was seeded at a rate of 2,000 seeds m 22 (186 seeds ft 22 ) in fall 2005. Removing perennial grasses nearly doubled medusahead density and biomass compared with any other removal treatment. The second highest density and biomass of medusahead occurred from removing rhizomatous forbs (phlox). We found perennial grasses played a relatively more significant role than other species in minimizing invasion by medusahead. We suggest that the most effective basis for establishing medusahead-resistant plant communities is to establish 2 or 3 highly productive grasses that are complementary in niche and that overlap that of the invading species. Nomenclature: Medusahead, Taeniatherum caput-medusae (L.) Nevski TACA8; bluebunch wheatgrass, Pseudo

Manipulating plant litter to direct successional trajectories is rarely considered as a management strategy. Our objective was to determine the influence of litter from an intact native plant community on a community dominated by an invasive species within the same habitat type as well as the influence of litter from a community dominated by an invasive species on an intact native plant community. We hypothesized that litter amount, type (source), and fragment size would influence various functional groups within a native plant community differently than within a weed-dominated plant community. We used reciprocal plant litter exchanges between native and invasive plant–dominated grasslands to gain an initial understanding of litter's influence on the density and biomass of native grasses, native forbs, common St. Johnswort, and downy brome. Common St. Johnswort was not influenced by any treatment. Native grass density increased with application of low (454 g/m2) amounts of litter where the grasses were subordinate to common St. Johnswort, and adding native plant litter to the weedy site nearly doubled native grass biomass. Low amounts of finely fragmented litter and high amounts of coarse litter induced native forbs to produce about twice the biomass as found in the non–litter-amended controls. Our study suggests that plant litter may be a component of vegetation that can be managed to shift the plant community toward those plants that are desired.

Heterogeneity in disturbance regimes, propagule pools, and factors affecting plant performance are a ubiquitous feature of wildlands. We tested a conceptual framework, termed augmentative restoration, aimed at identifying and selectively repairing or replacing damaged processes based on their predicted influence on the three causes of succession: site availability, species availability, and species performance. This framework was tested at three sites each with a different cause of succession naturally occurring in an ephemeral wetland dominated by invasive plants that had varying levels of disturbance (site availability), remnant native plants (species availability), and water availability (species performance). Our hypotheses were (1) seeding combined with watering would augment meadow vole disturbance to increase desired species composition, (2) shallow tilling combined with watering would augment remnant native species, and (3) shallow tilling combined with seeding would augment mesic soils to increase desired species composition. Shallow tilling, watering, and seeding were applied in a factorial arrangement at all three sites. These eight treatment combinations were applied in a split-plot design with four replications to generate 32 whole plots (2 m2). The herbicide 2,4-D was applied on half of each whole plot to influence relative species performance. In two of the three sites, using augmentative restoration to guide our management approaches improved our decision as to the treatment combinations that would maximize seedling establishment. Selectively augmenting successional processes that remain intact by repairing or replacing processes occurring at inadequate levels can improve implementation of successional management and provide a refined process-based framework for restoration across heterogeneous landscapes. Besides the clear economic advantages of lower management inputs associated with augmentative restoration, avoiding unnecessary management inputs has the additional advantage of minimizing unintended negative impacts on ecosystem processes.

Invasion by annual grasses, such as cheatgrass, into the western U.S. sagebrush-steppe is a major concern of ecologists and resource managers. Maintaining or improving ecosystem health depends on our ability to protect or re-establish functioning, desired plant communities. In frequently disturbed ecosystems, nutrient status and the relative ability of species to acquire nutrients are important drivers of invasion, retrogression, and succession. Thus, these processes can potentially be modified to direct plant community dynamics toward a desired plant community. The overall objective of this review paper is to provide the ecological background of invasion by exotic plants and propose a concept to facilitate the use of soil nitrogen (N) management to achieve desired plant communities that resist invasion. Based on the literature, we propose a model that predicts the outcome of community dynamics based on N availability. The model predicts that at low N levels, native mid- and late-seral species ar...

Restoring range sites dominated by western juniper is central to maintaining healthy functioning shrub–steppe ecosystems. On sites without adequate species composition to respond favorably to juniper controlled by fire, revegetation is necessary. We tested the following two hypotheses related to restoration after juniper control: (1) higher seeding rates would translate into higher density of seeded species, and (2) a rich seeding mixture would provide higher density and biomass than monocultures. Western juniper control was done by cutting 25% of the trees in 2002, allowing cut trees to dry for one year, followed by a broadcast fire applied in October 2003 which killed the remaining live juniper trees. Seeding treatments were applied in 2003 and included seeding six native species in monocultures, seeding a mixture of all six species at four rates (16.8, 22.4, 28.0, or 33.6 kg ha−1of pure live seeds), and a nonseeded control. Treatments were applied on a Sagebrush/bunchgrass and Snowberry/fescue site. We found that bluebunch wheatgrass, Idaho fescue, big bluegrass, and western yarrow density ranged from 450 to 700 plants m−2, which was over six-fold that of the control in 2004 at both sites. Only arrowleaf balsamroot did not establish successfully. The density of big bluegrass nearly doubled from 2004 to 2005. The highest plant density resulted from the highest seeding rate. The highest biomass production was combination seeding at 22.4 kg ha−1on the Sagebrush/bunchgrass site and 33.6 kg ha−1on the Snowberry/fescue site. Seeding a combination of species resulted in a moderate to high density of plants and optimized plant diversity and richness over seeding monocultures.

Controlling invasive plant infestations is very costly and often unsuccessful. Preventing invasions is more cost-effective than controlling invasive plants after they are established. Because prevention guidelines do not suggest any tools or methods to limit wind dispersal of invasive plant seeds, we investigated the influence of neighboring vegetation height on seed dispersal of a wind-dispersed (yellow salsify) and nonwind-dispersed (medusahead) species. To examine the influence of neighboring vegetation height on dispersal, seeds of both species were released in front of an artificial stand of desert wheatgrass in a modified wind tunnel. Treatments were a complete factorial design with two species, four vegetation heights (10, 30, 40, and 60 cm), three wind speeds (3, 5.5, and 10 km h−1), and three release distances from the neighboring vegetation (0, 15, and 30 cm). The ability of medusahead and yellow salsify seeds to disperse was influenced by the height of neighboring vegetation. Increasing height of neighboring vegetation decreased the number of yellow salsify seeds dispersing across neighboring vegetation. The greatest percentage of medusahead seeds dispersed across the neighboring vegetation was at the shortest height. Based on these results, we suggest that maintaining or promoting tall vegetation neighboring invasive plant infestations may reduce wind dispersal of seeds. More research is needed to investigate the influence of varying heights, densities, structural attributes, and composition of vegetation neighboring infestations and the dispersal of invasive plants.

Species richness and diversity are important indicators of ecosystem function and may be related to plant community resistance to invasion by nonindigenous species. Our specific objective was to determine the influence of clopyralid plus 2,4-D, glyphosate, and fosamine, at different application rates and timings, on richness and diversity of total species, total native species, and total nonnative species within a Russian knapweed–infested plant community. Twenty-eight treatments (3 herbicides by 3 rates by 3 application timings, and an untreated control) were applied to two sites located along the Missouri River riparian corridor in Montana. Clopyralid plus 2,4-D, glyphosate, and fosamine were applied in June (spring rosette stage of Russian knapweed), July (bud to bloom stage of Russian knapweed), and August (flowering stage of Russian knapweed). Herbicide rates were clopyralid plus 2,4-D at 0.08 (clopyralid) + 0.42 (2,4-D), 0.13 + 0.67, and 0.18 + 0.92 kg ai ha−1; glyphosate at 0.6, 1.2, and 1.8 kg ai ha−1; fosamine at 3.6, 7.2, and 10.8 kg ai ha−1. Density of each species was recorded during June and August of 2001 and 2002. Species richness and Simpson's Reciprocal Index (1/D) were calculated. By August 2002, only the glyphosate treatment (4.6 species m−2) yielded greater total richness over that of the control (3.5 species m−2). At that time, diversity after applying clopyralid plus 2,4-D remained similar to that of the control (1.4), but glyphosate (2.3) and fosamine (2.0) increased total species diversity. Nonnative grasses and forbs accounted for the increases in richness and diversity. Glyphosate may be appropriate for enhancing ecosystem function and possibly niche occupation to preempt reinvasion by Russian knapweed, but restoring native species seems unlikely using any of these herbicides alone.

The objective of this study was to test the potential for using a single-entry procedure to revegetate Russian knapweed– and green rabbitbrush–infested rangeland. I hypothesized that simultaneously applying an herbicide and seeding in the fall would produce the highest establishment and growth of desired species. For Russian knapweed, treatments included three seeding rates (zero, low, and high) and two herbicides (with and without clopyralid plus 2,4-D) applied in either the spring (2003) or fall (2004). The seed mixture included pubescent wheatgrass, Siberian wheatgrass, and alfalfa. Grasses were seeded on October 21, 2003 (fall dormant), and April 5, 2004 (spring), using a no-till rangeland drill. The seeding rates consisted of 3.4, 6.8, and 9.3 (low) or 5.0, 9.0, and 12.3 (high) kg ha 21 of alfalfa, Siberian wheatgrass, and pubescent wheatgrass, respectively. For green rabbitbrush, treatments included two seeding levels (13.5 kg ha 21 of Siberian wheatgrass and not seeded) and three herbicides (clopyralid, dicamba, and 2,4-D) and a control applied in July. Clopyralid plus 2,4-D (0.21 plus 1.12 kg ha 21 ) and dicamba at 2.1 kg ha 21 gave 61 and 66% control of Russian knapweed and green rabbitbrush, respectively. Herbicides interacted with seeding to provide the highest density of seeded species on the Russian knapweed site. These data support the hypothesis that simultaneously applying an herbicide and seeding in the fall would produce the highest establishment and growth of desired species. Conversely, only seeding affected Siberian wheatgrass establishment on the rabbitbrush sites. It may be reasonable to seed Siberian wheatgrass without controlling rabbitbrush if forage production is the primary objective. Nomenclature: Clopyralid; 2,4-D; green rabbitbrush, Ericameria teretifolia (Dur. & Hilg.) Jepson ERITE; Russian knapweed, Acroptilon repens (L.) DC. ACRRE; alfalfa, Medicago sativa L.; pubescent wheatgrass, Agropyron intermedium var. trichophorum (Link) Halac; Siberian wheatgrass, Agropyron fragile (Roth) Candargy.

Invasive plant species have adversely affected rangelands throughout the world and continue to invade previously uninfested lands at an alarming rate. Previous efforts have focused on eradication and control; however, recent efforts have recognized that preventing invasive plant species from infesting new areas is more cost-effective and efficient than trying to restore the system after it is infested. One of the major components of prevention is limiting the introduction of the invasive plant to uninfested areas. Guidelines to limit the introduction of invasive plants into new areas are usually general and not developed to address differences in dispersal vectors among invasive plants. To limit the dispersal of invasive plants, land managers need a framework that assists them in identifying major spatial dispersal vectors and management strategies based on those vectors. We propose an initial conceptual framework that integrates the ecology of invasive plant dispersal with prevention management. The framework identifies major potential vectors by incorporating invasive plant seed adaptations for dispersal through space and infestation locations relative to vector pathways. The framework then proposes management strategies designed to limit dispersal by those specific vectors. The framework ALSo identifies areas where research could improve the effectiveness of dispersal-prevention strategies by providing additional management tools.

Land managers typically use herbicides, biological controls, fire, grazing, and revegetation to manage and restore rangeland dominated by invasive plants. Without careful planning and implementation, these tools may temporarily control the weeds but may ultimately have minimal influence on ecological processes, fail over the long term, and lead to weed reinvasion. This can result from the lack of a broad ecological perspective. Successional management provides a process-based framework for weed ecologists to develop and test integrated weed management strategies and for land managers to organize implementation of these strategies in a way that adequately addresses ecological processes. This framework offers land managers practical methods for modifying ecological processes to direct plant community composition away from invasive species and toward desired plant assemblages. To date, successional management has not gained widespread application because, in part, it has not been conceptually linked to other successional models. Therefore, we illustrate how other successional models can be incorporated within the framework. Incorporating other prevailing successional models will further elucidate ecological processes, offer additional management strategies, and widen the possibilities for ecologically based management of rangeland weeds. Approaching management of weed-infested rangeland through this process-based framework will enable managers to implement strategies that maximize the likelihood of success because these methods will be integrated based on ecological principles. Successional management should be adjusted as we gain a better understanding of the factors that drive succession.

Invasive weed managers are presented with a complicated and ever-enlarging set of management alternatives. Identifying the optimal weed management strategy for a given set of conditions requires predicting how candidate strategies will affect plant community composition. Although field experiments have advanced our ability to predict postmanagement composition, extrapolation problems limit the prediction accuracy achieved by interpreting treatment means as predictions. Examples of extrapolation problems include nonlinear relationships between competing plants, site-to-site variation in plant population growth rates, and the carrying capacities of desired species and weeds. Our objective was to develop a model that improves predictions of weed management outcomes by overcoming a subset of these problems. To develop the model, we used data from two field experiments in which four Kentucky bluegrass, six western wheatgrass, and six invasive plant (i.e., leafy spurge) densities were combined in field plots. Graphs of our model's predictions vs. observed field experiment data indicate that the model predicted the data accurately. Our model may improve predictions of plant community response to invasive weed management actions.

The impact of invasive weed management on plant community composition is highly dependent on location-specific factors. Therefore, treatment means from experiments conducted at a given set of locations will not reliably predict community response to weed management elsewhere. We developed a model that rescales treatment means to better match local conditions. The goal of this paper was to determine if this rescaling improves predictions. We used our model to predict leafy spurge stem length density and grass biomass data from field experiments. The experiments consisted of herbicide-treated plots, untreated controls, and, in some cases, grass seeding treatments. When herbicides suppressed leafy spurge, the model explained 21 to 48% more variation in grass response than did mean grass response to the same or similar herbicide treatments applied at other sites. When herbicides killed grass, the model explained 53% more variation in leafy spurge response than did mean leafy spurge response to the same herbicide treatment applied at other sites. We regressed model predictions against observed data and tested the null hypothesis that resulting slope terms were equal to 1.0. Because the null hypothesis was rejected in two of four tests, the model may systematically over- or underpredict in some situations. However, measurement error in the observed data, unintended herbicide injury, or an inaccurate allometric relationship may account for a major proportion of the systematic deviations, and these factors would not cause prediction error in some management applications. Because the model tends to be better than the means from experiments at predicting plant community composition, we conclude that the model could advance managers' ability to predict plant community responses to invasive weed management.

Herbicides are an important tool for managing weeds where prescribed fire is used for rangeland improvement. Understanding how the season of herbicide application relates to prescribed burning is important. Our objective was to determine the effect of picloram and chlorsulfuron on Dalmatian toadflax cover, density, and biomass, where these herbicides were applied in the fall before burning or in the spring before or after burning. Six herbicide treatments and an untreated check were applied in a randomized complete block design with four replications to a prescribed burn at two sites infested with Dalmatian toadflax in Montana, United States. Herbicides were applied in the fall preburn, spring preburn, and spring postburn. Site 1 was treated in 1999 and 2000, and site 2 was treated in 2000 and 2001. Cover, biomass, and density of Dalmatian toadflax were sampled in September 2000, 2001, and 2002 at site 1 and September 2001 and 2002 at site 2. At site 1, cover, biomass, and density of Dalmatian toadflax were at least 76% lower compared with the check in both spring-applied picloram treatments, whereas the fall picloram treatment had similar Dalmatian toadflax cover, biomass, and density compared with the check 3 yr after application. By 2002, chlorsulfuron reduced Dalmatian toadflax cover, biomass, and density by at least 79% compared with the check in all timings of application at site 1. At site 2, Dalmatian toadflax cover, biomass, and density were reduced by at least 86% for all picloram and chlorsulfuron treatments in 2002, 2 yr after application. Chlorsulfuron applied in the fall or the spring and picloram applied in the spring effectively suppressed Dalmatian toadflax cover, biomass, and density for up to 3 yr.

Developing ecological principles applicable to invasive plant management is central to implementing sustainable strategies. We tested portions of a potentially useful successional-based management framework to further our understanding of the relationship between disturbance and colonization during revegetation of invasive weed-dominated grasslands. We hypothesized (1) intermediate wheatgrass density and biomass would be greatest at highest seeding rates, (2) control and tillage procedures that increase the availability of safe sites would increase wheatgrass abundance, and (3) spotted knapweed density and biomass would be lowest in treatments with highest wheatgrass density and biomass. Treatments included three disturbance levels: (1) no disturbance, (2) application of glyphosate, and (3) fall tillage. Colonization treatments were seeding intermediate wheatgrass of 0, 500, 2,500, and 12,500 seeds m−2. Treatments were factorially applied in a randomized complete-block design with four replicatio...

Sustainable invasive weed management must address treatment effects on desired vegetation. Our objective was to determine the influence of clopyralid plus 2,4-D, glyphosate, and fosamine, at various application rates and timing, on the density and biomass of Russian knapweed and desired plant groups growing in association with this invasive weed. In a randomized complete block design with four replications, three herbicides by three herbicide rates by three herbicide application timings and a nontreated control were factorially applied to two sites located along the Missouri River riparian corridor in Montana. Clopyralid plus 2,4-D, glyphosate, and fosamine were applied during the spring rosette stage of Russian knapweed (June), the bud to bloom stage of Russian knapweed (July), or the flowering stage of Russian knapweed (August). Herbicide rates were considered low, medium, and high based on label rates of clopyralid plus 2,4-D, glyphosate, or fosamine. Density and biomass of all species were sampled 3 y...

Successful control of invasive plants can have unexpected effects on native plants and wildland systems. Therefore, it is important for managers of invasive species to be concerned with ecological mechanisms and processes like invasion resistance, environmental heterogeneity, and succession that direct plant community dynamics. Augmentative restoration is a management approach for restoring desired species on wildlands dominated by invasive plants, where functioning ecological processes are maintained by selectively augmenting only those processes that are not operating sufficiently. The study was conducted within the Mission Valley, Montana, in an area where meadow vole disturbance provided site availability for colonization. In a split-plot design with four replications, eight factorial treatment combinations from three factors (shallow tilling, watering, and seeding) were applied to whole plots, and 2,4-D was applied to subplots. Cover and density of seeded species, spotted knapweed, and sulfu...

It is becoming increasingly clear that prescriptions for rangeland weed control are not sustainable because they treat the symptoms of weeds rather than their cause. Future restoration of invasive plant–infested rangeland must be based on ecological principles and concepts that provide for predictable outcomes. A generalized objective for ecologically based weed management is to develop and maintain a healthy plant community that is largely invasion resistant. Successional management based on ecological principles involves modifying the processes controlling the three general causes of succession: disturbance, colonization, and species performance. The processes controlling plant community dynamics can be modified to allow predictable successional trajectories. Successional management can lead to biomass optimization models for grazing management, spread vector analysis, and using resource availability to direct weedy plant communities toward those that are desired. Our challenge is to develop ec...

Reduction of weed competition using herbicides is critical to the successful revegetation of weed-infested rangeland. However, little is known about the influence of persistent broadleaf herbicides on the establishment of many desired grasses. The objective of this study was to determine the influence of various rates and times of application of picloram and clopyralid before seeding on two native and two nonnative grasses important to western United States rangeland. The study was conducted in 1997 and 1998 in Wyoming, Montana, and Washington. Five herbicide treatments (none, picloram at 0.14, 0.28, 0.56 kg ai/ha, clopyralid at 0.14, 0.56 kg ai/ha), three timings of herbicide application (44, 24, and 0 d prior to seeding[DPS]), and four grass species (Idaho fescue, bluebunch wheatgrass, crested wheatgrass, and pubescent wheatgrass) were factorially arranged in a randomized split-plot design with three blocks. The whole-plots were the seeded grass species, and the subplots were the herbicide treatments an...

Our objective was to maximize Canada thistle control and plant community diversity in a waterfowl production area administered by the U.S. Fish and Wildlife Service. We tested three rates (1.5, 3.0, and 4.5 ai/ha) of glyphosate applied during spring, summer, or fall using two application methods. The lowest rate of glyphosate decreased the Canada thistle density by about 30% relative to the control. Glyphosate applied in the fall decreased Canada thistle density below that of the control more consistently than when applied in spring or summer. Wick application generally resulted in less Canada thistle biomass than did broadcast application. Species richness was generally higher when glyphosate was wick applied, and all rates of this application method increased species richness when compared with the control. We recommend fall wick application of glyphosate at 1.5 kg ai/ha to control Canada thistle near the riparian areas. This application provided optimum Canada thistle control, while maintaining species...

Meadow foxtail is a rhizomatous grass widely grown for hay and pasture in wet meadows of the western United States and Canada. Two sulfonylurea herbicides, chlorsulfuron and metsulfuron-methyl, were evaluated for their effects on meadow foxtail biomass. Both herbicides were applied at four doses, 0.035, 0.070, 0.105, and 0.140 kg ai/ha, together with a control at each of two sites in October 2003. Treatments were replicated four times at each site and arranged in a randomized complete block design and sampled in July 2004 and 2005. Meadow foxtail biomass depended on site (P = 0.001) or year (P = 0.001), but not herbicide treatment (P = 0.182). No biomass production losses resulted from applying up to 0.14 kg/ha of either chlorsulfuron or metsulfuron-methyl on meadow foxtail, even in relatively high-pH soils. Nomenclature: Chlorsulfuron; metsulfuron-methyl; meadow foxtail, Alopecurus pratensis L.

Competition between bluebunch wheatgrass and spotted knapweed was quantified using three addition series experiments in an environmental chamber. Using two density matrices, spotted knapweed plants were thinned by either 45 or 90% of their original densities 40 d after emergence. Another matrix of density combinations was not thinned. All plant material was harvested 90 d after thinning. Spotted knapweed was about four times more competitive than bluebunch wheatgrass. Reducing spotted knapweed by 45% did not alter the competitive relationship between the two species. We believe remaining knapweed individuals captured the majority of the newly available resources. Ninety percent reduction was necessary to shift the competitive relationship in favor of bluebunch wheatgrass. Successful integrated spotted knapweed management must exploit key mechanisms and processes directing plant community dynamics, in conjunction with weed density reduction, if communities are to be shifted toward those that are desired.

An educational software program was developed to demonstrate the interactions among weed population demographic processes, plant community interactions, and grassland noxious weed manage- ment options. The software includes an example model parameterized to simulate yellow starthistle population dynamics and graphic output. The software allows students to change weed and competing grass population demographic parameters, herbicide, and biological control management options. The graphic output allows the user to observe the outcome of integration of weed biology and various weed management options. Nomenclature: Yellow starthistle, Centaurea solstitialis L. #3 CENSO. Additional index words: Biological control, integrated weed management, weed interference model.

Sustainable rangeland management will require successional strategies to deal with the expanding weed problem. These strategies must be consistent with the view that plant communities are dynamic and technology is used to enhance the natural processes and mechanisms that direct vegetation change. The goal is to shift the dynamics toward a desired plant community. A unified conceptual model is necessary to direct the development and application of successional weed management systems. We propose using a resource management model as a conceptual basis for successional weed management. This model is based on the primary causes of succession: site availability, differential species availability, and differential species performance. This model provides the mechanistic framework necessary for developing successional weed management systems and it is meant to enhance communication among rangeland weed managers and scientists.

Effectiveness of the common soil fungusSclerotinia sclerotiorumas a biological control for spotted knapweed and its effect on competitive interactions between spotted knapweed and bluebunch wheatgrass were evaluated in a growth chamber study using addition series methods. Total seeding densities ranged from 2000 to 60 000 seeds/m2. Mean bluebunch wheatgrass plant weight was 3.5 times greater than spotted knapweed weight per plant, respectively. Coefficient ratios estimating species interaction showed bluebunch wheatgrass density had a greater influence than spotted knapweed density on both bluebunch wheatgrass and spotted knapweed weights (2.11 and 0.51, respectively) when not under the influence ofS. sclerotiorum. Niche differentiation ratios indicated a lack of resource partitioning between species (1.11).S. sclerotiorumreduced spotted knapweed density by 68 to 80% without reducing bluebunch wheatgrass density. Spotted knapweed weight per plant also was reduced by the addition of 5.sclerotiorum(1.4 to 1.2 mg) but there was not a corresponding increase in bluebunch wheatgrass weight.S. sclerotiorumdecreased competition between spotted knapweed and bluebunch wheatgrass. This study provides evidence that establishment of bluebunch wheatgrass on spotted knapweed infested rangeland may be improved by combiningS. sclerotiorumwith high grass seeding rates.

Rapid germination and root growth have been identified as important plant characteristics for establishment in arid environments. This study evaluates germination, radicle elongation, and root growth rates of two annual grasses, two forage species, and yellow starthistle. Yellow starthistle germination, radicle elongation, and root growth were faster than those of medusahead and hedgehog dogtailgrass. Subterranean clover possessed rapid seedling growth characteristics similar to those of yellow starthistle, whereas Palestine orchardgrass was most delayed.