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Level TE9c of the Sima del Elefante site (Sierra de Atapuerca, Spain) is one of the oldest sites with evidence of human occupation in western Europe. We began excavating level TE9c in 2003, and the work there continues today. The studies of the archaeology, palaeontology and geology from this locality have provided an indispensable dataset with which to capture a picture in the scenario of the origin of humans in Europe. Based on these data, we raise and discuss several topics, such as the possible origin of the lineage of the first hominins that inhabited western Europe; their capacity to have active hunting or scavenging abilities; whether their subsistence strategies were successful; and what the environment and habitats where these hominin groups settled was like. The aim of this paper is to present the results and discussions obtained from the research team and to establish the primary features of early human occupations in southwestern Europe. Tentatively, we may conclude, based on the events recorded at TE9c, that the first humans were in the Iberian peninsula at around 1.2 Ma they used the caves of the Sierra de Atapuerca as shelters probably during their hunting activities; the cavities were surrounded by Mediterranean forest, rivers and water ponds, and varied habitats as suggested by the rich and diverse assemblage of fossils of vertebrates (fish, amphibians and reptiles, birds, large and small mammals); where humans possibly caught what they found in the surroundings.

Received 16 June 2009 / Received in revised form 16 February 2010 / Accepted 16 February 2010 / Available online 23 March 2010 © 2010 Elsevier Ltd. All rights reserved. Corresponding author. Tel.: +34 947 255006. E-mail address: jesus.rodriguez@cenieh.es (J. Rodríguez). Supplementary material associated with this article can be found in the online version at doi:10.1016/j.quascirev.2010.02.021., The present paper analyses the evidence provided by three sites (Sima del Elefante, Gran Dolina, and Galería) located in the Trinchera del Ferrocarril of the Sierra de Atapuerca. These three sites are cave infillings that contain sediments deposited from approximately 1.2 Ma to 200 kyr. Pollen, herpetofauna, and small and large mammal remains are used as proxies to obtain a general picture of the environmental changes that occurred at the Sierra de Atapuerca throughout the one million-year period represented at these sites. Similarly, cultural changes are tracked analyzing the evidence of human behavior obtained from the study of several bone and lithic assemblages from these three sites. At least three periods with different cultural features, involving technology, subsistence and behavior, are determined from the available evidence. The first two periods correspond to the Mode 1 technology and Homo antecessor: the first is dated around 1.2 to 1.0 Ma and reflects opportunistic behavior both in technology and subsistence. The second period is around 800 kyr BP. Mode 1 technology is still maintained, but subsistence strategies include systematic hunting and the use of base camps. The third period is dated between 500 ka and 200 ka and corresponds to the Mode 2 technology and the acquisition of directional hunting and other organizational strategies by Homo heidelbergensis. A transition from Mode 2 to Mode 3 seems to appear at the end of this time-range, and may reflect the early phases of a fourth cultural change. With regard to the environment, our main conclusion is that there was an absence of extremely harsh conditions at Atapuerca throughout this time period. The presence of Mediterranean taxa was constant and the dominant landscape was a savannah-like open environment, probably with small forest patches. An alternation of Mediterranean and mesic species as the dominant component of the tree storey was induced by the climatic cycles, and steppes spread across th, Ministerio de Ciencia e Innovación (España), Ministerio de Ciencia e Innovación (España), Ministerio de Ciencia e Innovación (España), Ministerio de Educación y Ciencia (España), Junta de Castilla y León, Ministerio de Ciencia e Innovación (España), Depto. de Geodinámica, Estratigrafía y Paleontología, Fac. de Ciencias Geológicas, TRUE, pub

Palaeobotanical (pollen, charcoal) data from Gorham's Cave reveals a diversified landscape in the Gibraltar region during the Middle (c. 32 560-23 780 year BP) and Upper Palaeolithic (c. 18 440-10 880 BP). Inferred vegetation types include oak, pine, juniper, and mixed woodlands and savannahs, grasslands with heaths, heliophytic matorrals, phreatophytic formations (e.g. riverine forests, wetlands), and thermomediterranean coastal scrub. A revision of palaeoecological data suggests that patches of trees persisted even in northern and continental territories of the Iberian Peninsula during the cold stages of OIS3 and OIS2. However, a southern Mediterranean coastal shelf extending from Gibraltar to Málaga, and probably further north up to Murcia, was unique in its combination of thermo-, meso-, and supramediterranean plant and animal species. Given the composition of these assemblages, this shelf and its adjacent mountains represented a crucial reservoir of biodiversity during the Upper Pleistocene. It is within this physiographically complex context with its diversity of resources where the last Neanderthals extraordinarily survived until c. 24 000 BP, that is over 10 000 years later than the disappearance of Neanderthals from elsewhere in temperate Europe. © 2008 Elsevier Ltd. All rights reserved.