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Abstract Cretoxyrhina mantelli was a large pelagic lamniform shark geographically widespread during the Late Cretaceous, and well known because of several nearly complete skeletons from the Western Interior Seaway of North America. Here we report 15 partial skeletons belonging to lamniform sharks from the 'lastame' lithozone of the Upper Cretaceous Scaglia Rossa Formation of the Lessini Mountains (northeastern Italy). Seven partial but articulated skeletons include tooth sets that allow a confident attribution to Cretoxyrhina mantelli based on dental morphologies. We review the taxonomic history of C. mantelli , evidencing that the taxon was erected by Agassiz (1835) and tracing back four of the original syntypes. Based on calcareous plankton biostratigraphy, the rock in which the Italian skeletal remains are embedded is constrained to the middle-upper Turonian. Total length estimates of the specimens suggest that the sample includes the largest specimen of Cretoxyrhina mantelli (615–650 cm estimated total length) known to date. The placoid scale morphology indicates that C. mantelli most likely was a fast swimmer with a similar ecology as the extant white shark, Carcharodon carcharias. The associated skeletal elements of the specimens represent large chondrichthyan deadfalls and the cadavers decayed on the seafloor where they remained exposed for several months, as indicated by bioerosional traces, some of which are interpreted as a product of bone-eating worm activities and other bioerosional traces with Gastrochaenolites -like structure. The Cretoxyrhina mantelli remains described herein provide new information about the ʻlastameʼ vertebrate assemblage, which seemingly was strongly dominated by chondrichthyans, especially lamniform sharks. Highlights • Description of partial skeletal remains of the ʻginsuʼ shark Cretoxyrhina mantelli. • Nomenclatural Remarks about the species C. mantelli. • Emendation of the diagnosis of the species C. mantelli. • Paleobiological and taphonomic Remarks about the deadfalls of C. mantelli. • Micropaleontological analyses on the C. mantelli specimen matrix. [ABSTRACT FROM AUTHOR]

Sclerorhynchiform sawfishes are a diverse and extinct clade of elasmobranchs that is restricted to the Cretaceous. Most taxa are known only by isolated rostral spines, whereas skeletal remains are rare and have been reported from a small number of Upper Cretaceous localities. Here, we describe skeletal remains of the giant sclerorhynchiform Onchosaurus pharao for the first time, which provides new morphological information. The single specimen comes from middle-basal upper Turonian strata of the Lessini Mountains in northeastern Italy and represents the first record of this genus from Italy. The specimen consists of unidentifiable cranial remains, several diagnostic rostral spines, the rostrum with fragments of tessellated calcified cartilage, and 87 disarticulated vertebrae. The rostrum preserves the characteristic sensory system of sclerorhynchiforms. It is devoid of any lateral sockets indicating that rostral spines were attached laterally to its surface. This pattern is identical to most sclerorhynchiforms and extant pristiophoriformes implying also similar replacement patterns as in most other sclerorhynchiforms with the exception for Schizorhiza . Additionally, the bases of two longitudinally arranged rows of ventral rostral spines are identifiable concurring with patterns seen in Sclerorhynchus . The axial skeleton is partly preserved. Re-arranging the disarticulated vertebrae according to their life position in combination with measures of the size and thickness of preserved vertebral centra, and the ratio rostrum length/body size depending on the number of vertebral centra indicate that the specimen was ca. 450 cm long. Growth rings in the vertebral centra show that the specimen was about four years old and thus probably not yet fully sexual mature when it died. This age assumption corresponds well with the calculated size when compared with complete skeletons of extinct sclerorhynchiforms and extant pristiforms. The size of the specimen and its occurrence in hemipelagic rocks corroborates previous assumptions that this sclerorhynchiform was a large and pelagic sawfish. [ABSTRACT FROM AUTHOR]

The Coccondontoidea is a distinctive superfamily of pycnodont fishes characterized by a series of derived characters, including thick and well-ossified cranial bones supporting horns and spines and a hypertrophied pectoral girdle sutured to the skull forming a sort of cephalo-torax. A new distinctive coccodontoid species, Gladiopycnodus byrnei n. sp., is described from the Cenomanian locality of Hjoûla, Lebanon. This new species exhibits a unique combination of features that clearly support its inclusion within the coccodontoid family Gladiopycnodontidae, including an extremely elongate snout forming a sword-shaped rostrum, thick dermal bones ornamented with rounded tubercles, large supracleithrum and cleithrum extensively sutured to the skull, and irregularly imbricated scales covering the entire body. The new taxon differs from type species G. karami by having different meristic counts and body proportions, dermal bones strongly ornamentated, skull roof with partially fused bones, scales ornamented with small tubercles, and rounded scales covering the caudal peduncle absent. The presence of pectoral fins in our exquisitely well-preserved specimen allowed us to redefine the morphology of the genus Gladiopycnodus, also modifying the diagnosis of the family. In order to interpret the patterns of morphospace occupation and quantify the morphological diversification of pycnodonts through time, the geometric morphometric approach was used. The morphospace analysis revealed that a significant increase in morphological disparity of pycnodonts during the Late Cretaceous was related, at least in part, to the appearance of the representatives of the superfamily Coccodontoidea. The rapid evolutionary radiation of these well-armored pycnodonts was interpreted as an adaptive solution in response to the large predatory-prey escalation in the context of the Mesozoic Marine Revolution. [ABSTRACT FROM AUTHOR]

The Molí del Baró-1 site (north-eastern Iberia) has yielded one of the richest fossil assemblages of the continental upper Maastrichtian from western Europe. It includes abundant plant (charophytes, sporomorphs, angiosperm leaves, seeds and logs), invertebrate (mollusc shells, partial insect exoskeletons and eggs) and vertebrate (teeth, bones, eggshells) fossils. The fossil assemblage is composed of both parautochthonous and allochthonous remains that mostly deposited in a pool formed at the edge of a meandering river. Plant megafossil evidence suggest that at least three angiosperm species (a willow-like dicot, palms and a likely typhacean monocot) formed the plant community growing in this fluvial environment. Tooth remains indicate that crocodylomorphs were diverse (including atoposaurids, ziphosuchians and eusuchians) which together with small theropod dinosaurs likely consumed prey carcasses or scavenged near the depositional setting. Prismatoolithid eggshells also suggest the presence of nearby nesting area of theropods. On the other hand, hadrosauroids apparently dominated phytophagous faunas including lambeosaurines as well as more basal forms. The taxonomic composition of the Molí del Baró-1 site matches the expected for the timespan and environment and represents a relevant locality to know the diversity of the latest Cretaceous terrestrial ecosystems in south western Europe. [ABSTRACT FROM AUTHOR]

Abstract: In this study, we erect Sauroniops pachytholus gen. et sp. nov., a large-bodied theropod dinosaur from the Cenomanian (Upper Cretaceous) of Morocco, on the basis of an almost complete frontal showing a unique combination of features including a naso–frontal suture extended along 40% of the frontal length, a thick dome in the anterolateral corner of the dorsal surface, a trapezoidal prefrontal facet that is restricted to the anterodorsal margin of the lateral surface of the frontal with no participation in the orbital roof and separated from the lacrimal facet by a narrow vertical lamina, a hypertrophied ‘D-shaped’ lacrimal facet that is four times the anterior depth of the postorbital facet, and a raised posteromedial margin of the dorsal surface describing a saddle with the anterolateral dome and confluent with a series of anteromedial rugosities. Phylogenetic analysis found robust support for placing Sauroniops among the basal carcharodontosaurids and related to Eocarcharia, showed that some of the unusual features of the new theropod were convergently acquired by abelisaurids, and revealed a mosaic pattern in the evolution of the carcharodontosaurid skull table. The frontals of Sauroniops and Carcharodontosaurus, both from the ‘Kem Kem compound assemblage’ of Morocco, show comparable size but differ in the extent of the naso–frontal articulation, the shape and disposition of the prefrontal and lacrimal articulations, the development of dorsal ornamentation and the morphology of the supratemporal fossa. Among carcharodontosaurids, the skull table developed unique configurations among each lineage and appears diagnostic at the species-level. The dome-like frontal in Sauroniops may indicate head-butting behaviour in this taxon or evolved for visual display. [Copyright &y& Elsevier]

Abstract: An isolated theropod dinosaur tooth discovered in 1984 near Nalut in northwestern Libya is the only known dinosaur record from the Aptian–Albian (Lower Cretaceous) Chicla Formation in the Jabal Nafusah region of the country. The tooth was re-examined in an effort to better ascertain its taxonomic affinities. A stepwise discriminant function analysis compared the Libyan tooth to the dentitions of 24 other theropods and classified the tooth with cranial material from the Late Cretaceous of India previously referred to the abelisaurid Indosuchus. The temporal and paleogeographic “distance” separating the Libyan specimen from “Indosuchus” indicates that the former cannot pertain to that genus. However, the results of the analysis and synapomorphies of Abelisauridae present on the Libyan crown indicate that it can be referred to the clade, thus contributing to the growing record of Abelisauridae in the Cretaceous of mainland Africa. [Copyright &y& Elsevier]