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A detailed understanding of decadal to millennial-scale climate changes requires seasonal-scale (summer-winter) reconstructions of past precipitation and temperature fluctuations. Comparing seasonally resolved varve records with pollen-based sum of growing degree-days (GDD) reconstructions from Lake Nautajärvi, we examined the intra-annual nature of climate variability in central southern Finland during the Holocene. The organic varve record and the GDD reconstruction show roughly comparable trends supporting the interpretation that both proxies predominantly reflect summer temperatures in the study area. The records suggest low but rising early-Holocene (9500 to 8500 cal. yr BP) summer temperatures. The Holocene Thermal Maximum (HTM) in the GDD record dates to about 7500 to 4500 cal. yr BP, but the organic varve record along with reconstructed changes in vegetation composition, notably a peak of Tilia pollen percentages, indicate that during the HTM there was a trend towards a more continental climate with maximum mid-summer temperatures reached at 6500 to 4500 cal. yr BP. Both records reflect the start of the post-HTM cooling at about 4500 cal. yr BP, simultaneously with an increase of the amount of catchment erosion and mineral matter influx into the lake, suggesting gradually colder and/or longer winters with high net accumulation of snow. The organic varve record and the GDD record start to diverge at 2000 cal. yr BP, possibly owing to the human influence on catchment processes. The reconstructed mid-Holocene summer temperature peak deviates from the regional climate model outputs, which suggest highest summer temperatures during the early Holocene. [ABSTRACT FROM AUTHOR]