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Recent analyses have demonstrated that personality affects political behavior. According to the mediation hypothesis, the effect of personality on political participation is mediated by classical predictors, such as political interest, internal efficacy, political discussion, or the sense that voting is a civic duty. This paper outlines various paths that link personality traits to two participatory activities: voter turnout in European Parliament elections and participation in protest actions. The hypotheses are tested with data from a large, nationally representative, face-to-face survey of the Spanish population conducted before and after the 2009 European Parliament elections using log-linear path models that are well suited to study indirect relationships. The results clearly confirm that the effects of personality traits on voter turnout and protest participation are sizeable but indirect. They are mediated by attitudinal predictors. [ABSTRACT FROM AUTHOR]