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Oxygen isotopes in animal tissues are directly related to body water composition and thus the environment. Accurate measurement of animal tissue δ18O provides information about local climate, an animal's geographical origin and subsequent movements, with wide applications in palaeobiology and forensic science. The genesis and evolution of tissue-based oxygen isotopes within species and within individuals are complex. We present the first data, for non-human primates, rhesus macaques (Macaca mulatta), on the relationship between oxygen isotope sources in bio-apatite (PO4 and PCO3) and hair taken from six sample sites in Asia, ranging from western India to northern Vietnam. The range of values is similar within each tissue type, with good correlation between tissues (r = 0.791 to 0.908), allowing cross-tissue extrapolations. This is important when the availability of suitable tissues is limited. Biological interpretation of the small data set is difficult: macaque diets are eclectic, and the samples are from various locations. However, factors such as overall climate, precipitation quantity and source, and altitude are clearly influencing the results for each discrete geographical grouping. Future work could be aimed at assessing δ18O tissue associations for other species as the relationships appear to be species-specific.

1. The brown bear, Ursus arctos, was Holocene Britain’s largest carnivoran and has appeared in recent rewilding discussions. 2. Despite widespread interest, we know very little about the species in Holocene Britain as few studies have been undertaken. This paper draws together information on the brown bear to examine the presence and extinction of bears through examination of the archaeological and palaeontological evidence. 3. Data were collected from published literature and museum catalogues. Information on the chronological date of the sites, number of specimens, and the body parts present were particularly sought. 4. A total of 86 sites were identified, 57 are well dated, 27 are of uncertain date but likely to be Holocene, and two have uncertain identifications. Very few specimens from non-archaeological sites (i.e. fens and caves) are well-dated, skewing the data towards anthropogenic sites such as settlements, castles, and graves. 5. Analysis of body part representation shows that the bear bones found in the Bronze Age, Iron Age and most of the early medieval period are from skins. All body parts are present during the Romano-British, medieval, and post-medieval period, indicating that live bears were present, and were probably imported for entertainment. 6. It seems the brown bear was rare throughout the Holocene, and based on current evidence two extinction scenarios can be put forward: extinction in the late Neolithic/early Bronze Age, or extinction in the early medieval period.

1. The brown bear, Ursus arctos, was Holocene Britain’s largest carnivoran and has appeared in recent rewilding discussions. 2. Despite widespread interest, we know very little about the species in Holocene Britain as few studies have been undertaken. This paper draws together information on the brown bear to examine the presence and extinction of bears through examination of the archaeological and palaeontological evidence. 3. Data were collected from published literature and museum catalogues. Information on the chronological date of the sites, number of specimens, and the body parts present were particularly sought. 4. A total of 86 sites were identified, 57 are well dated, 27 are of uncertain date but likely to be Holocene, and two have uncertain identifications. Very few specimens from non-archaeological sites (i.e. fens and caves) are well-dated, skewing the data towards anthropogenic sites such as settlements, castles, and graves. 5. Analysis of body part representation shows that the bear bones found in the Bronze Age, Iron Age and most of the early medieval period are from skins. All body parts are present during the Romano-British, medieval, and post-medieval period, indicating that live bears were present, and were probably imported for entertainment. 6. It seems the brown bear was rare throughout the Holocene, and based on current evidence two extinction scenarios can be put forward: extinction in the late Neolithic/early Bronze Age, or extinction in the early medieval period.

1. The brown bear, Ursus arctos, was Holocene Britain’s largest carnivoran and has appeared in recent rewilding discussions. 2. Despite widespread interest, we know very little about the species in Holocene Britain as few studies have been undertaken. This paper draws together information on the brown bear to examine the presence and extinction of bears through examination of the archaeological and palaeontological evidence. 3. Data were collected from published literature and museum catalogues. Information on the chronological date of the sites, number of specimens, and the body parts present were particularly sought. 4. A total of 86 sites were identified, 57 are well dated, 27 are of uncertain date but likely to be Holocene, and two have uncertain identifications. Very few specimens from non-archaeological sites (i.e. fens and caves) are well-dated, skewing the data towards anthropogenic sites such as settlements, castles, and graves. 5. Analysis of body part representation shows that the bear bones found in the Bronze Age, Iron Age and most of the early medieval period are from skins. All body parts are present during the Romano-British, medieval, and post-medieval period, indicating that live bears were present, and were probably imported for entertainment. 6. It seems the brown bear was rare throughout the Holocene, and based on current evidence two extinction scenarios can be put forward: extinction in the late Neolithic/early Bronze Age, or extinction in the early medieval period.

1. The brown bear, Ursus arctos, was Holocene Britain’s largest carnivoran and has appeared in recent rewilding discussions. 2. Despite widespread interest, we know very little about the species in Holocene Britain as few studies have been undertaken. This paper draws together information on the brown bear to examine the presence and extinction of bears through examination of the archaeological and palaeontological evidence. 3. Data were collected from published literature and museum catalogues. Information on the chronological date of the sites, number of specimens, and the body parts present were particularly sought. 4. A total of 86 sites were identified, 57 are well dated, 27 are of uncertain date but likely to be Holocene, and two have uncertain identifications. Very few specimens from non-archaeological sites (i.e. fens and caves) are well-dated, skewing the data towards anthropogenic sites such as settlements, castles, and graves. 5. Analysis of body part representation shows that the bear bones found in the Bronze Age, Iron Age and most of the early medieval period are from skins. All body parts are present during the Romano-British, medieval, and post-medieval period, indicating that live bears were present, and were probably imported for entertainment. 6. It seems the brown bear was rare throughout the Holocene, and based on current evidence two extinction scenarios can be put forward: extinction in the late Neolithic/early Bronze Age, or extinction in the early medieval period.

1. The brown bear, Ursus arctos, was Holocene Britain’s largest carnivoran and has appeared in recent rewilding discussions. 2. Despite widespread interest, we know very little about the species in Holocene Britain as few studies have been undertaken. This paper draws together information on the brown bear to examine the presence and extinction of bears through examination of the archaeological and palaeontological evidence. 3. Data were collected from published literature and museum catalogues. Information on the chronological date of the sites, number of specimens, and the body parts present were particularly sought. 4. A total of 86 sites were identified, 57 are well dated, 27 are of uncertain date but likely to be Holocene, and two have uncertain identifications. Very few specimens from non-archaeological sites (i.e. fens and caves) are well-dated, skewing the data towards anthropogenic sites such as settlements, castles, and graves. 5. Analysis of body part representation shows that the bear bones found in the Bronze Age, Iron Age and most of the early medieval period are from skins. All body parts are present during the Romano-British, medieval, and post-medieval period, indicating that live bears were present, and were probably imported for entertainment. 6. It seems the brown bear was rare throughout the Holocene, and based on current evidence two extinction scenarios can be put forward: extinction in the late Neolithic/early Bronze Age, or extinction in the early medieval period.

1. The brown bear, Ursus arctos, was Holocene Britain’s largest carnivoran and has appeared in recent rewilding discussions. 2. Despite widespread interest, we know very little about the species in Holocene Britain as few studies have been undertaken. This paper draws together information on the brown bear to examine the presence and extinction of bears through examination of the archaeological and palaeontological evidence. 3. Data were collected from published literature and museum catalogues. Information on the chronological date of the sites, number of specimens, and the body parts present were particularly sought. 4. A total of 86 sites were identified, 57 are well dated, 27 are of uncertain date but likely to be Holocene, and two have uncertain identifications. Very few specimens from non-archaeological sites (i.e. fens and caves) are well-dated, skewing the data towards anthropogenic sites such as settlements, castles, and graves. 5. Analysis of body part representation shows that the bear bones found in the Bronze Age, Iron Age and most of the early medieval period are from skins. All body parts are present during the Romano-British, medieval, and post-medieval period, indicating that live bears were present, and were probably imported for entertainment. 6. It seems the brown bear was rare throughout the Holocene, and based on current evidence two extinction scenarios can be put forward: extinction in the late Neolithic/early Bronze Age, or extinction in the early medieval period.

1. The brown bear, Ursus arctos, was Holocene Britain’s largest carnivoran and has appeared in recent rewilding discussions. 2. Despite widespread interest, we know very little about the species in Holocene Britain as few studies have been undertaken. This paper draws together information on the brown bear to examine the presence and extinction of bears through examination of the archaeological and palaeontological evidence. 3. Data were collected from published literature and museum catalogues. Information on the chronological date of the sites, number of specimens, and the body parts present were particularly sought. 4. A total of 86 sites were identified, 57 are well dated, 27 are of uncertain date but likely to be Holocene, and two have uncertain identifications. Very few specimens from non-archaeological sites (i.e. fens and caves) are well-dated, skewing the data towards anthropogenic sites such as settlements, castles, and graves. 5. Analysis of body part representation shows that the bear bones found in the Bronze Age, Iron Age and most of the early medieval period are from skins. All body parts are present during the Romano-British, medieval, and post-medieval period, indicating that live bears were present, and were probably imported for entertainment. 6. It seems the brown bear was rare throughout the Holocene, and based on current evidence two extinction scenarios can be put forward: extinction in the late Neolithic/early Bronze Age, or extinction in the early medieval period.