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A new species, Megarthrus heise Zhang, Cuccodoro, Chen & Liu sp. nov., resembling Megarthrus nitidulus Kraatz, 1857 is described from Qinghai, Sichuan, Gansu, Shaanxi, Shanxi provinces and Beijing in China. The record of M. nitidulus in China was apparently based on a misidentification, and the species is thus removed from the list of Chinese Megarthrus. Phoresy of acari and teratological modification of the abdomen are also reported for the first time in the genus.

The monospecific clavigerite genus Archiclaviger Heller, 1936 (Coleoptera: Staphylinidae: Pselaphinae), previously only known from Java, is reported to occur in China. A new species, A. gaofani sp. nov., was collected in nests of Nylanderia Emery, 1906 ants (Hymenoptera: Formicidae) in Jiangsu Province, and is described here. The new species can be separated from A. overbecki Heller, 1936 by different proportions of the antennomeres, and much sparser setae at the apex of the elytra and base of the abdomen. A revised diagnosis of Archiclaviger is provided, and a lectotype is designated for A. overbecki.

Two new species of Megarthrus are described from cloud forests of the Mexican state of Veracruz: M. cavianae Rodríguez, Navarrete-Heredia & Arriaga-Varela sp. nov., and Chiapas: M. chiapas Cuccodoro sp. nov. They differ from the two hitherto known Mexican species M. altivagans Bernhauer, 1929, and M. alatorreorum Rodríguez & Navarrete-Heredia, 2015, both from temperate forests of the Transmexican Volcanic Belt, by having synapomorphic features of the M. inaequalis-supergroup of species. This lineage includes all the Central and South American members of the genus, with the inclusion of these species, the distribution of the group is extend by more than 5 degree of latitude to the North. Within this lineage, the two new species share a very peculiar morphology of the male abdominal sternite VIII found elsewhere in the genus only in M. flavosignatus Bierig, 1940, and M. zunilensis Sharp, 1887, with which they form the M. zunilensis-group of species defined here.

A new fossil genus and species of the subfamily Pselaphinae (Coleoptera: Staphylinidae), †Cretobrachygluta laurasiensis gen. et sp. nov., is described based on an exquisitely preserved specimen in mid-Cretaceous amber from northern Myanmar. The new genus possesses a series of characters diagnostic of the extant Brachyglutini: Brachyglutina (supertribe Goniaceritae), but it also exhibits several plesiomorphic traits that suggest a basal position within the tribe. The discovery of the first and oldest brachyglutine beetle in Burmese amber considerably improves our understanding of the early morphological evolution of Pselaphinae during the Mesozoic Era.

Achilia crassicornis Jeannel, 1962 Figs 1-11, 21, 24, 31 Achilia crassicornis crassicornis Jeannel, 1962: 399, figs 144 (habitus) and 146 (aedeagus). ��� Franz, 1996: 114, fig. 61 (aedeagus). Achilia crassicornis antarctica Jeannel, 1962: 399, fig. 145 (ae- deagus). ��� Franz, 1996: 114 syn. nov. Achilia obscura Jeannel, 1962: 401 syn. nov. Type material (41 ex.): SOUTHERN CHILI: Regi��n Magallanes y de la Ant��rtica Chilena: Ant��rtica Chilena prov.: MHNS; 1 ♂ (holotype of Achilia crassicornis antarctica n�� 1623); Navarino Island, Puerto Williams; 31.I.1957; G. Kuschel. ��� MNHN; 12 ♂ and 18 ♀ (paratypes of Achilia crassicornis antarctica); Navarino Island, Puerto Williams; 54�� 56���S; 31.I.1957; G. Kuschel; Nothofagus betuloides and Nothofagus pumilio forest. ��� ��ltima Esperanza prov.: MHNS; 1 ♂ (holotype of Achilia crassicornis crassicornis n�� 1575); Puerto Eden; 06.XII.1958; G. Kuschel. ��� MNHN; 6 ♂ and 2 ♀ (paratypes of Achilia crassicornis crassicornis); Wellington Island, Puerto Eden, Carlos Islet; 49�� 09���S; 06.XII.1958; G. Kuschel; Nothofagus betuloides dense forest. ��� MNHN; 1 ♀ (holotype of Achilia obscura; according to the original description the holotype of A. obscura should have been deposited in MHNS, however we found it in MNHN); Wellington Island, Puerto Eden, Carlos Islet; 49�� 09���S; 600 m; 6.XII.1958; Nothofagus betuloides forest. Additional material (2166 ex.): See Appendix 1. Description: Body 1.50-1.75 mm long, reddish brown with head and abdomen sometimes slightly darker and palpi yellowish. Pubescence decumbent with dense and long setae, uniform on entire body. Head wider than long; frontal lobe short with rounded sides; surface smooth, shiny, with some minute punctures; vertexal sulcus deeply impressed and narrowed in middle; vertexal foveae shallow and large; eyes protruding, longer than convex temples. Pronotum wider than long and wider than head; posterior portion of lateral outlines sinuate; disc smoothly convex, shiny, with some small punctures; basal margin bordered with row of contiguous shallow impressions; median antebasal fovea smaller than lateral foveae. Elytra together wider than long with protruding humeri; disc smooth, shiny, with some small punctures; generally four basal foveae (two lateral foveae very close) or occasionally three (lateral foveae consisting of two combined foveae); sutural stria entire; discal stria extending to about elytral midlength. Legs slender. Abdomen smooth, shiny, with some minute punctures; tergite I with basal striae slightly diverging, extending to about one-third of paratergal length, separated at base by about onethird of tergal width, with short and sparse setal brush between striae. Male: Head as in Figs 21 & 24, with occiput strongly convex. Antennae (Fig. 11) with scape distinctly longer than wide; pedicel slightly longer than wide; antennomeres III-VIII small and slightly tranverse; antennomere IX strongly transverse with protruding mesal margin; antennomere X strongly thickened, wider and longer than XI, medial side truncate with broad subtriangular fairly flat area entirely delimited by sharp low ridge and covered with short dense pubescence. Metasternum convex; ventral margin of mesotrochanters with small acute lateral spine; all tibiae unarmed. Abdominal tergites unmodified; ventrites IV-V slightly flattened at middle. Aedeagus (Figs 1-2) 0.39- 0.41 mm long; dorsal plate ovoid with rounded sides and diverging dorsal strips starting from middle of dorsal plate; copulatory pieces each divided into two simple or bifurcated spreading branches slightly variable in shape (Figs 1-10). Parameres with outer outline only slightly convex or fairly straigth at level of middle seta; tips narrow recurved internally. Female: Similar to male except: head with occiput broadly convex, not swollen; antennomeres X and XI less wide than in male, X with medial side not truncate and shorter than XI; mesotrochanters and abdominal ventrites unmodified. Collecting data: Collected from September to March, mainly in Nothophagus forests, but also in Araucaria, Saxegothaea conspicua and Fitzroya cupressoides forests, where it was found in remnants of forests or at their edges at elevations ranging from sea level up to 1500 m and the treeline. Most specimens came from sifted samples of leaf and log litter, moss, dead trunks, vegetable debris, and sometimes mushrooms, but other collecting techniques include car netting, flight intercept (window) traps, malaise traps, carrion traps, pan traps, and screen sweeping. Jeannel (1962) mentions specimens collected wandering on the sandy shores of Lacar Lake (Argentina, Neuqu��n province). Distribution: Achilia crassicornis is distributed (Fig. 31) from the southernmost regions of Chile and Argentina to Central Chile (northernmost province: ��uble) and Central Western Argentina (northernmost province: Neuqu��n). Records from Chepu (Chilo�� prov.) come from Jeannel (1962). According to Jeannel (1962) it is the only species of Achilia to be so widespread in the Valdivian and Magellanes forests. Comments: The holotype and only known specimen of A. obscura should be in the MHNS collection according to Jeannel (1962: 401, but it is in the MNHN collection. Moreover, in the catalog of the MHNS holotypes of insects (Camousseight, 1980) this taxon is not mentioned. Jeannel (1962) described A. antarctica as a subspecies of A. crassicornis that was characterized by having the pronotum barely transverse, the aedeagus larger with shorter paramere apices, and the internal sac with stouter copulatory pieces. He also described the new species A. obscura to accomodate specimens differing from A. crassicornis by having the body darker and slightly longer, as well as the basal striae of abdominal tergite I more narrowed. However, after examining abundant materials we concluded that these differences were overestimated and pertain to intraspecific variation, and consequently place here both A. crassicornis antarctica Jeannel, 1962 and A. obscura Jeannel, 1962 as junior synonyms of A. crassicornis Jeannel, 1962 (syn. nov.). Two males from Chilo�� Island (San Pedro and Cucao) have the antennomeres X narrower with their medial side not truncate, but their aedeagi are similar to that of the other males of A. crassicornis that we examined, and the shape of their copulatory pieces (Figs 8-9) fall within the range of intraspecific variation that we observed for this structure., Published as part of Giorgio Sabella, Sergey A. Kurbatov & Giulio Cuccodoro, 2017, A revision of the Chilean Brachyglutini - Part 2. Revision of Achilia Reitter, 1890: A. crassicornis, A. tumidifrons, A. bifossifrons, and A. lobifera species groups (Coleoptera: Staphylinidae: Pselaphinae), pp. 119-140 in Revue suisse de Zoologie 124 (1) on pages 120-122, DOI: 10.5281/zenodo.322671, {"references":["Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale (pp. 295 - 479). In: Deboutteville C. D., Rapoport E. (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris.","Franz H. 1996. Neue Beitrage zur Kenntnis der Pselaphidenfauna von Chile und Argentinien (Coleoptera: Pselaphidae). Koleopterologische Rundschau 66: 83 - 146.","Camousseight A. 1980. Catalogo de los Tipos de Insecta depositados en la coleccion del Museo Nacional de Historia Natural (Santiago, Chile). Publicaciones ocasionales del Museo Nacional de Historia Natural 32: 1 - 45."]}

Achilia nahuelbutae Franz, 1996 Achilia nahuelbutae Franz, 1996: 115, fig. 62 (aedeagus). Comments: We have examined the type series of this species, which is housed in NHMW. The holotype appears to be a male of the cosmoptera group, while paratypes belong to at least two other Achilia species of the kindermanni and the grandiceps groups, and includes a specimen of the tribe Euplectini. All of these taxa except the member of the Euplectini will be dealt with in a later paper., Published as part of Giorgio Sabella, Sergey A. Kurbatov & Giulio Cuccodoro, 2017, A revision of the Chilean Brachyglutini - Part 2. Revision of Achilia Reitter, 1890: A. crassicornis, A. tumidifrons, A. bifossifrons, and A. lobifera species groups (Coleoptera: Staphylinidae: Pselaphinae), pp. 119-140 in Revue suisse de Zoologie 124 (1) on page 123, DOI: 10.5281/zenodo.322671, {"references":["Franz H. 1996. Neue Beitrage zur Kenntnis der Pselaphidenfauna von Chile und Argentinien (Coleoptera: Pselaphidae). Koleopterologische Rundschau 66: 83 - 146."]}

Achilia bifossifrons (Reitter, 1883) Figs 17, 23, 26, 33 Bryaxis bifossifrons Reitter, 1883: 50, pl. 1 fig. 9; Reitter 1885: 325, 329. Achilia bifossifrons, Jeannel, 1962: 404, figs 151 (head and an- tenna), 152 (aedeagus). Type material (1 ex.): CENTRAL CHILI: Región Los Ríos: Valdivia prov.: MNHN (coll. Raffray); 1 ♂ (holotype of Achilia bifossifrons here fixed); Chili. Additional material (1907 ex.): See Appendix 1. Description: Body 1.25-1.40 mm long, reddish with dark head, pronotum and abdomen, palpi yellowish. Pubescence on head short and suberect, long and decumbent over rest of body. Head wider than long; frontal lobe flattened with rounded sides; surface smooth, shiny, with some minute punctures; vertexal foveae small; eyes protruding, longer than convex temples. Pronotum wider than long and as wide as head, width maximal at middle; posterior portion of lateral outlines sinuate; disc strongly convex, smooth and shiny; median antebasal fovea slightly smaller than lateral foveae; basal margin bordered with row of contiguous shallow impressions. Elytra together wider than long with protruding humeri; disc smooth, shiny, with some small punctures; generally four basal elytral foveae (two lateral foveae very close and sometimes merged); sutural stria entire; discal stria extending to about elytral midlength. Legs slender. Abdomen smooth, with some minute punctures; tergite I with basal striae slightly diverging, extending to about onethird of paratergal length, separated at base by about one-third of tergal width, with short and sparse setal brush between striae. Male: Head as in Figs 23 & 26, with occiput very swollen, its anterior margin falls steeply to deep transverse sulcus. Antennae with scape short, as long as wide; pedicel slightlylongerthanwide; antennomeresIII–VIIIsmalland subglobose; antennomere IX transverse with protruding mesal margin; antennomere X transverse and larger than IX, with protruding mesal margin; antennomere XI ovoid, longer than wide and as long as VII-X combined. Metasternum with a large median impression occupying 2/3 of its surface; mesotibiae enlarged at middle and shallowly emarginate subapically. Abdominal tergites unmodified; all abdominal ventrites slightly flattened at middle. Aedeagus (Fig. 17) 0.28-0.29 mm long; dorsal plate elongate with sides sinuate and dorsal strips long and divergent; copulatory pieces consisting of pair of long sclerites curved apically. Parameres with middle seta on distinct lobe, that seta thin, reduced compared to that of the other species treated here; tips broad, recurved posteriorly. Female: Similar to male except: head with occiput barely swollen; metasternum convex; abdominal ventrites not flattened at middle; mesotibiae unmodified. Collecting data: Collected from September to April, mainly in Nothofagus-Araucaria forest and secondary and disturbed Valdivian rainforest at elevations ranging from sea level up to about 800 m. Most specimens came from sifted samples of leaf and log litter, moss on forest floor and trees, dead trunks, vegetational debris, and bracket fungi, but also by flight intercept (window) traps, screen sweeping, and car traps. Distribution: Achilia bifossifrons is distributed in Central Chile from Aysén northward to Ñuble provinces (Fig. 33). Comments: Reitter (1883) described A. bifossifrons based on what he thought was a unique female from Valdivia. However the description is definitely that of a male, so we recognized the only male we found in Raffray’s collection as the holotype of this species, and labelled it accordingly., Published as part of Giorgio Sabella, Sergey A. Kurbatov & Giulio Cuccodoro, 2017, A revision of the Chilean Brachyglutini - Part 2. Revision of Achilia Reitter, 1890: A. crassicornis, A. tumidifrons, A. bifossifrons, and A. lobifera species groups (Coleoptera: Staphylinidae: Pselaphinae), pp. 119-140 in Revue suisse de Zoologie 124 (1) on pages 127-132, DOI: 10.5281/zenodo.322671, {"references":["Reitter E. 1883. Beitrag zur Kenntniss der Pselaphiden-Fauna von Valdivia. Deutsche Entomologische Zeitschrift 27: 47 - 54, pl. I.","Reitter E. 1885. Beitrag zur Kenntniss der Pselaphiden-Fauna von Valdivia, Zweiter Theil. Deutsche Entomologische Zeitschrift 29: 321 - 332, pl. II.","Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale (pp. 295 - 479). In: Deboutteville C. D., Rapoport E. (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris."]}

Achilia parvula Jeannel, 1962 Achilia parvula Jeannel, 1962: 401. Comments: According to the original description the holotype and only known specimen of A. parvula should have been deposited in the MHNS, however we found it in the MNHN. In the catalog of the MHNS holotypes of insects (Camousseight, 1980) this taxon is not mentioned. It appears to be a female of the humidula group, which will be dealt with in a later paper., Published as part of Giorgio Sabella, Sergey A. Kurbatov & Giulio Cuccodoro, 2017, A revision of the Chilean Brachyglutini - Part 2. Revision of Achilia Reitter, 1890: A. crassicornis, A. tumidifrons, A. bifossifrons, and A. lobifera species groups (Coleoptera: Staphylinidae: Pselaphinae), pp. 119-140 in Revue suisse de Zoologie 124 (1) on page 122, DOI: 10.5281/zenodo.322671, {"references":["Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale (pp. 295 - 479). In: Deboutteville C. D., Rapoport E. (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris.","Camousseight A. 1980. Catalogo de los Tipos de Insecta depositados en la coleccion del Museo Nacional de Historia Natural (Santiago, Chile). Publicaciones ocasionales del Museo Nacional de Historia Natural 32: 1 - 45."]}

Achilia antennalis Jeannel, 1962 Figs 12, 19-20, 28, 30, 35 Achilia antennalis Jeannel, 1962: 405, 406 Type material (1 ex.): CENTRAL ARGENTINA: Neuqu��n prov.: MNHN; 1 ♂ (holotype of Achilia antennalis); Saint Martin de los Andes, Lanin reserve; about 40�� S; 1000 m; III.1959; specimens wandering on the sandy shores of Lacar Lake; C. Delamare. Additional material (124 ex.): See Appendix 1. Description: Body 1.30-1.45 mm long, reddish with darker head and abdomen, with yellowish maxillary palpi. Pubescence decumbent with dense and long setae, sparser on head and pronotum. Head wider than long; surface smooth, shiny, with some minute punctures; vertexal foveae shallow and large; eyes protruding, longer than convex temples. Pronotum wider than long and wider than head, with maximal width on anterior half; posterior portion of lateral outlines sinuate; disc convex, smooth and shiny; median antebasal fovea smaller than lateral foveae; basal margin bordered with row of contiguous shallow impressions. Elytra together wider than long with very protruding humeri; disc smooth, shiny, with some minute punctures; four basal elytral foveae (two lateral foveae very close); sutural stria entire; discal stria extending to about elytral midlength. Legs slender. Abdomen smooth, with some minute punctures; tergite I with basal striae subparallel and very short, extending to less than 1/6 paratergal length, separated at base by more than one-third of tergal width, with short and sparse setal brush between striae. Male: Head as in Figs 28 and 30. Antennae (Fig. 12) with scape longer than wide; pedicel slightly longer than wide; antennomere III small and transverse; antennomere IV wider than long with medial margin enlarged, anterior margin thicker than posterior, dorsal surface slightly concave and covered with numerous very little bristles, antennomeres V-X wider than long: antennomere XI distinctly longer than wide, longer than VIII-X combined. Metasternum bearing large median sulcus with pubescent sides; mesotibiae forming stout subapical spur. Abdominal tergites and ventrites unmodified. Aedeagus (Figs 19-20) 0.28-0.29 mm long; dorsal plate large with sides sinuate; dorsal strips long and divergent; copulatory pieces consisting of pair of large sclerites that are apically enlarged and trifid, laterally ending in four tips. Parameres with middle seta on distinct lobe, that seta very stout; tips broad recurved posteriorly. Female: Similar to male except: head with occiput only slightly swollen, frontal region flattened with slightly convergent sides and impressed vertexal sulcus; antennae unmodified, with antennomeres III as long as wide and IV-V slightly longer than wide; metasternum convex; mesotibiae unmodified. Collecting data: Collected from December and March in Nothofagus forest that is sometimes with Chusquea, in Saxegothea forest with Drimys, and in Araucaria araucana forest at elevations ranging from 300 m up to about 1500 m. Most specimens came from sifted samples of leaf and log litter, but also were taken by flight intercept (window) and malaise traps. Jeannel (1962: 407) reports that specimens were collected wandering on the sandy shores of Lacar Lake (Argentina, Neuqu��n province). Distribution: Achilia antennalis is distributed for Central Western Argentina (Neuqu��n province) and Central Chile from Osorno to the Maule region (Talca province) (Fig. 35). Comments: In the original description Jeannel (1962) mentioned five males (holotype and paratypes) collected in Saint Martin de los Andes. He also stated that A. antennalis and A. lobifera have the same external and aedeagal morphology, and differ only by the shape of the male head and antennae, however, the aedeagi are really distinctive, notably in examination of the copulatory pieces (Figs 18-19)., Published as part of Giorgio Sabella, Sergey A. Kurbatov & Giulio Cuccodoro, 2017, A revision of the Chilean Brachyglutini - Part 2. Revision of Achilia Reitter, 1890: A. crassicornis, A. tumidifrons, A. bifossifrons, and A. lobifera species groups (Coleoptera: Staphylinidae: Pselaphinae), pp. 119-140 in Revue suisse de Zoologie 124 (1) on page 133, DOI: 10.5281/zenodo.322671, {"references":["Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale (pp. 295 - 479). In: Deboutteville C. D., Rapoport E. (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris."]}

Oberea fuscipennis (Chevrolat, 1852) species group is revised based on morphology and DNA barcode data. Oberea diversipes Pic, 1919 and O. infratestacea Pic, 1936 are restored from synonymy. The following two new synonymies are proposed: Oberea fuscipennis ssp. fairmairei Breuning, 1962 = Oberea diversipes Pic, 1919; and Oberea hanoiensis Pic, 1923 = O . fuscipennis (Chevrolat, 1852)

Colilodion schulzi new species Figs 1 A, 3 Holotype: ♀, labeled ���PAL-09/08: PHILIPPINES: Palawan, Puerto Princesa Region, Sabang, Mt. Bloomfield, 10 �� 11 ��� 37 ������N, 118 �� 52 ��� 21 ������E, 500-700 m, primary forest, 10.XII. 2009, leg. A. Schulz. PAL-09/08 (printed) / MHNG ENTO 0 0 0 0 8847 (accession number) (printed) / Holotype ♀, Colilodion schulzi sp. n., det. Yin & Cuccodoro, 2015 (hand written)���. Differential diagnosis: Colilodion schulzi shares with C. concinnus Besuchet and C. inopinatus Besuchet the broadened antennomeres III with the impressed dorsal surface indicated by divided smooth fields. It can be readily separated from these two species by the antennomeres III being strongly broadened throughout their entire length, with a slightly narrowed base, a stouter pronotum with coarser discal punctation, and a relatively much broader elytral base. Both C. concinnus and C. inopinatus have the antennomeres III much narrower at the base than at the apex, their pronota are less stout, the discal punctation is finer, and the elytral base is relatively much narrower. Description: Length 2.37 mm. Body and appendages reddish brown (Fig. 1 A). Pubescence of body short and recumbent. Head longer than wide, HL 0.43 mm, HW 0.34 mm.Vertex (Fig. 3 A) strongly raised dorsally, narrowed apically, surface of raised area densely and roughly punctate, lateral area (Fig. 3 B) vertical, very finely punctate, dorsal margin convex in lateral view, situated slightly below level of pronotum, bearing thick, posteriorly-oriented dorsolateral and dorsomedian trichomes; posterior edge of vertex narrowed to become keel-like below, bearing two fairly long, diverging trichomes oriented posteriorly. Frons sparsely but roughly punctate, pubescence fine. Each eye composed of about 22 facets, unevenly divided by thick lateral ocular carina (Fig. 3 B), dorsal part with about 20 facets, ventral part with 2 facets. Gular ridge (Fig. 3 C) broad, thickened at middle, anterolateral surface sparsely and roughly punctate, with short pubescence. Occipital constriction impunctate, shiny, vertical at ventral margin. Antennae (Fig. 3 D) three-segmented; antennomere I visible in dorsal view, slightly transverse, roughly punctate on ventral surface, with short, thick setae; antennomere II distinctly transverse, wider and shorter than antennomere I, surface roughly punctate, with short setae; antennomere III 0.92 mm long, 0.39 mm wide, strongly broadened throughout length, impressed dorsal surface indicated by divided smooth fields, sparsely covered with short setae; setae of anterior margin longer and thicker; apical sensilla with raised margin, with two short setae. Pronotum trapezoidal, PL 0.63 mm, PW 0.53 mm (at base), gradually narrowed apically; apical portion (Fig. 3 A) broadly notched, notch deeper at middle; median groove evenly narrow, sharply delimited, extending posteriorly near pronotal base and anteriorly to posterior edge of apical notch; sub-anterolateral areas broadly concave; dorsum extremely coarsely punctate, distinctly margined laterally at anterior third, sub-basal area smooth; lateral surface (Fig. 3 B) extremely finely punctate and with fine setae; posterolateral angles distinct, posterior margin smoothly sinuate; anterolateral edges slightly oblique, each bearing conspicuous trichome forming rim oriented dorso-anteriorly and pointed dorsally; dorsal anterior edge with shorter and thinner trichomes oriented anteriorly and curved dorsally near tip. Elytra (Fig. 3 E) wider than long, EL 0.73 mm, EW 0.95 mm; elytral disc flattened, slightly raised, sparsely covered with coarse punctation, densely microsculptured, with short, recumbent setae; each elytron with seven longitudinal striae, sutural and pair of inner striae complete, pair of central striae with inner stria extending from base to half of elytral length, and outer one much fainter, pair of outer striae with inner stria complete, outer one extending from basal third to posterior margin; anterolateral margin round, area nearby finely punctate and lacking microsculpture, punctation and setae of lateral area similar to those of disc, interval between punctures smooth, lacking microsculpture; posterior area with row of sparse, long, and thick setae, with bunch of thick setae at posterolateral margin. Prosternum (Fig. 3 C) with triangular process rising from anterior margin, and large vertical process rising from posterior margin; median area with several setae, lacking obvious trichome, lateral surface of process distinctly microsculptured. Anterior mesoventral edge (Fig. 3 F) slightly raised, pointed at middle, lateral areas with big and shallow punctures, inner sides with microsculpture. Metaventrite (Fig. 3 F) raised at middle, anterior half and lateral area of basal half with shallow, large punctures and short setae. Abdomen transverse, AL 0.58 mm, AW 0.89 mm; first visible tergite (Fig. 3 G) broadly and deeply impressed between elongate basolateral ridges, with dense setae along posterior margin of impression; disc finely punctate and lacking microsculpture at basal half, with shallow, large punctures and microsculptured interval at apical half, pubescence mostly fine, with two rows of thick, erect setae at apical portion, with truncate apex; paratergites well-demarcated, with few long, erect setae; second tergite with row of long, erect apical setae, with distinct lateral tubercles. First visible sternite (morphologically sternite III) (Fig. 3 H) with coarse punctation and dense microsculpture, setae fine; second sternite long, punctation and microsculpture much more denser at middle than at lateral portion; following sternites similarly microsculptured and with short setae. Tibiae (Figs 3 I-K) distinctly sculptured, narrowed at basal third, apical two-thirds abruptly thickened, bearing conspicuous rows of erect setae on dorsal side. Key to Colilodion species (modified from L��bl, 1994) (Figs 1-3) 1 Antenna four-segmented........................................................................................................................................... 2 - Antenna three-segmented......................................................................................................................................... 3 2 Antennomere IV evenly curved laterally throughout whole length (Fig. 2 B); body size smaller, 2.15 mm. (southern China: Hainan).................................................................................................................. C. tetramerus L��bl, 1998 - Antennomere IV markedly curved laterally at basal one-fifth (Fig. 2 C); body size larger, 2.54 mm. (central Vietnam: Thua Thien Hue)............................................................................................. C. thienmu Nomura & Sugaya, 2007 3 Antennomere III strongly broadened, with impressed dorsal surface indicated by divided smooth fields............. 4 - Antennomere III subcylindrical or dorsally flattened, lacking impressed fields...................................................... 6 4 Antennomere III broadened throughout whole length, base slightly narrowed (Figs 1 A, 3 D); posterior margin of elytra with long, thick golden setae (Fig. 3 E). (Philippines: Palawan)............................... C. schulzi, new species - Antennomere III much more narrowed at base than at apex (Figs 1 B, 1 D); posterior margin of elytra lacking long, thick setae................................................................................................................................................................ 5 5 Punctation of tergite IV distinct, similar to that of elytra and pronotum (Fig. 1 B). (Indonesia: western Sumatra)....................................................................................................................................... C. concinnus Besuchet, 1991 - Punctation of tergite IV obsolete, much finer than that of elytra and pronotum (Fig. 1 D). (East Malaysia: Sabah).................................................................................................................................... C. inopinatus Besuchet, 1991 6 Antennomere III flattened dorsally (Fig. 2 D); elytral marginal carina short, disappear before reaching mid-length of elytron. (West Malaysia: Pahang)........................................................................................ C. wuesti L��bl, 1994 - Antennomere III subcylindrical (Figs 1 C, 2 A); elytral marginal carina extended posteriorly beyond mid-length of elytron...................................................................................................................................................................... 7 7 Antennomere III barely curved, shorter than half of body length (Fig. 1 C). (East Malaysia: Sabah).......................................................................................................................................................... C. incredibilis Besuchet, 1991 - Antennomere III distinctly curved, longer than half of body length (Fig. 2 A). (East Malaysia: Sabah).............................................................................................................................................................. C. mirus Besuchet, 1991 Biology: The single female was collected from a sample of sifted vegetable debris in a sparse coniferous forest that was subsequently processed using Winkler- Moczarski eclectors. The locality is a quite dry and hot place on a hilltop with lots of stones on the ground. The most common ant genera in that area were Camponotus, Paratrechina, and some other myrmecine genera (A. Schulz pers. comm.). Distribution: The new species is known only from the type locality. Etymology: The specific epithet is dedicated to Andreas Schulz, who collected the holotype., Published as part of Zi-Wei Yin & Giulio Cuccodoro, 2016, Colilodion schulzi sp. n. (Coleoptera: Staphylinidae: Pselaphinae) from Palawan, the Philippines, with habitus photographs and a revised key to all Colilodion species, pp. 153-158 in Revue suisse de Zoologie 123 (1) on pages 154-158, DOI: 10.5281/zenodo.46295, {"references":["Lobl I. 1994. The systematic position of Colilodionini with description of a new species (Coleoptera, Pselaphidae). Revue suisse de Zoologie 101: 289 - 297.","Lobl I. 1998. A new species of Colilodion (Coleoptera: Staphylinidae: Pselaphinae) from China. Bulletin de la Societe entomologique suisse 71: 467 - 470.","Nomura S., Sugaya H. 2007 A new species of the genus Colilodion Besuchet (Coleoptera: Staphylinidae: Pselaphinae) from Vietnam. Annales de la Societe Entomologique de France (Nouvelle serie) 43: 333 - 339.","Besuchet C. 1991. Revision chez les Clavigerinae (Coleoptera, Pselaphidae). Revue suisse de Zoologie 98: 499 - 515."]}

Ogmocerodes navigator Cuccodoro & Zi-Wei Yin, sp. n. Figs 1-4 Holotype: Cameroon; ♂, labeled ��� Cameroon, Centre Region, 100 m SW of Ebogo Tourist Center 04.IX. 2014, alt. 650 m, leg. G. Cuccodoro, # 8 sifting leaf litter in wooden canoe abandoned in flood forest on the left shore of Nyong River��� (in MHNG; accession number MHNG ENTO 00008839). Paratypes: 5 ♂♂, 2 ♀♀, same label data as the holotype (in MHNG; accession numbers MHNG ENTO 00008840���00008846). Differential diagnosis: Ogmocerodes navigator sp. n. shares markedly punctate pronotal disc with O. raffrayi Brauns, 1915, but the latter species lacks femoral spines in the male. The only other congener to possess metafemoral spines in the male is O. hustaerti Jeannel, 1949 b, but it has no mesofemoral spines. The conformation of the antennal modification, and the shape of the aedeagus are also diagnostic. Description: Male (Fig. 1). Length 4.41-4.79 mm. Head longer than wide, HL 1.00- 1.09 mm, HW 0.80- 0.83 mm, regularly punctate and setose on dorsal surface, with low, short frontal rostrum. Antennal tubercles slightly prominent and close, divided by a short, deep frontal sulcus; lacking frontal fovea, with deep, setose vertexal foveae (Fig. 2A); lacking postantennal notches at lateral margins; small areas just posterior to antennal tubercles glabrous, lacking setae and punctation; antennae (Fig. 4B) with eleven antennomeres, clubs formed by enlarged antennomeres VII-XI; scapes about as long as antennomeres II-VIII combined, distinctly sinuate near base, antennomeres II-VI each transverse, successively larger, antennomeres VII and VIII (Fig. 4C) strongly modified, each greatly enlarged and strongly concave at anteromedial margin of VII and posteromedial margin of VIII, antennomeres IX and X each transverse, narrowed at apex, XI as long as IX and X combined, with truncate base and round apex; reniform eyes each composed of about holotype male. Scale: 1.0 mm. 200 small facets; ocular-mandibular carinae (Fig. 2B) present; maxillary palpi with palpomere I short, II elongate, narrowed at base and widest at truncate apex, III transversely triangular, IV largest, ovoidal, with small palpal cone at apex; gula (Fig. 2C) considerably depressed posteromedially, indistinct gular sulcus present, each gular fovea has two openings, posterior pair of openings larger than anterior pair. Pronotum (Fig. 2D) about as long as wide, PL 0.85- 0.91 mm, PW 0.85-0.90 mm, with slightly constricted base, regularly round margins at middle, apex constricted to form apical collar; deep antebasal sulcus connecting lateral antebasal foveae, sulcus interrupted by short longitudinal basomedian carina, with short, shallow sulcus anterior to carina; punctation and setation of pronotal disc similar to that of head. Prosternite (Fig. 2E) with two pairs of lateral procoxal foveae; lacking paranotal sulci or carinae. Elytra (Fig. 2F) wider than long, EL 1.15-1.29 mm, EW 1.56-1.70 mm, disc more sparsely and finely punctate than pronotum, covered with recumbent setae; each elytron with two large, nude basal foveae; lacking subbasal fovea; sutural stria entire; discal stria extending to apical 4 / 5 of elytral length; lacking subhumeral foveae and marginal sulci. Mesoventrite (Fig. 3A) with single, large median mesoventral fovea, large lateral mesoventral foveae not forked; lateral mesocoxal foveae deep. Metaventrite (Fig. 3A) with large median fovea slightly forked inwards; with distinct projections (Fig. 4D); posterior margin convex medially. Legs roughly punctate, densely setose; meso- (Fig. 4F) and metafemora (Fig. 4G) each spinose at their ventral margin near apex; third tarsomeres longer than second tarsomeres, protarsomeres II (Fig. 4E) acutely lobed, extending beneath III; with single tarsal claws, and one transparent bristle-like seta. Abdomen (Fig. 3 B-D) stout, AL 1.41-1.50 mm, AW 1.57- 1.66 mm, coarsely punctate, regularly setose; tergite IV (first visible tergite, Fig. 3C) slightly longer than V, lateral margins sulcate, with broad basal impression, lacking mediobasal foveae, with one large and one tiny pair of basolateral foveae in large sockets at lateral margins of impression, lacking discal carinae; tergite V as long as VI, VII much shorter than previous tergite, tergites V-VII each with basal sulcus and one pair of basolateral foveae, and with sulcate lateral margins, tergite VIII roundly triangular, lacking fovea; paratergites accompanying tergites IV-VII (Fig. 3C). Sternite IV (second visible sternite, Fig. 3D) longest, with wide, densely setose basal sulcus, with three pairs of basolateral foveae, lateral pair smaller than mesal two pairs; sternites V-VII each with setose basal sulcus and two pairs of basolateral foveae, sternite VIII strongly transverse, lacking fovea, with broadly concaved posterior margin, sternite IX membranous, composed of lateral pair of triangular sclerites and one oval, elongate median plate. Aedeagus (Fig. 4H, I) stout, length 0.51 mm, with symmetric median lobe and parameres, with large ovoidal dorsal diaphragm opening; anterior margin of median lobe (Fig. 4J) with membranous, finely setose fringe; parameres (Fig. 4H, I) each with rows of sparse setae from near base to apex, apex (Fig. 4K) with two large setae. Female.Similartomaleingeneralappearance, withsimple antennomeres VII-VIII (Fig. 4A), protarsi, and meso- and metafemora, and metaventrite lacking projections; each eye with about 180 facets. Measurements: BL 4.41- 4.51 mm, HL 0.96-0.99 mm, HW 0.77-0.78 mm, PL 0.84-0.87 mm, PW 0.84-0.85 mm, EL 1.18-1.23 mm, EW 1.55-1.59 mm, AL 1.40-1.45 mm, AW 1.57-1.64 mm. Habitat: The type series was collected at an elevation of 650 m just before the main rainy season, from sifted leaf litter accumulated in an old wooden canoe abandoned in a flood forest some twenty meters from the waters of the Nyong River (Fig. 5). Distribution: The new species is known only from the type locality in central Cameroon approximately 10 km southeast of the city of Mbalmayo. Etymology: The epithet ��� navigator ��� means ���sailor��� in Latin, remembering that the type series was collected in a canoe., Published as part of Giulio Cuccodoro & Zi-Wei Yin, 2015, A new species of Goniacerini from Cameroon (Coleoptera: Staphylinidae: Pselaphinae), pp. 415-421 in Revue suisse de Zoologie 122 (2) on pages 416-421, DOI: 10.5281/zenodo.30011