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Etielloides Shibuya, 1928 Etielloides Shibuya, 1928: 121. Type species: Etielloides curvella Shibuya, 1928, by original designation. Diagnosis: Etielloides is diagnosed by the frons (Figs 1a, 1b, 2a, 2b) with a conical prominence of tufts; the antenna (Fig. 3) of the male with a deep sinus bearing a ridge of scales at the base of flagellum; the elongate and narrow forewing (Figs 1c, 2c) with R 3+4 and R 5 stalked for about 2/3 of their lengths, M 2 and M 3 closely approximated near base; the broad subtrapezoidal hindwing (Figs 1c, 2c) with Sc + R 1 and Rs stalked or closely approximated for about one-half of their length, M 2 and M 3 stalked about one-half of their length; the absence of transtilla, the inflated, fusiform apical process of gnathos, the clavate costa usually with a free, curved, pointed projection at apex, and the cylindrical aedeagus with one sword-like cornutus in the male genitalia (Fig. 5), and the well-developed ostium, mostly sclerotized ductus bursae and the slender ductus seminalis originating from posterior part of the corpus bursae in the female genitalia (Fig. 6). Biology. The larvae of Etielloides curvella Shibuya feed on leaves of Malus pumila var. dilcissima Koidz and Pyrus spp. (Inoue 1982). Distribution. China (Fujian, Henan, Guizhou, Guangxi, Jiangxi, Shaanxi), Korea, Japan, Far East of Russia. Remarks. The character of free, curved costal apex projection of the valva is a remarkable feature of the genus Etielloides, but there are exceptions: this projection is reduced in E. bipartitellus and absent in E. longipalpus. The similar costal apex projection is also present in Salebriaria Heinrich, 1956, but shorter and spine-like. The genus Etielloides also differs from Salebriaria by the sharply elongate triangular uncus in the male genitalia and the corpus bursae without spines in the female genitalia. In Salebriaria, the uncus is broadly rounded at apex and the corpus bursae covered with a dense matting of fine spines., Published as part of Yang, Linlin, Liu, Hongxia & Ren, Yingdang, 2021, One new species of the genus Etielloides Shibuya, 1928 from China (Lepidoptera Pyralidae, Phycitinae), pp. 361-368 in Zootaxa 4990 (2) on page 362, DOI: 10.11646/zootaxa.4990.2.9, http://zenodo.org/record/5026717, {"references":["Shibuya, J. (1928) Descriptions of one new genus and one new species, with four unrecorded species of Phycitinae from Japan (Lepid.). Insecta Matsumurana, 2 (3), 121 - 124.","Inoue, H. (1982) Pyralidae. 1 & 2. In: Inoue, H., Sugi, S., Kuroko, H., Moriuti, S., Kawabe, A. & Owada, M. (Eds.), Moths of Japan. Vols. 1 & 2. Kodansha, Tokyo, pp. 307 - 404 & 233 - 254. [in Japanese]","Heinrich, C. (1956) American moths of the subfamily Phycitinae. Bulletin of the United States National Museum, Washington, D. C., 207, i - viii + 1 - 581, figs. 1 - 1138. http: // biostor. org / reference / 184747"]}

Etielloides luniformis sp. nov. (Figs 1−6) http://zoobank.org/ urn:lsid:zoobank.org:act: 2FD2A663-0901-46E6-B6BF-10311F56150D Diagnosis: This new species resembles E. kogii Yamanaka, but differs from the latter by the costa of the same length as valva, the triangular clasper, the shield-shaped juxta 1.6 times length of width in the male genitalia, and the ostium represented by a pair of horned plates, the corpus bursae with a pair of ovate sclerotized patches posterolaterally in the female genitalia. In E. kogii, the costa is longer than the valva, the clasper is absent, and the juxta is rounded in the male genitalia; the ostium is funnel-shaped, and the corpus bursae has no sclerotized patch in the female genitalia. Description. Adult (N = 72): Wingspan 17.5−21.0 mm. Male (Figs 1, 3): Vertex pale yellow, frons with gray- ish brown bulged tufts. Antenna with scape grayish brown, three times as long as wide; flagellum with first five segments coalesced, forming a deep sinus, bearing a ridge of scale tuft that grayish anteriorly and whitish posteriorly, and a few short teeth concealed within the tuft. Labial palpus upturned, extending about twice length of head beyond it, segmental ratio 2:4.5:1; basal segment gray, terminal two segments white. Maxillary palpus yellowish white, in the form of an aigrette, about 4/5 length of second segment of labial palpus, segmental ratio 2.5:0.8:1. Patagium, tegula and thorax grayish brown. Forewing ground color grayish white, basal area diffused ocherous and chestnut brown, central area mottled ocherous and black, distal area mixed with yellowish white; antemedial line near middle of forewing, white, obscure at costa, bordered inwardly with an obscure ocherous yellow shade, edged outwardly with a black line; postmedial line grayish white, waved, edged black on both sides; discal spots black, fused into a single crescent spot; terminal line grayish black; cilia basally chestnut brown, distally grayish black, with white basal and middle lines. Hindwing semitransparent, grayish white, tinted with pale brown along termen; cilia yellowish white. Fore- and midlegs yellowish white on upper surface, blackish brown on lower surface except yellowish white at apices of tarsomeres; hindleg yellowish white, variegated grayish brown on tarsus. Female (Figs 2, 4): Head ocherous. Antenna with scape ocherous, flagellum normal, ocherous and blackish brown. Labial palpus segmental ratio 1.5:4:1, second segment slender than that of male, ocherous to brown. Maxillary palpus yellowish brown, squamous, about half length of second segment of labial palpus, three segments about equal length. Patagium, tegula and thorax chestnut brown. Forewing ground color brown, basal area overlaid chestnut brown, mixed with a few blackish brown scales; terminal line chestnut brown. Legs yellowish white on upper surface, blackish brown on lower surface except yellowish white at apices of tarsomeres, mid- and hindlegs mottled grayish white. Other characters same as those of males. Male genitalia (Fig. 5). Uncus triangular, 1.6 times as long as basal width, narrowly rounded towards apex. Apical process of gnathos as long as uncus, elongate, leaflike, protruded a small hook apically; gnathos arms arising from center of dorsal surface. Transtilla absent. Valva 5.0 times length of width, narrowly rounded apically, ventral 2/3 densely covered with long bristles; clasper triangular, rounded terminally; costa as long as valva, clubbed, distal 1/5 thorn like, separated from valva; sacculus about 2/5 length of valva, bone-shaped. Vinculum U-shaped, length equal to greatest width, with two semi rounded anterolateral processes. Juxta shield-shaped, 1.6 times length of width, lateral lobes absent. Aedeagus clavate, slightly shorter than valva, granulated in distal 1/5; cornutus about 2/5 length of aedeagus, with basal half curved in S-shape, distal half thornlike. Eighth abdominal tergite somewhat fan-shaped; eighth abdominal sternite inverted T-shaped, associated with four pairs of culcita. Female genitalia (Fig. 6). Eighth tergite with anterior margin straight, posterior margin triangularly concave to middle of eighth tergite. Apophysis anterioris basally inflated, 1.3 times length of apophysis posterioris. Ostium funnel shaped, with a pair of elongate, horned plates, arranged in V-shaped. Ductus bursae very short, membranous, about 2/5 length of corpus bursae. Corpus bursae membranous, 1.7 times as long as wide, elliptically rounded, with a curved ribbon-like plate posteriorly, and a pair of ovate sclerotized patches posterolaterally, each patch half-length of corpus bursae, comprising of small, honeycombed plates. Ductus seminalis from corpus bursa near junction with ductus bursae. Type material. Holotype ♂, China: Mt. Leigong (28°01’N 111°40’E), Leishan county, Qiandongnan Miao and Dong autonomous prefecture, Guizhou, 1200 m, 7.v.2012, coll. Yingdang Ren & Xiaoguang Liu, gen. slide No. RYD20120007. Paratypes: China: Guizhou: 31 ♂, 40 ♀, same data as for holotype, gen. slide Nos RYD 20120005m, RY- D 2014025m, RYD 2014026m, RYD 2014027m, RYD 2014158m, DNAYLL 18047m, DNAYLL 18060m, RY- D 2014032m, DNAYLL 18085m, DNAYLL 18086m, DNAYLL 18088m, RYD20120006f, RYD20120008f, RYD2014028f, RYD2014029f, RYD2014030f, RYD2014031f, RYD2014152f, RYD2014153f, RYD2014154f, RYD2014157f, DNAYLL18061f, DNAYLL18087f, DNAYLL18090f, DNAYLL18091f. DNA Barcode. Five DNA barcodes of paratypes was generated and deposited in the GenBank (accession numbers: MT 648697 – MT 648701). The intraspecific distances of the new species ranges from 0 to 0.46%, and interspecific distances between the new species and its closest relative E. kogii (accession numbers: MT 648693 – MT 648696) ranges from 4.40% to 4.89 %. Distribution: China (Guizhou). Etymology: The specific name is derived from the Latin word luniformis, meaning crescent, referring to the crescent discal spot on the forewing.

Etielloides luniformis sp. nov. (Figs 1���6) http://zoobank.org/ urn:lsid:zoobank.org:act: 2FD2A663-0901-46E6-B6BF-10311F56150D Diagnosis: This new species resembles E. kogii Yamanaka, but differs from the latter by the costa of the same length as valva, the triangular clasper, the shield-shaped juxta 1.6 times length of width in the male genitalia, and the ostium represented by a pair of horned plates, the corpus bursae with a pair of ovate sclerotized patches posterolaterally in the female genitalia. In E. kogii, the costa is longer than the valva, the clasper is absent, and the juxta is rounded in the male genitalia; the ostium is funnel-shaped, and the corpus bursae has no sclerotized patch in the female genitalia. Description. Adult (N = 72): Wingspan 17.5���21.0 mm. Male (Figs 1, 3): Vertex pale yellow, frons with gray- ish brown bulged tufts. Antenna with scape grayish brown, three times as long as wide; flagellum with first five segments coalesced, forming a deep sinus, bearing a ridge of scale tuft that grayish anteriorly and whitish posteriorly, and a few short teeth concealed within the tuft. Labial palpus upturned, extending about twice length of head beyond it, segmental ratio 2:4.5:1; basal segment gray, terminal two segments white. Maxillary palpus yellowish white, in the form of an aigrette, about 4/5 length of second segment of labial palpus, segmental ratio 2.5:0.8:1. Patagium, tegula and thorax grayish brown. Forewing ground color grayish white, basal area diffused ocherous and chestnut brown, central area mottled ocherous and black, distal area mixed with yellowish white; antemedial line near middle of forewing, white, obscure at costa, bordered inwardly with an obscure ocherous yellow shade, edged outwardly with a black line; postmedial line grayish white, waved, edged black on both sides; discal spots black, fused into a single crescent spot; terminal line grayish black; cilia basally chestnut brown, distally grayish black, with white basal and middle lines. Hindwing semitransparent, grayish white, tinted with pale brown along termen; cilia yellowish white. Fore- and midlegs yellowish white on upper surface, blackish brown on lower surface except yellowish white at apices of tarsomeres; hindleg yellowish white, variegated grayish brown on tarsus. Female (Figs 2, 4): Head ocherous. Antenna with scape ocherous, flagellum normal, ocherous and blackish brown. Labial palpus segmental ratio 1.5:4:1, second segment slender than that of male, ocherous to brown. Maxillary palpus yellowish brown, squamous, about half length of second segment of labial palpus, three segments about equal length. Patagium, tegula and thorax chestnut brown. Forewing ground color brown, basal area overlaid chestnut brown, mixed with a few blackish brown scales; terminal line chestnut brown. Legs yellowish white on upper surface, blackish brown on lower surface except yellowish white at apices of tarsomeres, mid- and hindlegs mottled grayish white. Other characters same as those of males. Male genitalia (Fig. 5). Uncus triangular, 1.6 times as long as basal width, narrowly rounded towards apex. Apical process of gnathos as long as uncus, elongate, leaflike, protruded a small hook apically; gnathos arms arising from center of dorsal surface. Transtilla absent. Valva 5.0 times length of width, narrowly rounded apically, ventral 2/3 densely covered with long bristles; clasper triangular, rounded terminally; costa as long as valva, clubbed, distal 1/5 thorn like, separated from valva; sacculus about 2/5 length of valva, bone-shaped. Vinculum U-shaped, length equal to greatest width, with two semi rounded anterolateral processes. Juxta shield-shaped, 1.6 times length of width, lateral lobes absent. Aedeagus clavate, slightly shorter than valva, granulated in distal 1/5; cornutus about 2/5 length of aedeagus, with basal half curved in S-shape, distal half thornlike. Eighth abdominal tergite somewhat fan-shaped; eighth abdominal sternite inverted T-shaped, associated with four pairs of culcita. Female genitalia (Fig. 6). Eighth tergite with anterior margin straight, posterior margin triangularly concave to middle of eighth tergite. Apophysis anterioris basally inflated, 1.3 times length of apophysis posterioris. Ostium funnel shaped, with a pair of elongate, horned plates, arranged in V-shaped. Ductus bursae very short, membranous, about 2/5 length of corpus bursae. Corpus bursae membranous, 1.7 times as long as wide, elliptically rounded, with a curved ribbon-like plate posteriorly, and a pair of ovate sclerotized patches posterolaterally, each patch half-length of corpus bursae, comprising of small, honeycombed plates. Ductus seminalis from corpus bursa near junction with ductus bursae. Type material. Holotype ♂, China: Mt. Leigong (28��01���N 111��40���E), Leishan county, Qiandongnan Miao and Dong autonomous prefecture, Guizhou, 1200 m, 7.v.2012, coll. Yingdang Ren & Xiaoguang Liu, gen. slide No. RYD20120007. Paratypes: China: Guizhou: 31 ♂, 40 ♀, same data as for holotype, gen. slide Nos RYD 20120005m, RY- D 2014025m, RYD 2014026m, RYD 2014027m, RYD 2014158m, DNAYLL 18047m, DNAYLL 18060m, RY- D 2014032m, DNAYLL 18085m, DNAYLL 18086m, DNAYLL 18088m, RYD20120006f, RYD20120008f, RYD2014028f, RYD2014029f, RYD2014030f, RYD2014031f, RYD2014152f, RYD2014153f, RYD2014154f, RYD2014157f, DNAYLL18061f, DNAYLL18087f, DNAYLL18090f, DNAYLL18091f. DNA Barcode. Five DNA barcodes of paratypes was generated and deposited in the GenBank (accession numbers: MT 648697 ��� MT 648701). The intraspecific distances of the new species ranges from 0 to 0.46%, and interspecific distances between the new species and its closest relative E. kogii (accession numbers: MT 648693 ��� MT 648696) ranges from 4.40% to 4.89 %. Distribution: China (Guizhou). Etymology: The specific name is derived from the Latin word luniformis, meaning crescent, referring to the crescent discal spot on the forewing., Published as part of Yang, Linlin, Liu, Hongxia & Ren, Yingdang, 2021, One new species of the genus Etielloides Shibuya, 1928 from China (Lepidoptera Pyralidae, Phycitinae), pp. 361-368 in Zootaxa 4990 (2) on pages 363-367, DOI: 10.11646/zootaxa.4990.2.9, http://zenodo.org/record/5026717

Dusungwua paradichromella (Yamanaka, 1980) (Figs 1, 9, 20, 31) Eurhodope paradichromella Yamanaka, 1980: 69. TL: Japan. TD: Coll. HY. Eurhotope [sic] paradichromella: Lu (ed.), 1994: 30. Eurhodope (Eurhophaea) paradichromella: Xu et al., 2005, In: Yang & Jin (eds.): 323. Trachycera paradichromella: Roesler, 1987: 316. Furcata paradichromella: Du, Sung & Wu, 2005: 103. Trachycera nipponella Yamanaka, 2000: 233. Syn. nov. Trachycera yakushimensis Yamanaka, 2000: 235. Syn. nov. Trachycera vicinella Yamanaka, 2000: 235. Syn. nov. Dusungwua paradichromella: Kemal, Kizildağ & Koçak, 2020: 2. Diagnosis. Adult (Fig. 9) with wingspan 21.0̅23.0 mm. This species is characterized by the forewing has a small, blackish brown costal spot on inner margin of antemedial line, and a large, subtriangular, blackish brown posterior patch on outer margin of lower half of antemedial line; the uncus is gradually narrowed, the gnathos is clubbed, dentate apically, the transtilla has right-angled triangle postero-lateral lobe in the male genitalia (Fig. 20); the antrum is semicircular, the signum situates at posterior 2/5 of the corpus bursae in the female genitalia (Fig. 31). It is related to D. dichromella, but differs as noted in the diagnosis of the latter. Material examined. 97 male and 77 female specimens in different localities of China are examined. Distribution. China (Anhui, Beijing, Guangdong, Gansu, Guizhou, Henan, Hubei, Shandong, Shaanxi, Sichuan, Tianjin, Yunnan, Zhejiang), Japan. Remarks. Three species Trachycera nipponella, T. yakushimensis and T. vicinella described by Yamanaka (2000) from Japan are synonymized with this species, because the differences among them are too tiny to be interspecific: the uncus is broader or narrower apically, the postero-lateral lobe is shorter or not, the valva is a little narrower or not. We treated these as intraspecific differences. Among these species, T. nipponella has the greatest diffrences with D. paradichromella, as is noted in the original description: nipponella ‘…apical process of gnathos nearly crescent-shaped, not stick-like as in paradichromella, and apex pointed, slightly hooked, apico-dorsal portion minutely indented…’. These differences, however, might be caused by mounting slides: crescent, slightly hooked at apex when mounted laterally; clubbed, minutely indented at apex when mounted dorsal-ventrally., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on pages 350-353, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857, {"references":["Yamanaka, H. (1980) Descriptions of two new species of Eurhodope and synonymic notes of two known species belonging to Eurhodope and Assara from Japan (Lepidoptera: Pyralidae: Phycitinae). Transactions of the Lepidopterological Society of Japan, 31 (1 - 2), 66 - 73.","Roesler, R. U. (1987) Die bisher als Rhodophaea gelaufige Gattung Trachycera Ragonot 1893 (Lepidoptera, Pyralidae, Phycitinae) in der Palaarktis-Taxonomische Neuorientierung und Beschreibung neuer Taxa. Deutsche Entomologische Zeitschrift, Neue Folge, 97 (21), 305 - 319.","Du, Y. L., Sung, S. M. & Wu, C. S. (2005) A new genus in the subfamily Phycitinae (Lepidoptera: Pyralidae) from China. Annales Zoologici, 55 (1), 99 - 105.","Yamanaka, H. (2000) The Japanese species of the genus Trachycera Ragonot (Lepidoptera, Pyralidae, Phycitinae), with descriptions of three new species. Tinea, 16 (4), 232 - 239.","Kemal, M., Kizildag, S. & Kocak, A. O. (2020) On the nomenclature of a generic name in the Phycitinae of East Asia (Lepidop- tera, Pyraloidea). Miscellaneous Papers, 205, 1 - 2. [https: // www. researchgate. net / publication / 339415662]"]}

Dusungwua strictivinculum Ren & Li, sp. nov. (Figs 1, 13, 24, 34) Type material. Holotype. [CHINA] ♂, Mt. Tianzhu, Xiamen (24°36’N 117°57’E), Fujian, 220 m, 24-VIII-2010, coll. Bingbing Hu & Jing Zhang. Paratypes. [CHINA] Fujian: 84 ♂♂, 83 ♀♀, same data as for holotype, except dated 13-VII̅ 24-VIII-2010, gen. slide Nos. LJY 10862m, LJY 10493 m, TKJ 14080 m, TKJ14079 f, LJY11372 f, TKJ 14091 m, LHX14176 f; 2 ♀♀, Xiamen University (24°26’N 118°07’E), 85 m, 30-V-2010, 19-VI-2010, coll. Jing Zhang, gen. slide No. LLX11289 f; Guangxi: 1 ♂, 1 ♀, Jiman, Anchui (25°20’N 109°19’E), Rongshui County, 650 m, 2004-VII-16, coll. Jiasheng Xu, gen. slide Nos. LJY09168 m, LJY09167 f; 1 ♂, 1 ♀, Huaping, 950 m, 8- VIII-2006, coll. Weichun Li; 10 ♂♂, 8 ♀♀, Mt. Yuanbao, 700 m, 11̅ 12-VIII-2006, coll. Weichun Li, gen. slide No. LJY 10524m; Hunan: 2 ♂♂, 3 ♀♀, Chengguan Town (28°31’N 112°09’E), Taojiang County, 26-VII-2004, coll. Yunli Xiao; 2 ♂♂, 4 ♀♀, Maqiao, Baituo (27°49’N 112°56’E), Xiangtan city, 30-VII-2004, coll. Yunli Xiao, gen. slide Nos. RYD04819 m, RYD04820 f; 1 ♀, Yantang, Weishan, Xinhua County (27°36’N 111°18’E), 6-VIII- 2004, coll. Yunli Xiao; 2 ♂♂, 3 ♀♀, Cangxi town, Xinhua County, 8-VIII-2004, coll. Yunli Xiao, gen. slide No. RYD04291 m; 1 ♀, Mt. Jinpen (25°00’N 114°51’E), Jiangxi, 18-VII-2006, coll. Jiasheng Xu & Weichun Li. Diagnosis. This new species can be differentiated from its allies by the viculum conspicuously constricted on anterior 1/ 3 in the male genitalia. It resembles D. paradichromella superficially, but differ in the male scape is 1.5 times as long as wide, the bell-shaped uncus, the finger-like postero-lateral lobe of the transtilla in the male genitalia, and the corpus bursae is double-kerneled peanut shaped in the female genitalia. In D. paradichromella, the male scape is 2.5 times as long as wide, the uncus is triangular, the postero-lateral lobe of the transtilla is right-angled triangle shaped, and the corpus bursae is elongate bag-shaped. Description. Adult (Fig. 13) with wingspan 16.5̅22.0 mm. Head fuscous brown. Antenna brown, scape about 1.5 times as long as wide; male flagellum shallowly incurved basally, with a cluster of smaller scales. Labial palpus upturned just beyond vertex; first segment grayish white, second and third segments fuscous brown except whitetipped; second segment 1.7 times as long as third segment. Maxillary palpus brown. Patagium, tegula and thorax grayish brown. Forewing grayish brown, mottled with grayish white; antemedial line grayish white, from costal 1/3 from slightly oblique outward to dorsal 2/5, incurved at A, bordered outwardly by a black line and inwardly by an ovate, blackish brown patch near dorsum; discal spots blackish brown, fused, crescent; postmedial line grayish white, incurved at M 1 and CuA 2, edged outwardly with a grayish brown line and inwardly an indistinct broader band; terminal line black; cilia brown. Hindwing brown, cilia grayish brown. Foreleg blackish brown, mid and hindleg grayish brown mottled grayish white. Abdomen blackish brown, each segment darker in base than apex. Male genitalia (Fig. 24). Uncus bell-shaped, about same length as width at proximal base. Gnathos about 5/8 length of uncus, tapered, apex slightly hooked. Transtilla produced in U shape posteriorly, postero-lateral lobe fingerlike, uniform in width, twice as long as width. Valva four times as long as wide, slightly narrower distally; clasper short and stout; costa clubbed, about 5/6 length of valva, apical 1/3 narrowed; sacculus 2/7 length of valva, sinuate in S shape, sharp at end. Juxta V-shaped; lateral lobe short, clubbed, as long as gnathos, three times as long as wide. Vinculum somewhat goblet-shaped, 1.4 times as long as maximum width, conspicuously constricted on anterior 1/3. Phallus stout, about same length as valva, with fascicle-shaped crimples. Female genitalia (Fig. 34). Papillae anales large and wide, rounded at apex, with dense setae. Apophyses posteriores expanded at base, as long as apophyses anteriores. Eighth segment 1/3 as long as width. Antrum bowl-like, weakly sclerotized. Ductus bursae about half length of corpus bursae, smooth in posterior half, slightly broadened and granulated in anterior half. Corpus bursae twice as long as maximum width, double-kerneled peanut shaped, granulated near junction with ductus bursae; signum a rounded, invaginated cup consisting of concentrically arranged granules, situated at anterior 2/5 of corpus bursae. Ductus seminalis posterior margin of corpus bursae. Distribution. China (Fujian, Guangxi, Hunan, Jiangxi). Etymology. The specific name is derived from the Latin prefix strict-, meaning constricted, and the terminology vinculum, referring to the vinculum of this species is conspicuously constricted on anterior 1/3.

Dusungwua karenkolla (Shibuya, 1928) (Figs 1, 7, 18, 29) Eurhodope karenkolla Shibuya, 1928: 90. TL: China (Taiwan). TD: EIHU. Eurhodope (Eurhophaea) karenkolla Shibuya: Xu et al., 2005: 323. Furcata karenkolla: Du, Sung & Wu, 2005: 102. Dusungwua karenkolla: Kemal, Kizildağ & Koçak, 2020: 2. Diagnosis. Adult (Fig. 7) with wingspan 20.0̅26.0 mm. This species is recognized by the forewing with antemedial line from costal 1/3 obliquely outward to dorsal 2/5, broadened on upper half; the clubbed gnathos slightly bifurcate apically, the transtilla with two short, stout, apically truncate postero-lateral lobes that concave in V shape and forming an acute angle of about 60°, the subovate juxta and the phallus with a tapered spinose patch near apex in the male genitalia (Fig. 18); the lip-like antrum, the corpus bursae weakly sclerotized at junction with ductus bursae, and the absence of signum in the female genitalia (Fig. 29). It is related to D. dichromella, but in the latter species the uncus is sub-triangular basally and tubular apically, the gnathos is dilated apically, and postero-lateral lobe of the transtilla is longer and rounded apically in the male genitalia, and the signum is present in the female genitalia. Material examined. 132 male and 59 female specimens in different localities of China are examined. Distribution. China (Anhui, Fujian, Gansu, Guangxi, Guizhou, Henan, Hubei, Hunan, Shaanxi, Shanxi, Sichuan, Taiwan, Yunnan, Zhejiang), Korea., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on pages 349-350, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857, {"references":["Shibuya, J. (1928) The systematic study on the formosan Pyralidae. Journal of the Faculty of Agriculture, Hokkaido Imperial University, Sapporo, 22 (1), 1 - 300, pls. 1 - 9.","Du, Y. L., Sung, S. M. & Wu, C. S. (2005) A new genus in the subfamily Phycitinae (Lepidoptera: Pyralidae) from China. Annales Zoologici, 55 (1), 99 - 105.","Kemal, M., Kizildag, S. & Kocak, A. O. (2020) On the nomenclature of a generic name in the Phycitinae of East Asia (Lepidop- tera, Pyraloidea). Miscellaneous Papers, 205, 1 - 2. [https: // www. researchgate. net / publication / 339415662]"]}

Dusungwua ohkunii (Shibuya, 1928) (Figs 1, 8, 19, 30) Eurhodope ohkunii Shibuya, 1928: 90. TL: China (Taiwan). TD: EIHU. Furcata ohkunii: Ren & Li, 2012: 373. Dusungwua ohkunii: Kemal, Kizildağ & Koçak, 2020: 2. Diagnosis. Adult (Fig. 8) with wingspan 18.0̅24.0 mm. This species is featured by the following characters: the forewing has a small, blackish brown costal spot on inner margin of antemedial line, and a large, triangular, blackish brown posterior patch on outer margin of lower 3/4 of antemedial line; in the male genitalia (Fig. 19), the uncus is broad-triangular, rounded at apex, the clubbed gnathos is slightly broader at middle, the short postero-lateral lobes of the transtilla form an acute angle of about 45°, the valva is rather narrowed, slightly broadened apically, and the lateral lobe of the juxta is hillock-shaped; in the female genitalia (Fig. 30), the antrum is rectangular, the corpus bursae is three times as long as wide, and the signum situated at posterior 1/3 of the corpus bursae. It is similar to D. paradichromella except the triangular posterior patch is bigger, and covered with much more grayish white scales in median area of the forewing, the valva is narrower, the postero-lateral lobe of the transtilla is shorter, and the lateral lobe of the juxta is smaller in the male genitalia than the latter. Material examined. 674 male and 132 female specimens in different localities of China are examined. Distribution. China (Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangxi, Liaoning, Shaanxi, Shandong, Shanxi, Sichuan, Taiwan, Zhejiang)., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on page 350, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857, {"references":["Shibuya, J. (1928) The systematic study on the formosan Pyralidae. Journal of the Faculty of Agriculture, Hokkaido Imperial University, Sapporo, 22 (1), 1 - 300, pls. 1 - 9.","Li, H. H. et al. (2012) Microlepidoptera of Qinling Mountains (Insecta: Lepidoptera). Science Press, Beijing, xviii + 1271 pp., 35 col. pls.","Kemal, M., Kizildag, S. & Kocak, A. O. (2020) On the nomenclature of a generic name in the Phycitinae of East Asia (Lepidop- tera, Pyraloidea). Miscellaneous Papers, 205, 1 - 2. [https: // www. researchgate. net / publication / 339415662]"]}

Dusungwua quadrangula (Du, Sung & Wu, 2005) (Figs 1, 11, 22, 32) Furcata quadrangula Du, Sung et Wu, 2005: 101. TL: China (Fujian). TD: IZCAS. Dusungwua quadrangula: Kemal, Kizildağ & Koçak, 2020: 2. Diagnosis. This species can be separated from its congeners by the forewing has bigger and separated discal spots, the uncus is quadrate, postero-lateral lobe of the transtilla is short, fingerlike, forming an obtuse angle of about 120°, lateral lobe of the juxta is short, bean-shaped, and the phallus has dense fascicle-shaped microtrichia and crimples, and a sclerotized sword-like structure bearing dense spinules in the male genitalia (Fig. 22), and the sub-rounded corpus bursae has a round, invaginated cup-shaped signum situated at middle in the female genitalia. It is quite similar to the new species D. similiquadrangula Ren & Li, sp. nov., but differs as noted in the description of the latter. Description. Adult (Fig. 11) with wingspan 21.0̅24.0 mm. Female genitalia (Fig. 32). Papillae anales large and wide, shield-like, rounded at apex, with dense setae. Apophyses posteriores expanded at base, slightly longer than apophyses anteriores. Eighth segment broad and short. Antrum sub-rectangular. Ductus bursae membranous, less than half length of corpus bursae. Corpus bursae membranous, sub-rounded, 1.4 times as long as maximum width, granulated on inner wall; signum round, invaginated cup consisting of concentrically arranged granules, situated at middle of corpus bursae. Ductus seminalis from corpus bursae near ductus bursae. Material examined. 39 male and 20 female specimens in different localities of China are examined. Distribution. China (Anhui, Fujian, Guangdong, Guangxi, Jiangxi, Zhejiang). Remarks. The female of this species is described for the first time in science., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on page 354, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857, {"references":["Du, Y. L., Sung, S. M. & Wu, C. S. (2005) A new genus in the subfamily Phycitinae (Lepidoptera: Pyralidae) from China. Annales Zoologici, 55 (1), 99 - 105.","Kemal, M., Kizildag, S. & Kocak, A. O. (2020) On the nomenclature of a generic name in the Phycitinae of East Asia (Lepidop- tera, Pyraloidea). Miscellaneous Papers, 205, 1 - 2. [https: // www. researchgate. net / publication / 339415662]"]}

Genus Dusungwua Kemal, Kizildağ & Koçak, 2020 Dusungwua Kemal, Kizildağ & Koçak, 2020: 1. Furcata Du, Sung & Wu, 2005: 99. Junior homonym of Furcata Werby, 1938. Type species: Rhodophaea dichromella Ragonot, 1893. Generic characters. Adult with vertex flat. Antenna filiform, flagellum with fine cilia ventrally; male flagellum usually simple, with basal segments slightly constricted and incurved, or strongly constricted into arched shape in some species. Ocellus present; chaetosema well-developed. Labial palpus upturned. Maxillary palpus tiny, fanshaped at tip. Proboscis well-developed. Forewing without ridge of raised scales; antemedial line usually with a large black spot posteriorly on its inner side; with 11 veins (Fig. 2): R 2 free, from upper edge; R 3+4 and R 5 stalked in basal half, from upper angle of cell; M 2 and M 3 separated, M 3 from lower angle of cell; CuA 1 and CuA 2 free; cell about 3/5 length of forewing. Hindwing with 10 veins (Fig. 2): Sc approximated to Rs in basal 2/5; M 1 and Rs shortly stalked, from upper angle of cell; M 2 and M 3 shortly stalked or approximated in basal 1/5 to1/2, both from lower angle of cell; CuA 1 and CuA 2 free; cell about 1/2 length of hindwing. Male genitalia (Figs 13 ̅22). Uncus variously shaped, subtriangular, or bell-shaped, or trapezoidal. Gnathos clubbed, slightly bifid apically, in some species constricted at middle, enlarged in distal half. Transtilla well developed, produced in a U or V shape, and forming a pair of postero-lateral lobes. Valva narrow; clasper undeveloped, usually papillary; costa shorter than valva; sacculus shorter than half length of valva. Juxta U-shaped or V-shaped, lateral lobes stout. Vinculum U-shaped or V-shaped, sometimes with lateral sides incurved anteriorly. Phallus with flocculent crimples. Culcita five bunches, inner pair with slim scales and reed-like, outer pair with broad scales, petal-like, middle bunch respectively consisting of cup-like, triangular and style-shaped scales. Female genitalia (Figs 23 ̅31). Papillae anales broad, rounded at apex. Apophyses anteriores and posteriores short and subequal. Eighth segment broad and short. Antrum flat, sclerotized weakly. Ductus bursae and corpus bursae membranous, granulated on inner surface. Signum present or absent, usually invaginated and consisting of several granules. Ductus seminalis usually from posterior margin of corpus bursae, seldom from ductus bursae. Diagnosis. This genus is similar to Eurhodope H̹bner, 1825, but can be recognized by the male genitalia with the transtilla extending postero-laterally to form a pair of lateral lobes, which is not in the latter. Distribution. China, Japan and Korea. The distribution of this genus in China is shown in Fig. 1. Key to species based on male genitalia 1. Uncus elongate, longer than width at proximal base.......................................................... 2 - Uncus broad and short, shorter or about equal length as width at proximal base.................................... 7 2. Uncus abruptly narrowed from middle or distal 1/3 (Fig. 16)....................................... D. dichromella - Uncus gradually narrowed from base to apex............................................................... 3 3. Valva narrowly elongate, parallel sided.................................................................... 4 - Valva broad, sub-rectangular, or expanded and arched ventro-medially........................................... 5 4. Gnathos slightly expanded from middle to before apex; lateral lob of juxta rounded at apex (Fig. 19)........... D. ohkunii - Gnathos clubbed; lateral lobe of juxta truncate at apex (Fig. 21)............................... D. paripalpa sp. nov. 5. Valva broad sub-rectangular; transtilla ovately widened at middle (Fig. 14).................... D. antennalveata sp. nov. - Valva expanded and arched ventro-medially; transtilla not widened at middle..................................... 6 6. Transtilla produced in V shape posteriorly; vincullum conspicuously concave on lateral margins at anterior 1/3 (Fig. 15)....................................................................................... D. basinigra sp. nov. - Transtilla produced in U shape posteriorly; vinculum gradually narrowed anteriorly (Fig. 20).......... D. paradichromella 7. Uncus broad triangular or bell-shaped, rounded at apex....................................................... 8 - Uncus subquadrate, shallowly concave at middle at apex.................................................... 10 8. Vinculum constricted at anterior 1/3 (Fig. 24)......................................... D. strictivinculum sp. nov. - Vinculum gradually narrowed anteriorly, not constricted...................................................... 9 9. Transtilla produced in U shape posteriorly; lateral lobe of juxta slightly incurved distally; phallus with fascicular spines (Fig. 17)............................................................................. D. fascecornuta sp. nov. - Transtilla produced in V shape posteriorly; lateral lobe of juxta not incurved distally; phallus without fascicular spines (Fig. 18)....................................................................................... D. karenkolla 10. Valva sub-rectangular, with a triangular clasper (Fig. 22).......................................... D. quadrangula - Valva rectangular, with a short fingerlike clasper (Fig. 23)............................. D. similiquadrangula sp. nov. Key to species based on female genitalia 1. Signum present...................................................................................... 2 - Signum absent (Fig. 29)..................................................................... D. karenkolla 2. Corpus bursae shorter than ductus bursae.................................................................. 3 - Corpus bursae longer than ductus bursae.................................................................. 4 3. Signum placed posterior 1/3 of corpus bursae (Fig. 25)................................... D. antennalveata sp. nov. - Signum placed near anterior end of corpus bursae (Fig. 28)................................. D. fascecornuta sp. nov. 4. Signum placed at about middle.......................................................................... 5 - Signum placed before or beyond middle................................................................... 7 5. Ductus bursae with anterior 3/5 granulate on inner wall; antrum a narrow transverse band (Fig. 26).... D. basinigra sp. nov. - Ductus bursae smooth on inner wall; antrum not band-like.................................................... 6 6. Antrum sub-rectangular (Fig. 32)............................................................ D. quadrangula - Antrum dome-like (Fig. 33)..................................................... D. similiquadrangula sp. nov. 7. Signum placed before middle........................................................................... 8 - Signum placed beyond middle.......................................................................... 9 8. Corpus bursae elongate, 3.5 times as long as wide (Fig. 27)........................................ D. dichromella - Corpus bursae double-kerneled peanut shaped, 2.5 times as long as wide (Fig. 34).............. D. strictivinculum sp. nov. 9. Antrum semicircular; ductus bursae with inner wall granulated on anterior 3/4 (Fig. 31).............. D. paradichromella - Antrum rectangular; ductus bursae with inner wall granulated on anterior half (Fig. 30)..................... D. ohkunii, Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on pages 342-343, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857, {"references":["Kemal, M., Kizildag, S. & Kocak, A. O. (2020) On the nomenclature of a generic name in the Phycitinae of East Asia (Lepidop- tera, Pyraloidea). Miscellaneous Papers, 205, 1 - 2. [https: // www. researchgate. net / publication / 339415662]","Du, Y. L., Sung, S. M. & Wu, C. S. (2005) A new genus in the subfamily Phycitinae (Lepidoptera: Pyralidae) from China. Annales Zoologici, 55 (1), 99 - 105.","Werby, H. J. (1938) A new genus of Acanthocephala with forked lemnisci. Transactions of the American Microscopical Society, 57, 204 - 212. https: // doi. org / 10.2307 / 3222750","Ragonot, E. L. (1893) Monographie des Phycitinae et des Galleriinae. In: Romanoff, N. M., Memoires sur les Lepidopteres, St. Ptersburg, VII, pp. i-lvi + 1 - 658 pp., pls. 1 - 23."]}

Dusungwua similiquadrangula Ren & Li, sp. nov. (Figs 1, 12, 23, 33) Type material. Holotype. [CHINA] ♂, Rare Botanical Garden, Ruili (24°00’N 97°50’E), Yunnan, 1000 m, 6-VIII- 2005, coll. Yingdang Ren, gen. slide No. RYD04729. Paratypes. [CHINA] 2 ♂♂, 2 ♀♀, same data as for holotype, gen. slide Nos LHX 14117 m, RYD04730 f. Diagnosis. This new species resembles D. quadrangula (Du, Sung & Wu, 2005). It differs in the transtilla produced in U shape posteriorly and the postero-lateral lobe is three times as long as wide, and in the sub-rectangular valva that has a triangular clasper in the male genitalia. In D. quadrangula, the transtilla is produced in a V shape and postero-lateral lobe is twice as long as wide, and the triangular valva has a short fingerlike clasper in the male genitalia. In the female genitalia, the antrum is dome-like and the ductus bursae is broader in the new species; while the antrum is rectangular and the ductus bursae is narrower in the latter species. Description. Adult (Fig. 12) with wingspan 20.0̅ 24.5 mm. Head grayish brown, occiput creamy with a few grayish brow scales. Antenna brown, scape about 1.5 times as long as wide; male flagellum simple, neither constricted nor incurved basally. Labial palpus upturned just beyond vertex; first segment grayish brown mixed with grayish white, second and third segments brown; second segment 1.2 times as long as third segment. Maxillary palpus brown. Patagium, tegula and thorax grayish brown, with scattered grayish white scales. Forewing gray mottled grayish white; antemedial line at basal 1/3, indistinct, grayish white, arched, bordered by blackish brown patch near costa on its outer margin and near dorsum on its inner margin respectively; discal spots black, ovate, distinct, separated; postmedial line grayish white, sinuate, incurved at M 1 and near CuA 2, edged with black; terminal line black; cilia grayish brown. Hindwing and cilia grayish brown. Foreleg black, midleg grayish brown, hindleg grayish white. Abdomen with dorsal surface grayish brown, with grayish white at end of each segment, ventral surface grayish white basally and blackish brown distally on each segment. Male genitalia (Fig. 23). Uncus somewhat quadrate, about same length as width at proximal base, apex slightly concave at middle. Gnathos clubbed, gradually narrowed to weakly bifurcate apex, about 3/5 length of uncus. Transtilla produced in U shape posteriorly, postero-lateral lobe fingerlike, three times as long as wide, uniform in width. Valva four times as long as wide, sub-rectangular, slightly narrower distally; clasper triangular, sinuate along outer margin, with sparse setae; costa clubbed, about 4/5 length of valva, gradually narrowed to apex; sacculus 1/3 length of valva. Juxta V-shaped; lateral lobe short, broad, bean-shaped, twice as long as wide, about 4/5 length of gnathos. Vinculum U-shaped, same length as maximum width. Phallus cylindrical, about same length as valva, with fine, sclerotized fascicle-shaped crimples and a sclerotized sword-like structure bearing dense spinules. Female genitalia (Fig. 33). Papillae anales large and wide, shield-like, rounded at apex, with dense setae. Apophyses posteriores expanded at base, slightly longer than apophyses anteriores. Eighth segment broad and short. Antrum dome-like. Ductus bursae membranous, smooth on inner wall, about half length of corpus bursae. Corpus bursae membranous, oblong, 1.7 times as long as maximum width, granulated on inner wall; signum a rounded, invaginated cup consisting of concentrically arranged granules, situated at posterior 2/5 of corpus bursae. Ductus seminalis from junction between ductus bursae and corpus bursae. Distribution. China (Yunnan). Etymology. The specific name is derived from the Latin prefix simili-, meaning similar, and the specific name of another species, quadrangula, referring to their similarities.

Dusungwua paripalpa Ren & Li, sp. nov. (Figs 1, 10, 21) Type material. Holotype. [CHINA] ♂, Zhangjiajie (29°07΄N 110°29΄E), Hunan, 650 m, 11-VIII-2001, coll. Shulian Hao, gen. slide No. RYD04803. Paratype. [CHINA] 1 ♂, same data as for holotype. Diagnosis. This new species is similar to D. ohkunii. It can be distinguished by the second and third segments of the labial palpus about equal length; the forewing with a few scattered grayish white scales not forming a pattern; the gnathos constricted near middle, slightly expanded in distal half, and the lobate juxta with the lateral lobe slightly incurved distally in the male genitalia. In D. ohkunii, the second segment of the labial palpus is 1.5 times length of third segment; the forewing is covered with numerous grayish white scales that form a wedge-shaped grayish white blotch between the antemedial and the postmedial lines; the clubbed gnathos is slightly broader at middle, and the hillock-shaped lateral lobe of the juxta is not incurved distally in the male genitalia. Description. Adult (Fig. 10) with wingspan 16.0̅18.0 mm. Head grayish brown. Antenna brown, scape about 1.5 times as long as wide; male flagellum simple, neither constricted nor incurved basally. Labial palpus upturned beyond vertex; first segment grayish white; second and third segments brown, about equal length. Maxillary palpus grayish brown. Patagium, tegula and thorax blackish brown. Forewing fuscous brown, with scattered grayish white scales; antemedial line grayish white, from costal 2/5 obliquely outward to dorsal 3/7, its inner margin bordered with an indistinct, ovate, blackish brown patch near dorsum; discal spots blackish brown, fused, crescent; postmedial line grayish white, indistinct, incurved at M 1 and CuA 2; terminal line brown; cilia grayish brown to blackish brown. Hindwing and cilia grayish brown. Fore leg grayish brown, midleg and hindleg grayish white; tarsi fuscous brown, white at apex of each tarsomere. Abdomen with dorsal surface brown, darker in basal half of each segment, ventral surface yellowish brown. Male genitalia (Fig. 21). Uncus triangular, about same length as width at proximal base, gradually narrowed to rounded apex. Gnathos about 4/7 length of uncus, constricted near middle, slightly expanded from middle to before apex; apex attenuated, bifid. Transtilla produced in U shape posteriorly; postero-lateral lobe three times as long as wide, fingerlike, uniform in width, rounded at apex. Valva narrow, four times as long as wide, almost parallel-sided; clasper an indistinct mastoid process at middle near base; costa extending to near end of valva; sacculus 2/5 length of valva, slightly sinuate, narrowed from somewhat broad base to pointed apex. Juxta semicircular; lateral lobe slightly shorter than gnathos, slightly incurved distally, truncate apically, bearing long setae. Vinculum U-shaped, same length as maximum width, almost straight on anterior margin. Phallus cylindrical, slightly curved, about 4/5 length of valva, with sclerotized crimples in distal 2/3. Female. Unknown. Distribution. China (Hunan). Etymology. The specific name is derived from the Latin prefix pari -, meaning parity, and the morphological term palpus, referring to the second and third segments of the labial palpus about equal length.

Dusungwua basinigra Ren & Li, sp. nov. (Figs 1, 2, 4, 15, 26) Type material. Holotype. [CHINA] ♂, Heishuihe, Baxianshan (40°03΄N 117°24΄E), Jixian, Tianjin, 550 m, 8-VII- 2009, coll. Zhipin Liang. Paratypes. [CHINA] Anhui: 1 ♂, Tanqiaozhen (29°42΄N 118°20΄E), Huangshan, 6-VIII- 2004, coll. Jiasheng Xu & Jialiang Zhang; Beijing: 1 ♂, 2 ♀♀, Shandongzhuang, Pingguqu (40°11΄N 117°09΄E), 1080 m, 17̅ 20-VIII-2009, coll. Aihuan Zhang & Hongmei Chen; 13 ♂♂, 2 ♀♀, Mutianyu, Huairou (40°26΄N 116°34΄E), 23-VII-2010, coll. Aihuan Zhang & Hongmei Chen, gen. slide No. LJY 10650 m; 3 ♀♀, Xiaolongmen Forest Yard (40°25΄N 117°06΄E), 1080 m, 28̅ 30-VII-2006, coll. Hui Zhen & Denghui Kuang; 3 ♀♀, same place as former, 31-VII, 14-VIII-2009, coll. Aihuan Zhang; 4 ♂♂, 2 ♀♀, Dakezhuang, Songshan (40°01΄N 115°26΄E), VII-13,1- VIII-2010, coll. Aihuan Zhang; Fujian: 1 ♂, 2 ♀♀, Guadun (27°43΄N 117°38΄E), Mt. Wuyi, 1100 m, 30̅ 31-VII-2008, coll. Weichun Li, Yongling Sun & Haiyan Bai, gen. slide No. LJY 10534 m, LJY 10535 m; 1 ♀, Sangang (27°45΄N 117°41΄E), Mt. Wuyi, 740 m, 25-VII-2008, coll. Weichun Li, Yongling Sun & Haiyan Bai; Gansu: 1 ♂, Bifenggou, Wenxian, 360 m, 13-VII-2005, coll. Haili Yu; Guangdong: 1 ♀, Mt. Babao (23°30΄N 113°31΄E), Nanling, 1070 m, 23-VIII-2010, coll. Shulian Hao; Guangxi: 3 ♀♀, Huaping (24°51΄N 106°22΄E), 950 m, 6̅ 8-VIII-2006, coll. Weichun Li, gen. slide No. LJY10873; 1 ♀, Mt. Yuanbao (42°02΄N 119°17΄E), 700 m, 11- VIII-2006, coll. Weichun Li; Guizhou: 1 ♂, Dashahe, Daozhen, 1350 m, 25-V-2004, coll. Shulian Hao, gen. slide No. RYD04333; 1 ♀, Chengjiashan, Daozhen, 1300 m, 19-VIII-2004, coll. Yunli Xiao, gen. slide No. RYD04270; 1 ♀, Huguosi, Fanjingshan, 1300 m, 1-VIII-2001, coll. Houhun Li & Xinpu Wang; 1 ♀, Kuankuoshui, Suiyang, 840 m, 9-VI-2010, coll. Linlin Yang; Hebei: 1 ♀, Mt. Qishi (39°12΄N 114°43΄E), Laiyuan, 1300 m, 21-VII-2000, coll. Haili Yu, gen. slide No. DYL00696; 2 ♂♂, Mt. Xiantai (38°07΄N 113°50΄E), Jingxing, 1200 m, 23-VII-2000, coll. Haili Yu, gen. slide No. DYL00684; 1 ♂, Suanzaoping (37°22΄N 114°03΄E), Neiqiu, 670 m, 29-VII-2000, coll. Haili Yu, gen. slide No. DYL00697; 1 ♂, Lianhuachi, Mt. Wuling (40°03΄N 117°29΄E), Xinglong, 1800 m, 1-VIII- 2000, coll. Yanli Du & Zhendong Li; 3 ♂♂, 4 ♀♀, Mt. Wuling, Xinglong, 800 m, 2̅ 4-VII-2009, coll. Qing Jin; 4 ♂♂, 3 ♀♀, Mt. Wuling, Xinglong, 800 m, 20̅ 29-VIII-2009, coll. Aihuan Zhang & Xueling Tian; 2 ♂♂, 2 ♀♀, Mt. Wuling, Xinglong, 800 m, 15̅ 28-VII-2010, coll. Aihuan Zhang & Xiaoyu Shi; Henan: 1 ♂, Mt. Wangwu, Jiyuan, 700 m, 4-VI-2000, coll. Haili Yu, gen. slide No. DYL00608; 1 ♂, Songshan (34°31΄N 113°00΄E), Dengfeng, 800 m, 9-VI-2000, coll. Meicai Wei; 1 ♀, Huangshian (33°40΄N 111°38΄E), Xixia, 890 m, 18-VII-1998, coll. Houhun Li; 6 ♂♂, 2 ♀♀, Mt. Jigong (31°05΄N 114°05΄E), Xinyang, 700 m, 13-VII-2001, coll. Dandan Zhang; 1 ♂, Shiziping (33°47΄N 110°53΄E), Lushi, 1200 m, 19-VII-2001, coll. Dandan Zhang; 8 ♂♂, 8 ♀♀, Baotianman, Neixiang, 1200 m, 20-V̅ 4-VI-2006, coll. Jinmei Lv & Xu Zhang; 1 ♂, 2 ♀♀, Shibanyan (36°10΄N 113°44΄E), Linzhou, 550 m, 23-VII-2006, coll. Hui Zhen & Denghui Kuang; 28 ♂♂, 10 ♀♀, Guanshan, Huixian, 550 m, 25̅ 26-VII-2006, coll. Hui Zhen & Denghui Kuang, gen. slide No. LJY11367 f; 56 ♂♂, 36 ♀♀, Mt. Wangwu, Jiyuan, 800 m, 28̅ 30-VII- 2006, coll. Hui Zhen & Denghui Kuang; 16 ♂♂, 14 ♀♀, Mt. Huaguo, Yiyang, 1000 m, 1-VIII-2006, coll. Hui Zhen & Denghui Kuang; 1 ♂, 2 ♀♀, Dakuaidi, Lushi, 1100 m, 8-VIII-2006, coll. Hui Zhen & Denghui Kuang; Hubei: 1 ♂, Pingbaying (29°02΄N 108°59΄E), Xianfeng, 1280 m, 21-VII-1999, coll. Houhun Li, gen. slide No. RYD04030; 5 ♂♂, Bajiaomiao, Shennongjia, 1100 m, 18̅ 19-VII-2003, coll. Shulian Hao; 8 ♂♂, 3 ♀♀, Songbaizhen, Shennongjia, 1200̅ 1400 m, 15̅ 16-VII-2003, coll. Shulian Hao; 1 ♂, 3 ♀♀, Wujiashan (31°01΄N 115°05΄E), Yingshan, 7-VII-2008, coll. Yunli Xiao; 1 ♂, Zhangjiajie (29°07΄N 110°29΄E), Hunan, 650 m, 10-VIII-2001, coll. Houhun Li & Xinpu Wang; Jiangxi: 2 ♀♀, Mt. Lu (29°40΄N 115°59΄E), VIII-1975; 1 ♀, Sanqingsan (28°54΄N 118°04΄E), Yushan, 610 m, 21-VIII-1985, gen. slide No. DYL00897; 1 ♀, Mt. Jiulian (24°33΄N 114°37΄E), 21-VII-2006, coll. Jiasheng Xu & Weichun Li; Liaoning: 4 ♂♂, Laotudingzi, Huanren (41°19΄N 124°53΄E), 7̅ 9-VIII-2009, coll. Weichun Li & Jiayu Liu; 1 ♀, Mt. Baiyun (40°34΄N 123°01΄E), Haicheng, 15-VII-2010, Jiayu Liu & Yanpeng Cai; 1 ♀, Mt. Taibai, Shaanxi, 840 m, 11-VII-2005, coll. Ping You; Shandong: 8 ♂♂, 1 ♀, Mt. Yimeng (35°00΄N 118°40΄E), Linyi, 27̅ 30-VII-2009, coll. Qing Jin; 1 ♂, Beijiushui, Laoshan (36°12΄N 120°03΄E), Qingdao, 2-VIII-2009, coll. Qing Jin; Sichuan: 2 ♂♂, 5 ♀♀, Labahe (30°11΄N 102°26΄E), Tianquan, 1300 m, 29-VII-2004, coll. Yingdang Ren, gen. slide Nos RYD04364 m, RYD04365 f; 4 ♂♂, 4 ♀♀, Caopingcun (30°57΄N 103°19΄E), Wenchuan, 1557 m, 9̅ 11-VII-2014, coll. Kaijian Teng et al.; Tianjin: 2 ♂♂, Mt. Baxian, Jixian, 550 m, 23-VI-2001, coll. Houhun Li; 3 ♂♂, Mt. Baxian, Jixian, 500 m, 08-IV, 20-VII-2004, coll. Houhun Li; 5 ♂♂, 2 ♀♀, Mt. Baxian, Jixian, 560 m, 12̅ 15-VII-2005, coll. Houhun Li, gen. slide No. KDH05090 m; 6 ♂♂, 1 ♀, Mt. Pan (40°05΄N 117°16΄E), Jixian, 170 m, 20-VII-2005, coll. Houhun Li; 1 ♂, 3 ♀♀, Mt. Baxian, Jixian, 500 m, 16-VII, 1-IX-2005, coll. Houhun Li; 1 ♀, Mt. Baxian, Jixian, 300̅ 600 m, 4-VIII-2007, coll. Houhun Li; 1 ♀, Mt. Baxian, Jixian, 600 m, 4-VIII-2007, coll. Bingbing Hu & Mingfeng Cao; 41 ♂♂, 111 ♀♀, Mt. Baxian, Jixian, 550 m, 8, 14, 19, 28-VII, 2, 6, 11, 31-VIII, 13-IX-2009, coll. Bingbing Hu, Jingzhang & Zhipin Liang, gen. slide Nos LJY11085 W, LJY09155 f, LJY09153 f, LJY09154 f, LJY09150 f; 41 ♂♂, 37 ♀♀, Heishuihe, Mt. Baxian, Jixian, 600 m, 4-VI̅ 29-VIII-2010, coll. Yinghui Mou & Shurong Liu; 3 ♀♀, Heishuihe, Mt. Baxian, Jixian, 550 m, 20-IX-2009, coll. Shurong Liu; Zhejiang: 2 ♂♂, Kaishanlaodian, Mt. Tianmu (30°01΄N 117°29΄E), 1140 m, 17-VII-1999, coll. Houhun Li; 1 ♂, Chanyuansi Mt. Tianmu, 350 m, 15-VIII-1999, coll. Houhun Li; 1 ♂, Houshanmen, Mt. Tianmu, 500 m, 16-VIII-1999, coll. Houhun Li; 1 ♂, Mt. Qingliang (30°07΄N 119°02΄E), Lin’an, 800 m, 12-VIII-2005, coll. Yunli Xiao; 2 ♀♀, Mt. Wuyan (27°21΄N 119°56΄E), Taishun, 250̅ 1000 m, 3̅ 4-VIII-2005, coll. Yunli Xiao; 1 ♂, 1 ♀, Mt. Tianmu, Lin’an, 350 m, 9-VIII-2007, coll. Qing Jin; 1 ♂, Sanmuping, Mt. Tianmu, 400 m, 29-VII-2011, coll. Linlin Yang & Na Chen, slide No. LHX14123 W; 2 ♂♂, 2 ♀♀, Tianmucun, Mt. Tianmu, 1-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 5 ♂♂, 5 ♀♀, Laoan, Mt. Tianmu, 555 m, 3̅ 6-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 1 ♀, Sanmuping, Mt. Tianmu, 789 m, 14-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 3 ♀♀, Xiguan, Mt. Tianmu, 17̅ 19-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 1 ♂, Pinggang, Mt. Longxu (30°25΄N 119°33΄E), 754 m, 20-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 2 ♀♀, Qianqingtang, Mt. Qingliang, 1059 m, 29-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 1 ♂, 1 ♀, Sanmuping, Mt. Tianmu, 789 m, 9-VIII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 1 ♂, 1 ♀, Laoan, Mt. Tianmu, 555 m, 16-VIII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 1 ♂, Longxushancun, Mt. Longxu, 778 m, 20-VIII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang. Diagnosis. This new species can be separated from its congeners by the forewing black from the base to the inner margin of the antemedial line. The genitalia structures are similar to those of D. dichromella, differing in the male genitalia by the triangular uncus gradually narrowed to apex, the clubbed gnathos tapered to a hooked apex; and in the female genitalia by the subtriangular corpus bursae twice as long as the maximum width and the signum situated in the middle of the corpus bursae. In D. dichromella, the uncus is subtriangular in the basal half, sharply narrowed at middle and tubular in the distal half, the gnathos is broadened in distal half and the apex is not hooked in the male genitalia; the elongate bag-shaped corpus bursae is 3.5 times as long as its maximum width, and the signum is situated at the anterior 2/7 of the corpus bursae in the female genitalia. Description. Adult (Fig. 4) with wingspan 21.5̅ 27.5 mm. Head grayish brown. Antenna grayish white, scape about twice as long as wide, male with basal several flagellomeres constricted, incurved, without scale tufts; female flagellum simple. Labial palpus upturned beyond vertex; first segment grayish brown mixed with grayish white; second and third segments about equal length, grayish brown to blackish brown. Maxillary palpus grayish brown mixed with grayish white. Patagium, tegula and thorax yellowish brown to brownish fuscous. Forewing black in basal area, from grayish white gradually to grayish brown in medial area, grayish brown in distal area; antemedial line white, from costal 1/3 obliquely outward to dorsal 2/7; discal spots indistinct, black, fused into a club shape; postmedial line grayish white, sinuate, incurved at M 1 and CuA 2; terminal line black; cilia grayish brown. Hindwing pale grayish brown; cilia brown basally, pale brown distally. Foreleg blackish brown with scattered white scales, midleg grayish brown mixed with white, hindleg grayish white; tarsi black except white at apex of each tarsomere. Abdomen with dorsal surface blackish brown except yellow at posterior margin of each segment, ventral surface yellowish brown. Male genitalia (Fig. 15). Uncus triangular, longer than width at proximal base, gradually narrowed to rounded apex. Gnathos about 2/3 length of uncus, clubbed, tapered to hooked and slightly bifurcate apex, apico-dorsal portion minutely serrated. Transtilla produced in V shape posteriorly; postero-lateral lobe three times length of its medial width, tapered from base to narrowly rounded apex, forming an acute angle of about 45°. Valva three times as long as wide, narrowed at base, gradually broadened to middle, then narrowed to rounded apex; clasper an indistinct mastoid process at basal 1/7, bearing short setae; costa about 5/6 length of valva; sacculus sinuate, clubbed, 2/7 length of valva. Vinculum U-shaped posteriorly, 1.3 times as long as wide, concave on lateral margins at anterior 1/3, rounded anteriorly. Juxta broad V-shaped; lateral lobes about 4/5 length of gnathos, clubbed, slightly widened to rounded apex, bearing sparse setae. Phallus slightly longer than valva, with sclerotized crimples, crimples granulate in distal half. Female genitalia (Fig. 26). Papillae anales large and wide, with sparse setae, rounded at apex. Apophyses anteriores slightly longer and stouter than apophyses posteriores. Eighth segment about 2.5 times as wide as long. Antrum a narrow transverse band. Ductus bursae short, about 1/4 length of corpus bursae, gradually broadened, posterior 2/5 smooth, anterior 3/5 granulate on inner wall. Corpus bursae not easily differentiated from ductus bursae, subtriangular, twice as long as maximum width, granulate on inner wall in posterior 1/5; signum a rounded, invaginated cup consisting of concentrically arranged granules, situated at middle of corpus bursae. Ductus seminalis originating from posterior 1/5 of corpus bursae. Distribution. China (Anhui, Beijing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hebei, Henan, Hubei, Hunan, Jiangxi, Liaoning, Shaanxi, Shandong, Sichuan, Tianjin, Zhejiang). Etymology. The specific name is derived from the Latin prefix basi -, meaning basal, and the Latin word niger, meaning black, referring to the forewing black in the basal area., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on pages 346-348, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857

Dusungwua dichromella (Ragonot, 1893) (Figs 1, 5, 16, 27) Rhodophaea dichromella Ragonot, 1893: 75. TL: Japan. TD: CNC. Eurhodope dichromella: Shibuya, 1932: 331. Acrobasis dichromella: Roesler, 1985: 29. Trachycera dichromella: Roesler, 1987: 316. Furcata dichromella: Du, Sung & Wu, 2005: 101. Eurhodope pseudodichromella Yamanaka, 1980: 67. Syn. nov. Acrobasis pseudodichromella: Roesler, 1985: 29. Trachycera pseudodichromella: Roesler, 1987: 316. Eurhodope (Eurhophaea) pseudodichromella Yamanaka: Xu et al., 2005: 324. Furcata pseudodichromella: Du, Sung & Wu, 2005: 102. Dusungwua dichromella: Kemal, Kizildağ & Koçak, 2020: 2. Diagnosis. Adult (Fig. 5) with wingspan 18.0̅24.0 mm. This species can be recognized by the following characters: in the male genitalia (Fig. 16), the uncus is tapered abruptly near middle from broad base, with rather narrow and tubular distal part, the gnathos is expanded in distal half and has a small bifid apex, postero-lateral lobe of the transtilla is digitate, form an acute angle of about 60°, and the phallus has an ovate spinose patch near apex; in the female genitalia (Fig. 27), the antrum is rounded-triangular, the corpus bursae is elongate, 3.5 times as long as wide, and the signum situated at anterior 2/7 of the corpus bursae. It is superficially quite similar to D. paradichromella, but can be separated from the latter by the forewing diffused with much more white scales in median area, the uncus tapered abruptly near middle, the gnathos with distal half dilated in the male genitalia, and the signum situated at anterior 2/7 of the corpus bursae; in D. paradichromella, the forewing is mottled with less white scales in median area, the uncus is gradually narrowed, the gnathos is clubbed in the male genitalia, and the signum situated at posterior 2/5 of the corpus bursae in the female genitalia. Material examined. 251 male and 86 female specimens in different localities of China are examined. Distribution. China (Anhui, Beijing, Fujian, Gansu, Guangxi, Guizhou, Hebei, Henan, Hubei, Hunan, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Sichuan, Tianjin, Zhejiang), Japan. Discussion. Variations occur in both male and female genitalia. In the male genitalia (Fig. 16 a-d), the distal narrow part of the uncus varies from 1/3 to 1/2 of the total length; the distal half of the gnathos is expanded to varying degrees; postero-lateral lobes of the transtilla form an acute angle of 45° to 60°, shallowly or deeply within individuals. In the female genitalia, the corpus bursae varies in shape: it is usually elongate, slightly widening toward anterior margin; in some species, the distal 1/3 is wider or dilated subroundly (Fig. 27a). Yamanaka (1980) described D. pseudodichromella (Yamanaka, 1980) from Japan. Yamanaka stated that D. pseudodichromella differed from D. dichromella in the antemedian line of the forewing more gently curved, the postero-lateral lobes of the transtilla are narrower and more deeply concave in between. We regard these as variations within species. Hence, we treat D. pseudodichromella as a synonym of D. dichromella., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on page 348, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857, {"references":["Ragonot, E. L. (1893) Monographie des Phycitinae et des Galleriinae. In: Romanoff, N. M., Memoires sur les Lepidopteres, St. Ptersburg, VII, pp. i-lvi + 1 - 658 pp., pls. 1 - 23.","Shibuya, J. (1932) List of the Japanese Phycitinae. Insect world, Gifu, 35, 255 ‾ 258 + 295 ‾ 298 + 331 ‾ 335.","Roesler, R. U. (1985) Neue Resultate in der Benennung von Termini bei Phycitinae (Lepidoptera, Pyraloidea) mit Neunachweisen f ʾ r Europa. Neue Entomologische Nachrichten, 17, 29 - 38.","Roesler, R. U. (1987) Die bisher als Rhodophaea gelaufige Gattung Trachycera Ragonot 1893 (Lepidoptera, Pyralidae, Phycitinae) in der Palaarktis-Taxonomische Neuorientierung und Beschreibung neuer Taxa. Deutsche Entomologische Zeitschrift, Neue Folge, 97 (21), 305 - 319.","Du, Y. L., Sung, S. M. & Wu, C. S. (2005) A new genus in the subfamily Phycitinae (Lepidoptera: Pyralidae) from China. Annales Zoologici, 55 (1), 99 - 105.","Yamanaka, H. (1980) Descriptions of two new species of Eurhodope and synonymic notes of two known species belonging to Eurhodope and Assara from Japan (Lepidoptera: Pyralidae: Phycitinae). Transactions of the Lepidopterological Society of Japan, 31 (1 - 2), 66 - 73.","Kemal, M., Kizildag, S. & Kocak, A. O. (2020) On the nomenclature of a generic name in the Phycitinae of East Asia (Lepidop- tera, Pyraloidea). Miscellaneous Papers, 205, 1 - 2. [https: // www. researchgate. net / publication / 339415662]"]}

Ren, Yingdang, Yang, Linlin, Liu, Hongxia, Li, Houhun (2020): Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms. Zootaxa 4894 (3): 341-365, DOI: https://doi.org/10.11646/zootaxa.4894.3.3

Dusungwua fascecornuta Ren & Li, sp. nov. (Figs 1, 6, 17, 28) Type material. Holotype. [CHINA] ♂, Xiannvdong (28°52΄N 107°37΄E), Daozhen, Guizhou, 600 m, 17-V-2004, coll. Yunli Xiao, gen. slide No. RYD04203. Paratypes. [CHINA] 2 ♂♂, 1 ♀, same data as for holotype; 5 ♂♂, Xiasicun, Kuankuoshui, Suiyang, 840 m, 8̅ 9-VI-2010, coll. Linlin Yang; Henan: 1 ♀, Mt. Huaguo, Yiyang, 1000 m, 3-VIII-2006, coll. Denghui Kuang & Hui Zhen; Hubei: 1 ♂, Pingbaying, Xianfeng, 1280 m, 21-VII-1999, coll. Houhun Li & Yanli Du; Sichuan: 3 ♂♂, Baoxing, 1100 m, 1-VIII-2004, coll. Yingdang Ren; Guizhou: 1 ♂, Dashahe, Daozhen, 1450 m, 23-V-2004, coll. Shulian Hao; 1 ♀, Suoluo (28°27΄N 105°59΄E), Chishui, 240 m, 21- IX-2000, coll. Haili Yu. Diagnosis. This new species is quite similar to the new species D. paripalpa sp. nov. It can be recognized by the valva not parallel-sided and conspicuously convex before middle on the ventral margin, and the phallus with fascicular spines in the male genitalia. In D. paripalpa sp. nov., the valva is almost parallel-sided except slightly convex before the middle on the ventral margin, and the phallus has no spines in the male genitalia. Description. Adult (Fig. 6) with wingspan 18.5̅22.0 mm. Head grayish fuscous brown. Antenna coppery brown, scape about 1.5 times as long as wide; male flagellum simple, neither constricted nor incurved basally. Labial palpus upturned beyond vertex; first segment grayish white, second and third segments brown fuscous, scattered with a few grayish white scales on inner surface; second segment 1.5 times as long as third segment. Maxillary palpus grayish brown. Patagium, tegula and thorax grayish brown. Forewing gray powdered with white; antemedial line grayish white, from costal 2/7 obliquely outward to dorsal 2/5, its outer margin bordered by a triangular, black costal patch, its inner margin bordered by an ovate, blackish brown patch along posterior half; discal spots blackish brown, fused, reniform; postmedial line grayish white, incurved at M 1 and CuA 2; terminal line black, interrupted; cilia grayish brown. Hindwing and cilia grayish brown. Foreleg blackish brown, midleg and hindleg brown mixed with white. Abdomen blackish brown basally and yellowish brown distally on each segment. Male genitalia (Fig. 17). Uncus broad-triangular, shorter than width at proximal base, apex narrowly rounded. Gnathos about 3/5 length of uncus, constricted near middle, slightly expanded from middle to before apex, apex attenuated, bifid. Transtilla produced in broad U shape posteriorly; postero-lateral lobe twice as long as wide, fingerlike, uniform in width, rounded at apex. Valva three times as long as maximum wide, convex near middle on ventral margin, rounded at apex; costa straight, narrowly banded, about 5/6 length of valva; clasper a short, fingerlike process at middle near base; sacculus sinuate, 1/3 length of valva. Juxta V-shaped, lateral lobe about same length as gnathos, lobate, slightly incurved apically, bearing sparse setae. Vinculum U-shaped, same length as maximum width. Phallus cylindrical, slightly curved, about 1.2 times length of valva, with sclerotized crimples and fascicular spines in basal 3/4. Female genitalia (Fig. 28). Papillae anales large and wide, with dense setae, rounded at apex. Apophyses anteriores slightly longer than apophyses posteriores. Eighth segment about half length of wide. Antrum sectorshaped. Ductus bursae about 2.2 times length of corpus bursae, posterior 3/5 narrow, smooth, anterior 2/5 gradually broadened toward corpus bursae, granulated on inner wall. Corpus bursae somewhat rounded, granulated; signum an ovate, invaginated cup consisting of concentrically arranged granules, situated at anterior end of corpus bursae. Ductus seminalis originating from anterior 1/3 of corpus bursae. Distribution. China (Guizhou, Henan, Hubei, Sichuan). Etymology. The specific name is derived from the Latin prefix fasce -, meaning fascicular, and the morphological term cornutus, referring to the male phallus with cornuti consisting of fascicular spines., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on page 349, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857

Dusungwua fascecornuta Ren & Li, sp. nov. (Figs 1, 6, 17, 28) Type material. Holotype. [CHINA] ♂, Xiannvdong (28°52΄N 107°37΄E), Daozhen, Guizhou, 600 m, 17-V-2004, coll. Yunli Xiao, gen. slide No. RYD04203. Paratypes. [CHINA] 2 ♂♂, 1 ♀, same data as for holotype; 5 ♂♂, Xiasicun, Kuankuoshui, Suiyang, 840 m, 8̅ 9-VI-2010, coll. Linlin Yang; Henan: 1 ♀, Mt. Huaguo, Yiyang, 1000 m, 3-VIII-2006, coll. Denghui Kuang & Hui Zhen; Hubei: 1 ♂, Pingbaying, Xianfeng, 1280 m, 21-VII-1999, coll. Houhun Li & Yanli Du; Sichuan: 3 ♂♂, Baoxing, 1100 m, 1-VIII-2004, coll. Yingdang Ren; Guizhou: 1 ♂, Dashahe, Daozhen, 1450 m, 23-V-2004, coll. Shulian Hao; 1 ♀, Suoluo (28°27΄N 105°59΄E), Chishui, 240 m, 21- IX-2000, coll. Haili Yu. Diagnosis. This new species is quite similar to the new species D. paripalpa sp. nov. It can be recognized by the valva not parallel-sided and conspicuously convex before middle on the ventral margin, and the phallus with fascicular spines in the male genitalia. In D. paripalpa sp. nov., the valva is almost parallel-sided except slightly convex before the middle on the ventral margin, and the phallus has no spines in the male genitalia. Description. Adult (Fig. 6) with wingspan 18.5̅22.0 mm. Head grayish fuscous brown. Antenna coppery brown, scape about 1.5 times as long as wide; male flagellum simple, neither constricted nor incurved basally. Labial palpus upturned beyond vertex; first segment grayish white, second and third segments brown fuscous, scattered with a few grayish white scales on inner surface; second segment 1.5 times as long as third segment. Maxillary palpus grayish brown. Patagium, tegula and thorax grayish brown. Forewing gray powdered with white; antemedial line grayish white, from costal 2/7 obliquely outward to dorsal 2/5, its outer margin bordered by a triangular, black costal patch, its inner margin bordered by an ovate, blackish brown patch along posterior half; discal spots blackish brown, fused, reniform; postmedial line grayish white, incurved at M 1 and CuA 2; terminal line black, interrupted; cilia grayish brown. Hindwing and cilia grayish brown. Foreleg blackish brown, midleg and hindleg brown mixed with white. Abdomen blackish brown basally and yellowish brown distally on each segment. Male genitalia (Fig. 17). Uncus broad-triangular, shorter than width at proximal base, apex narrowly rounded. Gnathos about 3/5 length of uncus, constricted near middle, slightly expanded from middle to before apex, apex attenuated, bifid. Transtilla produced in broad U shape posteriorly; postero-lateral lobe twice as long as wide, fingerlike, uniform in width, rounded at apex. Valva three times as long as maximum wide, convex near middle on ventral margin, rounded at apex; costa straight, narrowly banded, about 5/6 length of valva; clasper a short, fingerlike process at middle near base; sacculus sinuate, 1/3 length of valva. Juxta V-shaped, lateral lobe about same length as gnathos, lobate, slightly incurved apically, bearing sparse setae. Vinculum U-shaped, same length as maximum width. Phallus cylindrical, slightly curved, about 1.2 times length of valva, with sclerotized crimples and fascicular spines in basal 3/4. Female genitalia (Fig. 28). Papillae anales large and wide, with dense setae, rounded at apex. Apophyses anteriores slightly longer than apophyses posteriores. Eighth segment about half length of wide. Antrum sectorshaped. Ductus bursae about 2.2 times length of corpus bursae, posterior 3/5 narrow, smooth, anterior 2/5 gradually broadened toward corpus bursae, granulated on inner wall. Corpus bursae somewhat rounded, granulated; signum an ovate, invaginated cup consisting of concentrically arranged granules, situated at anterior end of corpus bursae. Ductus seminalis originating from anterior 1/3 of corpus bursae. Distribution. China (Guizhou, Henan, Hubei, Sichuan). Etymology. The specific name is derived from the Latin prefix fasce -, meaning fascicular, and the morphological term cornutus, referring to the male phallus with cornuti consisting of fascicular spines.

Dusungwua strictivinculum Ren & Li, sp. nov. (Figs 1, 13, 24, 34) Type material. Holotype. [CHINA] ♂, Mt. Tianzhu, Xiamen (24°36’N 117°57’E), Fujian, 220 m, 24-VIII-2010, coll. Bingbing Hu & Jing Zhang. Paratypes. [CHINA] Fujian: 84 ♂♂, 83 ♀♀, same data as for holotype, except dated 13-VII̅ 24-VIII-2010, gen. slide Nos. LJY 10862m, LJY 10493 m, TKJ 14080 m, TKJ14079 f, LJY11372 f, TKJ 14091 m, LHX14176 f; 2 ♀♀, Xiamen University (24°26’N 118°07’E), 85 m, 30-V-2010, 19-VI-2010, coll. Jing Zhang, gen. slide No. LLX11289 f; Guangxi: 1 ♂, 1 ♀, Jiman, Anchui (25°20’N 109°19’E), Rongshui County, 650 m, 2004-VII-16, coll. Jiasheng Xu, gen. slide Nos. LJY09168 m, LJY09167 f; 1 ♂, 1 ♀, Huaping, 950 m, 8- VIII-2006, coll. Weichun Li; 10 ♂♂, 8 ♀♀, Mt. Yuanbao, 700 m, 11̅ 12-VIII-2006, coll. Weichun Li, gen. slide No. LJY 10524m; Hunan: 2 ♂♂, 3 ♀♀, Chengguan Town (28°31’N 112°09’E), Taojiang County, 26-VII-2004, coll. Yunli Xiao; 2 ♂♂, 4 ♀♀, Maqiao, Baituo (27°49’N 112°56’E), Xiangtan city, 30-VII-2004, coll. Yunli Xiao, gen. slide Nos. RYD04819 m, RYD04820 f; 1 ♀, Yantang, Weishan, Xinhua County (27°36’N 111°18’E), 6-VIII- 2004, coll. Yunli Xiao; 2 ♂♂, 3 ♀♀, Cangxi town, Xinhua County, 8-VIII-2004, coll. Yunli Xiao, gen. slide No. RYD04291 m; 1 ♀, Mt. Jinpen (25°00’N 114°51’E), Jiangxi, 18-VII-2006, coll. Jiasheng Xu & Weichun Li. Diagnosis. This new species can be differentiated from its allies by the viculum conspicuously constricted on anterior 1/ 3 in the male genitalia. It resembles D. paradichromella superficially, but differ in the male scape is 1.5 times as long as wide, the bell-shaped uncus, the finger-like postero-lateral lobe of the transtilla in the male genitalia, and the corpus bursae is double-kerneled peanut shaped in the female genitalia. In D. paradichromella, the male scape is 2.5 times as long as wide, the uncus is triangular, the postero-lateral lobe of the transtilla is right-angled triangle shaped, and the corpus bursae is elongate bag-shaped. Description. Adult (Fig. 13) with wingspan 16.5̅22.0 mm. Head fuscous brown. Antenna brown, scape about 1.5 times as long as wide; male flagellum shallowly incurved basally, with a cluster of smaller scales. Labial palpus upturned just beyond vertex; first segment grayish white, second and third segments fuscous brown except whitetipped; second segment 1.7 times as long as third segment. Maxillary palpus brown. Patagium, tegula and thorax grayish brown. Forewing grayish brown, mottled with grayish white; antemedial line grayish white, from costal 1/3 from slightly oblique outward to dorsal 2/5, incurved at A, bordered outwardly by a black line and inwardly by an ovate, blackish brown patch near dorsum; discal spots blackish brown, fused, crescent; postmedial line grayish white, incurved at M 1 and CuA 2, edged outwardly with a grayish brown line and inwardly an indistinct broader band; terminal line black; cilia brown. Hindwing brown, cilia grayish brown. Foreleg blackish brown, mid and hindleg grayish brown mottled grayish white. Abdomen blackish brown, each segment darker in base than apex. Male genitalia (Fig. 24). Uncus bell-shaped, about same length as width at proximal base. Gnathos about 5/8 length of uncus, tapered, apex slightly hooked. Transtilla produced in U shape posteriorly, postero-lateral lobe fingerlike, uniform in width, twice as long as width. Valva four times as long as wide, slightly narrower distally; clasper short and stout; costa clubbed, about 5/6 length of valva, apical 1/3 narrowed; sacculus 2/7 length of valva, sinuate in S shape, sharp at end. Juxta V-shaped; lateral lobe short, clubbed, as long as gnathos, three times as long as wide. Vinculum somewhat goblet-shaped, 1.4 times as long as maximum width, conspicuously constricted on anterior 1/3. Phallus stout, about same length as valva, with fascicle-shaped crimples. Female genitalia (Fig. 34). Papillae anales large and wide, rounded at apex, with dense setae. Apophyses posteriores expanded at base, as long as apophyses anteriores. Eighth segment 1/3 as long as width. Antrum bowl-like, weakly sclerotized. Ductus bursae about half length of corpus bursae, smooth in posterior half, slightly broadened and granulated in anterior half. Corpus bursae twice as long as maximum width, double-kerneled peanut shaped, granulated near junction with ductus bursae; signum a rounded, invaginated cup consisting of concentrically arranged granules, situated at anterior 2/5 of corpus bursae. Ductus seminalis posterior margin of corpus bursae. Distribution. China (Fujian, Guangxi, Hunan, Jiangxi). Etymology. The specific name is derived from the Latin prefix strict-, meaning constricted, and the terminology vinculum, referring to the vinculum of this species is conspicuously constricted on anterior 1/3., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on pages 355-356, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857

Dusungwua paripalpa Ren & Li, sp. nov. (Figs 1, 10, 21) Type material. Holotype. [CHINA] ♂, Zhangjiajie (29°07΄N 110°29΄E), Hunan, 650 m, 11-VIII-2001, coll. Shulian Hao, gen. slide No. RYD04803. Paratype. [CHINA] 1 ♂, same data as for holotype. Diagnosis. This new species is similar to D. ohkunii. It can be distinguished by the second and third segments of the labial palpus about equal length; the forewing with a few scattered grayish white scales not forming a pattern; the gnathos constricted near middle, slightly expanded in distal half, and the lobate juxta with the lateral lobe slightly incurved distally in the male genitalia. In D. ohkunii, the second segment of the labial palpus is 1.5 times length of third segment; the forewing is covered with numerous grayish white scales that form a wedge-shaped grayish white blotch between the antemedial and the postmedial lines; the clubbed gnathos is slightly broader at middle, and the hillock-shaped lateral lobe of the juxta is not incurved distally in the male genitalia. Description. Adult (Fig. 10) with wingspan 16.0̅18.0 mm. Head grayish brown. Antenna brown, scape about 1.5 times as long as wide; male flagellum simple, neither constricted nor incurved basally. Labial palpus upturned beyond vertex; first segment grayish white; second and third segments brown, about equal length. Maxillary palpus grayish brown. Patagium, tegula and thorax blackish brown. Forewing fuscous brown, with scattered grayish white scales; antemedial line grayish white, from costal 2/5 obliquely outward to dorsal 3/7, its inner margin bordered with an indistinct, ovate, blackish brown patch near dorsum; discal spots blackish brown, fused, crescent; postmedial line grayish white, indistinct, incurved at M 1 and CuA 2; terminal line brown; cilia grayish brown to blackish brown. Hindwing and cilia grayish brown. Fore leg grayish brown, midleg and hindleg grayish white; tarsi fuscous brown, white at apex of each tarsomere. Abdomen with dorsal surface brown, darker in basal half of each segment, ventral surface yellowish brown. Male genitalia (Fig. 21). Uncus triangular, about same length as width at proximal base, gradually narrowed to rounded apex. Gnathos about 4/7 length of uncus, constricted near middle, slightly expanded from middle to before apex; apex attenuated, bifid. Transtilla produced in U shape posteriorly; postero-lateral lobe three times as long as wide, fingerlike, uniform in width, rounded at apex. Valva narrow, four times as long as wide, almost parallel-sided; clasper an indistinct mastoid process at middle near base; costa extending to near end of valva; sacculus 2/5 length of valva, slightly sinuate, narrowed from somewhat broad base to pointed apex. Juxta semicircular; lateral lobe slightly shorter than gnathos, slightly incurved distally, truncate apically, bearing long setae. Vinculum U-shaped, same length as maximum width, almost straight on anterior margin. Phallus cylindrical, slightly curved, about 4/5 length of valva, with sclerotized crimples in distal 2/3. Female. Unknown. Distribution. China (Hunan). Etymology. The specific name is derived from the Latin prefix pari -, meaning parity, and the morphological term palpus, referring to the second and third segments of the labial palpus about equal length., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on page 353, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857

Dusungwua basinigra Ren & Li, sp. nov. (Figs 1, 2, 4, 15, 26) Type material. Holotype. [CHINA] ♂, Heishuihe, Baxianshan (40°03΄N 117°24΄E), Jixian, Tianjin, 550 m, 8-VII- 2009, coll. Zhipin Liang. Paratypes. [CHINA] Anhui: 1 ♂, Tanqiaozhen (29°42΄N 118°20΄E), Huangshan, 6-VIII- 2004, coll. Jiasheng Xu & Jialiang Zhang; Beijing: 1 ♂, 2 ♀♀, Shandongzhuang, Pingguqu (40°11΄N 117°09΄E), 1080 m, 17̅ 20-VIII-2009, coll. Aihuan Zhang & Hongmei Chen; 13 ♂♂, 2 ♀♀, Mutianyu, Huairou (40°26΄N 116°34΄E), 23-VII-2010, coll. Aihuan Zhang & Hongmei Chen, gen. slide No. LJY 10650 m; 3 ♀♀, Xiaolongmen Forest Yard (40°25΄N 117°06΄E), 1080 m, 28̅ 30-VII-2006, coll. Hui Zhen & Denghui Kuang; 3 ♀♀, same place as former, 31-VII, 14-VIII-2009, coll. Aihuan Zhang; 4 ♂♂, 2 ♀♀, Dakezhuang, Songshan (40°01΄N 115°26΄E), VII-13,1- VIII-2010, coll. Aihuan Zhang; Fujian: 1 ♂, 2 ♀♀, Guadun (27°43΄N 117°38΄E), Mt. Wuyi, 1100 m, 30̅ 31-VII-2008, coll. Weichun Li, Yongling Sun & Haiyan Bai, gen. slide No. LJY 10534 m, LJY 10535 m; 1 ♀, Sangang (27°45΄N 117°41΄E), Mt. Wuyi, 740 m, 25-VII-2008, coll. Weichun Li, Yongling Sun & Haiyan Bai; Gansu: 1 ♂, Bifenggou, Wenxian, 360 m, 13-VII-2005, coll. Haili Yu; Guangdong: 1 ♀, Mt. Babao (23°30΄N 113°31΄E), Nanling, 1070 m, 23-VIII-2010, coll. Shulian Hao; Guangxi: 3 ♀♀, Huaping (24°51΄N 106°22΄E), 950 m, 6̅ 8-VIII-2006, coll. Weichun Li, gen. slide No. LJY10873; 1 ♀, Mt. Yuanbao (42°02΄N 119°17΄E), 700 m, 11- VIII-2006, coll. Weichun Li; Guizhou: 1 ♂, Dashahe, Daozhen, 1350 m, 25-V-2004, coll. Shulian Hao, gen. slide No. RYD04333; 1 ♀, Chengjiashan, Daozhen, 1300 m, 19-VIII-2004, coll. Yunli Xiao, gen. slide No. RYD04270; 1 ♀, Huguosi, Fanjingshan, 1300 m, 1-VIII-2001, coll. Houhun Li & Xinpu Wang; 1 ♀, Kuankuoshui, Suiyang, 840 m, 9-VI-2010, coll. Linlin Yang; Hebei: 1 ♀, Mt. Qishi (39°12΄N 114°43΄E), Laiyuan, 1300 m, 21-VII-2000, coll. Haili Yu, gen. slide No. DYL00696; 2 ♂♂, Mt. Xiantai (38°07΄N 113°50΄E), Jingxing, 1200 m, 23-VII-2000, coll. Haili Yu, gen. slide No. DYL00684; 1 ♂, Suanzaoping (37°22΄N 114°03΄E), Neiqiu, 670 m, 29-VII-2000, coll. Haili Yu, gen. slide No. DYL00697; 1 ♂, Lianhuachi, Mt. Wuling (40°03΄N 117°29΄E), Xinglong, 1800 m, 1-VIII- 2000, coll. Yanli Du & Zhendong Li; 3 ♂♂, 4 ♀♀, Mt. Wuling, Xinglong, 800 m, 2̅ 4-VII-2009, coll. Qing Jin; 4 ♂♂, 3 ♀♀, Mt. Wuling, Xinglong, 800 m, 20̅ 29-VIII-2009, coll. Aihuan Zhang & Xueling Tian; 2 ♂♂, 2 ♀♀, Mt. Wuling, Xinglong, 800 m, 15̅ 28-VII-2010, coll. Aihuan Zhang & Xiaoyu Shi; Henan: 1 ♂, Mt. Wangwu, Jiyuan, 700 m, 4-VI-2000, coll. Haili Yu, gen. slide No. DYL00608; 1 ♂, Songshan (34°31΄N 113°00΄E), Dengfeng, 800 m, 9-VI-2000, coll. Meicai Wei; 1 ♀, Huangshian (33°40΄N 111°38΄E), Xixia, 890 m, 18-VII-1998, coll. Houhun Li; 6 ♂♂, 2 ♀♀, Mt. Jigong (31°05΄N 114°05΄E), Xinyang, 700 m, 13-VII-2001, coll. Dandan Zhang; 1 ♂, Shiziping (33°47΄N 110°53΄E), Lushi, 1200 m, 19-VII-2001, coll. Dandan Zhang; 8 ♂♂, 8 ♀♀, Baotianman, Neixiang, 1200 m, 20-V̅ 4-VI-2006, coll. Jinmei Lv & Xu Zhang; 1 ♂, 2 ♀♀, Shibanyan (36°10΄N 113°44΄E), Linzhou, 550 m, 23-VII-2006, coll. Hui Zhen & Denghui Kuang; 28 ♂♂, 10 ♀♀, Guanshan, Huixian, 550 m, 25̅ 26-VII-2006, coll. Hui Zhen & Denghui Kuang, gen. slide No. LJY11367 f; 56 ♂♂, 36 ♀♀, Mt. Wangwu, Jiyuan, 800 m, 28̅ 30-VII- 2006, coll. Hui Zhen & Denghui Kuang; 16 ♂♂, 14 ♀♀, Mt. Huaguo, Yiyang, 1000 m, 1-VIII-2006, coll. Hui Zhen & Denghui Kuang; 1 ♂, 2 ♀♀, Dakuaidi, Lushi, 1100 m, 8-VIII-2006, coll. Hui Zhen & Denghui Kuang; Hubei: 1 ♂, Pingbaying (29°02΄N 108°59΄E), Xianfeng, 1280 m, 21-VII-1999, coll. Houhun Li, gen. slide No. RYD04030; 5 ♂♂, Bajiaomiao, Shennongjia, 1100 m, 18̅ 19-VII-2003, coll. Shulian Hao; 8 ♂♂, 3 ♀♀, Songbaizhen, Shennongjia, 1200̅ 1400 m, 15̅ 16-VII-2003, coll. Shulian Hao; 1 ♂, 3 ♀♀, Wujiashan (31°01΄N 115°05΄E), Yingshan, 7-VII-2008, coll. Yunli Xiao; 1 ♂, Zhangjiajie (29°07΄N 110°29΄E), Hunan, 650 m, 10-VIII-2001, coll. Houhun Li & Xinpu Wang; Jiangxi: 2 ♀♀, Mt. Lu (29°40΄N 115°59΄E), VIII-1975; 1 ♀, Sanqingsan (28°54΄N 118°04΄E), Yushan, 610 m, 21-VIII-1985, gen. slide No. DYL00897; 1 ♀, Mt. Jiulian (24°33΄N 114°37΄E), 21-VII-2006, coll. Jiasheng Xu & Weichun Li; Liaoning: 4 ♂♂, Laotudingzi, Huanren (41°19΄N 124°53΄E), 7̅ 9-VIII-2009, coll. Weichun Li & Jiayu Liu; 1 ♀, Mt. Baiyun (40°34΄N 123°01΄E), Haicheng, 15-VII-2010, Jiayu Liu & Yanpeng Cai; 1 ♀, Mt. Taibai, Shaanxi, 840 m, 11-VII-2005, coll. Ping You; Shandong: 8 ♂♂, 1 ♀, Mt. Yimeng (35°00΄N 118°40΄E), Linyi, 27̅ 30-VII-2009, coll. Qing Jin; 1 ♂, Beijiushui, Laoshan (36°12΄N 120°03΄E), Qingdao, 2-VIII-2009, coll. Qing Jin; Sichuan: 2 ♂♂, 5 ♀♀, Labahe (30°11΄N 102°26΄E), Tianquan, 1300 m, 29-VII-2004, coll. Yingdang Ren, gen. slide Nos RYD04364 m, RYD04365 f; 4 ♂♂, 4 ♀♀, Caopingcun (30°57΄N 103°19΄E), Wenchuan, 1557 m, 9̅ 11-VII-2014, coll. Kaijian Teng et al.; Tianjin: 2 ♂♂, Mt. Baxian, Jixian, 550 m, 23-VI-2001, coll. Houhun Li; 3 ♂♂, Mt. Baxian, Jixian, 500 m, 08-IV, 20-VII-2004, coll. Houhun Li; 5 ♂♂, 2 ♀♀, Mt. Baxian, Jixian, 560 m, 12̅ 15-VII-2005, coll. Houhun Li, gen. slide No. KDH05090 m; 6 ♂♂, 1 ♀, Mt. Pan (40°05΄N 117°16΄E), Jixian, 170 m, 20-VII-2005, coll. Houhun Li; 1 ♂, 3 ♀♀, Mt. Baxian, Jixian, 500 m, 16-VII, 1-IX-2005, coll. Houhun Li; 1 ♀, Mt. Baxian, Jixian, 300̅ 600 m, 4-VIII-2007, coll. Houhun Li; 1 ♀, Mt. Baxian, Jixian, 600 m, 4-VIII-2007, coll. Bingbing Hu & Mingfeng Cao; 41 ♂♂, 111 ♀♀, Mt. Baxian, Jixian, 550 m, 8, 14, 19, 28-VII, 2, 6, 11, 31-VIII, 13-IX-2009, coll. Bingbing Hu, Jingzhang & Zhipin Liang, gen. slide Nos LJY11085 W, LJY09155 f, LJY09153 f, LJY09154 f, LJY09150 f; 41 ♂♂, 37 ♀♀, Heishuihe, Mt. Baxian, Jixian, 600 m, 4-VI̅ 29-VIII-2010, coll. Yinghui Mou & Shurong Liu; 3 ♀♀, Heishuihe, Mt. Baxian, Jixian, 550 m, 20-IX-2009, coll. Shurong Liu; Zhejiang: 2 ♂♂, Kaishanlaodian, Mt. Tianmu (30°01΄N 117°29΄E), 1140 m, 17-VII-1999, coll. Houhun Li; 1 ♂, Chanyuansi Mt. Tianmu, 350 m, 15-VIII-1999, coll. Houhun Li; 1 ♂, Houshanmen, Mt. Tianmu, 500 m, 16-VIII-1999, coll. Houhun Li; 1 ♂, Mt. Qingliang (30°07΄N 119°02΄E), Lin’an, 800 m, 12-VIII-2005, coll. Yunli Xiao; 2 ♀♀, Mt. Wuyan (27°21΄N 119°56΄E), Taishun, 250̅ 1000 m, 3̅ 4-VIII-2005, coll. Yunli Xiao; 1 ♂, 1 ♀, Mt. Tianmu, Lin’an, 350 m, 9-VIII-2007, coll. Qing Jin; 1 ♂, Sanmuping, Mt. Tianmu, 400 m, 29-VII-2011, coll. Linlin Yang & Na Chen, slide No. LHX14123 W; 2 ♂♂, 2 ♀♀, Tianmucun, Mt. Tianmu, 1-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 5 ♂♂, 5 ♀♀, Laoan, Mt. Tianmu, 555 m, 3̅ 6-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 1 ♀, Sanmuping, Mt. Tianmu, 789 m, 14-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 3 ♀♀, Xiguan, Mt. Tianmu, 17̅ 19-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 1 ♂, Pinggang, Mt. Longxu (30°25΄N 119°33΄E), 754 m, 20-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 2 ♀♀, Qianqingtang, Mt. Qingliang, 1059 m, 29-VII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 1 ♂, 1 ♀, Sanmuping, Mt. Tianmu, 789 m, 9-VIII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 1 ♂, 1 ♀, Laoan, Mt. Tianmu, 555 m, 16-VIII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang; 1 ♂, Longxushancun, Mt. Longxu, 778 m, 20-VIII-2014, coll. Aihui Yin, Xuemei Hu & Qingyun Wang. Diagnosis. This new species can be separated from its congeners by the forewing black from the base to the inner margin of the antemedial line. The genitalia structures are similar to those of D. dichromella, differing in the male genitalia by the triangular uncus gradually narrowed to apex, the clubbed gnathos tapered to a hooked apex; and in the female genitalia by the subtriangular corpus bursae twice as long as the maximum width and the signum situated in the middle of the corpus bursae. In D. dichromella, the uncus is subtriangular in the basal half, sharply narrowed at middle and tubular in the distal half, the gnathos is broadened in distal half and the apex is not hooked in the male genitalia; the elongate bag-shaped corpus bursae is 3.5 times as long as its maximum width, and the signum is situated at the anterior 2/7 of the corpus bursae in the female genitalia. Description. Adult (Fig. 4) with wingspan 21.5̅ 27.5 mm. Head grayish brown. Antenna grayish white, scape about twice as long as wide, male with basal several flagellomeres constricted, incurved, without scale tufts; female flagellum simple. Labial palpus upturned beyond vertex; first segment grayish brown mixed with grayish white; second and third segments about equal length, grayish brown to blackish brown. Maxillary palpus grayish brown mixed with grayish white. Patagium, tegula and thorax yellowish brown to brownish fuscous. Forewing black in basal area, from grayish white gradually to grayish brown in medial area, grayish brown in distal area; antemedial line white, from costal 1/3 obliquely outward to dorsal 2/7; discal spots indistinct, black, fused into a club shape; postmedial line grayish white, sinuate, incurved at M 1 and CuA 2; terminal line black; cilia grayish brown. Hindwing pale grayish brown; cilia brown basally, pale brown distally. Foreleg blackish brown with scattered white scales, midleg grayish brown mixed with white, hindleg grayish white; tarsi black except white at apex of each tarsomere. Abdomen with dorsal surface blackish brown except yellow at posterior margin of each segment, ventral surface yellowish brown. Male genitalia (Fig. 15). Uncus triangular, longer than width at proximal base, gradually narrowed to rounded apex. Gnathos about 2/3 length of uncus, clubbed, tapered to hooked and slightly bifurcate apex, apico-dorsal portion minutely serrated. Transtilla produced in V shape posteriorly; postero-lateral lobe three times length of its medial width, tapered from base to narrowly rounded apex, forming an acute angle of about 45°. Valva three times as long as wide, narrowed at base, gradually broadened to middle, then narrowed to rounded apex; clasper an indistinct mastoid process at basal 1/7, bearing short setae; costa about 5/6 length of valva; sacculus sinuate, clubbed, 2/7 length of valva. Vinculum U-shaped posteriorly, 1.3 times as long as wide, concave on lateral margins at anterior 1/3, rounded anteriorly. Juxta broad V-shaped; lateral lobes about 4/5 length of gnathos, clubbed, slightly widened to rounded apex, bearing sparse setae. Phallus slightly longer than valva, with sclerotized crimples, crimples granulate in distal half. Female genitalia (Fig. 26). Papillae anales large and wide, with sparse setae, rounded at apex. Apophyses anteriores slightly longer and stouter than apophyses posteriores. Eighth segment about 2.5 times as wide as long. Antrum a narrow transverse band. Ductus bursae short, about 1/4 length of corpus bursae, gradually broadened, posterior 2/5 smooth, anterior 3/5 granulate on inner wall. Corpus bursae not easily differentiated from ductus bursae, subtriangular, twice as long as maximum width, granulate on inner wall in posterior 1/5; signum a rounded, invaginated cup consisting of concentrically arranged granules, situated at middle of corpus bursae. Ductus seminalis originating from posterior 1/5 of corpus bursae. Distribution. China (Anhui, Beijing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hebei, Henan, Hubei, Hunan, Jiangxi, Liaoning, Shaanxi, Shandong, Sichuan, Tianjin, Zhejiang). Etymology. The specific name is derived from the Latin prefix basi -, meaning basal, and the Latin word niger, meaning black, referring to the forewing black in the basal area.

Dusungwua similiquadrangula Ren & Li, sp. nov. (Figs 1, 12, 23, 33) Type material. Holotype. [CHINA] ♂, Rare Botanical Garden, Ruili (24°00’N 97°50’E), Yunnan, 1000 m, 6-VIII- 2005, coll. Yingdang Ren, gen. slide No. RYD04729. Paratypes. [CHINA] 2 ♂♂, 2 ♀♀, same data as for holotype, gen. slide Nos LHX 14117 m, RYD04730 f. Diagnosis. This new species resembles D. quadrangula (Du, Sung & Wu, 2005). It differs in the transtilla produced in U shape posteriorly and the postero-lateral lobe is three times as long as wide, and in the sub-rectangular valva that has a triangular clasper in the male genitalia. In D. quadrangula, the transtilla is produced in a V shape and postero-lateral lobe is twice as long as wide, and the triangular valva has a short fingerlike clasper in the male genitalia. In the female genitalia, the antrum is dome-like and the ductus bursae is broader in the new species; while the antrum is rectangular and the ductus bursae is narrower in the latter species. Description. Adult (Fig. 12) with wingspan 20.0̅ 24.5 mm. Head grayish brown, occiput creamy with a few grayish brow scales. Antenna brown, scape about 1.5 times as long as wide; male flagellum simple, neither constricted nor incurved basally. Labial palpus upturned just beyond vertex; first segment grayish brown mixed with grayish white, second and third segments brown; second segment 1.2 times as long as third segment. Maxillary palpus brown. Patagium, tegula and thorax grayish brown, with scattered grayish white scales. Forewing gray mottled grayish white; antemedial line at basal 1/3, indistinct, grayish white, arched, bordered by blackish brown patch near costa on its outer margin and near dorsum on its inner margin respectively; discal spots black, ovate, distinct, separated; postmedial line grayish white, sinuate, incurved at M 1 and near CuA 2, edged with black; terminal line black; cilia grayish brown. Hindwing and cilia grayish brown. Foreleg black, midleg grayish brown, hindleg grayish white. Abdomen with dorsal surface grayish brown, with grayish white at end of each segment, ventral surface grayish white basally and blackish brown distally on each segment. Male genitalia (Fig. 23). Uncus somewhat quadrate, about same length as width at proximal base, apex slightly concave at middle. Gnathos clubbed, gradually narrowed to weakly bifurcate apex, about 3/5 length of uncus. Transtilla produced in U shape posteriorly, postero-lateral lobe fingerlike, three times as long as wide, uniform in width. Valva four times as long as wide, sub-rectangular, slightly narrower distally; clasper triangular, sinuate along outer margin, with sparse setae; costa clubbed, about 4/5 length of valva, gradually narrowed to apex; sacculus 1/3 length of valva. Juxta V-shaped; lateral lobe short, broad, bean-shaped, twice as long as wide, about 4/5 length of gnathos. Vinculum U-shaped, same length as maximum width. Phallus cylindrical, about same length as valva, with fine, sclerotized fascicle-shaped crimples and a sclerotized sword-like structure bearing dense spinules. Female genitalia (Fig. 33). Papillae anales large and wide, shield-like, rounded at apex, with dense setae. Apophyses posteriores expanded at base, slightly longer than apophyses anteriores. Eighth segment broad and short. Antrum dome-like. Ductus bursae membranous, smooth on inner wall, about half length of corpus bursae. Corpus bursae membranous, oblong, 1.7 times as long as maximum width, granulated on inner wall; signum a rounded, invaginated cup consisting of concentrically arranged granules, situated at posterior 2/5 of corpus bursae. Ductus seminalis from junction between ductus bursae and corpus bursae. Distribution. China (Yunnan). Etymology. The specific name is derived from the Latin prefix simili-, meaning similar, and the specific name of another species, quadrangula, referring to their similarities., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on pages 354-355, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857, {"references":["Du, Y. L., Sung, S. M. & Wu, C. S. (2005) A new genus in the subfamily Phycitinae (Lepidoptera: Pyralidae) from China. Annales Zoologici, 55 (1), 99 - 105."]}

Dusungwua antennalveata Ren & Li, sp. nov. (Figs 1, 3, 14, 25) Type material. Holotype. [CHINA] ♂, Dashahe (28°52΄N 107°36΄E), Daozhen, Guizhou, 1420 m, 22-V-2004, coll. Shulian Hao, gen. slide No. RYD04331. Paratypes. [CHINA] Gansu: 1 ♂, Douba Forest Yard (33°15΄N 105°29΄E), Kangxian, 1200 m, 04-VI-1995, coll. Aishaer Mamati; Guizhou: 1 ♀, Huguosi, Fanjingshan (27°55΄N 108°04΄E), 1300 m, 03-VII-2001, coll. Houhun Li & Xinpu Wang; 1 ♂, Dashahe (28°52΄N 107°37΄E), Daozhen, 1450 m, 23-V- 2004, coll. Shulian Hao; 1 ♀, same data as for holotype, gen. slide No. RYD04332; 2 ♂♂, Kuankuoshui (28°14΄N 107°01΄E), Suiyang, 840 m, 9-VI-2010, coll. Linlin Yang, gen. slide No. LJY10413; Henan: 7 ♂♂, Baotianman (33°31΄N 111°56΄E), Neixiang, 1200 m, 20-V̅ 4-VI-2006, coll. Hui Zhen & Denghui Kuang; 13 ♂♂, 2 ♀♀, Guanshan (35°30΄N 113°35΄E), Huixian, 550 m, 25-VII-2006, coll. Hui Zhen & Denghui Kuang; 43 ♂♂, 10 ♀♀, Wangwushan (35°8΄N 112°17΄E), Jiyuan, 800 m, 28̅ 30-VII-2006, coll. Hui Zhen & Denghui Kuang, gen. slide Nos. LJY 11362 m, LHX14127 f; 8 ♂♂, 2 ♀♀, Huaguoshan (34°19΄N 111°53΄E), Yiyang, 1000 m, 3-VIII-2006, coll. Hui Zhen & Denghui Kuang; 1 ♂, Dakuaidi (33°45΄N 110°59΄E), Lushi County, 1100 m, 6-VIII-2006, coll. Hui Zhen & Denghui Kuang; Hubei: 3 ♂♂, 2 ♀♀, Songbaizhen (31°45΄N 110°39΄E), Shennongjia, 1200̅ 1400 m, 15̅ 17-VII- 2003, coll. Shulian Hao; 14 ♂♂, 6 ♀♀, Bajiaomiao (31°46΄N 110°34΄E), Shennongjia, 1100 m, 18-VII-2003, coll. Shulian Hao, gen. slide Nos. RYD04806 m, RYD04807 f; Shaanxi: 7 ♂♂, 6 ♀♀, Hualongshan (32°01΄N 109°22΄E), Ankang, 800 m, 2̅ 5-VII-2003, coll. Haili Yu, gen. slide Nos. RYD04557 m, RYD04558 f; 5 ♂♂, 3 ♀♀, Xunangba (33°33΄N 108°33΄E), Ningshan, 1360 m, 26-VI̅ 3-VII-2007, coll. Haili Yu; Sichuan: 2 ♂♂, Baoxing (30°22΄N 102°49΄E), 1100 m, 1-VIII-2004, coll. Yingdang Ren, gen. slide No. RYD04440; 1 ♀, Yonghong, Mabian (28°33΄N 103°25΄E), 1500 m, 23-VII-2004, coll. Yingdang Ren, gen. slide No. RYD04547. Diagnosis. This new species can be distinguished from its congeners by the male flagellum having a deep notch at base containing a cluster of smaller scales that are arranged in helix shape. The male genitalia resemble those of D. paradichromella, but differ in the transtilla being ovate at connection and the lateral lobes of the juxta truncate at apex. In D. paradichromella, the transtilla is band-like at connection, and the juxta lateral lobe is rounded. Description. Adult (Fig. 3) with wingspan 17.5̅27.0 mm. Head yellowish white to golden yellow. Antenna brown to blackish brown, male scape about 2.5 times as long as wide, basal several flagellomeres incurved, forming a deep notch containing a cluster of smaller scales arranged in helix shape; female scape shorter, flagellum simple. Labial palpus upturned just beyond vertex, grayish brown to blackish brown; third segment slightly shorter than second segment. Maxillary palpus blackish brown. Patagium, tegula and thorax grayish brown to blackish brown. Forewing grayish brown to blackish brown, individual scales grayish white-tipped; antemedial line ill-defined, grayish white, from costal 1/4 sinuate to dorsal 2/5, incurved at CuP, bordered by ill-defined sub-triangular black patch on its outer margin near costa and on its inner margin near dorsum; discal spots black, separated; postmedial line grayish white, sinuate, incurved at M 1 and near CuA 2, edged with black; terminal line black; cilia grayish brown to blackish brown. Hindwing pale grayish brown to fuscous brown; cilia brown. Legs grayish brown to blackish brown, midleg and hindleg grayish white at apex of each tarsomere. Abdomen yellowish brown to blackish brown, edged with grayish yellow on posterior margin of each segment, sometimes light-colored on ventral surface. Male genitalia (Fig. 14). Uncus triangular, longer than width at proximal base, rounded at apex. Gnathos clubbed, about half length of uncus. Transtilla produced in V shape posteriorly; postero-lateral lobe three times length of its medial width, tapered from base to narrowly rounded apex, forming an acute angle of about 45°. Valva 2.5 times as long as wide, gradually broadened toward rounded apex; clasper an indistinct mastoid process at basal 1/5, bearing short setae; costa about 4/5 length of valva, slightly narrowed distally; sacculus 2/7 length of valva, basal 1/3 subtriangular, smooth, distal 2/3 spatulate, with long setae. Juxta V-shaped, lateral lobes slightly shorter than gnathos, digitate, truncate at apex, bearing sparse setae in distal half. Vinculum U-shaped, length same as maximum width. Phallus about 1.2̅1.4 times length of valva, with sclerotized crimples and a bundle of spines in distal half. Female genitalia (Fig. 25). Papillae anales large and wide, with sparse setae, rounded at apex. Apophyses posteriores and anteriores short, about equal length. Eighth segment broad and short. Antrum lip-shaped. Ductus bursae membranous, uniform in width, about 1.5 times length of corpus bursae, granulate on inner wall except near antrum. Corpus bursae ovate, 1.5 times as long as wide, granulated in posterior half; signum a rounded, invaginated cup consisting of concentrically arranged granules, situated at posterior 1/3 of corpus bursae. Ductus seminalis originating from posterior 2/5 of corpus bursae. Distribution. China (Gansu, Guizhou, Henan, Hubei, Shaanxi, Sichuan). Etymology. The specific name is derived from the Latin prefix antenn -, meaning antenna, and the Latin word alveatus, meaning notched, referring to the deep notch at base of flagellum in male., Published as part of Ren, Yingdang, Yang, Linlin, Liu, Hongxia & Li, Houhun, 2020, Taxonomic review of the genus Dusungwua Kemal, Kizildağ & Koçak, 2020 (Lepidoptera: Pyralidae), with descriptions of six new species and propositions of synonyms, pp. 341-365 in Zootaxa 4894 (3) on pages 343-346, DOI: 10.11646/zootaxa.4894.3.3, http://zenodo.org/record/4315857

1. Thiallela dolokensis Roesler et K��ppers, 1981 Thiallela dolokensis ROeSLeR et K��PPeRS, 1981: 75; ROeSLeR, 1983: 96. TL: INDONeSIA (SUMATRA). TD: LNK. DISTRIbUTION: INDONeSIA (SUMATRA)., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on page 438, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645, {"references":["Roesler, R. U. & Kuppers, P. V. (1981) Beitrage zur Kenntnis der Insektenfauna Sumatras. Teil 9. Die Phycitinae (Lepidoptera: Pyralidae) von Sumatra; Taxonomie Teil B, Okologie und Geobiologie. Beitrage zur Naturkundlichen Forschung in Sudwestdeutschland, 4, 1 - 282, 4 text abbildung, 42 tafel.","Roesler, R. U. (1983) Die Phycitinae von Sumatra (Lepidoptera: Pyralidae). Heterocera Sumatrana, Keltern, 3, 1 - 136, pls. 1 - 69."]}

2. Thiallela eduardi Roesler et K��ppers, 1981 Thiallela eduardi ROeSLeR et K��PPeRS, 1981: 73; ROeSLeR, 1983: 95. TL: INDONeSIA (SUMATRA). TD: LNK. DISTRIbUTION: INDONeSIA (SUMATRA)., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on page 438, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645, {"references":["Roesler, R. U. & Kuppers, P. V. (1981) Beitrage zur Kenntnis der Insektenfauna Sumatras. Teil 9. Die Phycitinae (Lepidoptera: Pyralidae) von Sumatra; Taxonomie Teil B, Okologie und Geobiologie. Beitrage zur Naturkundlichen Forschung in Sudwestdeutschland, 4, 1 - 282, 4 text abbildung, 42 tafel.","Roesler, R. U. (1983) Die Phycitinae von Sumatra (Lepidoptera: Pyralidae). Heterocera Sumatrana, Keltern, 3, 1 - 136, pls. 1 - 69."]}

8. Thiallela naevilla Li et Ren, 2006 Thiallela naevilla LI et ReN, 2006: 325. TL: CHINA (HUbeI, SICHUAN). TD: NKUM. DISTRIbUTION: CHINA (HUbeI, SICHUAN)., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on page 439, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645

3. Thiallela endochralis Hampson, 1908 Thiallela endochralis HAMPSON, 1908: 266. TL: SRI LANKA. TD: BMNH. DISTRIbUTION: SRI LANKA., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on page 438, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645

10. Thiallela rhodoptila Meyrick, 1932 Thiallela rhodoptila MeYRICK, 1932: 239. TL: FIjI. TD: BMNH. DISTRIbUTION: FIjI., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on page 439, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645, {"references":["Meyrick, E. (1932) Exotic Microlepidoptera. Vol. 4. Thornhanger, Marlborough, Wilts, 32 pp. [pp. 225 - 256]"]}

Thiallela celadontis sp. nov. (FIGS. 1, 4, 11, 15) Diagnosis: THIS NeW SPeCIeS IS SIMILAR TO T. ligeralis SUPeRFICIALLY, bUT CAN be DISTINGUISHeD FROM THe LATTeR bY THe GeNITALIA STRUCTUReS: IN THe MALe, THe CLASPeR eXTeNDS TO NeAR eND OF SACCULUS, HAS A FINGeRLIKe FLAP AT bASe, AND THe AeDeAGUS HAS FIVe CORNUTI; IN THe FeMALe, THe ANTRUM IS ReCTANGULAR, AND THe SIGNUM IS LARGe, STRAIGHT, CLUbbeD. IN T. ligeralis, THe CLASPeR eXTeNDS beYOND eND OF SACCULUS, HAS A TRIANGULAR FLAP AT bASe, AND THe AeDeAGUS HAS TWO CORNUTI IN THe MALe GeNITALIA; THe ANTRUM IS TRAPeZOIDAL, AND THe SIGNUM IS SMALLeR AND beNT IN THe FeMALe GeNITALIA. Description. WINGSPAN 20.0��23.0 MM (N = 19) (FIG. 4). VeRTeX LIGHT YeLLOWISH bROWN, SCALeS YeLLOWISH WHITe- TIPPeD; FRONS YeLLOWISH WHITe. ANTeNNA YeLLOWISH WHITe ALTeRNATeD WITH bROWNISH FUSCOUS, MALe FLAGeLLUM SIMPLe, bASAL SeGMeNTS NOT DILATeD, WITHOUT NOTCH, CILIA AbOUT 2/3 LeNGTH OF DIAMeTeR OF FLAGeLLUM. LAbIAL PALPUS WITH FIRST SeGMeNT LIGHT YeLLOWISH WHITe, SeCOND SeGMeNT DARK bROWNISH FUSCOUS, YeLLOWISH WHITe AT eND, THIRD SeGMeNT AbOUT HALF LeNGTH OF SeCOND SeGMeNT, YeLLOWISH WHITe, MIXeD WITH bROWNISH FUSCOUS AT bASe. OCCIPUT, PATAGIUM, TeGULA AND THORAX GRAYISH GReeN OR GRAYISH bROWN, SCALeS GRAYISH WHITe-TIPPeD. FOReWING WITH bASAL MAjORITY ReGION GRAYISH GReeN, SCATTeReD A FeW bLACK SCALeS, WITH AN DISTINCT, LARGe, TRIANGULAR, RUSTY bROWN PATCH INWARDLY SUb- ADjACeNT TO ANTeMeDIAL LINe; CeNTRAL ReGION GRAYISH GReeN NeAR ANTeMeDIAL LINe, GRAYISH WHITe NeAR POSTMeDIAL LINe, SUFFUSeD LOTS OF RUSTY bROWN SCALeS, DIFFUSeD bLACK AROUND VeINS beLOW DISCAL SPOTS; DISTAL ReGION GRAYISH bROWN, DIFFUSeD MORe WHITe SCALeS AT APeX, bROWNISH FUSCOUS ALONG VeINS; ANTeMeDIAL LINe AT bASAL 2/5 OF FOReWING, SINUATe, bLACK, INWARDLY eDGeD WITH AN INDISTINCT GRAYISH WHITe LINe; POSTMeDIAL LINe AT DISTAL 1/5 OF FOReWING, DISTINCT, ARCUATe, SLIGHTLY CONCAVe AT M1 AND CUA 2, bLACK, eDGeD OUTWARDLY WITH AN ObSCURe WHITe LINe, AND A bROWNISH FUSCOUS SHADe ADjACeNT TO WHITe LINe; DISCAL SPOTS NARROWLY FUSeD, ObSCURe, bLACK; TeRMINAL LINe bLACK; CILIA LIGHT YeLLOW. HINDWING GRAY, FUSCOUS ALONG VeINS AND MARGINS; CILIA LIGHT bROWN. FOReLeG WHITe DORSALLY, bLACK VeNTRALLY, TARSOMeReS YeLLOWISH WHITe AT eND; MID AND HIND LeGS WHITe WITH SCATTeReD bLACK SCALeS ON COXA, FeMUR AND TIbIA, TARSUS MOTTLeD WITH bLACKISH bROWN. Male genitalia (FIG. 11). UNCUS HOODLIKe, bROADeR THAN LONG, ROUNDeD AT APeX, SeTOSe IN DISTAL HALF ON DORSAL SURFACe. APICAL PROCeSS OF GNATHOS TAPeReD, WITH A SMALL HOOK DISTALLY, AbOUT 2/3 LeNGTH OF UNCUS. TRANSTILLA WITH TeRMINAL PART FUSeD INTO A SCLeROTIZeD ARCeD PLATe, WITH A PAIR OF STRONGLY SCLeROTIZeD, INWARDLY beNT, AURIFORM POSTeROLATeRAL PROCeSSeS. VALVA GRADUALLY bROADeNeD TO ROUNDeD APeX; COSTA A NARROW bAND, eXTeNDING TO eND OF VALVA, NeARLY eqUAL WIDTH eXCePT SLIGHTLY TAPeReD DISTALLY; SACCULUS AbOUT HALF LeNGTH OF VALVA, CUNIFORM, bROAD AT bASe, TAPeReD TO A DISSOCIATeD, POINTeD APeX; CLASPeR A NARROWeD, ARCHeD ARM, ARISING FROM NeAR bASe OF COSTA ObLIqUe OUTWARDLY TO eND OF SACCULUS, WITH A FINGeRLIKe FLAP AT bASe, beARING SPAReS SeTAe. JUXTA U-SHAPeD, LATeRAL LObeS WeLL DeVeLOPeD, AS LONG AS APICAL PROCeSS OF GNATHOS, FINGeRLIKe, beARING A FeW SeTAe. VINCULUM U-SHAPeD, MODeRATeLY LONG WITH TRANSVeRSe POSTeRIOR MARGIN. AeDeAGUS STOUT, GRADUALLY bROADeNeD TO APeX, SLIGHTLY SHORTeR THAN VALVA; WITH ONe TAPeReD PLATe NeAR MIDDLe, FOUR SHARP, CONICAL CORNUTI NeAR APeX, AND ONe LARGe, CLUbbeD CORNUTA beARING SMALL THORNS AT APeX. EIGHTH AbDOMINAL TeRGITe FAN-SHAPeD; eIGHTH AbDOMINAL STeRNITe SMALL, TRIANGULAR, ASSOCIATeD WITH A PAIR OF CULCITA. Female genitalia (FIG. 15). PAPILLAe ANALIS bROAD, ReCTANGULAR IN bASAL HALF, SeMICIRCULAR IN DISTAL HALF, WITH VeSTURe OF THIN HAIRS. EIGHTH AbDOMINAL SeGMeNT SHORT, 1/3 AS LONG AS WIDe; A TRIANGULAR SCLeRITe POSTeRIOR TO eIGHTH AbDOMINAL SeGMeNT. ANTRUM WeLL DeVeLOPeD, ReCTANGULAR, FOUR TIMeS AS WIDe AS LONG. DUCTUS bURSAe VeRY SHORT, AbOUT 1.2 TIMeS LeNGTH OF ANTRUM, PUNCTATe ON INNeR WALL AT jUNCTION WITH CORPUS bURSAe. CORPUS bURSAe MeMbRANOUS, OVATe, TWICe AS LONG AS WIDe, DeCORATeD WITH COARSe SCObINATIONS POSTeRIORLY TO FORM A SOMeWHAT SOLe-SHAPeD AReA; SIGNUM A STRONG, SOMeWHAT STRAIGHT, TAPeReD, CLUbbeD HORN ON A bROAD, FORKeD bASe, SITUATeD AT AbOUT MIDDLe OF CORPUS bURSAe. DUCTUS SeMINALIS INCePTeD POSTeRIOR 1/3 OF CORPUS bURSAe, SAMe SIDe WITH SIGNUM. Type material. Holotype ♂, China: NATIONAL FOReST PARK, TAIYANGHe (22��40'N 101��01'E), PU���eR, YUNNAN, 1450 M, 3.VII.2015, COLL. KAIjIAN TeNG (NKUM). Paratypes: China: Yunnan: 9 ♂♂, 1 ♀, SAMe DATA AS FOR HOLOTYPe, GeN. SLIDe NO. YLL15178M, YLL1590M, YLL 15191 F (NKUM); 5 ♂♂, SAMe PLACe AS FOR HOLOTYPe, 8.VII.2013, COLL. LINLIN YANG, GeN. SLIDe NO. RYD2013100, RYD2014218 (HAASM); 2 ♂♂, SAMe PLACe AS FOR HOLOTYPe, 31.III. 2014, COLL. ZHeNGUO ZHANG, GeN. SLIDe NOS YLL15181, YLL15183 (NKUM); 1 ♂, NANKANG (24��49'N 98��47'E), BAOSHAN, 2009 M, 20.VII.2015, COLL. KAIjIAN TeNG & XIA BAI, GeN. SLIDe NO. YLL15192 (NKUM). Distribution. CHINA (YUNNAN). Etymology. THe SPeCIFIC NAMe IS DeRIVeD FROM THe LATIN WORD celadontis, MeANING GRAYISH GReeN, ReFeRRING TO THe FOReWING IS GRAYISH GReeN IN bASAL MAjORITY ReGION., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on pages 447-448, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645

Thiallela ligeralis Walker, 1863 (FIGS. 1, 3, 6��8, 10, 14) Myelois ligeralis Walker, 1863: 37. TL: Sri Lanka. TD: BMNH. Thiallela signifera Walker, 1863: 120. TL: Borneo. TD: BMNH. As synonym of Thiallela ligeralis (Walker) by Roesler & K��ppers, 1981: 72. Luconia pallidobasella Ragonot, 1888: 7; Ragonot, 1893: 1. TL: Borneo. TD: BMNH. As synonym of Thiallela signifera Walker by Hampson, 1896: 106. As synonym of Thiallela ligeralis (Walker) by Roesler & K��ppers, 1981: 72. Acrobasis ligeralis: Ragonot, 1893: 95. Rhodophaea ligeralis: Hampson, 1896: 99. Thiallela ligeralis: Roesler & K��ppers 1981: 72; Roesler, 1983: 95. Redescription. WINGSPAN 20.5��24.0 MM (N = 21) (FIG. 3). VeRTeX AND FRONS SILVeR-WHITe TO YeLLOWISH WHITe. ANTeNNA YeLLOWISH WHITe ALTeRNATeD WITH bROWN, MALe FLAGeLLUM SIMPLe, bASAL SeGMeNTS NOT DILATeD, WITHOUT NOTCH, WITH CILIA AbOUT SAMe LeNGTH AS DIAMeTeR OF FLAGeLLUM. LAbIAL PALPUS ReDDISH bROWN MIXeD WITH YeLLOWISH WHITe; THIRD SeGMeNT AbOUT HALF LeNGTH OF SeCOND SeGMeNT. OCCIPUT, PATAGIUM, TeGULA AND THORAX PALe WHITe TINGeD WITH GRAYISH GReeN, WITH SPARSe ReDDISH bROWN AND DARK bROWN SCALeS. FOReWING WITH bASAL MAjORITY ReGION PALe GRAYISH bROWN TO GRAYISH GReeN, WITH A FeW bLACK SCALeS NeAR bASe, DIFFUSeD LOTS OF GRAYISH WHITe SCALeS NeAR ANTeMeDIAL LINe, WITH AN ObSCURe, ARCHeD, SUbTRIANGULAR, RUSTY bROWN PATCH INWARDLY ADjACeNT TO ANTeMeDIAL LINe; CeNTRAL ReGION YeLLOWISH bROWN TO GRAYISH GReeN NeAR ANTeMeDIAL LINe, GRAYISH WHITe SUFFUSeD WITH ReDDISH bROWN NeAR POSTMeDIAL LINe; DISTAL ReGION GRAYISH WHITe, SUFFUSeD WITH ReDDISH bROWN, bROWNISH FUSCOUS ALONG VeINS; ANTeMeDIAL LINe beYOND MIDDLe OF FOReWING, DISTINCT, ARCHeD, bLACK, INWARDLY eDGeD WITH AN INDISTINCT GRAYISH WHITe LINe; POSTMeDIAL LINe AT DISTAL 1 /6 OF FOReWING, DISTINCT, ARCUATe, SLIGHTLY CONCAVe AT M1 AND CUA2, bLACK, eDGeD OUTWARDLY WITH AN ObSCURe WHITe LINe, AND A RUSTY bROWN SHADe ADjACeNT TO WHITe LINe FROM M1 TO DORSUM; DISCAL SPOTS FUSeD, CReSCeNT, ReDDISH bROWN; TeRMINAL LINe RUSTY bROWN; CILIA bROWN. HINDWING SeMITRANSPAReNT, PALe GRAY, FUSCOUS ALONG VeINS AND TeRMeN; CILIA LIGHT bROWN. LeGS GRAYISH WHITe MIXeD WITH RUSTY ReD, TARSUS bLACKISH bROWN AT bASe, WHITe AT eND. Male genitalia (FIG. 10). UNCUS HOODLIKe, SLIGHTLY bROADeR THAN LONG, ROUNDeD AT APeX, SeTOSe IN DISTAL HALF ON DORSAL SURFACe. APICAL PROCeSS OF GNATHOS AbOUT 3/4 LeNGTH OF UNCUS, bROAD AT bASe, TAPeReD TO 2/3, SLeNDeR AND HOOKeD IN DISTAL 1/3. TRANSTILLA WITH TeRMINAL PART FUSeD INTO A WeLL SCLeROTIZeD SADDLe-SHAPeD PLATe, WITH A PAIR OF ROUNDeD POSTeROLATeRAL PROCeSSeS. VALVA GRADUALLY bROADeNeD TO ROUNDeD APeX; COSTA A NARROWeD bAND, eXTeNDING TO eND OF VALVA, NeARLY eqUAL WIDTH eXCePT SLIGHTLY TAPeReD DISTALLY; SACCULUS 3/5 LeNGTH OF VALVA, TRIANGULAR, bROAD AT bASe, TAPeReD TO A DISSOCIATeD, POINTeD APeX; CLASPeR NARROW, eLONGATe, SLIGHTLY SINUATe, ARISING FROM bASe OF COSTA ObLIqUe OUTWARDLY TO DISTAL 1/3 OF POSTeRIOR MARGIN, beYOND eND OF SACCULUS, WITH A TRIANGULAR FLAP AT bASe, beARING SPAReS SeTAe. JUXTA V-SHAPeD, LATeRAL LObeS WeLL DeVeLOPeD, AbOUT 3/4 LeNGTH OF APICAL PROCeSS OF GNATHOS, FINGeRLIKe, SLIGHTLY TWISTeD, SWOLLeN bASALLY, WITH A FeW SeTAe APICALLY. VINCULUM V-SHAPeD, LeNGTH SLIGHTLY SHORTeR THAN TRANSVeRSe POSTeRIOR MARGIN. AeDeAGUS CYLINDRICAL, 3/4 LeNGTH OF VALVA; WITH TWO CLUbbeD CORNUTI: ONe AbOUT HALF LeNGTH OF AeDeAGUS, ITS DISTAL PART SLIGHTLY beNT, PROTRUDeD, FORMING TWO MINUTe TINeS, ONe SLIGHTLY SHORTeR, DeNTATe ON DISTAL MARGIN. EIGHTH AbDOMINAL TeRGITe FAN-SHAPeD; eIGHTH AbDOMINAL STeRNITe TRIANGULAR, ASSOCIATeD WITH A PAIR OF CULCITA. Female genitalia (FIG. 14). PAPILLAe ANALIS SeMIeLLIPTICAL, WITH VeSTURe OF LONG, THIN HAIRS. APOPHYSIS POSTeRIORIS SLIGHT SLeNDeR AND SHORTeR THAN ANTeRIORIS. EIGHTH AbDOMINAL SeGMeNT SHORT AND bROAD, TeRGITe V-SHAPeD CONCAVe ON POSTeRIOR MARGIN; A LIP-LIKe SCLeRITe POSTeRIOR TO eIGHTH AbDOMINAL SeGMeNT. ANTRUM WeLL DeVeLOPeD, TRAPeZOIDAL, POSTeRIOR MARGIN AbOUT 1.5 TIMeS AS WIDe AS ANTeRIOR MARGIN, WITH A PAIR OF CALAbASH-SHAPeD HeAVILY SCLeROTIZeD AReA ON LATeRAL SIDeS. DUCTUS bURSAe MeMbRANOUS, AbOUT SAMe LeNGTH AS ANTRUM, PUNCTATe ON INNeR WALL. CORPUS bURSAe MeMbRANOUS, OVATe, 1.6 TIMeS AS LONG AS WIDe, WITH A qUADRATe PLATe NeAR jUNCTION WITH DUCTUS bURSAe, AND A qUADRATe AReA DeCORATeD WITH COARSe SCObINATIONS; SIGNUM A STRONG beNDING HORN ON A bROAD bASe, SITUATeD AT AbOUT MIDDLe OF DUCTUS bURSAe, PUNCTATe AROUND. DUCTUS SeMINALIS INCePTeD POSTeRIOR MARGIN OF CORPUS bURSAe, SAMe SIDe WITH SIGNUM. Material examined. CHINA: Hainan: 1 ♀, WUZHISHAN (18��46'N 109��31'E), 740 M, 14.IV.2009, COLL. QING JIN & BINGbING HU (NKUM); 1 ♂, MT. DIAOLUO (18��39'N 109��54'E), LINGSHUI, 980 M, 24.IV.2014, COLL. TeNGTeNG LIU, WeI GUAN & XUeMeI HU, GeN. SLIDe NO. LHX14112 (NKUM); 1 ♂, MT. JIANFeNG (18��43'N 108��53'E), LeDONG, 1050 M, 27.IV.2014, COLL. TeNGTeNG LIU, WeI GUAN & XUeMeI HU, GeN. SLIDe NO. RYD2014211 (NKUM); 4 ♂♂, 2 ♀♀, MT. JIANFeNG, 960 M, 13-15.IV.2013, COLL. YINGDANG ReN & XIAOGUANG LIU, GeN. SLIDe NOS RYD014212M, RYD2014213f (HAASM); 4 ♂♂, 3 ♀♀, MT. DIAOLUO, 900 M, 9.IV.2013, COLL. YINGDANG ReN & XIAOGUANG LIU, GeN. SLIDe NOS RYD2014214M, RYD2014215F (HAASM). Yunnan: 1 ♀, SIMAO (22��48'N 100��58'E), 1320 M, 24.IV.1995, COLL. GUANGYUN YAN, GeN. SLIDe NO. RYD04739 (NKUM); 1 ♀, RARe BOTANICAL GARDeN (24��00��N 97��51��E), RUILI, 1000 M, 06.VIII.2005, COLL. YINGDANG ReN, GeN. SLIDe NO. LHX14111 (NKUM); 2 ♂♂, TAIYANGHe (22��40'N 101��01'E), 1450 M, 18.V, 11.VI.2014, COLL. ZHeNGUO ZHANG, GeN. SLIDe NO. YLL15180 (NKUM); 1 ♂, NATIONAL FOReST PARK, TAIYANGHe, 1450 M, 6.VII.2015, COLL. KAIjIAN TeNG, GeN. SLIDe NO. YLL15179 (NKUM). Distribution: CHINA (HAINAN, YUNNAN), ASSAM, SRI LANKA, BURMA, MALAYSIA, BORNeO, SUMATRA, NeW GUINeA, AUSTRALIA, FIjI. Notes. Thiallela ligeralis IS ReCORDeD FOR THe FIRST TIMe FROM CHINA. We CHeCKeD TYPeS OF ligeralis (FIG. 6), signifera (FIG. 7) AND pallidobasella (FIG. 8) DePOSITeD IN BMNH UNDeR THe HeLP OF MR. DAVID Lee. DeSPITe THe SLIGHTLY DAMAGeD WING PATTeRN, We CAN STILL ReCOGNIZe THAT THeSe SPeCIMeNS RePReSeNT THe SAMe SPeCIeS, SO We AGRee WITH ROeSLeR & K��PPeRS���S VIeW (1981) THAT signifera AND pallidobasella ARe SYNONYMS OF ligeralis. THe CHINeSe SPeCIMeNS ARe DARKeR IN COLOR THAN THeSe TYPeS, AND SUFFUSeD WITH MORe GRAYISH GReeN SCALeS IN bASAL HALF, bUT THe GeNITALIA CHARACTeRS ARe IDeNTICAL WITH T. ligeralis STATeD AND DRAWN bY ROeSLeR & K��PPeRS (1981, FIGS. 55.120)., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on pages 441-447, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645, {"references":["Walker, F. (1863) Crambites & Tortricites. In: List of Specimens of Lepidopterous Insects in the Collection of the British Museum. Vol. 27. Printed by order of the Trutees, London, pp. 1 - 286.","Roesler, R. U. & Kuppers, P. V. (1981) Beitrage zur Kenntnis der Insektenfauna Sumatras. Teil 9. Die Phycitinae (Lepidoptera: Pyralidae) von Sumatra; Taxonomie Teil B, Okologie und Geobiologie. Beitrage zur Naturkundlichen Forschung in Sudwestdeutschland, 4, 1 - 282, 4 text abbildung, 42 tafel.","Ragonot, E. L. (1888) Nouveaux genres et especes de Phycitidae & Galleriidae. 52 pp. Paris.","Ragonot, E. L. (1893) Monographie des Phycitinae et des Galleriinae. In: Romanoff, N. M. (Ed.), Memoires sur les Lepidopteres VII 7. St. Petersburg. pp. i - lvi, 1 - 658, pls 1 - 23.","Roesler, R. U. (1983) Die Phycitinae von Sumatra (Lepidoptera: Pyralidae). Heterocera Sumatrana, Keltern, 3, 1 - 136, pls. 1 - 69."]}

Thiallela celadontis sp. nov. (FIGS. 1, 4, 11, 15) Diagnosis: THIS NeW SPeCIeS IS SIMILAR TO T. ligeralis SUPeRFICIALLY, bUT CAN be DISTINGUISHeD FROM THe LATTeR bY THe GeNITALIA STRUCTUReS: IN THe MALe, THe CLASPeR eXTeNDS TO NeAR eND OF SACCULUS, HAS A FINGeRLIKe FLAP AT bASe, AND THe AeDeAGUS HAS FIVe CORNUTI; IN THe FeMALe, THe ANTRUM IS ReCTANGULAR, AND THe SIGNUM IS LARGe, STRAIGHT, CLUbbeD. IN T. ligeralis, THe CLASPeR eXTeNDS beYOND eND OF SACCULUS, HAS A TRIANGULAR FLAP AT bASe, AND THe AeDeAGUS HAS TWO CORNUTI IN THe MALe GeNITALIA; THe ANTRUM IS TRAPeZOIDAL, AND THe SIGNUM IS SMALLeR AND beNT IN THe FeMALe GeNITALIA. Description. WINGSPAN 20.0¯23.0 MM (N = 19) (FIG. 4). VeRTeX LIGHT YeLLOWISH bROWN, SCALeS YeLLOWISH WHITe- TIPPeD; FRONS YeLLOWISH WHITe. ANTeNNA YeLLOWISH WHITe ALTeRNATeD WITH bROWNISH FUSCOUS, MALe FLAGeLLUM SIMPLe, bASAL SeGMeNTS NOT DILATeD, WITHOUT NOTCH, CILIA AbOUT 2/3 LeNGTH OF DIAMeTeR OF FLAGeLLUM. LAbIAL PALPUS WITH FIRST SeGMeNT LIGHT YeLLOWISH WHITe, SeCOND SeGMeNT DARK bROWNISH FUSCOUS, YeLLOWISH WHITe AT eND, THIRD SeGMeNT AbOUT HALF LeNGTH OF SeCOND SeGMeNT, YeLLOWISH WHITe, MIXeD WITH bROWNISH FUSCOUS AT bASe. OCCIPUT, PATAGIUM, TeGULA AND THORAX GRAYISH GReeN OR GRAYISH bROWN, SCALeS GRAYISH WHITe-TIPPeD. FOReWING WITH bASAL MAjORITY ReGION GRAYISH GReeN, SCATTeReD A FeW bLACK SCALeS, WITH AN DISTINCT, LARGe, TRIANGULAR, RUSTY bROWN PATCH INWARDLY SUb- ADjACeNT TO ANTeMeDIAL LINe; CeNTRAL ReGION GRAYISH GReeN NeAR ANTeMeDIAL LINe, GRAYISH WHITe NeAR POSTMeDIAL LINe, SUFFUSeD LOTS OF RUSTY bROWN SCALeS, DIFFUSeD bLACK AROUND VeINS beLOW DISCAL SPOTS; DISTAL ReGION GRAYISH bROWN, DIFFUSeD MORe WHITe SCALeS AT APeX, bROWNISH FUSCOUS ALONG VeINS; ANTeMeDIAL LINe AT bASAL 2/5 OF FOReWING, SINUATe, bLACK, INWARDLY eDGeD WITH AN INDISTINCT GRAYISH WHITe LINe; POSTMeDIAL LINe AT DISTAL 1/5 OF FOReWING, DISTINCT, ARCUATe, SLIGHTLY CONCAVe AT M1 AND CUA 2, bLACK, eDGeD OUTWARDLY WITH AN ObSCURe WHITe LINe, AND A bROWNISH FUSCOUS SHADe ADjACeNT TO WHITe LINe; DISCAL SPOTS NARROWLY FUSeD, ObSCURe, bLACK; TeRMINAL LINe bLACK; CILIA LIGHT YeLLOW. HINDWING GRAY, FUSCOUS ALONG VeINS AND MARGINS; CILIA LIGHT bROWN. FOReLeG WHITe DORSALLY, bLACK VeNTRALLY, TARSOMeReS YeLLOWISH WHITe AT eND; MID AND HIND LeGS WHITe WITH SCATTeReD bLACK SCALeS ON COXA, FeMUR AND TIbIA, TARSUS MOTTLeD WITH bLACKISH bROWN. Male genitalia (FIG. 11). UNCUS HOODLIKe, bROADeR THAN LONG, ROUNDeD AT APeX, SeTOSe IN DISTAL HALF ON DORSAL SURFACe. APICAL PROCeSS OF GNATHOS TAPeReD, WITH A SMALL HOOK DISTALLY, AbOUT 2/3 LeNGTH OF UNCUS. TRANSTILLA WITH TeRMINAL PART FUSeD INTO A SCLeROTIZeD ARCeD PLATe, WITH A PAIR OF STRONGLY SCLeROTIZeD, INWARDLY beNT, AURIFORM POSTeROLATeRAL PROCeSSeS. VALVA GRADUALLY bROADeNeD TO ROUNDeD APeX; COSTA A NARROW bAND, eXTeNDING TO eND OF VALVA, NeARLY eqUAL WIDTH eXCePT SLIGHTLY TAPeReD DISTALLY; SACCULUS AbOUT HALF LeNGTH OF VALVA, CUNIFORM, bROAD AT bASe, TAPeReD TO A DISSOCIATeD, POINTeD APeX; CLASPeR A NARROWeD, ARCHeD ARM, ARISING FROM NeAR bASe OF COSTA ObLIqUe OUTWARDLY TO eND OF SACCULUS, WITH A FINGeRLIKe FLAP AT bASe, beARING SPAReS SeTAe. JUXTA U-SHAPeD, LATeRAL LObeS WeLL DeVeLOPeD, AS LONG AS APICAL PROCeSS OF GNATHOS, FINGeRLIKe, beARING A FeW SeTAe. VINCULUM U-SHAPeD, MODeRATeLY LONG WITH TRANSVeRSe POSTeRIOR MARGIN. AeDeAGUS STOUT, GRADUALLY bROADeNeD TO APeX, SLIGHTLY SHORTeR THAN VALVA; WITH ONe TAPeReD PLATe NeAR MIDDLe, FOUR SHARP, CONICAL CORNUTI NeAR APeX, AND ONe LARGe, CLUbbeD CORNUTA beARING SMALL THORNS AT APeX. EIGHTH AbDOMINAL TeRGITe FAN-SHAPeD; eIGHTH AbDOMINAL STeRNITe SMALL, TRIANGULAR, ASSOCIATeD WITH A PAIR OF CULCITA. Female genitalia (FIG. 15). PAPILLAe ANALIS bROAD, ReCTANGULAR IN bASAL HALF, SeMICIRCULAR IN DISTAL HALF, WITH VeSTURe OF THIN HAIRS. EIGHTH AbDOMINAL SeGMeNT SHORT, 1/3 AS LONG AS WIDe; A TRIANGULAR SCLeRITe POSTeRIOR TO eIGHTH AbDOMINAL SeGMeNT. ANTRUM WeLL DeVeLOPeD, ReCTANGULAR, FOUR TIMeS AS WIDe AS LONG. DUCTUS bURSAe VeRY SHORT, AbOUT 1.2 TIMeS LeNGTH OF ANTRUM, PUNCTATe ON INNeR WALL AT jUNCTION WITH CORPUS bURSAe. CORPUS bURSAe MeMbRANOUS, OVATe, TWICe AS LONG AS WIDe, DeCORATeD WITH COARSe SCObINATIONS POSTeRIORLY TO FORM A SOMeWHAT SOLe-SHAPeD AReA; SIGNUM A STRONG, SOMeWHAT STRAIGHT, TAPeReD, CLUbbeD HORN ON A bROAD, FORKeD bASe, SITUATeD AT AbOUT MIDDLe OF CORPUS bURSAe. DUCTUS SeMINALIS INCePTeD POSTeRIOR 1/3 OF CORPUS bURSAe, SAMe SIDe WITH SIGNUM. Type material. Holotype ♂, China: NATIONAL FOReST PARK, TAIYANGHe (22°40'N 101°01'E), PU’eR, YUNNAN, 1450 M, 3.VII.2015, COLL. KAIjIAN TeNG (NKUM). Paratypes: China: Yunnan: 9 ♂♂, 1 ♀, SAMe DATA AS FOR HOLOTYPe, GeN. SLIDe NO. YLL15178M, YLL1590M, YLL 15191 F (NKUM); 5 ♂♂, SAMe PLACe AS FOR HOLOTYPe, 8.VII.2013, COLL. LINLIN YANG, GeN. SLIDe NO. RYD2013100, RYD2014218 (HAASM); 2 ♂♂, SAMe PLACe AS FOR HOLOTYPe, 31.III. 2014, COLL. ZHeNGUO ZHANG, GeN. SLIDe NOS YLL15181, YLL15183 (NKUM); 1 ♂, NANKANG (24°49'N 98°47'E), BAOSHAN, 2009 M, 20.VII.2015, COLL. KAIjIAN TeNG & XIA BAI, GeN. SLIDe NO. YLL15192 (NKUM). Distribution. CHINA (YUNNAN). Etymology. THe SPeCIFIC NAMe IS DeRIVeD FROM THe LATIN WORD celadontis, MeANING GRAYISH GReeN, ReFeRRING TO THe FOReWING IS GRAYISH GReeN IN bASAL MAjORITY ReGION.

Thiallela epicrociella (Strand, 1918) (FIGS. 1, 2, 9, 13) Phycita (?) epicrociella Strand, 1918: 273. TL: China (Taiwan, Alishan). TD: DEI. Numonia epicrociella: Shibuya, 1928: 18. Acrobasis epicrociella: Yamanaka & Yoshiyasu, 1992: 93. Thiallela epicrociella: Roesler, 1993: 240. Redescription. WINGSPAN 20.5¯22.5 MM (N = 9) (FIG. 2). VeRTeX AND FRONS CReAMY YeLLOW. ANTeNNA KHAKI, MALe FLAGeLLUM WITH FIRST TO FIFTH SeGMeNTS DILATeD, SIXTH SeGMeNT DISTINCTLY NOTCHeD, CILIA SHORTeR THAN 1/3 LeNGTH OF DIAMeTeR OF FLAGeLLUM. LAbIAL PALPUS YeLLOWISH WHITe, FIRST AND SeCOND SeGMeNTS MIXeD WITH GRAYISH bLACK; THIRD SeGMeNT AbOUT HALF LeNGTH OF SeCOND SeGMeNT. OCCIPUT, PATAGIUM, TeGULA AND THORAX YeLLOWISH OCHeROUS WITH SPARSe GRAYISH bLACK AND bROWN SCALeS. FOReWING WITH bASAL MAjORITY ReGION YeLLOWISH OCHeROUS, WITH A FeW SCATTeReD bLACK SCALeS, AND A LARGe, ARCHeD, SUbTRIANGULAR GRAYISH bLACK PATCH ADjACeNT TO DORSAL MARGIN AND INWARDLY SUb- ADjACeNT TO ANTeMeDIAL LINe; CeNTRAL ReGION ASH GRAY, MOTTLeD LOTS OF GRAYISH bLACK AND PALe WHITe SCALeS, TINTeD WITH SeVeRAL bLACK, WHITe AND RUSTY bROWN SCALeS, bLACK ALONG VeINS; DISTAL ReGION GRAY; ANTeMeDIAL LINe NeAR MIDDLe OF FOReWING, bLACK, ARCHeD, INTeRRUPTeD, INWARDLY eDGeD WITH AN INDISTINCT YeLLOWISH WHITe LINe; POSTMeDIAL LINe AT DISTAL 1/6 OF FOReWING, bLACK, ARCUATe, SLIGHTLY CONCAVe AT M1 AND CUA2, eDGeD OUTWARDLY WITH AN ObSCURe GRAYISH WHITe LINe, AND A bROWN SHADe ADjACeNT TO GRAYISH WHITe LINe; DISCAL SPOTS FUSeD, ObSCURe, bLACKISH bROWN; TeRMINAL LINe bLACK, INTeRRUPTeD, DISTINCT; CILIA bROWN. HINDWING SeMITRANSPAReNT, PALe GRAYISH WHITe, DARK bROWN ALONG MARGINS AND VeINS; CILIA GRAYISH WHITe. LeGS YeLLOWISH WHITe, FOReLeG MOTTLeD WITH NUMeROUS bLACKISH GRAY, MID TIbIA WITH AN ObLIqUe bLACKISH GRAY PATCH NeAR eND, TARSUS MIXeD WITH bLACKISH GRAY, HIND TARSUS MIXeD WITH bLACKISH bROWN. Male genitalia (FIG. 9). UNCUS HOODLIKe, bROADeR THAN LONG, ROUNDeD AT APeX, SeTOSe IN DISTAL HALF OF DORSAL SURFACe. APICAL PROCeSS OF GNATHOS STRONGLY SCLeROTIZeD, SPATULATe, APeX TRUNCATe, AbOUT SAMe LeNGTH AS UNCUS. TRANSTILLA WITH TeRMINAL PART FUSeD INTO AN ARCeD PLATe, WITH A PAIR OF STRONGLY SCLeROTIZeD TRIANGULAR POSTeROLATeRAL PROCeSSeS. VALVA GRADUALLY bROADeNeD TO ROUNDeD APeX; COSTA A NARROWeD bAND, eXTeNDING TO eND OF VALVA, NeARLY eqUAL WIDTH eXCePT SLIGHTLY TAPeReD DISTALLY; SACCULUS 2/3 LeNGTH OF VALVA, TRIANGULAR, bASe bROAD, TAPeReD TO APeX, APICAL 1/3 FRee; CLASPeR A NARROWeD bAND, ARISING FROM NeAR bASe OF COSTA ObLIqUe OUTWARDLY TO eND OF SACCULUS, bASAL HALF ARCHeD, DISTAL HALF WeAKLY SCLeROTIZeD, WITH A MASTOID FLAP AT bASe, beARING SPAReS SeTAe. JUXTA V-SHAPeD, LATeRAL LObeS WeLL DeVeLOPeD, FUSIFORM, 4/5 LeNGTH OF APICAL PROCeSS OF GNATHOS, beARING SPARSe SeTAe TeRMINALLY. VINCULUM U-SHAPeD, LeNGTH SHORTeR THAN TRANSVeRSe POSTeRIOR MARGIN. AeDeAGUS CYLINDRICAL, 3/4 LeNGTH OF VALVA, WITH AN IRReGULAR PLATe NeAR bASe, NUMeROUS SMALL CONICAL CORNUTI IN DIFFeReNT SIZe IN DISTAL 3/5. EIGHTH AbDOMINAL TeRGITe FAN-SHAPeD; eIGHTH AbDOMINAL STeRNITe bOWL-SHAPeD, ASSOCIATeD WITH A PAIR OF CULCITA. Female genitalia (FIG. 13). PAPILLAe ANALIS SUbTRIANGULAR, APeX ROUNDeD, WITH VeSTURe OF THIN HAIRS. APOPHYSIS POSTeRIORIS SLIGHT LONGeR THAN ANTeRIORIS. EIGHTH AbDOMINAL SeGMeNT SHORT, A THIRD AS LONG AS WIDe. ANTRUM STRONGLY SCLeROTIZeD, SOMeWHAT ReCTANGULAR, THRee TIMeS AS WIDe AS LONG, ITS POSTeRIOR MARGIN INTeRMeDIATeLY V-SHAPeD CONCAVe. DUCTUS bURSAe WeAKLY SCLeROTIZeD, AbOUT 1/3 WIDTH, TWICe LeNGTH OF ANTRUM, PUNCTATe ON INNeR WALL IN POSTeRIOR HALF. CORPUS bURSAe OVATe, 1.5 TIMeS AS LONG AS WIDe; SIGNUM STRONGLY SCLeROTIZeD, A LARGe L-SHAPeD beNDING HORN ON A NARROW bASe, SITUATeD AT AbOUT POSTeRIOR 1/3 OF CORPUS bURSAe, PUNCTATe AROUND. DUCTUS SeMINALIS INCePTeD POSTeRIOR MARGIN OF CORPUS bURSAe, OPPOSITe TO SIGNUM. Material examined. CHINA: Guangdong: 1 ♀, LIANHUASHAN (22°58'N 113°30'E), DONGGUAN, 21.VI.2002, COLL. GUILIN LIU, GeN. SLIDe NO. RYD 04659 (NKUM). Guangxi: 1 ♀, DONGZHONG YARD (21°37'N 107°31'E), FANGCHeNGGANG, 640 M, 8.IV.2002, COLL. SHULIAN HAO & HUAIjUN XUe, GeN. SLIDe NO. RYD 04514 (NKUM). Hainan: 1 ♂, YAjIA (19°05'N 109°07'E), MT. BAWANG, 261 M, 20.VII.2015, COLL. QINGYUN WANG et al., GeN. SLIDe NO. YLL 15193 M (NKUM). Yunnan: 3 ♂♂, 3 ♀♀, YeXIANGGU (22°10'N 100°52'E), XISHUANGbANNA, 762 M, 17¯ 19.VII.2014, COLL. KAIjIAN TeNG et al., GeN. SLIDe NOS YLL 15185 M, YLL15187M, YLL15188F (NKUM). Distribution. CHINA (GUANGDONG, GUANGXI, HAINAN, TAIWAN, YUNNAN). Notes. THe MALe OF T. epicrociella IS DeSCRIbeD FOR THe FIRST TIMe., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on pages 440-441, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645, {"references":["Strand, E. (1918) H. Sauter's Formosa-Ausbeute: Pyralididae, Subfam. Galleriinae, Crambinae, Schoenobiinae, Anerastiinae und Phycitinae. Stettiner Entomologische Zeitung, 79 (2), 248 - 276.","Shibuya, J. (1928) The systematic study on the formosan Pyralidae. Journal of the Faculty of Agriculture, Hokkaido Imperial University, Sapporo, 22 (1), 1 - 300, pls. 1 - 9.","Roesler, R. U. (1993) Quadrifine Acrobasiina. 2. Teilband der Phycitinae. In: Amsel, H. G., Gregor, F., Reisser, H. & Roesler, R. U. (Eds.), Microlepidoptera Palaearctica. 8 (1 - 2). Braun, G., Karlsruhe, pp. i - xxii, 1 - 305, pls. 1 - 82."]}

6. Thiallela hiranoi Yamanaka, 2002 Thiallela hiranoi YAMANAKA, 2002: 83; LI & ReN, 2006: 330; YAMANAKA, 2013: 345. TL: JAPAN (HONSHU). TD: HOLOTYPe IN NSMT; PARATYPeS IN COLL. HY. DISTRIbUTION: CHINA (ZHejIANG), JAPAN., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on page 438, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645, {"references":["Yamanaka, H. (2002) Descriptions of two new species of the Phycitinae (Lepidoptera: Pyralidae) from Japan. Tinea, 17 (2), 81 - 85.","Yamanaka, H. (2013) Pyralidae. In: Hirowatari, T., Nasu, Y., Sakamaki, Y. & Kishida, Y. (Eds.), The Standard of Moths in Japan. Vol. 4. Gakken Education Publishing, Tokyo, pp. 314 - 373."]}

Thiallela sulciflagella sp. nov. (FIGS. 1, 5, 12, 16) Diagnosis. THIS NeW SPeCIeS ReSeMbLeS TO T. epicrociella, bUT DIFFeRS FROM THe LATTeR bY THe MALe FLAGeLLUM WITH FIRST THRee SeGMeNTS SLIGHTLY DILATeD, FOURTH TO SIXTH SeGMeNTS DISTINCTLY NOTCHeD; IN THe MALe GeNITALIA, THe APICAL PROCeSS OF THe GNATHOS TAPeReD, THe TRANSTILLA WITH A PAIR OF HILLOCK-SHAPeD POSTeROLATeRAL PROCeSSeS, AND THe AeDeAGUS WITH FIVe HORN-LIKe CORNUTI DISTALLY; IN THe FeMALe GeNITALIA, THe CORPUS bURSAe WITH AN IRReGULAR SCLeROTIZeD AReA beARING DeNSe THORNS. IN T. epicrociella, THe MALe FLAGeLLUM IS DILATeD IN bASAL FIVe SeGMeNTS, NOTCHeD IN SIXTH SeGMeNT; IN THe MALe GeNITALIA, THe APICAL PROCeSS OF THe GNATHOS IS SPATULATe, THe TRANSTILLA HAS A PAIR OF TRIANGULAR POSTeROLATeRAL PROCeSSeS, AND THe AeDeAGUS WITHOUT HORN-LIKe CORNUTI; IN THe FeMALe GeNITALIA, THe ANTRUM IS THRee TIMeS AS WIDe AS LONG, AND THe CORPUS bURSAe HAS NO DeCORATION OF THORNS. Description. WINGSPAN 17.5��21.0 MM (N = 12) (FIG. 5). VeRTeX AND FRONS SILVeR-WHITe. ANTeNNA YeLLOWISH WHITe TINTeD WITH GOLDeN GLOSSY; SCAPe WITH LONG-NARROW SCALeS; MALe FLAGeLLUM WITH FIRST THRee SeGMeNTS SLIGHTLY DILATeD, FOURTH TO SIXTH SeGMeNTS DISTINCTLY NOTCHeD, CILIA SHORTeR THAN 1/3 LeNGTH OF DIAMeTeR OF FLAGeLLUM. LAbIAL PALPUS WITH bASAL TWO SeGMeNTS bROWNISH FUSCOUS MIXeD WITH LIGHT YeLLOW AND YeLLOWISH WHITe SCALeS, THIRD SeGMeNT AbOUT 1/3 LeNGTH OF SeCOND SeGMeNT, YeLLOWISH WHITe. OCCIPUT, PATAGIUM, TeGULA AND THORAX YeLLOWISH WHITe WITH SCATTeReD GRAYISH bLACK. FOReWING WITH bASAL MAjORITY ReGION GRAYISH YeLLOW, MOTTLeD A FeW bLACK SCALeS, TINGeD WITH GRAYISH bROWN ALONG COSTA, WITH AN INDISTINCT, ARCHeD, GRAYISH bLACK PATCH INWARDLY SUb-ADjACeNT TO ANTeMeDIAL LINe; CeNTRAL ReGION ASHY GRAY TINTeD WITH SeVeRAL GRAYISH bLACK AND RUSTY bROWN SCALeS; DISTAL ReGION ASHY GRAY; ANTeMeDIAL LINe beYOND MIDDLe OF FOReWING, SLIGHTLY ARCHeD, bLACK, eDGeD INWARDLY bY YeLLOWISH WHITe; POSTMeDIAL LINe AT DISTAL 1/6 OF FOReWING, bLACK, SLIGHTLY CONCAVe AT M1 AND CUA 2, eDGeD OUTWARDLY WITH AN ObSCURe GRAYISH WHITe LINe, AND A bROWN SHADe ADjACeNT TO GRAYISH WHITe LINe; DISCAL SPOTS FUSeD, ObSCURe, GRAYISH bLACK; TeRMINAL LINe bLACK; CILIA GRAYISH bROWN. HINDWING SeMITRANSPAReNT, GRAY TINTeD WITH COPPeRY GLOSSY, FUSCOUS ALONG VeINS AND TeRMeN; CILIA LIGHT YeLLOW. LeGS YeLLOWISH WHITe MOTTLeD WITH NUMeROUS GRAYISH bLACK. Male genitalia (FIG. 12). UNCUS HOODLIKe, LONGeR THAN bROAD, TRUNCATe AT APeX, SeTOSe IN DISTAL HALF ON DORSAL SURFACe. APICAL PROCeSS OF GNATHOS TAPeReD, SLIGHTLY SHORTeR THAN UNCUS, WITH A SMALL HOOK DISTALLY. TRANSTILLA WITH TeRMINAL PART FUSeD INTO A SCLeROTIZeD, ROUNDeD-ReCTANGULAR PLATe, WITH A PAIR OF SHORT, HILLOCK-SHAPeD POSTeROLATeRAL PROCeSSeS. VALVA SLIGHTLY bROADeNeD TO ROUNDeD APeX; COSTA A NARROWeD bAND, eXTeNDING TO eND OF VALVA, NeARLY eqUAL WIDTH eXCePT SLIGHTLY TAPeReD DISTALLY; SACCULUS AbOUT HALF LeNGTH OF VALVA, CUNIFORM, bROAD AT bASe, TAPeReD TO APeX, DISTAL 1/5 FRee; CLASPeR A NARROWeD ARM, SLIGHTLY SINUATe, ARISING FROM bASe OF COSTA ObLIqUe OUTWARDLY TO DISTAL 1/4 OF VeNTRAL MARGIN, beYOND eND OF SACCULUS, WITH A NARROWeD, TRIANGULAR FLAP AT bASe, beARING SPAReS SeTAe, WITH A SMALL SPINe DORSOAPICALLY, SLIGHTLY INFLATeD VeNTROAPICALLY. JUXTA U-SHAPeD, LATeRAL LObeS WeLL DeVeLOPeD, AS LONG AS APICAL PROCeSS OF GNATHOS, FINGeRLIKe, CURVeD OUTWARD, beARING SPARSe SeTAe. VINCULUM U-SHAPeD, MODeRATeLY LONG WITH TRANSVeRSe POSTeRIOR MARGIN. AeDeAGUS CYLINDRICAL, 3/4 LeNGTH OF VALVA; WITH AN IRReGULAR PLATe AT bASe, CORNUTI RePReSeNT bY NUMeROUS SMALL, CONICAL THORNS MeDIALLY, AND FIVe CONICAL HORN DISTALLY. EIGHTH AbDOMINAL TeRGITe SOMeWHAT ReCTANGULAR; eIGHTH AbDOMINAL STeRNITe SMALL, bOWL-SHAPeD, ASSOCIATeD WITH A PAIR OF CULCITA. Female genitalia (FIG. 16). PAPILLAe ANALIS bROAD, SUbTRIANGULAR, ROUNDeD AT APeX, WITH VeSTURe OF THIN HAIRS. EIGHTH AbDOMINAL SeGMeNT 2.2 TIMeS AS WIDe AS LONG; A TRIANGULAR SCLeRITe POSTeRIOR TO eIGHTH AbDOMINAL SeGMeNT. ANTRUM WeLL DeVeLOPeD, ReCTANGULAR, 1.4 TIMeS AS WIDe AS LONG, DeNSeLY PUNCTATe ANTeRIORLY. DUCTUS bURSAe VeRY SHORT, AbOUT 1/3 LeNGTH OF ANTRUM, PUNCTATe ON INNeR WALL. CORPUS bURSAe MeMbRANOUS, THRee TIMeS AS LONG AS WIDe, WITH AN IRReGULAR SCLeROTIZeD AReA beARING DeNSe CONICAL THORNS IN DIFFeReNT SIZe; SIGNUM A LARGe, STRONGLY SCLeROTIZeD, bOMMeRANG-SHAPeD HORN, SITUATeD AT AbOUT MIDDLe OF CORPUS bURSAe, OUTSIDe PROTRUDeD SeCTION HANDLe- LIKe, INSIDe PROTRUDeD SeCTION FALCIFORM. DUCTUS SeMINALIS INCePTeD POSTeRIOR 1/3 OF CORPUS bURSAe. Type material. Holotype ♂, China: YAjIA (19��05'N 109��07'E), MT. BAWANG, HAINAN, 261 M, 22.VII.2015, COLL. QINGYUN WANG et al. (NKUM). Paratypes: China: Guangxi: 1 ♂, LONGRUI NATIONAL ReSeRVe (22��44'N 110��55'E), 19.III. 2011, COLL. MUCHUN Cheng, gen. slide No. RYD2014216 (HAASM); 2 ♀♀, Nonggang National reserve (22��26'N 106��57'E), LONGZHOU, 20.VIII.2011, COLL. DANDAN ZHANG & JINWeI LI, GeN. SLIDe NO. RYD2014217 (HAASM). Hainan: 2 ♂♂, 2 ♀♀, SAMe DATA AS FOR HOLOTYPe, GeN. SLIDe NOS YLL 15194 M, YLL15195F (NKUM); 2 ♂♂, HONGKAN (19��02'N 109��33'E), MT. YINGGe, 540 M, 23.VII.2015, COLL. QINGYUN WANG et al., GeN. SLIDe NO. YLL15196 (NKUM). Yunnan: 2 ♂♂, YeXIANGGU (22��10'N 100��52'E), XISHUANGbANNA, 762 M, 10.VII.2015, COLL. KAIjIAN TeNG & XIA BAI, GeN. SLIDe NO. YLL 15184 (NKUM). Distribution. CHINA (GUANGXI, HAINAN, YUNNAN). Etymology. THe SPeCIFIC NAMe IS DeRIVeD FROM THe LATIN PReFIX sulc -, MeANING NOTCHeD, AND MORPHOLOGICAL TeRM flagellum, ReFeRRING TO THe MALe FLAGeLLUM WITH FOURTH TO SIXTH SeGMeNTS DISTINCTLY NOTCHeD., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on pages 448-449, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645

5. Thiallela escigera (Meyrick, 1932) Phalobathra escigera MeYRICK, 1932: 234; WHALLeY, 1970: 56. TL: FIjI. TD: LeCTOTYPe IN BMNH. Thiallela escigera: ROeSLeR & KüPPeRS, 1981: 76; ROeSLeR, 1983: 96. DISTRIbUTION: FIjI, INDONeSIA (SUMATRA)., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on page 438, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645, {"references":["Meyrick, E. (1932) Exotic Microlepidoptera. Vol. 4. Thornhanger, Marlborough, Wilts, 32 pp. [pp. 225 - 256]","Whalley, P. E. S. (1970) A synonymic catalogue of the genera of Phycitinae (Lepidoptera: Pyralidae) of the world. Bulletin of the British Museum (Natural History) Entomology Series, 25 (2), 33 - 72.","Roesler, R. U. & Kuppers, P. V. (1981) Beitrage zur Kenntnis der Insektenfauna Sumatras. Teil 9. Die Phycitinae (Lepidoptera: Pyralidae) von Sumatra; Taxonomie Teil B, Okologie und Geobiologie. Beitrage zur Naturkundlichen Forschung in Sudwestdeutschland, 4, 1 - 282, 4 text abbildung, 42 tafel.","Roesler, R. U. (1983) Die Phycitinae von Sumatra (Lepidoptera: Pyralidae). Heterocera Sumatrana, Keltern, 3, 1 - 136, pls. 1 - 69."]}

Thiallela sulciflagella sp. nov. (FIGS. 1, 5, 12, 16) Diagnosis. THIS NeW SPeCIeS ReSeMbLeS TO T. epicrociella, bUT DIFFeRS FROM THe LATTeR bY THe MALe FLAGeLLUM WITH FIRST THRee SeGMeNTS SLIGHTLY DILATeD, FOURTH TO SIXTH SeGMeNTS DISTINCTLY NOTCHeD; IN THe MALe GeNITALIA, THe APICAL PROCeSS OF THe GNATHOS TAPeReD, THe TRANSTILLA WITH A PAIR OF HILLOCK-SHAPeD POSTeROLATeRAL PROCeSSeS, AND THe AeDeAGUS WITH FIVe HORN-LIKe CORNUTI DISTALLY; IN THe FeMALe GeNITALIA, THe CORPUS bURSAe WITH AN IRReGULAR SCLeROTIZeD AReA beARING DeNSe THORNS. IN T. epicrociella, THe MALe FLAGeLLUM IS DILATeD IN bASAL FIVe SeGMeNTS, NOTCHeD IN SIXTH SeGMeNT; IN THe MALe GeNITALIA, THe APICAL PROCeSS OF THe GNATHOS IS SPATULATe, THe TRANSTILLA HAS A PAIR OF TRIANGULAR POSTeROLATeRAL PROCeSSeS, AND THe AeDeAGUS WITHOUT HORN-LIKe CORNUTI; IN THe FeMALe GeNITALIA, THe ANTRUM IS THRee TIMeS AS WIDe AS LONG, AND THe CORPUS bURSAe HAS NO DeCORATION OF THORNS. Description. WINGSPAN 17.5¯21.0 MM (N = 12) (FIG. 5). VeRTeX AND FRONS SILVeR-WHITe. ANTeNNA YeLLOWISH WHITe TINTeD WITH GOLDeN GLOSSY; SCAPe WITH LONG-NARROW SCALeS; MALe FLAGeLLUM WITH FIRST THRee SeGMeNTS SLIGHTLY DILATeD, FOURTH TO SIXTH SeGMeNTS DISTINCTLY NOTCHeD, CILIA SHORTeR THAN 1/3 LeNGTH OF DIAMeTeR OF FLAGeLLUM. LAbIAL PALPUS WITH bASAL TWO SeGMeNTS bROWNISH FUSCOUS MIXeD WITH LIGHT YeLLOW AND YeLLOWISH WHITe SCALeS, THIRD SeGMeNT AbOUT 1/3 LeNGTH OF SeCOND SeGMeNT, YeLLOWISH WHITe. OCCIPUT, PATAGIUM, TeGULA AND THORAX YeLLOWISH WHITe WITH SCATTeReD GRAYISH bLACK. FOReWING WITH bASAL MAjORITY ReGION GRAYISH YeLLOW, MOTTLeD A FeW bLACK SCALeS, TINGeD WITH GRAYISH bROWN ALONG COSTA, WITH AN INDISTINCT, ARCHeD, GRAYISH bLACK PATCH INWARDLY SUb-ADjACeNT TO ANTeMeDIAL LINe; CeNTRAL ReGION ASHY GRAY TINTeD WITH SeVeRAL GRAYISH bLACK AND RUSTY bROWN SCALeS; DISTAL ReGION ASHY GRAY; ANTeMeDIAL LINe beYOND MIDDLe OF FOReWING, SLIGHTLY ARCHeD, bLACK, eDGeD INWARDLY bY YeLLOWISH WHITe; POSTMeDIAL LINe AT DISTAL 1/6 OF FOReWING, bLACK, SLIGHTLY CONCAVe AT M1 AND CUA 2, eDGeD OUTWARDLY WITH AN ObSCURe GRAYISH WHITe LINe, AND A bROWN SHADe ADjACeNT TO GRAYISH WHITe LINe; DISCAL SPOTS FUSeD, ObSCURe, GRAYISH bLACK; TeRMINAL LINe bLACK; CILIA GRAYISH bROWN. HINDWING SeMITRANSPAReNT, GRAY TINTeD WITH COPPeRY GLOSSY, FUSCOUS ALONG VeINS AND TeRMeN; CILIA LIGHT YeLLOW. LeGS YeLLOWISH WHITe MOTTLeD WITH NUMeROUS GRAYISH bLACK. Male genitalia (FIG. 12). UNCUS HOODLIKe, LONGeR THAN bROAD, TRUNCATe AT APeX, SeTOSe IN DISTAL HALF ON DORSAL SURFACe. APICAL PROCeSS OF GNATHOS TAPeReD, SLIGHTLY SHORTeR THAN UNCUS, WITH A SMALL HOOK DISTALLY. TRANSTILLA WITH TeRMINAL PART FUSeD INTO A SCLeROTIZeD, ROUNDeD-ReCTANGULAR PLATe, WITH A PAIR OF SHORT, HILLOCK-SHAPeD POSTeROLATeRAL PROCeSSeS. VALVA SLIGHTLY bROADeNeD TO ROUNDeD APeX; COSTA A NARROWeD bAND, eXTeNDING TO eND OF VALVA, NeARLY eqUAL WIDTH eXCePT SLIGHTLY TAPeReD DISTALLY; SACCULUS AbOUT HALF LeNGTH OF VALVA, CUNIFORM, bROAD AT bASe, TAPeReD TO APeX, DISTAL 1/5 FRee; CLASPeR A NARROWeD ARM, SLIGHTLY SINUATe, ARISING FROM bASe OF COSTA ObLIqUe OUTWARDLY TO DISTAL 1/4 OF VeNTRAL MARGIN, beYOND eND OF SACCULUS, WITH A NARROWeD, TRIANGULAR FLAP AT bASe, beARING SPAReS SeTAe, WITH A SMALL SPINe DORSOAPICALLY, SLIGHTLY INFLATeD VeNTROAPICALLY. JUXTA U-SHAPeD, LATeRAL LObeS WeLL DeVeLOPeD, AS LONG AS APICAL PROCeSS OF GNATHOS, FINGeRLIKe, CURVeD OUTWARD, beARING SPARSe SeTAe. VINCULUM U-SHAPeD, MODeRATeLY LONG WITH TRANSVeRSe POSTeRIOR MARGIN. AeDeAGUS CYLINDRICAL, 3/4 LeNGTH OF VALVA; WITH AN IRReGULAR PLATe AT bASe, CORNUTI RePReSeNT bY NUMeROUS SMALL, CONICAL THORNS MeDIALLY, AND FIVe CONICAL HORN DISTALLY. EIGHTH AbDOMINAL TeRGITe SOMeWHAT ReCTANGULAR; eIGHTH AbDOMINAL STeRNITe SMALL, bOWL-SHAPeD, ASSOCIATeD WITH A PAIR OF CULCITA. Female genitalia (FIG. 16). PAPILLAe ANALIS bROAD, SUbTRIANGULAR, ROUNDeD AT APeX, WITH VeSTURe OF THIN HAIRS. EIGHTH AbDOMINAL SeGMeNT 2.2 TIMeS AS WIDe AS LONG; A TRIANGULAR SCLeRITe POSTeRIOR TO eIGHTH AbDOMINAL SeGMeNT. ANTRUM WeLL DeVeLOPeD, ReCTANGULAR, 1.4 TIMeS AS WIDe AS LONG, DeNSeLY PUNCTATe ANTeRIORLY. DUCTUS bURSAe VeRY SHORT, AbOUT 1/3 LeNGTH OF ANTRUM, PUNCTATe ON INNeR WALL. CORPUS bURSAe MeMbRANOUS, THRee TIMeS AS LONG AS WIDe, WITH AN IRReGULAR SCLeROTIZeD AReA beARING DeNSe CONICAL THORNS IN DIFFeReNT SIZe; SIGNUM A LARGe, STRONGLY SCLeROTIZeD, bOMMeRANG-SHAPeD HORN, SITUATeD AT AbOUT MIDDLe OF CORPUS bURSAe, OUTSIDe PROTRUDeD SeCTION HANDLe- LIKe, INSIDe PROTRUDeD SeCTION FALCIFORM. DUCTUS SeMINALIS INCePTeD POSTeRIOR 1/3 OF CORPUS bURSAe. Type material. Holotype ♂, China: YAjIA (19°05'N 109°07'E), MT. BAWANG, HAINAN, 261 M, 22.VII.2015, COLL. QINGYUN WANG et al. (NKUM). Paratypes: China: Guangxi: 1 ♂, LONGRUI NATIONAL ReSeRVe (22°44'N 110°55'E), 19.III. 2011, COLL. MUCHUN Cheng, gen. slide No. RYD2014216 (HAASM); 2 ♀♀, Nonggang National reserve (22°26'N 106°57'E), LONGZHOU, 20.VIII.2011, COLL. DANDAN ZHANG & JINWeI LI, GeN. SLIDe NO. RYD2014217 (HAASM). Hainan: 2 ♂♂, 2 ♀♀, SAMe DATA AS FOR HOLOTYPe, GeN. SLIDe NOS YLL 15194 M, YLL15195F (NKUM); 2 ♂♂, HONGKAN (19°02'N 109°33'E), MT. YINGGe, 540 M, 23.VII.2015, COLL. QINGYUN WANG et al., GeN. SLIDe NO. YLL15196 (NKUM). Yunnan: 2 ♂♂, YeXIANGGU (22°10'N 100°52'E), XISHUANGbANNA, 762 M, 10.VII.2015, COLL. KAIjIAN TeNG & XIA BAI, GeN. SLIDe NO. YLL 15184 (NKUM). Distribution. CHINA (GUANGXI, HAINAN, YUNNAN). Etymology. THe SPeCIFIC NAMe IS DeRIVeD FROM THe LATIN PReFIX sulc -, MeANING NOTCHeD, AND MORPHOLOGICAL TeRM flagellum, ReFeRRING TO THe MALe FLAGeLLUM WITH FOURTH TO SIXTH SeGMeNTS DISTINCTLY NOTCHeD.

7. Thiallela ligeralis (Walker, 1863) Myelois ligeralis WALKeR, 1863: 37. TL: SRI LANKA. TD: BMNH. Thiallela signifera WALKeR, 1863: 120. TL: BORNeO. TD: BMNH. AS SYNONYM OF Thiallela ligeralis (WALKeR) bY ROeSLeR & K��PPeRS, 1981: 72. Luconia pallidobasella RAGONOT, 1888: 7; RAGONOT, 1893: 1. TL: BORNeO. TD: BMNH. AS SYNONYM OF Thiallela signifera WALKeR bY HAMPSON, 1896: 106. AS SYNONYM OF Thiallela ligeralis (WALKeR) bY ROeSLeR & K��PPeRS, 1981: 72. Acrobasis ligeralis: RAGONOT, 1893: 95. Rhodophaea ligeralis: HAMPSON, 1896: 99. Thiallela ligeralis: ROeSLeR & K��PPeRS 1981: 72; ROeSLeR, 1983: 95. DISTRIbUTION: CHINA (HAINAN, YUNNAN), ASSAM, SRI LANKA, BURMA, MALAYSIA, BORNeO, SUMATRA, NeW GUINeA, AUSTRALIA, FIjI., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on page 438, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645, {"references":["Walker, F. (1863) Crambites & Tortricites. In: List of Specimens of Lepidopterous Insects in the Collection of the British Museum. Vol. 27. Printed by order of the Trutees, London, pp. 1 - 286.","Roesler, R. U. & Kuppers, P. V. (1981) Beitrage zur Kenntnis der Insektenfauna Sumatras. Teil 9. Die Phycitinae (Lepidoptera: Pyralidae) von Sumatra; Taxonomie Teil B, Okologie und Geobiologie. Beitrage zur Naturkundlichen Forschung in Sudwestdeutschland, 4, 1 - 282, 4 text abbildung, 42 tafel.","Ragonot, E. L. (1888) Nouveaux genres et especes de Phycitidae & Galleriidae. 52 pp. Paris.","Ragonot, E. L. (1893) Monographie des Phycitinae et des Galleriinae. In: Romanoff, N. M. (Ed.), Memoires sur les Lepidopteres VII 7. St. Petersburg. pp. i - lvi, 1 - 658, pls 1 - 23.","Roesler, R. U. (1983) Die Phycitinae von Sumatra (Lepidoptera: Pyralidae). Heterocera Sumatrana, Keltern, 3, 1 - 136, pls. 1 - 69."]}

Key to species of Thiallela based on male genitalia 1. Lateral lobes of juxta obviously longer than apical process of gnathos........................................... 3 - Lateral lobes of juxta not longer than apical process of gnathos................................................ 4 2. Transtilla with middle area of posterior margin protruded in hillock shape, posterolateral processes narrowed and elongate (Li & Ren 2006, Fig. 5)........................................................................... T. naevilla - Transtilla with middle area of posterior margin short-fingerlike protruded, posterolateral processes short and broad (Li & Ren 2006, Fig. 6)............................................................................... T. platantra 3. Apical process of gnathos trapezoidal (Yamanaka 2002, Fig. 4)......................................... T. hiranoi - Apical process of gnathos tapered or spatulate............................................................. 5 4. Clasper extending to or near end of sacculus............................................................... 6 - Clasper extending beyond end of sacculus................................................................ 7 5. Apical process of gnathos spatulate; posterolateral processes of transtilla triangular (Fig. 9)............... T. epicrociella - Apical process of gnathos tapered; posterolateral processes of transtilla auriform, bent inwardly (Fig. 11).................................................................................................... T. celadontis sp. nov. 6. Clasper not inflated ventroapically, without small spine dorsoapically; aedeagus with two clubbed cornuti (Fig. 10).................................................................................................... T. ligeralis - Clasper slightly inflated ventroapically, with a small spine dorsoapically; aedeagus with numerous small, conical thorn-like and five horn-like cornuti (Fig. 12)..................................................... T. sulciflagella sp. nov., Published as part of Yang, Linlin & Ren, Yingdang, 2018, Two new species, one unrecorded species, and one newly reported male of genus Thiallela Walker, 1863 from China (Lepidoptera: Pyralidae: Phycitinae), pp. 436-450 in Zootaxa 4387 (3) on page 439, DOI: 10.11646/zootaxa.4387.3.2, http://zenodo.org/record/1187645, {"references":["Yamanaka, H. (2002) Descriptions of two new species of the Phycitinae (Lepidoptera: Pyralidae) from Japan. Tinea, 17 (2), 81 - 85."]}

Cathyalia Ragonot, 1888 Cathyalia Ragonot, 1888: 7; Roesler & K��ppers, 1981: 15; Roesler, 1983: 78; Robinson, Tuck & Shaffer, 1994: 159; Shaffer, Nielsen & Horak, 1996: 177. Type species: Cathyalia fulvella Ragonot, 1888. General characters. Vertex flat, with rough scales. Antenna simple and shortly pubescent on both sexes. Ocellus present. Chaetosema well developed. Labial palpus upturned beyond head, with third segment slightly shorter than second, apex pointed. Maxillary palpus short, no longer than second segment of labial palpus. Proboscis well developed. Forewing with R 2 and R 3+4 +R 5 close at base or shortly stalked, R 3+4 and R 5 stalked for about half length of R 5, M 2 close to M 3 at base, from lower angle of cell, CuA 1 and CuA 2 free; cell about 2/3 length of forewing. Hindwing with Rs and Sc+R 1 long-stalked, M 1 and Sc+Rs+R 1 stalked for 2/5 of M 1, M 2 and M 3 close at base, from lower angle of cell, CuA 1 and CuA 2 free; cell very short, only 1/5 length of hindwing. Male genitalia. Uncus bell-shaped. Apical process of gnathos small, triangular or clubbed, with a small hook distally. Transtilla fused, usually a narrow arched band. Valva with clasper; costa and sacculus strongly sclerotized, extended to end of valva. Vinculum U-shaped, longer than width. Juxta usually U-shaped, lateral lobes various in shape, globular or finger like, or fused and extremely extended to middle of uncus. Aedeagus slender, with one sinuate sclerite; cornuti a cluster of spines or thorns. Culcita present. Female genitalia. Anal papillae triangular. Eighth abdominal segment short and wide, collar-shaped. Apophyses anteriores and posteriores short, nearly equal in length. Antrum rectangular. Ductus bursae mostly sclerotized. Corpus bursae membranous, elongate elliptic. Signa usually comprised of a transverse row of flag-shaped or conical spines. Ductus seminalis incepted at junction of ductus bursae and corpus bursae. Distribution. China, Japan, Thailand, Malaysia, Indonesia, Brunei, India, Sri Lanka, Australia, Papua New Guinea. Key to Chinese Cathyalia species (based on male genitalia). 1. Forewing with a broad white streak from basal 1/5 to apex along costa (Figs 1, 5); transtilla without process, clasper simple in the male genitalia; ostium at intersegmental membrane between seventh and eighth segments, ductus bursae without thorn in female genitalia............................................................................................................................................................................................ 2 Forewing without broad white streak along costa (Fig. 3); transtilla with three short processes on posterior margin, clasper complex in the male genitalia (Fig. 8); ostium at center of seventh segment, ductus bursae with two sharp thorns posterolaterally in female genitalia (Fig. 11)................................................................................................................................................................ C. fulvella 2. Labial palpus with terminal two segments about equal length (Fig. 1a); apical process of gnathos small triangular, juxta with lateral lobes fused and extremely extended to middle of uncus in male genitalia (Fig. 7); ductus bursae about half length of corpus bursae, strongly sclerotized except with a narrow membranous ring at 1/3, corpus bursae not scobinate in female genitalia (Fig. 10).................................................................................................................................................................................................. C. okinawana Labial palpus with second segment about 2.2 times length of third segment (Fig. 5a); apical process of gnathos long clubbed, juxta with lateral lobes separated, not extended backward in male genitalia (Fig. 9); ductus bursae about same length of corpus bursae, sclerotized in posterior 4/5, membranous in anterior 1/5, corpus bursae scobinate posterolaterally in female genitalia (Fig. 12)............................................................................................................................................................................................... C. pallicostella, Published as part of Ren, Yingdang & Yang, Linlin, 2017, Genus Cathyalia Ragonot, 1888 (Lepidoptera: Pyralidae), new record phycitine to China, pp. 514-520 in Zoological Systematics 42 (4) on pages 514-515, DOI: 10.11865/zs.201724, http://zenodo.org/record/4617043

Ren, Yingdang, Yang, Linlin (2017): Genus Cathyalia Ragonot, 1888 (Lepidoptera: Pyralidae), new record phycitine to China. Zoological Systematics 42 (4): 514-520, DOI: 10.11865/zs.201724, URL: http://zoobank.org/0b9c4c6c-c8b7-4ca0-969b-177ac60391dd

Cathyalia fulvella Ragonot, 1888 new record to China (Figs 3���4, 8, 11) Cathyalia fulvella Ragonot, 1888: 7; Roesler & K��ppers, 1981: 14; Roesler, 1983: 79; Robinson, Tuck & Shaffer, 1994: 159; Shaffer, Nielsen & Horak, 1996: 177. Spatulipalpia erythrina Hampson, 1896: 103. Synonymized by Roesler & K��ppers, 1981: 14. Cryptoblabes rufimarginella Pagenstecher, 1900: 166. Synonymized by Shaffer, Nielsen & Horak, 1996: 177. Trissonca clytopa Turner, 1947: 52. Synonymized by Shaffer, Nielsen & Horak, 1996: 177. Adult (Figs 3���4). Wingspan 12.0���15.0 mm. Vertex yellowish brown, frons ocherous brown. Antenna with scape yellowish brown, flagellum yellowish brown to grayish brown. Labial palpus (Fig. 3a) with basal segment grayish white, terminal two segments about same length, pale yellow on inner side, grayish brown tined with ocherous brown on outer side. Maxillary palpus yellowish brown. Thorax, tegula and patagium purplish brown. Forewing blackish brown, tined with ocherous brown; markings indistinctive, antemedial line, discal spot, postmedial line obscure; a large, rounded-triangular patch on dorsum between antemedial and postmedial lines, colored yellowish white mixed with brown; an arched, yellowish brown stripe aside postmedial line, parallel with termen; terminal line black; cilia grayish brown. Hindwing pale orangey yellow, grayish yellow along costa till apex, orangey yellow along veins; cilia orangey yellow. Foreleg with coxa yellowish white on ventral surface, femur yellowish brown mixed with blackish brown, tibia and tarsus blackish brown; midleg and hindleg yellowish white on inner side, blackish brown on outer side, gray to dark red at end of each tarsal segment. Abdomen orangey yellow. Male genitalia (Fig. 8). Uncus bell-shaped. Apical process of gnathos clubbed, 1.2 times length of uncus, hooked at apex. Transtilla n-shaped on anterior margin, with three short processes at middle and lateral sides of posterior margin. Valva broad in basal half, apical half about 1/2 width of basal half, apex rounded, densely setose in apical 2/5; costa slightly sinuate, triangularly protruded at basal 1/3, humped at basal 2/3; clasper well developed, presented as a smooth sclerite extending from basal 1/3 of costa to end of sacculus, then angled and outwardly stretched to apical 1/6 of valva along ventral margin, with a lobate, setose process at base; sacculus stout, clubbed, 3/5 length of valva, rounded-triangularly protruded near end. Vinculum 1.8 times as long as wide. Lateral lobes of juxta stout, with globular, setose apex. Aedeagus slender, about same length of valva; cornuti consisted of a twist sclerotized crimple at base, and three or four clusters of spinule distally. Culcita one pair. Female genitalia (Fig. 11). Ostium at center of seventh segment, surrounded sections strongly sclerotized in flabellate shape. Antrum semicircular. Ductus bursae slightly broader, membranous, twist in anterior 1/3 slightly; narrowed, strongly sclerotized in posterior 2/3, with two slender lateral thorns at posterior margin in V shape. Corpus bursae elongate, oval, slightly longer than ductus bursae, crimpled at junction with ductus bursae. Signa about 10 flag-shaped sclerites located at sclerotized ovate plates, obliquely arranged in posterior 2/5 of corpus bursae. Material examined. Hainan, 4♂ 9♀ (NKUM); Yunnan, 2♂ 4♀ (NKUM). Distribution. China (Hainan, Yunnan), Thailand, Malaysia, Indonesia, Brunei, India, Sri Lanka, Australia, Papua New Guinea., Published as part of Ren, Yingdang & Yang, Linlin, 2017, Genus Cathyalia Ragonot, 1888 (Lepidoptera: Pyralidae), new record phycitine to China, pp. 514-520 in Zoological Systematics 42 (4) on page 517, DOI: 10.11865/zs.201724, http://zenodo.org/record/4617043

Ren, Yingdang, Yang, Linlin, Li, Houhun (2015): Taxonomic review of the genus Indomyrlaea Roesler & Küppers 1979 of China, with descriptions of five new species (Lepidoptera: Pyralidae: Phycitinae). Zootaxa 4006 (2): 311-329, DOI: http://dx.doi.org/10.11646/zootaxa.4006.2.4

Indomyrlaea fastigipalpa Ren & Li sp. nov. (Figs 6, 7, 22, 27) Diagnosis. This new species is similar to I. sutasoma, but can be distinguished from the latter by the labial palpus with second segment 3 times length of the diameter of compound eye in the male; the gnathos with medial lobe twice length of lateral lobe, the aedeagus with dense short thorns forming two triangular areas near apex in the male genitalia; and the antrum 2.5 times as wide as long, H-shaped in the female genitalia. In I. sutasoma, the second segment of the labial palpus is twice length of the diameter of compound eye in the male; the medial lobe of the process of gnathos is the same length as the lateral lobe, the aedeagus has sparse dentations near apex in the male genitalia; and the antrum is 1.5 times as wide as long, almost rectangular in the female genitalia. Description. Wingspan 23.0���24.0 mm. Male (Fig. 6): Vertex with two cluster brownish long scales surrounded by antenna, and with yellowish brown scales between compound eyes. Antenna with scape twice as long as wide, blackish brown on dorsal surface, yellowish white on ventral surface; flagellum dark brown, sinus shallow, covered with vertically arranged black scale tufts, and a row of short, black toothlet beneath scale tufts. Labial palpus yellowish white; second segment elongate, vertically upturned, 3.0 times length of diameter of compound eye; third segment short, bend forward, about 1 / 15 length of second segment. Maxillary palpus yellowish white. Patagium, tegula and thorax coppery brown. Forewing ground color yellowish brown, mixed with blackish, whitish and ocherous brown scales; basal 1 / 4 with dense, thick scales, its anterior 1 / 3 black, posterior 2 / 3 grayish black, yellowish white at borderline; antemedian line and discal spot absent; postmedial line grayish white, sinuate, margined with black on both sides, angled inward at R 4 and CuA 2; terminal line black; cilia yellowish brown. Hindwings grayish brown, opaque; cilia yellowish white. Foreleg ocherous brown; midleg yellowish white, tarsus mixed with ocherous brown; hindleg pale yellowish brown, tibia with long scales on dorsal surface. Female (Fig. 7): Vertex with grayish brown lobate scales. Antenna with scape twice as long as wide, yellowish brown; flagellum yellowish brown and ocherous brown. Labial palpus yellowish white; second segment 1.5 times length of diameter of compound eye; third segment about 2 / 5 length of second segment, tinged with reddish brown on outer surface. Maxillary palpus yellowish white. Patagium, tegula and thorax fuscous grayish brown. Forewing ochreous brown, mixed with numerous reddish brown and blackish brown scales on lower half and between postmedial line and terminal line; antemedian line absent; discal spot faint, blackish brown; postmedial line grayish brown, sinuate, concave at R 4 and CuA 2, margined with blackish brown; terminal line black; cilia brown. Hindwing grayish brown, opaque; cilia yellowish brown. Legs gray, mottled dark purple. Male genitalia (Fig. 22). Uncus bell-shaped, rounded at apex. Gnathos trilobate, anchor-shaped, lateral lobes crescent, medial lobe cuniform, twice length of lateral lobe. Transtilla weakly sclerotized. Valva broad in basal 2 / 5, narrowed in distal 3 / 5 and about half width of basal 2 / 5, rounded apically; bearing dense long hair medially from basal 1 / 5 to apex; costa strongly sclerotized, about half width of valva at base, tapered to apex; sacculus strongly sclerotized, narrowed, clubbed, 3 / 4 length of valva, straight in basal 2 / 3, curvedly concave in distal 1 / 3, with a strong hooked process distally. Vinculum U-shaped, 1.4 times as long as wide. Juxta U-shaped, shield-shaped on median area, with a pair of fingerlike lateral lobes bearing sparse setae. Aedeagus cylindrical, stout, strongly sclerotized, about 1.3 times length of valva, with dense short thorns forming two triangular areas near apex; cornuti comprised of numerous spinule, small triangular thorns and sclerotized crimples: an arched, spined band at base, followed by a bunch of needlelike thorns, a bunch of subuliform thorns from basal 1 / 3 to middle, numerous granular spines from middle to apex. Culcita four pairs, three-dimensional arranged. Female genitalia (Fig. 27). Antrum 2.5 times as wide as long, H-shaped. Ductus bursae sclerotized and punctate, membranous and smooth at junction of ductus bursae and corpus bursae. Corpus bursae twice length of ductus bursae, circled dense long bodkin-shaped spines at middle, anterior half membranous and smooth, posterior half thickened and weakly sclerotized. Caudal bursa bearing minute spines distally. Ductus seminalis incepted end of caudal bursa. Type material. Holotype ♂, China: Mt. Jianfeng (18 �� 50 'N 108 �� 43 'E), Hainan, 960 m, 13���15 -IV- 2013, coll. Yingdang Ren & Xiaoguang Liu, gen. slide No. RYD 2012155 (HAASM). Paratypes: China: Hainan: 1 ♀, same data as for holotype, gen. slide No. RYD 20120256 (HAASM); 2 ♀, Mt. Diaoluo (18 �� 39 'N 109 �� 54 'E), 930 m, 9 -IV- 2013, coll. Yingdang Ren & Xiaoguang Liu, gen. slide Nos. RYD 20120258 ♀, RYD 20120261 ♀ (HAASM); 1 ♂, Mt. Limu (19 ��09'N 109 �� 28 'E), Qiongzhong, 640 m, 5 -V- 2014, coll. Tengteng Liu, Wei Guan & Xuemei Hu, gen. slide No. LHX 14189 (NKUM). Distribution. China (Hainan). Etymology. The specific name is derived from the Latin prefix fastigatus -, meaning upturned, rising to a point, and morphological term palpus, referring to the upturned labial palpus with third segment pointed at apex in male., Published as part of Ren, Yingdang, Yang, Linlin & Li, Houhun, 2015, Taxonomic review of the genus Indomyrlaea Roesler & K��ppers 1979 of China, with descriptions of five new species (Lepidoptera: Pyralidae: Phycitinae), pp. 311-329 in Zootaxa 4006 (2) on pages 319-323, DOI: 10.11646/zootaxa.4006.2.4, http://zenodo.org/record/244837

Indomyrlaea sutasoma Roesler & K��ppers, 1979 (Figs. 2, 3, 19, 20) Indomyrlaea sutasoma Roesler & K��ppers, 1979: 119; Roesler, 1983: 55. Sandrabatis crassiella Ragonot, 1893 (nec): Hampson, 1896: 75; Roesler & K��ppers, 1979: 107; Roesler, 1983: 53. Misidentification. Diagnosis. Wingspan 22.0���24.5 mm (Figs. 2, 3). Indomyrlaea sutasoma can be recognized by the following characters: the base of the forewing with dense and thick scales; the triangular uncus pointed at apex, the valva with broadened, rounded apex, the terminal of costa bluntly protruded, the sacculus stretching to 2 / 5 of the valva, with a small hook distally, the stout aedeagus bearing sparse dentations apically, cornuti comprising of numerous spinule and sclerotized crimples in the male genitalia (Fig. 19); the gourd-shaped corpus bursae with lots of long bodkin-shaped spines surrounding near middle, and a caudal bursa protruding on posterior margin in the female genitalia (Fig. 20). Material examined. CHINA: Yunnan: 1 ♂, 1 ♀, Bubang (21 �� 36 'N 101 �� 35 'E), Mengla, 650 m, 24 -VIII- 2005, coll. Yingdang Ren, gen. slide Nos. RYD04713 m, RYD04714 f (NKUM); 1 ♀, Rare Botanical Garden (24 ��00��N 97 �� 50 ��E), Ruili, 1000 m, 06-VIII- 2005, coll. Yingdang Ren, gen. slide No. RYD04764 (NKUM). Xizang: 1 ♀, Yadong (27 �� 19 ���N 88 �� 35 ���E), 2950 m, 27 -VIII- 2003, coll. Xinpu Wang & Huaijun Xue (NKUM). Distribution. China (Xizang, Yunnan), Indonesia (Sumatra), India, Sikkim, Bhutan, Sir Lanka. Remarks. Indomyrlaea sutasoma is recorded for the first time from China., Published as part of Ren, Yingdang, Yang, Linlin & Li, Houhun, 2015, Taxonomic review of the genus Indomyrlaea Roesler & K��ppers 1979 of China, with descriptions of five new species (Lepidoptera: Pyralidae: Phycitinae), pp. 311-329 in Zootaxa 4006 (2) on page 315, DOI: 10.11646/zootaxa.4006.2.4, http://zenodo.org/record/244837, {"references":["Roesler, R. U. & Kuppers, P. V. (1979) Die Phycitinae (Lepidoptera: Pyralidae) von Sumatra; Taxonomie Teil A. Beitrage zur Naturkundlichen Forschung in Sudwestdeutschland, 3, 1 - 249.","Roesler, R. U (1983) Die Phycitinae von Sumatra (Lepidoptera: Pyralidae). Heterocera Sumatrana, 3, 1 - 136, pls. 1 - 69.","Ragonot, E. L. (1893) Monographie des Phycitinae et des Galleriinae. In: Romanoff, N. M. (Ed.), Memoires sur les Lepidopteres, VII, pp. i - lvi, 1 - 658, pls 1 - 23. [St. Ptersburg] http: // dx. doi. org / 10.5962 / bhl. title. 10956","Hampson, G. F. (1896) Moths. The Fauna of British India, including Ceylon and Burma, 4, i - xxviii, 1 - 594. http: // dx. doi. org / 10.5962 / bhl. title. 100797"]}

3. Indomyrlaea kalmasapada Roesler & Küppers, 1979 Indomyrlaea kalmasapada Roesler & Küppers, 1979: 120; Roesler, 1983: 56. TL: Sumatra. TD: Holotype (♂) in LNK. Distribution: Indonesia (Sumatra).

Indomyrlaea bannensis Ren & Li sp. nov. (Figs. 4, 5, 21, 26) Diagnosis. This new species is similar to I. kalmasapada, but can be distinguished from the latter by the valva with the costa produced to a small pointed process apically, the sacculus with a minute pointed end separated from the valva, and the aedeagus with two cornuti in the male genitalia. In I. kalmasapada, the costa is not protruded apically, the sacculus is neither pointed nor separated from the valva at end, and the aedeagus has one cornutus in the male genitalia. Description. Wingspan 21 ���22.5 mm. Male (Fig. 4): Vertex yellowish white except grayish brown on inner side of antenna. Antenna with scape 1.3 times as long as wide, yellowish white; flagellum blackish brown, sinus shallow, covered with vertically arranged yellowish brown scale tufts, and a row of short, black toothlet beneath scale tufts. Labial palpus yellowish white except brown at base of second segment; second segment 1.5 times length of diameter of compound eye; third segment short, slightly bend forward, about 1 / 3 length of second segment. Maxillary palpus yellow. Occiput, patagium and tegula dark reddish brown, thorax gray. Forewing brown, with sparse yellowish white scales, with two white spots at middle of dorsum, inner one bigger than outer one; antemedian line and discal spot absent; postmedial line grayish white, faint, sinuate, interrupted; terminal line black, interrupted; cilia brown. Hindwing pale yellow, with a tuft of yellowish bristle-like scales along Sc+Rs; cilia yellowish brown. Foreleg and midleg pale brown, hindleg yellowish white, tibia with a tuft of yellowish white long hairs at base on outer surface, tarsus blackish brown, margined yellowish white distally. Abdomen yellowish brown on dorsal surface, yellowish white on ventral surface of third and fourth segments, with two yellowish white anal tufts. Female (Fig. 5): Vertex with forward directed, fuscous brown lobate scales. Antenna with scape twice as long as wide, yellowish white mixed with fuscous brown; flagellum yellowish brown and blackish brown. Labial palpus yellowish white; second segment tinged with fuscous brown on outer surface; third segment about 1 / 3 length of second segment, fuscous brown on basal half. Maxillary palpus yellowish white. Patagium, tegula and thorax fuscous grayish brown. Forewing ochreous brown, mixed with yellowish white; antemedian line, discal spot absent; postmedial line faint, sinuate; terminal line black; cilia brown. Hindwing gray; cilia yellowish brown. Legs yellow on dorsal surface, yellowish brown on ventral surface; tarsomeres yellowish white at end. Male genitalia (Fig. 21). Uncus triangular. Gnathos anchor-shaped, lateral lobes crescent, medial lobe clubbed, about 1.5 times length of lateral lobe. Transtilla weakly sclerotized. Valva with costal and ventral margins parallel sided in basal 1 / 3, protruded at middle on ventral margin, slightly broadened in distal 1 / 4; bearing dense long hair in distal 1 / 4 and upper margin of sacculus, with stout subuliform spines along distal 1 / 5 of ventral margin; costa strongly sclerotized, about half width of valva, slightly narrowed toward apex, protruded a small pointed process apically; sacculus strongly sclerotized, half length of valva, cuniform, pointed and separated from valva at end. Vinculum U-shaped, 1.2 times as long as wide. Juxta U-shaped, membranous on median area, with a pair of well-developed fingerlike lateral lobes bearing sparse setae. Aedeagus cylindrical, slightly longer than valva, stout in basal 1 / 4, spinose distally; cornuti comprised of one long, clubbed thorn, about 3 / 5 length of aedeagus, and one short, sinuate spine near apex. Culcita five pairs, four pairs present on 8 th segment, three-dimensional arranged, another pair on 7 th segment. Female genitalia (Fig. 26). Antrum twice wider than long, bowl-shaped. Ductus bursae membranous in posterior 1 / 4, weakly sclerotized in anterior 3 / 4, with a trapezoidal, punctate area on inner wall. Corpus bursae pyriform, with dense long bodkin-shaped spines in posterior 2 / 5, numerous small, stout spines near junction with ductus bursae. Caudal bursa punctate on inner wall caudally. Ductus seminalis incepted end of caudal bursa. Type material. Holotype ♂, China: Menglun (21 �� 42 ��N 101 �� 23 ��E), Mengla, Xishuangbanna, Yunnan, 640 m, 13 -VIII- 2010, coll. Yunhui Sun & Lixia Li, gen. slide No. LHX 12615 (NKUM). Paratypes: China: Yunnan: 1 ♀, same data as for holotype except dated 12 -VIII- 2010, gen. slide No. LHX 11038 (NKUM); 1 ♂, same place as for holotype, 21-30 -IV- 1974, coll. Yao Zhou, Feng Yuan & Yinyue Hu, gen. slide No. RYD04763 (NKUM). Distribution. China (Yunnan). Etymology. The specific name is derived from the type locality Xishuangbanna, and Latin postfix - iensis., Published as part of Ren, Yingdang, Yang, Linlin & Li, Houhun, 2015, Taxonomic review of the genus Indomyrlaea Roesler & K��ppers 1979 of China, with descriptions of five new species (Lepidoptera: Pyralidae: Phycitinae), pp. 311-329 in Zootaxa 4006 (2) on pages 318-319, DOI: 10.11646/zootaxa.4006.2.4, http://zenodo.org/record/244837

5. Indomyrlaea phaeella (Hampson, 1903) comb. nov. Sandrabatis phaeella Hampson, 1903: 25. TL: Sikkim. TD: BMNH. Distribution: Sikkim., Published as part of Ren, Yingdang, Yang, Linlin & Li, Houhun, 2015, Taxonomic review of the genus Indomyrlaea Roesler & K��ppers 1979 of China, with descriptions of five new species (Lepidoptera: Pyralidae: Phycitinae), pp. 311-329 in Zootaxa 4006 (2) on page 314, DOI: 10.11646/zootaxa.4006.2.4, http://zenodo.org/record/244837, {"references":["Hampson, G. F. (1903) The moths of India. Supplementary paper to the volumes in \" The fauna of British India. \" Series II. Part IX, X. Journal of the Bombay Natural History Society, 15, 19 - 37 (part IX) & 206 - 226 (part X)."]}

Indomyrlaea sinuapalpa Ren & Li sp. nov. (Figs 9, 25) Diagnosis. This new species is similar to I. nigra Ren & Li sp. nov., but can be distinguished by the labial palpus with second segment sinuate in S shape in the male, the valva with distal part hammerheaded, and the aedeagus with a conic thorn-like cornutus near basal 3 / 7 in the male genitalia. In I. nigra Ren & Li sp. nov., the second segment of labial palpus is not sinuate in S shape, the distal part of the valva is acinaciform, and the conic thornlike cornutus is near apex of the aedeagus. Description. Male (Fig. 9): Wingspan 23.5���24.0 mm. Vertex with scales yellowish brown in basal half, grayish brown in distal half. Antenna with scape twice as long as wide, ferruginous brown on dorsal surface, yellowish brown on ventral surface; flagellum dark brown, sinus gentle, with yellowish brown, vertical arranged scale tufts, and a row of short, black toothlet under scale tufts. Labial palpus yellowish white; second segment elongate, 2.5 times length of diameter of compound eye, vertically upturned, sinuate in S shape; third segment short, slightly bend forward, about 1 / 15 length of second segment. Maxillary palpus yellowish white. Patagium, tegula and thorax ocherous brown. Forewing ground color yellowish brown, gradually deeper from base toward apex, mixed with numerous ferruginous brown scales, with black scales along veins; scales white-tipped between postmedial line and terminal line; basal 1 / 4 with an elliptic area bearing dense, thick scales from lower margin of cell to dorsum; antemedian line absent; discal spots fused, black; postmedial line yellowish white, sinuate, margined with blackish brown on both sides, angled inward at R 4 and CuA 2; terminal line black; cilia ferruginous brown. Hindwings yellowish brown, opaque; cilia yellowish white. Foreleg and midleg blackish brown mixed with ocherous brown; hindleg ocherous brown on dorsal surface, blackish brown mixed with ocherous brown on ventral surface, tibia with long yellowish brown scales on dorsal surface. Male genitalia (Fig. 25). Uncus bell-shaped, blunt at apex, evenly bearing dense, short setae in distal half on dorsal surface. Gnathos trilobate, anchor-shaped, lateral lobes crescent, medial lobe fingerlike, half length of lateral lobe. Transtilla weakly sclerotized. Valva slightly narrowed to middle, somewhat hammerheaded in distal part: narrowed and about half width of base from middle to distal 1 / 6, broadened in distal 1 / 6, rounded dorsal-apically, triangularly protruded ventral-apically; bearing dense long hair distally and along dorsal margin of sacculus from basal 1 / 5 to end; costa strongly sclerotized, about half width of valva, base and apex broader than middle, archedly concave medially; sacculus narrowed, 3 / 4 length of valva, straight in basal 4 / 5, deep concave in distal 1 / 5, with a slender, hooked process distally. Vinculum U-shaped, 1.5 times as long as wide. Juxta U-shaped, shield-shaped on median area, with a pair of fingerlike lateral lobes bearing sparse setae. Aedeagus cylindrical, stout, strongly sclerotized, about 1.4 times length of valva, with dense short thorns forming a quadrangular and an arched band near apex; cornuti comprised of numerous spinule and sclerotized crimples, arranging in wheatear-shape: an arched, spined band at base, a cluster of spines near this, a conic thorn at basal 3 / 7, and a helical sclerotized crimple surrounded by dense granulae from basal 2 / 5 to apex. Culcita four pairs, three-dimensional arranged. Female unknown. Type material. Holotype ♂, China: Taiyanghe (30 �� 26 'N 119 �� 34 'E), Puer, 1600 m, Yunnan, 11 -VII- 2013, coll. Zhenguo Zhang, gen. slide No. LHX 14195 (NKUM). Paratype: China: Yunnan: 1 ♂, same data as for holotype, gen. slide No. LHX 13372 (NKUM); 2 ♂, Baihualing, Mt. Gaoligong, Baoshan, 1473 m, 29 -VII- 2013, coll. Linlin Yang, gen. slide Nos. RYD 20120252, RYD 20120253 (HAASM). Distribution. China (Yunnan). Etymology. The specific name is derived from the Latin prefix sinua -, meaning sinuate, and morphological term palpus, referring to the condition that the second segment of labial palpus is sinuate and in S shape in male., Published as part of Ren, Yingdang, Yang, Linlin & Li, Houhun, 2015, Taxonomic review of the genus Indomyrlaea Roesler & K��ppers 1979 of China, with descriptions of five new species (Lepidoptera: Pyralidae: Phycitinae), pp. 311-329 in Zootaxa 4006 (2) on pages 326-329, DOI: 10.11646/zootaxa.4006.2.4, http://zenodo.org/record/244837

Indomyrlaea Roesler & Küppers, 1979 Indomyrlaea Roesler & Küppers, 1979: 118; Roesler, 1983: 55. Type species: Indomyrlaea sutasoma Roesler & Küppers, 1979, by original designation and monotypy. Diagnosis. This genus is sexually dimorphic: in the male, the flagellum basely has a shallow sinus, on which a row of black toothlet is covered by rough scale tufts, the labial palpus is elongate and obviously upturned beyond vertex, and the base of the forewing usually has dense and thick scales; in the female, the flagellum is common, the labial palpus is not elongate, and the base of the forewing is not thickened. The genus is also characterized by the male genitalia with the process of the gnathos trilobate, somewhat anchor-like, and the female genitalia with posterior half of corpus bursae weakly sclerotized, and ornamented with numerous long spines. Generic characters. Adults (Figs 2−12). Frons rounded, with rough scales. Ocellus present. Chaetosema well developed. Antenna (Fig. 14) extends about half forewing; scape elongate; male flagellum with a gentle sinus, bearing a rough scale tuft and a row of black toothlet hiding in the tuft. Labial palpus upturned; in male (Fig. 15), second segment stout and vertically upturned beyond head, with a groove on inner surface; third segment very short and slightly bent forward; in female, second segment gently bend, without groove, third segment slightly longer than male, acuminate apically. Maxillary palpus in male (Fig. 16) pappi-shaped, same length and hiding in groove of second segment of labial palpus; in female cylindrical, fanned out distally. Tongue well developed. Forewing (Figs 17, 18) with dense and thick scales at base in male; with 11 vines, R 2 stalked with R 3 + 4 in basal half, R 3 + 4 and R 5 stalked for about 2 / 3 length of R 5, M 2 and M 3 shortly stalked, CuA 1 and M 2 + 3 from lower angle of cell in male, from lower margin but close to lower angle of cell in female, CuA 2 free; cell longer than half length of forewing. Hindwing with 10 vines: Sc close to or shortly stalked with Rs, M 1 from upper angle of cell, shortly stalked with Sc+Rs, M 2 and M 3 long stalked, CuA 1 and M 2 + 3 stalked beyond cell for half length of CuA 1, CuA 2 from lower angle of cell; cell shorter than 1 / 4 of hindwing; anal area in male bearing long scales. Legs with tarsi having two lines of black spines. Male genitalia (Figs 19, 21 − 25). Uncus triangular to trapezoidal. Apical process of gnathos trilobate, anchorshaped. Transtilla not or weakly sclerotized. Valva narrowly elongate, bearing dense long hair distally and near ventral margin; costa broad, strongly sclerotized, extending to end of valva, with or without distal process; sacculus narrow, strongly sclerotized, usually with a hooked process distally; clasper absent. Vinculum well developed, Ushaped, longer than width. Juxta U-shaped, weakly sclerotized on median area, with a pair of fingerlike lateral lobes. Aedeagus cylindrical, stout; cornuti present. Culcita present on 8 th segment of abdomen, usually four pairs, three-dimensional arranged. Female genitalia (Figs 20, 26−28). Anal papillae triangular. 8 th sternite short and wide, collar-shaped. Apophyses anteriores and apophyses posteriores nearly same length. Antrum strongly sclerotized. Ductus bursae shorter than corpus bursae, partly sclerotized. Corpus bursae elongate, oval, posterior half usually weakly sclerotized, with a caudal bursa near junction of ductus bursae and corpus bursae. Signum comprised of numerous long spines. Ductus seminalis incepted at end of caudal bursa. Distribution. China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Xizang, Yunnan, Zhejiang), Bhutan, India, Indonesia (Sumatra), Japan, Korea, Sikkim, Sri Lanka. Included species

Key to the species of Indomyrlaea based on male 1. Forewing with two distinct black discal spots, and a black spot near basal 1 / 3 of vein A (Fig. 12).............. I. phaeella - Forewing without spot described as above................................................................ 2 2. Juxta with median area almost membranous; aedeagus slender, with one or two clubbed cornuti, without microcornuti arranged in wheatear-shape.................................................................................... 3 - Juxta with median area strongly sclerotized; aedeagus stout, with numerous microcornuti arranged in wheatear-shape, without clubbed cornutus..................................................................................... 4 3. Costa of valva with a process apically; sacculus of valva pointed and separated from valva at end (Fig. 21).................................................................................................... I. bannensis sp. nov. - Costa of valva without process apically; sacculus is neither pointed nor separated from the valva at end (Roesler 1983, Tafel 36: 22.54)............................................................................... I. kalmasapada 4. Uncus triangular; the arms of gnathos connect to invert-V-shaped.............................................. 5 - Uncus bell-shaped or trapezoid; the arms of gnathos connect to arch-shaped...................................... 6 5. Sacculus extends no more than half of valva, with a small hook apically (Fig. 19)......................... I. sutasoma - Sacculus extends more than half of valva, with a large hook apically (Fig. 24).................. I. proceripalpa sp. nov. 6. Gnathos with medial lobe twice length of lateral lobe; aedeagus without conic, thorn-like cornutus (Fig. 22)................................................................................................ I. fastigipalpa sp. nov. - Gnathos with medial lobe about same length of lateral lobe; aedeagus with a conic, thorn-like cornutus................ 7 7. Distal part of valva acinaciform; aedeagus with the thorn-like cornutus near apex (Fig. 23).............. I. nigra sp. nov. - Distal part of valva hammerheaded; aedeagus with the thorn-like cornutus at basal 3 / 7 (Fig. 25)...... I. sinuapalpa sp. nov.

Indomyrlaea Roesler & K��ppers, 1979 Indomyrlaea Roesler & K��ppers, 1979: 118; Roesler, 1983: 55. Type species: Indomyrlaea sutasoma Roesler & K��ppers, 1979, by original designation and monotypy. Diagnosis. This genus is sexually dimorphic: in the male, the flagellum basely has a shallow sinus, on which a row of black toothlet is covered by rough scale tufts, the labial palpus is elongate and obviously upturned beyond vertex, and the base of the forewing usually has dense and thick scales; in the female, the flagellum is common, the labial palpus is not elongate, and the base of the forewing is not thickened. The genus is also characterized by the male genitalia with the process of the gnathos trilobate, somewhat anchor-like, and the female genitalia with posterior half of corpus bursae weakly sclerotized, and ornamented with numerous long spines. Generic characters. Adults (Figs 2���12). Frons rounded, with rough scales. Ocellus present. Chaetosema well developed. Antenna (Fig. 14) extends about half forewing; scape elongate; male flagellum with a gentle sinus, bearing a rough scale tuft and a row of black toothlet hiding in the tuft. Labial palpus upturned; in male (Fig. 15), second segment stout and vertically upturned beyond head, with a groove on inner surface; third segment very short and slightly bent forward; in female, second segment gently bend, without groove, third segment slightly longer than male, acuminate apically. Maxillary palpus in male (Fig. 16) pappi-shaped, same length and hiding in groove of second segment of labial palpus; in female cylindrical, fanned out distally. Tongue well developed. Forewing (Figs 17, 18) with dense and thick scales at base in male; with 11 vines, R 2 stalked with R 3 + 4 in basal half, R 3 + 4 and R 5 stalked for about 2 / 3 length of R 5, M 2 and M 3 shortly stalked, CuA 1 and M 2 + 3 from lower angle of cell in male, from lower margin but close to lower angle of cell in female, CuA 2 free; cell longer than half length of forewing. Hindwing with 10 vines: Sc close to or shortly stalked with Rs, M 1 from upper angle of cell, shortly stalked with Sc+Rs, M 2 and M 3 long stalked, CuA 1 and M 2 + 3 stalked beyond cell for half length of CuA 1, CuA 2 from lower angle of cell; cell shorter than 1 / 4 of hindwing; anal area in male bearing long scales. Legs with tarsi having two lines of black spines. Male genitalia (Figs 19, 21 ��� 25). Uncus triangular to trapezoidal. Apical process of gnathos trilobate, anchorshaped. Transtilla not or weakly sclerotized. Valva narrowly elongate, bearing dense long hair distally and near ventral margin; costa broad, strongly sclerotized, extending to end of valva, with or without distal process; sacculus narrow, strongly sclerotized, usually with a hooked process distally; clasper absent. Vinculum well developed, Ushaped, longer than width. Juxta U-shaped, weakly sclerotized on median area, with a pair of fingerlike lateral lobes. Aedeagus cylindrical, stout; cornuti present. Culcita present on 8 th segment of abdomen, usually four pairs, three-dimensional arranged. Female genitalia (Figs 20, 26���28). Anal papillae triangular. 8 th sternite short and wide, collar-shaped. Apophyses anteriores and apophyses posteriores nearly same length. Antrum strongly sclerotized. Ductus bursae shorter than corpus bursae, partly sclerotized. Corpus bursae elongate, oval, posterior half usually weakly sclerotized, with a caudal bursa near junction of ductus bursae and corpus bursae. Signum comprised of numerous long spines. Ductus seminalis incepted at end of caudal bursa. Distribution. China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Xizang, Yunnan, Zhejiang), Bhutan, India, Indonesia (Sumatra), Japan, Korea, Sikkim, Sri Lanka. Included species, Published as part of Ren, Yingdang, Yang, Linlin & Li, Houhun, 2015, Taxonomic review of the genus Indomyrlaea Roesler & K��ppers 1979 of China, with descriptions of five new species (Lepidoptera: Pyralidae: Phycitinae), pp. 311-329 in Zootaxa 4006 (2) on pages 313-314, DOI: 10.11646/zootaxa.4006.2.4, http://zenodo.org/record/244837, {"references":["Roesler, R. U. & Kuppers, P. V. (1979) Die Phycitinae (Lepidoptera: Pyralidae) von Sumatra; Taxonomie Teil A. Beitrage zur Naturkundlichen Forschung in Sudwestdeutschland, 3, 1 - 249.","Roesler, R. U (1983) Die Phycitinae von Sumatra (Lepidoptera: Pyralidae). Heterocera Sumatrana, 3, 1 - 136, pls. 1 - 69."]}

Indomyrlaea proceripalpa Ren & Li sp. nov. (Figs 10, 11, 14���18, 24, 28) Sandrabatis crassiella Ragonot, 1893 (nec): Inoue, 1955: 135; 1982, 1: 391, 2: 250; 2013, 4: 357; Choi, et. al., 1998: 25; Bae, 2004: 94; Bae, 2008: 294. Misidentification. Diagnosis. This new species is similar to I. fastigipalpa Ren & Li sp. nov., but can be distinguished by the forewing with dense scales area grayish black; the gnathos with medial lobe slightly longer than lateral lobe, the aedeagus with dense short thorns forming a semi-ring near apex in the male genitalia; and the corpus bursae with a long, strongly sclerotized, handle-like process on posterior margin in the female genitalia. In I. fastigipalpa Ren & Li sp. nov., the forewing dense scales area is black in anterior 1 / 3, grayish black in posterior 2 / 3, yellowish white at borderline; medial lobe of the gnathos is twice length of lateral lobe, the aedeagus has dense short thorns forming two triangular areas near apex in the male genitalia; and the corpus bursae has no process in the female genitalia. It is also extremely similar to I. sutasoma, but the hooked process at the end of sacculus of the valva, and the handlelike process on the corpus bursae distinguish the new species from I. sutasoma. Description. Wingspan 18.0���26.5 mm. Male (Fig. 10): Vertex with long, black scales, mixed with yellowish white on outside, arranged in two whorls. Antenna with scape twice as long as wide, basely with blackish brown long scale tufts on dorsal surface, yellowish white on ventral surface; flagellum coppery brown, sinus gentle, with black, vertical arranged scale tufts, and a row of short, black toothlet under scale tufts. Labial palpus yellowish white; second segment elongate, vertically upturned, 3.0 times length of diameter of compound eye; third segment short, slightly bend forward, about 1 / 15 length of second segment. Maxillary palpus yellowish white. Patagium, tegula and thorax grayish brown. Forewing ground color yellowish brown, mixed with blackish brown, white and reddish brown scales; basal 1 / 4 with dense, thick, grayish black scales; antemedian line and discal spot absent; postmedial line grayish white, sinuate, margined with black on both sides, angled inward at R 4 and CuA 2; terminal line black; cilia yellowish brown. Hindwings grayish brown, opaque; cilia yellowish white. Foreleg reddish brown, bearing a tuft of long scales; midleg grayish brown; hindleg yellowish white, tibia with long scales on dorsal surface. Abdominal segments yellowish brown on dorsal surface, yellowish white on ventral surface; anal tuft yellowish white. Female (Fig. 11): Vertex with yellowish white lobate scales, mixed with ochreous brown. Antenna with scape twice as long as wide, yellowish white to ochreous brown; flagellum yellowish brown to fuscous brown. Labial palpus yellowish white, mixed with reddish brown scales; second segment 1.6 times length of diameter of compound eye; third segment about half length of second segment, pointed at apex. Maxillary palpus yellowish white. Patagium yellowish white to reddish brown, tegula and thorax grayish brown to reddish brown. Forewing dark brown, mixed with numerous reddish brown scales on lower half and beyond postmedial line, tinged with yellowish white near dorsum; antemedian line absent; discal spots blackish brown, fused into club-shape; postmedial line grayish white, sinuate, margined with black on both sides, angled inward at R 4 and CuA 2; terminal line black; cilia brown. Hindwings grayish brown, opaque; cilia yellowish white. Legs gray, mottled grayish white; tarsi with two lines of black spines. Abdominal segments ocherous yellow, mixed with brown on dorsal surface. Male genitalia (Fig. 24). Uncus triangular, rounded at apex, evenly bearing dense, short setae. Gnathos trilobate, anchor-shaped, lateral lobes crescent, medial lobe slightly longer than lateral lobe. Transtilla weakly sclerotized. Valva broad in basal 2 / 5, narrowed from 2 / 5 to 4 / 5, slightly broadened in distal 1 / 5, triangularly protruded ventral-apically; bearing dense long hair in distal 1 / 5 and along dorsal margin of sacculus; costa strongly sclerotized, about half width of valva, archedly concave medially; sacculus strongly sclerotized, 3 / 5 length of valva, with a strong hooked process distally. Vinculum U-shaped, 1.3 times as long as wide. Juxta U-shaped, shield-shaped on median area, with a pair of fingerlike lateral lobes bearing sparse setae. Aedeagus cylindrical, stout, strongly sclerotized, about 1.2 times length of valva, with dense short thorns forming a semi-ring near apex; cornuti comprised of numerous spinule and sclerotized crimples, arranging in wheatear-shape: an arched, spined band at basal 1 / 4, a sinuate band bearing subuliform thorns from basal 1 / 3 to distal 1 / 3, one or two irregular sclerite at distal 1 / 4. Culcita four pairs, three-dimensional arranged. Female genitalia (Fig. 28). Antrum twice wider than long, protruded and punctate on lateral sides. Ductus bursae membranous in posterior 1 / 3, weakly sclerotized in anterior 2 / 3, punctate on inner wall. Corpus bursae membranous in anterior half, sclerotized in posterior half, with a line of long bodkin-shaped spines medially, dense small spines on junction between ductus bursae and corpus bursae, with a long, strongly sclerotized, handle-like process on posterior margin. Caudal bursa protruded between handle-like process and ductus bursae. Ductus seminalis incepted end of caudal bursa. Type material. Holotype ♂, China: Linan, Mt. Tianmu (30 �� 26 'N 119 �� 34 'E), Zhejiang, 350 m, 9 -VIII- 2007, coll. Qing Jin, gen. slide No. LHX 12167 (NKUM). Paratypes: China: Anhui: 2 ♀, Wenquan (30 �� 51 ��N 116 �� 21 ��E), Yuexi, 7-8 -VIII- 1995, coll. Xiaofu Hu, gen. slide No. KDH05122 (NKUM); 1 ♂, Tanqiao town (30 �� 10 ��N 118 ��07��E), Huangshan, 6 -VIII- 2004, coll. Jiasheng Xu & Jialiang Zhang, gen. slide No. RYD04715 (NKUM); 2 ♀, Kecun, Mt. Jiuhua, (30 �� 32 ��N 117 �� 47 ��E), 160m, 8 - VIII- 2004, coll. Jiasheng Xu & Jialiang Zhang, gen. slide No. KDH05120 (NKUM); 1 ♂, 4 ♀, Jiuhuajie, Mt. Jiuhua, 650m, 9 -VIII- 2004, coll. Jiasheng Xu & Jialiang Zhang, gen. slide No. LHX 14191 f (NKUM). Fujian: 1 ♂, Guadun (27 �� 44 ��N 117 �� 38 ��E), Mt. Wuyi, 1100m, 29 -VII- 2008, coll. Weichun Li, Yongling Sun & Haiyan Bai, gen. slide No. LHX 12065 (NKUM); 1 ♂, Mt. Daiyun (25 �� 40 ��N 118 �� 12 ��E), 902 m, 23 -V- 2012, coll. Jinwei Li (HAASM). Guangdong: 1 ♂, 1 ♀, Babaoshan (24 �� 24 ��N 113 ��08��E), Nanling, 1070 m, 23 -VIII- 2010, coll. Shulian Hao, gen. slide No. LHX 12227 (NKUM); 2 ♂, Yanshuitian (23 �� 37 ��N 111 �� 52 ��E), Fengkai, 2 -V- 2011, coll. Dandan Zhang & Bo Tong, gen. slide No. LHX 14193 (NKUM); 1 ♂, same place as former, 6 -X- 2011, coll. Bo Tong (NKUM); 1 ♀, same place as former, 3 -VIII- 2011, coll. Jiaqin Xie & Bo Tong (NKUM). Guangxi: 1 ♂, Dongzhong Yard (21 �� 37 ��N 107 �� 32 ��E), Fangchenggang, 370 m, 9 -IV- 2002, coll. Shulian Hao & Huaijun Xue, gen. slide No. LHX 14222 (NKUM); 1 ♂, Hongqi Yard (21 �� 54 ��N 107 �� 54 ��E), Shangsi, 260 m, 1 -IV- 2002, coll. Shulian Hao & Huaijun Xue (NKUM); 1 ♂, 1 ♀, Xiangfen, Rongshui (25 �� 20 'N 109 �� 19 'E), 650 m, 19 -VII- 2004, coll. Jiasheng Xu, gen. slide. No. RYD04641 (NKUM); 1 ♀, Huaping (24 �� 51 'N 106 �� 23 'E), 950m, 7 -VIII- 2006, coll. Weichun Li, gen. slide No. LHX 12235 (NKUM); 1 ♂, Mt. Daming (23 �� 24 'N 108 �� 30 'E), 1250m, 26 -V- 2011, coll. Linlin Yang & Yinghui Mou, gen. slide No. LHX 14223 (NKUM); 1 ♀, same data as former except dated 27 -V- 2011 (NKUM); 1 ♀, Shaoping Yard (22 ��08'N 106 �� 44 'E), Pingxiang, 280m, 22 -V- 2012, coll. Xiaofei Yang, gen. slide No. LHX 14224 (NKUM); 4 ♂, Nonggang, Longzhou (22 �� 28 'N 106 �� 57 'E), 271 m, 19 -IV- 2012, coll. Jinwei Li, gen. slide No. LHX 14194 (NKUM). Guizhou: 1 ♀, Mt. Chengjia, Daozhen (28 �� 61 ��N 107 �� 36 ��E), 1300 m, 19 - VIII- 2004, coll. Yunli Xiao, gen. slide No. RYD04268 (NKUM); 8 ♂, Sanchahe (25 �� 15 ��N 108 ��07��E), Libo, 500 m, 11���14 -V- 2012, coll. Yingdang Ren & Xiaoguang Liu (HAASM). Hainan: 1 ♀, Shuimanxiang (18 �� 53 ��N 109 �� 40 ��E), Mt. Wuzhi, 640 m, 21 -V- 2007, coll. Zhiwei Zhang & Weichun Li, gen. slide No. LHX 14197 (NKUM); 1 ♂, Shuimanxiang, Mt. Wuzhi, 1 -VI- 2010, coll. Bingbing Hu & Jing Zhang, gen. slide No. LHX 14190 (NKUM). Henan: 1 ♂, 1 ♀, Taiping, Xixia (33 �� 18 ��N 111 �� 29 ��E), 974 m, 7 -VII- 2013, coll. Chunyang Yi (HAASM); 2 ♂, 2 ♀, Mt. Liankang, Xinxian (31 �� 38 ��N 114 �� 48 ��E), 600 m, 6 -VIII- 2013, coll. Chunyang Yi (HAASM); 1 ♀, Baotianman (33 �� 12 ��N 111 �� 53 ��E), Neixiang, 1300 m, 1 -VIII- 2012, coll. Yingdang Ren & Chunyang Yi (HAASM). Hubei: 1 ♂, Songbai Town (31 �� 42 �� N 110 �� 36 �� E), Shennongjia, 260 m, 17 -VII- 2003, coll. Shulian Hao (NKUM); 6 ♂, Wujiashan (31 ��05�� N 115 �� 48 �� E), Yingshan, 8,10 - VIII- 2004, coll. Yunli Xiao, gen. slide Nos. LHX 14192, LHX 14198 (NKUM); 1 ♀, same data as former except dated 15 -VII- 2011, gen. slide No. LHX 12616 (NKUM); 1 ♀, Mt. Jiugong (29 �� 23 �� N 114 �� 39 �� E), Xianning, 26 -VII- 2011, coll. Yunli Xiao, gen. slide No. LHX 14210 (NKUM). Hunan: 2 ♀, Lianhe, Chengguan, Taojiang (28 �� 30 ��� N 112 ��09��� E), 26,27 - VII- 2004, coll. Yunli Xiao, gen. slide No. LHX 14220 (NKUM); 1 ♀, Shuikou, Weishan, Xinhua (27 �� 42 �� N 111 �� 18 �� E), 4 -VIII- 2004, coll. Yunli Xiao, gen. slide No. LHX 14221 (NKUM); 1 ♀, Baituowuqiao, Xiantan City (27 �� 45 ��N 112 �� 43 ��E), 29 -VII- 2004, coll. Yunli Xiao (NKUM). Jiangxi: 1 ♂, Taohongling (29 �� 51 ��N 116 �� 33 ��E), 420 m, 9 -VIII- 2012, coll. Chunyang Yi & Wenhao Nan (HAASM); 3 ♂, Mt. Qiyun (25 �� 52 ��N 114 ��00��E), Chongyi, 650 m, 5 -V- 2007, coll. Jiasheng Xu (NKUM); 1 ♀, Linkesuo, Ganzhou (25 �� 46 ��N 115 ��00��E), 16 -VII- 2006, coll. Jiasheng Xu & Weichun Li (NKUM); 1 ♀, Mt. Jiulian (28 �� 42 ��N 114 �� 58 ��E), 20 -VII- 2006, coll. Jiasheng Xu & Weichun Li, gen. slide No. LHX 14212 (NKUM); 1 ♂, Shangfuzhen, Fengxinxian (28 �� 39 ��N 115 ��00��E), 108 m, 21 -IX- 2012, coll. Jinwei Li, gen. slide No. LHX 14211 (NKUM). Shaanxi: 1 ♂, Tielingpu (33 �� 37 ��N 110 �� 31 ��E), Danfeng, 680 m, 29 -V- 1994, coll. Jin Zhou, gen. slide No. LHX 14225 (NKUM); 2 ♂, Zhanghe (32 ��06��N, 108 �� 56 ��E), Langao, 800 m, 15 -VII- 2013, coll. Xiaoguang Liu & Chunyang Yi (HAASM). Sichuan: 2 ♀, Haichao, Luxian (28 �� 58 ��N 105 �� 17 ��E), 25���27 -VII- 1995, coll. Yongxing Zeng, gen. slide No. RYD04607 (NKUM). Zhejiang: 1 ♂, same data as for holotype, gen. slide No. LHX 14214 (NKUM); 2 ♂, Xianrending, Mt. Tianmu, 1500 m, 18 -VIII- 1999, coll. Houhun Li, gen. slide Nos. DYL00145, RYD04505 (NKUM); 3 ♀, Chanyuansi, Mt. Tianmu, 350 m, 15,20 - VIII- 1999, coll. Houhun Li, gen. slide Nos. RYD04598, KDH05112 (NKUM); 1 ♀, Houshanmen, Mt. Tianmu, 500 m, 16 -VIII- 1999, coll. Houhun Li, gen. slide No. LHX 14215 (NKUM); 1 ♀, Mt. Wuyan (27 �� 21 ��N, 119 �� 55 ��E), Taishun, 680 m, 02-VIII- 2005, coll. Yunli Xiao, gen. slide No. LHX 14216 (NKUM); 2 ♂, 1 ♀, Zhonglieci, Mt. Tianmu, 400 m, 27 -VII, 2 -VIII- 2011, coll. Linlin Yang & Na Chen, gen. slide Nos. LHX 12308 m, LHX 12041 f (NKUM); 1 ♂, Xianrending, Mt. Tianmu, 1500 m, 25 -VII- 2011, coll. Linlin Yang & Na Chen (NKUM); 1 ♂, 1 ♀, Longtangshan (30 �� 21 ��N 118 �� 52 ��E), Mt. Qingliang, 500 m, 21, 22 -V- 2012, coll. LinlinYang & Zhenguo Zhang, gen. slide No. LHX 14217 m (NKUM). Distribution. China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Zhejiang), Japan, Korea. Etymology. The specific name is derived from the Latin prefix procer -, meaning elongate, and morphological term palpus, referring to the elongate labial palpus in male., Published as part of Ren, Yingdang, Yang, Linlin & Li, Houhun, 2015, Taxonomic review of the genus Indomyrlaea Roesler & K��ppers 1979 of China, with descriptions of five new species (Lepidoptera: Pyralidae: Phycitinae), pp. 311-329 in Zootaxa 4006 (2) on pages 324-326, DOI: 10.11646/zootaxa.4006.2.4, http://zenodo.org/record/244837, {"references":["Ragonot, E. L. (1893) Monographie des Phycitinae et des Galleriinae. In: Romanoff, N. M. (Ed.), Memoires sur les Lepidopteres, VII, pp. i - lvi, 1 - 658, pls 1 - 23. [St. Ptersburg] http: // dx. doi. org / 10.5962 / bhl. title. 10956","Inoue, H. (1955) Check List of the Lepidoptera of Japan. Vol 2. Rikusuisha, Tokyo, pp. 112 a - 112 b, 113 - 217.","Choi, H. Y., Paek, M. K. & Bae, Y. S. (1998) Taxonomic Notes on Nine Species of the Tribe Phycitini (Lepidoptera, Pyralidae, Phycitinae) from Korea (I). Insecta Koreana, 15, 23 - 39.","Bae, Y. S. (2004) Superfamily Pyraloidea II (Phycitinae & Crambinae etc.). Economic Insects of Korea 22. Insecta Koreana Supplement 29. National Institute of Agricultural Science and Technology, Suwon, 207 pp.","Bae, Y. S., Byun, B. K. & Paek, M. K. (2008) Pyralid Moths of Korea (Lepidoptera: Pyraloidea). Korea National Arboretum, SAMSUNGAD. Com., Seoul, 426 pp."]}