Search

Löbl, Ivan (2023): Assessing Earth's biotic diversity in natural history museums - gone with the wind? Phegea 51, No. 2: 50-51, DOI: 10.6084/m9.figshare.22722733, URL: http://dx.doi.org/10.6084/m9.figshare.22722733

Members of the scaphidiine tribe Scaphisomatini collected recently in Tianmushan Biosphere Reserve (Zhejiang) and Wuyishan National Nature Reserve (part of Wuyishan Biosphere Reserve, Fujian), China, are studied. Two species, Scaphisoma krali Löbl, sp. nov. and Scaphisoma sekerkai Löbl, sp. nov., are described from Wuyishan. Baeocera franzi (Löbl, 1973) and Scaphisoma mutator Löbl, 2000 are recorded for the first time from Zhejiang province, and Scaphisoma binhanum (Pic, 1922) and Scaphoxium intermedium Löbl, 1984 are recorded for the first time from Fujian province.

Many members of our society ask in formal and informal meetings what is going on in natural history museums and especially in insect collections: little activity, little motivation, and little support. The editor-in-chief of Phegea asked an expert in museum insect collections for his opinion. Dr. Ivan Löbl is based at the natural history museum in Geneva and edited an impressive series of catalogues of Palaearctic Coleoptera https://brill.com/display/serial/CPC

Sapitia versicolor (Pic, 1920) Material examined. 474, Sabah, Batu Punggul Resort env., 24.vi.̅1.vii.1996, 11c, FIT (MHNG); 3, Sabah, Danum Valley, 14̅ 16.ii.1968, G. de Rougemont (MHNG); 1, Sabah, Poring Hot Springs, 500 m, 11.v.1987, Burckhard & Löbl (MHNG); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Danum Valley Conservation Area, Borneo Rain Forest Lodge, N 05°02.682′ E 117°45.553′, 250 m, iii̅ iv.2005, FIT, primary dipterocarp forest, E. Slade & J. Villanuava lgt. OUMNH 2055 ­062 (OUMNH). Remarks. Thie species appears to be widely distributed and has been reported from Indonesia: Sumatra; East Malaysia: Sabah and Thailand (see LÖBL 2018). It is very similar to S. sumatrana Löbl, 1978 while it differs notably from S. lombokina by the nearly fused mesepimera and the stoud legs., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 16, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2018: Coleoptera: Staphylinidae: Scaphidiinae. World Catalogue of Insects. Volume XVI: i ‾ xvi + 1 ‾ 418 pp."]}

Scaphicoma rufa (Pic, 1923) Material examined. 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 450 m, 22̅ 23.xi.2005, primary dipterocarp forest, west trail area, first inferior FIT, D. Mann, E. Slade & J. Villanuava lgt., OUMNH 2006 ­051 (OUMNH); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, E. Slade & J. Villanuava lgt., selectively logged dipterocarp forest, Yayasan Sabah Logging Concesion, OUMNH 2005 ­052 (MHNG). Remarks. These specimens exhibit characters congruent with the redescription given by LÖBL & LESCHEN (2010). Notable are the widely interrupted basal striae of the elytra, the mesoventrite with distinct mesal ridge, the lateral parts of the metaventrite lacking coarser punctures, and the lateral parts of ventrite I, each bearing a subbasal patch of distinct punctures, in combination with the uniformly rufous, about 1.9 mm long body. The species is very similar to S. arcuatum Champion, 1927 and S. pulex Heller, 1917., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 17, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & LESCHEN R. A. B. 2010: Notes on the Toxidium group (Coleoptera: Staphylinidae: Scaphidiinae). Folia Heyrovskyana Series A 18 (1 - 3): 71 ‾ 93."]}

Pseudobironium rapax sp. nov. (Figs 8 ̅10) Type material. Holotype male, Malaysia, Sabah, Crocker Range, Mawar Waterfall env. 17.VI.1996, 9c, vegetation debris and forest floor litter accumulated along large fallen trees near river (MHNG). Description. Length 1.95 mm, width 1.30 mm. Head and most of body dark reddishbrown, apex of abdomen lighter, yellowish; femora and tibiae roughly as body, tarsi and antennae light brown. Maxillary palpi with palpomere IV narrow, tapering, about 4 times as long as wide. Length ratio of antennomeres as follows: III 15: IV 22: V 26: VI 21: VII 26: VIII 26: IX 27: X: 26: XI 35; antennomere XI about 3.5 times as long as wide. Pronotal punctation very fine, similar to that on frons, with punctures very shallow and not well delimited; middle of pronotal disc with puncture intervals about 4 to 10 times larger than puncture diameters. Elytra lacking lateral impressions, not flattened or impressed apically, lacking humps; discal punctation similar to pronotal. Hypomera impunctate, impressed posteriad. Mesoventrite lacking mesal ridge and lacking distinct setiferous punctures. Mesanepisternum very finely punctate, lacking microsculpture. Metaventrite lacking microsculpture; median area nearly flat, lacking impressions and tubercle, with two irregular rows of distinct admesal punctures. Lateral parts of metaventrite very finely and sparsely punctate. Metacoxal process impunctate, horizontal. Submesocoxal area short, about as fourth shortest interval to metacoxa. Submesocoxal line finely punctate. Mesotibiae and metatibiae slightly bent, ventral spur of mesotibiae straight. Ventrite Ilacking microsculpture, with very fine and scattered punctures; following ventrites and exposed tergites with punctulate microsculpture. Male. Aedeagus (Figs 8 ̅10) 0.74 mm long. Median lobe with basal bulb shorter than apical process, the latter slightly inflexed, with sinuate ventral margin (lateral view), and blunt tip; apical part of median lobe lacking distinct valves. Parameres nearly evenly wide, bent in dorsal view, straight in lateral view. Internal sac with pair of strongly sclerotized denticles and mesal sclerite, lacking flagellum and lacking apical spinose structures. Etymology. The species epithet is a Latin noun, referring to the canine­like denticles of the internal sac. Differential diagnosis. The species is as P. irregulare less than 2 mm long and thus conspicuous by its small body­size. It may be easily distinguished from P. irregulare by the evenly very fine elytral punctation, the narrow submesocoxal areas and ventrite I lacking microsculpture. The aedeagal characters suggest relationship with members of the P. subovatum group (see LÖBL & TANG 2013), though the shape of the admesal sclerites of the internal sac is unique. The species falls in the key species to couplet 27, to P. schuhi Löbl & Tang, 2013 from which it differs notably by the parameres evenly narrow, except at their basis. Remarks. The single available specimen has its forelegs broken, probably lost when collected., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 14-15, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Mordelloscaphium Pic, 1915 The genus is monospecific. It was established to accommodate a species possessing characters typical for the termitophilous members of the Baeoceridium group (see LESCHEN & LÖBL 2005) and thus is assumed to live on fungus gardens of termites., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LESCHEN R. A. B. & LOBL I. 2005: Phylogeny and classification of Scaphisomatini Staphylinidae: Scaphidiinae with notes on mycophagy, termitophily, and functional morphology. Coleopterists Society Monographs 3: 1 ‾ 63."]}

Bironium borneense Löbl, 1987 Material examined. 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 200 m, 27̅ 28.x.2005, primary forest, D. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, West trail 10 FIT, OUMNH 2006­051 (OUMNH); 1, with the same data but 21.X.2005, West trail area (MHNG); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, primary forest, Slade & J. Villanuava lgt. lowland mixed dipterocarp forest, Yayasan Sabah Logging Concession OUMNH 2005­062 (OUMNH). Remarks. The species is known only from East Malaysia (LÖBL et al. 2020). It was based on specimens found in the Gunung Mulu National Park in Sarawak and subsequently recorted from the Mount Kinabalu National Park, the vicinity of Tambunan and the Batu Punggul Resort in Sabah., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I., LESCHEN R. A. B. & KODADA J. 2020: Review of the Asian species and cladistic analysis of Bironium Csiki (Coleoptera: Staphylinidae: Scaphidiinae) with comments on biogeography. Annales Zoologici (Warszawa) 70 (4): 711 ‾ 736."]}

Vituratella Reitter, 1908 The genus encompasses 31 species, all Afrotropical but the Asian V. termitophila (Champion, 1927) and V. kistneri (Löbl, 1979)., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 18, DOI: 10.5281/zenodo.7501705

Scaphicoma Motschulsky, 1863 The genus comprises 19 Afrotropical, Oriental, Melanesian and Australian species. Several, described by J. Achard and M. Pick, still lack revisions. Asingle species is present in samples from Sabah., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 17, DOI: 10.5281/zenodo.7501705

Baeocera incisa (Löbl, 1973) Material examined. 1 male, Sabah, Lahad Datu Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57′ E 117°48′, 450 m, 1̅ 5.ix.2005, Slade & Villanuava lgt., lowland mixed dipterocarp forest. Nature trail area 1, FIT, OUMNH (OUMNH). Remarks. The species is widely distributed in Sabah and seems to be common in samples of forest floor litter (see LÖBL 2021: 28)., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2021: On the Bornean species of Scaphoxium Lobl, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beitage 53 (1): 85 ‾ 89."]}

Baeocera doriae (Pic, 1020) Material examined. 1 male, Sabah, Mt. Kinabalu N. P., 1700 m, 16.vii.02, Kurbatov & Zimina (MHNG); 1 male, Sabah, rd Kota Kinabalu ̅ Tambunan, km 52, 1600̅ 1800 m, 2̅ 5.vii.02, Kurbatov & Zimina (MHNG). Remarks. This species is known from Sarawak and Sabah only. It is poorly represented in collections, a single specimen has been reported from each of these states., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 9, DOI: 10.5281/zenodo.7501705

Baeocera hutan sp. nov. (Figs 1 ̅4) Type material. Holotype male, Sabah: Lahad Datu Ulu Segama forest Reserve, Danum Valley For. Centre, N 04°57.9′ E 117°48.1′, 200 m, 21.xi.2005, D. Mann, E. Slade & J. Villanuava lgt., lowland mix. dipterocarp forest. West trail area 1, FIT, OUMNH 2006­051 (OUMNH). Description. Length 1.28 mm, width 0.76 mm. Head and body nearly evenly reddishbrown, elytra hardly darker than pronotum, apical abdominal segments slightly lighter than pronotum, femora and tibiae as elytra, tarsi and antennae lighter. Head with interocular distance nearly as dorsoventral eye diameter. Head, thorax and elytra lacking microsculpture. Length/width ratios of antennomeres as follows: III 20 /5: IV 20 /6: V 30 /7: VI 27 /7: VII 33 /8: VIII 27 /7: IX 36 /11: X: 37/15: XI 48 /17. Pronotal and elytral punctation even, very fine. Lateral contours of pronotum and elytra continuously, very weakly arcuate. Pronotum with anterior margin bead broadly interrupted, lateral margins sinuate in lateral view, lateral margin striae impunctate. Margin of exposed part of scutellum rounded. Elytra with lateral margins nearly parallel, lateral margins in lateral view slightly sinuate, lateral margin carinae visible in dorsal view, lateral margin striae impunctate; sutural striae starting at base, parallel, deeply impressed, curved along pronotal lobe to form basal striae extended to humeral areas, broadly separated from lateral striae; discal punctation sparse, near base with puncture intervals about 8 to 10 larger than puncture diameters. Hind wing fully developed. Hypomera and mesanepisterna appearing impunctate. Mesoventrite with mesal ridge, punctate along lateral margins. Mesepimeron about 4 times as long as large and 5 times as long as interval to mesocoxa. Median part of metaventrite nearly flat, impunctate mesally, with dense, fairly coarse punctures on posterior third and laterally smooth mesal area, bearing conspicuous, long pubescence. Lateral parts of metaventrite with few scattered extremely fine punctures. Submesocoxal line parallel, margined with row of coarse elongate punctures extended posterior the inner part of mesepimeron. Submesocoxal area about 0.02 mm long. Metanepisterna large, about 0.10 mm wide, twice as long as wide, slightly convex, with inner margin broadly rounded anteriorly, nearly straight posteriad; inner suture impunctate. Protibiae straight. Mesotibiae and metatibiae slightly bent. Ventrite I lacking striae, with row of elongate basal punctures not interrupted in middle, extended laterally behind tip of metepimeron; punctation posterior basal row very fine and sparse. Male. Protarsomeres Ito III strongly widened, bearing long tenent setae; protarsomere I wider than apex of protibia. Ventrite VI with apical lobe subtriangular, about 0.04 mm long. Aedeagus (Figs 1 ̅4) 0.60 mm long. Median lobe and parameres strongly sclerotized, symmetrical, dorsal valves excepted. Articular process not prominent, distinct. Apical process of median lobe somewhat shorter than basal bulb, moderately inflexed, with weakly concave ventral margin, gradually narrowed apically, tip acute in lateral view, obtuse in dorsal view. Parameres robust, gradually narrowed apically in dorsal view, widened apical in lateral view, with broadly rounded apex. Internal sac with complex sclerites, lacking spinose and denticulate structures, with apical section of ejaculatory duct wide, weakly membranous, extruded in repos. Etymology. The species epithet is a noun meaning "forest̏ in Malay. Differential diagnosis. This species is unique in having broadly interrupted pronotal bead. Therefore, it is placed in Baeocera tentatively (the mouth­parts and thorax of the unique available specimen were not dissected). In addition, its aedeagal characters are unusual, in particular by the structure of the internal sac and the asymmetrical valves of the median lobe. The internal sac lacking a flagellar guide sclerite and possessing a membranous ejaculatory duct extruded in repos is shared with the Chinese B. xichangana Löbl, 1999. It differs notably from B. xichangana by the not prominent articular process, the two asymmetrical dorsal valves, and the much shorter, robust parameres. Both species are also very distinct in external characters, in particular by their body­shape and size (B. xichangana is 2.0̅ 2.15 mm long). Remarks. As to the Sabah species, only B. barbara Löbl, 1990 and B. doriae (Pic, 1920) share the wide metanepisterna. These two species have asymmetrical aedeagus and may be easily distinguished by their external characters. The antitheses 7 and 8 have been inversed in key to the Sabah Baeocera given by LÖBL & SMETANA (2021): the thesis 6 (Metanepisternum fused,...) should lead to ̤ 8., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 8-9, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & SMETANA A. 2021: On the Baeocera Erichson (Coleoptera: Staphylinidae: Scaphidiinae) of Sabah, Malaysia, and a tale on mystified biodiversity. Journal of Insect Biodiversity 23 (2): 23 ‾ 42."]}

4. Key to the Baeocera species of the People’s Republic of China 1 Lateral parts of metaventrite impunctate, or very finely and sparsely punctate...................2 - Lateral parts of metaventrite coarsely, usually densely punctate, or at least with several conspicuous, coarse punctures...........................................................................................14 2 Elytra with basal striae interrupted, not joined to lateral striae. Exposed part of scutellum large....................................................................................................................3 - Elytral with basal striae entire, joined to lateral striae. Scutellum concealed or minute, except in B. breveapicalis...................................................................................................6 3 Apical part of median lobe large, not overlapped, about as long as basal bulb. Male metatibiae each with subapical denticle........................................ B. satana NAKANE, 1963 - Apical part of median lobe small, to part or entirely overlapped, much shorter than basal bulb. Male metatibiae lacking denticles.....................................................................4 4 Aedeagus with parameres simple, hook-like and curved at apices....... B. gilloghyi LÖBL, 1973 - Aedeagus with parameres lobed, not hook-like at apices....................................................5 5 Left parameres with small dorsal lobe and ventrally inflexed apex; right parameres lobed ventrally........................................................................... B. yunnanensis LÖBL, 1999 - Left parameres with large dorsal lobe and hardly inflexed apex; right parameres not lobed............................................................................................ B. pubiventris LÖBL, 1990 6 Antennomere VI short and wide, about 2 times as long as wide and as long as half of antennomere VII..................................................................................................................7 - Antennomere VI long and narrow, about 3 to 5 times as longs as wide, slightly shorter than antennomere VII..........................................................................................................8 7 Aedeagus with outer margin of left paramere sinuate and inner lobe subtriangular............................................................................................................... B. breveapicalis (PIC, 1926) - Aedeagus with outer margin of left paramere convexly rounded and inner lobe blunt................................................................................................................ B. takizawai LÖBL, 1984 8 Aedeagus with parameres strongly expanded apico-dorsally to form large, strongly sclerotized lobes................................................................................. B. kubani LÖBL, 1999 - Aedeagus with parameres not lobed, or each with small subapical lobe.............................9 9 Aedeagus with apices of parameres distinctly widened (dorsal view)....................................................................................................................................... B. huanshana LÖBL, 1999 - Aedeagus with apices of parameres not or hardly widened (dorsal view).........................10 10 Aedeagus lacking strongly sclerotized guide-sclerite, parameres slightly shorter than median lobe................................................................................ B. xichanagna LÖBL, 1999 - Aedeagus lacking with strongly sclerotized guide-sclerite, parameres much shorter than median lobe.......................................................................................................................11 11 Aedeagus with guide-sclerite narrow, angulate basally...................... B. callida LÖBL, 1986 - Aedeagus with guide-sclerite robust, not angulate basally................................................12 12 Aedeagus with guide-sclerite tapering apicad........................................ B. freyi LÖBL, 1966 - Aedeagus with guide-sclerite not tapering apicad.............................................................13 13 Aedeagus with guide-sclerite straight, becoming stouter apicad........ B. cooteri LÖBL, 1999 - Aedeagus with guide-sclerite abruptly curved in middle, widened subapically, acute at tip................................................................................................. B. hammondi LÖBL, 1984 14 Elytra with sutural striae shortened, starting well behind level of scutellar tip, far from elytral base........................................................................................................................15 - Elytra with sutural striae starting at elytral base, usually curved along pronotal lobe and forming basal striae...........................................................................................................16 15 Pronotal punctation coarse and dense, centre of metaventrite with mesal row of coarse punctures and narrow impunctate admesal areas................................ B. karenovicsi nov.sp. - Pronotal punctation very fine and sparse, hardly visible, metaventrite lacking mesal row of coarse punctures....................................................................... B. proclinata nov.sp. 16 Elytra with basal striae widely separated from lateral striae.............................................17 - Elytra with basal striae entire, joined to lateral striae........................................................19 17 Elytral punctation coarse anterior mid-length, very fine posterior mid-length or in apical thirds. Aedeagus with notched parameres...............................................................18 - Elytral punctation coarse on entire discal surfaces. Aedeagus with straight, not notched parameres.................................................................................................. B. repleta nov.sp. 18 Internal sac of aedeagus bearing tuft of spines.................................. B. franzi (LÖBL, 1973) Internal sac of aedeagus lacking tuft of spines.......................... B. proseminata LÖBL, 2013 19 Pronotal and elytral punctation similar, very fine..............................................................20 - Pronotal punctation very fine, elytral punctation in part or entirely coarse.......................21 20 Aedeagus with flagellum curved and thickened basally. Lateroposterior part of metaventrite coarsely punctate................................................... B. sordidoides LÖBL, 1992 - Aedeagus with flagellum almost straight in dorsal view, not thicken basally. Lateroposterior part of metaventrite very finely punctate.................. B. coalita LÖBL, 2013 21 Metanepisterna separated from metaventrite by distinct sutures.......................................................................................................................................... B. serendibense (LÖBL, 1971) - Metanepisterna fused with metaventrite............................................................................22 22 Punctation coarse on most of basal half to two thirds of elytral disc, apical third to half of elytral disc very finely punctate, appearing impunctate................................................23 - Elytra entirely or nearly entirely coarsely punctate...........................................................24 23 Parameres narrow, not folded apically............................................... B. erroris LÖBL, 1990 - Parameres wide, folded apically................................................ B. pseudinculta LÖBL, 1990 24 Ventrite 1 very finely punctate posterior basal puncture row, appearing impunctate............................................................................................................. B. longicornis (LÖBL, 1971) - Ventrite 1 coarsely punctate posterior basal puncture row................................................25 25 Apical halves of parameres distinctly bent and with weakly sclerotized inner margins................................................................................................................... B. pigra (LÖBL, 1971) - - Parameres straight, with inner and outer margins nearly evenly sclerotized...................26 26 Parameres almost as long as median lobe. Internal sac without narrow lateral rod............................................................................................................................ B. vidua LÖBL, 1990 - Parameres much shorter than median lobe. Internal sac with narrow lateral rod.............................................................................................................................. B. lasciva LÖBL, 2013, Published as part of Löbl, Ivan, 2018, Supplement to the knowledge of the genera Baeocera ERICHSON, 1845 and Scaphobaeocera CSIKI, 1909 (Coleoptera, Staphylinidae, Scaphidiinae) of the People's Republic of China, pp. 1295-1303 in Linzer biologische Beiträge 50 (2) on pages 1299-1300, DOI: 10.5281/zenodo.5275102, {"references":["LOBL I. (1999): A review of the Scaphidiinae (Coleoptera: Staphylinidae) of the People's Republic of China, I. - Revue suisse zool. 106: 691 - 744.","LOBL I. (1992): The Scaphidiidae (Coleoptera) of the Nepal Himalaya. - Revue suisse zool. 99: 471 - 627.","LOBL I. (1971): Die Scaphidiidae (Coleoptera) Sudindiens. - Revue s uisse zool. 86: 77 - 129."]}

Pseudobironium subovatum Pic, 1920 Material examined. 1 male, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, primary forest, E. Slade & J. Villanuava lgt. Lowland mixed dipterocarp forest, Yayasan Sabah Logging Concession OUMNH 2005­062 (OUMNH). Remarks. The species seems to be rather widely distributed. It is currently known from Indonesia (Sumatra and Kalimantan), Malaysia (West Malaysia, Sabah and Sarawak), and Thailand, though only a single Sabah record is published from the Gunung Emas in the Crocket Range (LÖBL & TANG 2013). The species may be easily distinguished by the long protibial setae in males., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 11, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Mordelloscaphium testaceimembre Pic, 1915 Material examined. 184, Sabah, Batu Punggul Resort env., 24.vi. ̅ 1.vii.1996, FIT (MHNG); 1, with the same data but in debris accumulates around large trees near river (MHNG); 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 200 m, 22̅ 23.x.2005, primary forest, D. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, west trail area, FIT, OUMNH 2006­051 (OUMNH); 1, with the same data but 3̅13.xi., D.J. Mann & E. Slade lgt,, forest natural trail area OUMNH 2065­051 (OUMNH); 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Conservation Area, Borneo Rain Forest Lodge, N 05°02.682′ E 117°45.553′, 250 m, iii̅ iv.2005, E. Slade & J. Villaneuve lgt., FIT, primary dipterocarp forest, OUMNH 2065­062 (OUMNH). Remarks. The currently known specimens of this species are the syntypes from east Kalimantan, Indonesia., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 10-11, DOI: 10.5281/zenodo.7501705

Pseudobironium Pic, 1920 The genus is Asian in distribution, with most members found in the tropics and subtropics, remaining unknown south of the Wallace line. Currently, six species are known to occur in Sabah (LÖBL & TANG 2013): P. convexum Löbl & Tang, 2013, P. horaki Löbl & Tang, 2013, P. pubiventer Löbl & Tang, 2013, P. sparsepunctatum (Pic, 1915), P. subovatum Pic, 1920, and P. vitalisi (Achard, 1920). Anew record is given, and two additional species are described below., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 11, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Scaphoxium sinuatum sp. nov. Figs 11 ̅13 Type material. Holotype male, Malaysia, Sabah, Batu Punggul Resort env., 24.VI. ̅ 1.VII.1996, 11c. vegetation debris and forest floor litter accumulated around large trees near river (MHNG). Description. Length 1.18 mm, width 0.55 mm, dorsoventral diameter 0.62 mm. Thorax, elytra and most of abdomen uniformly reddish­brown, head somewhat lighter, apical abdominal segments light, nearly yellowish, femora and tibiae roughly as thorax, tarsi lighter, antennae light brown. Length/width ratios of antennomeres as follows: III 22 /5: IV 17 /4: V 22 /4: VI 23 /4: VII 28 /7: VIII 20 /7: IX 28/11: X: 28/11: XI 38 /12. Pronotal and elytral punctation similar, very fine, hardly visible at 40 times magnification. Scutellum concealed. Elytra with sutural striae parallel, shortened, starting about 0.15 mm posterior of margin of pronotal lobe, adsutural areas hardly raised. Hypomeron with scattered, very fine punctured and short oblique stria. Mesoventrite with mesal impression shallow, narrowed posteriad, lacking mesal ridge and lacking microsculpture. Margin of mesocoxal process rounded. Metaventrite lacking microsculpture, very finely punctate, convex in middle. Lateral stria oblique, nearly reaching convex, finely punctate submesocoxal line. Submesocoxal area about 0.03 mm, as long as half of the shortest interval to margin of metacoxa. Metanepisternum flat, with suture fairly narrow, punctate, somewhat rounded. Abdomen very finely punctate, median areas of ventrites with punctulate microsculpture. Male. Segments Ito III of protarsi hardly widened. Aedeagus (Figs 11 ̅13) 0.44 mm long. Parameres narrowed posterior of basal section, gradually weakly widened toward narrow apical section, not forming subapical lobe or process; outer margins of parameres concave and inner margins strongly sclerotized posterior of narrowed apical section. Internal sac bearing two pairs of sclerotized pieces, outer larger and more sclerotized than inner pair, expanded apically, inner pair widened proximally, with margins becoming indistinct apically; scale­like, spine­like or filamentous structures absent. Etymology. The species epithet is a Latin adjective meaning "sinuatȅ, referring to the outer margins of the parameres. Differential diagnosis. The body­size of this new species is significantly smaller than that of other Bornean species and the shape of its parameres is diagnostic for it. The new species is similar to S. bilobum Löbl, 2015 from Lombok. It shares with the latter the simple internal sac of the aedeagus lacking spine­like, denticulate or striate membranous structures in combination with the presence of two pairs of admesal sclerites. However, the outer pair of sclerites is much more expanded and the inner sclerites are much shorter and broader in S. sinuatum, which may be distinguished from B. bilobum also by its wide impression of the mesoventrite, finer punctation of the lateral parts of the metaventrite and shorter submesocoxal areas., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 17-18, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2015: On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Islands. Revue suisse de zoologie 122 (1): 75 ‾ 120."]}

Baeocera sarawakensis Löbl, 1987 Material examined. 1 male, Sabah, Lahad Datu Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°58′ E 117°42′, 450 m, viii.1998, D. Bebber lgt., lowland mixed dipterocarp forest, FIT, OUMNH 2003­050 (OUMNH). Remarks. As B. incisa, this species is widely distributed in Sabah and seems to be common in forest floor litter (see LÖBL 2021: 31)., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2021: On the Bornean species of Scaphoxium Lobl, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beitage 53 (1): 85 ‾ 89."]}

Sapitia Achard, 1920 The genus includes three South-East Asian species, all reported also from Indonesia though only one of them from the Lesser Sundas. Members of Sapitia are presumably associated with termites (Leschen & Löbl, 2005)., Published as part of Löbl, Ivan, 2015, On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Islands, pp. 75-120 in Revue suisse de Zoologie 122 (1) on page 92, DOI: 10.5281/zenodo.14582, {"references":["Achard J. 1920. Notes sur les Scaphidiidae de la faune indomalaise. Annales de la Societe Entomologique de Belgique 60: 123 - 136.","Leschen R. A. B., Lobl I. 2005. Phylogeny and classification of Scaphisomatini (Staphylinidae: Scaphidiinae) with notes on mycophagy, termitophily and functional morphology. Coleopterists Society Monographs, Patricia Vaurie series no. 3: 63 pp."]}

Pseudobironium irregulare sp. nov. (Figs 5 ̅7) Type material. Holotype male, SABAH: Poring Hot Springs 600 m 9.V.1987 Burckhardt ̅ Löbl (MHNG); Paratypes: 1 female, SABAH: Crocker Ra. 1350 m, km 60 Kota Kinabalu­Tambunan 17.V.1987 Burckhardt ̅ Löbl; female, BORNEO SABAH Tamis, Hwy AI 10 km NW Kinabalu Park Entr. 1100 m 24.V.87 A. Smetana; 1 female, SABAH: E Mt. Kinabalu 1150 m, rte Ranau­Kota Kinabalu 24.V.1987 Burckhardt ̅ Löbl (all in MHNG). Description. Length 1.78̅ 1.87 mm, width 1.20̅ 1.35 mm. Head and most of body dark reddish­brown, metaventrite somewhat darkened, apex of abdomen lighter, yellowish, femora and tibiae roughly as body, tarsi and antennae lighter, yellowish. Maxillary palpi with palpomere IV narrow, tapering, about 4 times as long as wide. Length ratio of antennomeres as follows: III 14: IV 25: V 25: VI 22: VII 26: VIII 26: IX 26: X: 26: XI 31; antennomere XI about 4 times as long as wide. Pronotal punctation about as fine as that on frons, with punctures very shallow and not well delimited; middle of pronotal disc with puncture intervals about 5 to 10 times larger than puncture diameters. Elytra lacking lateral impressions, not flattened or impressed apically, lacking humps; discal punctation on prevailing surface similar to pronotal punctation, conspicuous patch of rather coarse punctures present on anterolateral area. Puncture patch separated from lateral and basal margins by rather broad, very finely punctate areas. Lateral striae coarsely punctate. Hypomera impunctate, flattened along basal margin. Mesoventrite lacking mesal ridge, with distinct setiferous punctures at lateral margins of coxal process, setae long. Mesanepisternum very finely punctate, lacking microsculpture. Metaventrite with inconspicuous striate to punctulate microsculpture; median area nearly flat, lacking impressions and lacking tubercle, its punctation fine, arranged to form two sparse admesal rows joined posteriorly. Lateral parts of metaventrite very finely and sparsely punctate. Metacoxal process impunctate, inflexed. Submesocoxal area slightly shorter than half of shortest interval to metacoxa. Submesocoxal line coarsely punctate. Mesotibiae and metatibiae slightly bent, ventral spur of mesotibiae straight. Exposed ventrites and tergites with distinct punctulate microsculpture, setiferous punctures very fine and scattered. Male. Protarsus shorter than protibia, protarsomeres Ito III hardly widened, narrower than apex of tibiae, bearing short setae. Aedeagus (Figs 5 ̅7) 0.68 mm long, weakly sclerotized. Median lobe with basal bulb shorter than apical process, the latter hardly inflexed, with sinuate ventral margin (lateral view) and blunt tip; apical part of median lobe overlapped by two valves. Parameres widened basally, weakly bent. Internal sac lacking sclerotized pieces, without distinct flagellum, bearing apical spinose structures. Etymology. The species epithet is a Latin adjective meaning "irregulaȑ, referring to the elytral punctation. Differential diagnosis. This species may be readily distinguished from its congeners, P. impressipenne Löbl, 1973 and P. subglabrum Löbl, 1990 excepted, by elytra bearing each an anterolateral patch of coarse punctures. In addition, its small body­size and the structure of the internal sac are diagnostic. It may be readily distinguished from P. impressipenne and P. subglabrum by the parameres widened basally. It falls to the couplet 25 in the key to the species in LÖBL & TANG (2013), to P. feai Pic, though the latter species is 2.90̅ 3.20 mm long and has a black body. The aedeagal characters suggest relationships with members of the P. plagiferum group. It may be distinguished from the three species placed in that group (P. javanum Löbl & Tang, 2013, P. plagiferum Löbl, 1980, and P. spinipes Löbl & Tang, 2013) by the shape of the parameres and apex of the median lobe., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 11-14, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Vituratella kistneri (Löbl, 1979) Materiel examined. 50, Sabah, Batu Punggul Resort, 24.vi̅1.vii.1996, FIT (MHNG); 1, Sabah, Mt. Kinabalu Nat. Park, Poring Hot Springs, 495 m, 30.viii.1988, A. Smetana [B161] (MHNG); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, selectively logged lowland mixed dipterocarp forest, Yayasan Sabah Logging Concession, E. Slade & J. Villanuava lgt., OUMNH­2005­062 (OUMNH); 1, Sabah, Lahad Datu, Ulu Segama, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 200 m, 27̅ 28.x.2005, primary forest, D.J. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, west trail 10, forest interior FIT, OUMNH­2006­051 (OUMNH). Remarks. The species was based on several specimens found in North Sumatra, in fungus gardens of Odontotermes takensis Ahmad, 1965. It was not reported after its description while V. termitophila Champion, 1927 is widely distributed and has been referred to many times (see OGAWA & MAETO 2015). The colour pattern of the abdomen used as one of the diagnostic characters proves to be unreliable in the examined material., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 18, DOI: 10.5281/zenodo.7501705, {"references":["OGAWA R. & MAETO K. 2015: The termitophilous Scaphidiinae (Coleoptera: Staphylinidae) from Sulawesi, Indonesia. The Coleopterists Bulletin 69 (2): 301 ‾ 304."]}

Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1-2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705, {"references":["LESCHEN R.A.B. & LOBL I. 2005: Phylogeny and classification of Scaphisomatini Staphylinidae: Scaphidiinae with notes on mycophagy, termitophily, and functional morphology. Coleopterists Society Monographs 3: 1‾63.","LOBL I. 2015: On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Islands. Revue suisse de zoologie 122(1): 75‾120.","LOBL I. 2018: Coleoptera: Staphylinidae: Scaphidiinae. World Catalogue of Insects. Volume XVI: i‾xvi + 1‾418 pp.","LOBL I. 2021: On the Bornean species of Scaphoxium Lobl, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beitage 53(1): 85‾89.","LOBL I. & LESCHEN R.A.B. 2010: Notes on the Toxidium group (Coleoptera: Staphylinidae: Scaphidiinae). Folia Heyrovskyana Series A 18(1-3): 71‾93.","LOBL I., LESCHEN R.A.B. & KODADA J. 2020: Review of the Asian species and cladistic analysis of Bironium Csiki (Coleoptera: Staphylinidae: Scaphidiinae) with comments on biogeography. Annales Zoologici (Warszawa) 70(4): 711‾736.","LOBL I. & SMETANA A. 2021: On the Baeocera Erichson (Coleoptera: Staphylinidae: Scaphidiinae) of Sabah, Malaysia, and a tale on mystified biodiversity. Journal of Insect Biodiversity 23(2): 23‾42.","LOBL I. & TANG L. 2013:Areview of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665‾734.","OGAWA R. & LOBL I. 2016: Areview of the genus Xotidium Lobl, 1992 (Coleoptera, Staphylinidae, Scaphidiinae), with description of five new species. Deutsche entomologische Zeitschrift 63: 155‾169.","OGAWA R. & MAETO K. 2015: The termitophilous Scaphidiinae (Coleoptera: Staphylinidae) from Sulawesi, Indonesia. The Coleopterists Bulletin 69(2): 301‾304.","PROKOFIEV A.M. 2013: Contribution to the knowledge of the scaphidiine genus Scaphisoma Leach of the Bu Gia Map National Park, Vietnam. Calodema 245: 1‾6."]}

Löbl, Ivan (2022): On 'Scaphisoma tricolor Heller' from Japan and Errata to the Descriptions of Scaphisoma konvickai Löbl and Toxidium hartmanni Löbl (Coleoptera: Staphylinidae: Scaphidiinae). Japanese Journal of Systematic Entomology 28 (1): 85-88, DOI: http://doi.org/10.5281/zenodo.6787471, {"references":["Heller, K.M., 1917. Scaphidiidae von den Philippinen. Wiener entomologische Zeitung, 36: 41-50.","Hoshina, H., 2013. Scaphidiinae, pp. 124-127. In: Catalogue of Japanese Staphylinidae. Bulletin of the Kyushu University Museum, 11: 69-218.","Lobl, I., 1970. Uber einige Scaphidiidae (Coleoptera) aus der Sammlung des Museum National d'Histoire Naturelle de Paris. Mitteilungen der schweizerischen entomologischen Gesellschaft, 43: 125-132.","Lobl, I., 1971. Scaphidiidae der Noona Dan Expedition nach den Philippinen und Bismark Inseln (Insecta, Coleoptera). Steenstrupia, 1: 247-253.","Lobl, I., 1972. Beitrag zur Kenntnis der Scaphidiidae (Coleoptera) von den Philippinen. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 45: 79-109.","Lobl, I., 1981. Insects of Micronesia Coleoptera Scaphidiidae. Insects of Micronesia, 15: 69-80.","Lobl, I., 1982. Uber die Scaphidiidae (Coleoptera) der japanischen Ryukyu-Inseln. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 55: 101-105.","Lobl, I., 2018. Coleoptera: Staphylinidae: Scaphidiinae. World Catalogue of Insects. Volume 16. Brill, Leiden/Boston, xvi + 418 pp.","Lobl, I. & R. Ogawa, 2016. On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia Achard and Scaphisoma Leach. Linzer biologische Beitrage, 48(2): 1339-1492.","Lobl, I., 2022. On new collections of Scaphidiinae (Coleoptera: Staphylinidae) from China, with description of two new species. Zootaxa, 5092 (4): 487-492.","Morimoto, K., 1985. Scaphidiidae, pp. 252-258. pl. 45. In: Ueno, S., Kurosawa, Y. & Sato, M. (eds.): The Coleoptera of Japan in color. Vol. 2. Hoikusha Publ. Co., Osaka, viii + 479 pp., 80 pls [in Japanese].","Ogawa, R., 2015. Phylogeography of the subfamily Scaphidiinae (Coleoptera, Staphylinidae) in Sulawesi, with its systematic revision. Graduate School of Agricultural Science, Kobe University, 180 pp. [PhD thesis, unpublished].","Schulke, M. & A. Smetana, 2015. Staphylinidae, pp. 304-900. In: Lobl, I. & D. Lobl, (eds), Catalogue of Palaearctic Coleoptera. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Volume 1. Brill, Leiden/Boston, xxv + 900 pp. [Received: February 21, 2022; accepted: June 12, 2022]"]}

Löbl, Ivan (2022): Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas. Revue suisse de Zoologie 129 (1): 103-118, DOI: 10.35929/RSZ.0064

Baeocera sp. B Material examined: MHNG; 1 female; MALUC: Kai Besar Island, G. Dab, leaf litter, 400 m, 5.IX.1981, D. Agosti, F911046. Remarks: This is the second Moluccan species of Baeocera remaining unnamed, in absence of at least one male specimen (see “ Baeocera sp. A ” in Löbl, 2015a). It is similar to the Papuan B. curta Löbl, 2002, differs by the less arcuate suture of the metanepisterna and the ventrite I lacking basal striae. Thus, it may be an undescribed species. Baeocera curta has a complex aedeagus providing reliable specific characters which are quite unlike those of members of the Baeocera lenta group and of B. agostii. The specimen has slightly prominent apical inner apical angles of the elytra, a character noted for the first time in Baeocera. It may be sexual, as it is in some species of Scaphisoma Leach., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 112, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2015 a. Contribution to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas. Stuttgarter Beitrage zur Naturkunde A, Neue Serie 8: 165 - 187.","Lobl I. 2002. On Baeocera (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea. Mitteilungen der Schweizerischen entomologischen Gesellschaft 75: 1 - 20."]}

Baeocera punctigera sp. nov. Figs 14-16 Materiel examined: Holotype; male; MHNG; MALUC: Tanimbar Isl., Kai Besar, G. Dab, 400 m, 5.IX.81, D. Agosti F911046. Etymology: The species epithet is a Latin adjective meaning bearing punctures. Diagnosis: The species is defined by the following characters in combination: Body-length about 1.2 mm long, body dark brown; pronotum coarsely and densely punctate, with many punctures larger than puncture intervals; lateral margins of pronotum and elytra separately arcuate; entire elytron coarsely punctate, sutural stria shortened, basal stria absent; mesepimeron with puncture row along anterior margin; hypomeron, mesanepisternum, metaventrite and ventrite I conspicuously punctate; ventrite I without basal striae or wrinkles; aedeagus symmetrical, with apical process narrowed apically, bent near tip (lateral view), parameres straight and evenly narrow in dorsal view, sinuate and widened toward apical third in lateral view, internal sac with complex sclerites as in Fig. 16, lacking membranous scale or denticle-like structures. Description: Length 1.17 mm, width 0.80 mm. Head and body dark brown, femora and tibiae lighter, reddish-brown, tarsi and antennae yellowish. Head with interocular distance slightly larger than dorsoventral eye diameter. Antennomere II slightly longer than antennomere III. Length/width ratios of antennomeres as: III 28/7: IV 27/6: V 32/6: VI 28/6: VII 33/10: VIII 28/8: IX 35/13: X 35/14: XI 42/15. Pronotum and elytra separately arcuate. Pronotum not microsculptured, pronotal punctation conspicuously coarse and dense, distinct at 15 times magnification, consisting of sharply delimited punctures smaller than puncture intervals; lateral carinae of pronotum hardly visible in dorsal view, with punctures very dense, larger than puncture intervals. Point of scutellum exposed. Elytra covering abdomen, weakly narrowed apically, with lateral margins evenly, weakly rounded; adsutural area flat and narrow; sutural stria punctate, shortened, starting about 0.20 mm posterior of tip of scutellum; basal stria absent; lateral stria punctate. Elytral disc not microsculptured; punctation entirely coarse and dense, punctures larger than those on pronotum and nearly as well delimited, puncture intervals mostly larger than puncture diameters. Hind wings fully developed. Hypomeron not microsculptured, conspicuously punctate except on smooth areas along upper margin and near mesocoxa, with some punctures about as large as puncture intervals and as pronotal punctures. Mesoventrite punctate, without median ridge, fused with metaventral process. Mesanepisternum conspicuously punctate, punctures similar to those on hypomeron. Mesepimeron with row of coarse punctures along anterior margin, about three times as long as wide and three times as long as interval to coxa. Metaventrite flattened in middle, all over coarsely and densely punctate, narrow apical band excepted; punctures on lateral part of metaventrite well delimited, mostly larger than puncture intervals. Submesocoxal lines parallel, with marginal punctures smaller than punctures on lateral area of metaventrite and somewhat extended laterally along margin of metaventrite. Submesocoxal areas about 0.02 mm long, as fifth of shortest distance to metacoxae. Metanepisternum flat, fused with metaventrite, its margin indicated by outer puncture row of metaventrite. Tibiae straight. Abdomen not microsculptured. Ventrite I densely and coarsely punctate, punctures smaller than those on lateral area of metaventrite, basal punctures not enlarged and not elongate, basal striae or wrinkles absent. Following ventrites finely punctate. Male characters: Protarsomeres I to III slightly widened. Aedeagus (Figs 14-16) 0.29 mm long, weakly sclerotized, symmetrical.Apical process gradually narrowed apically, with tip bent and blunt in lateral view. Parameres straight and evenly narrow in dorsal view, sinuate and gradually widened toward apical third in lateral view. Internal sac with complex sclerites, as in Fig. 16 with flagellar guidesclerite widened apically, broad, nearly membranous accessory lateral sclerite, lacking membranous scale- or denticle-like structures. Habitat: South slope of G. Dab, secondary forest on limestone, soil and very humid leaf litter. Distribution: Indonesia: Moluccas, Tanimbar Islands. Remarks: This is another species of the B. lenta group, in external characters very similar to B. notata. It may be easily distinguished from the latter by the mesepimeron bearing a row of coarse punctures along its anterior margin and by the aedeagal characters, in particular by the parameres straight and narrow in dorsal view. In addition, the body-size of B. punctigera is smaller, the antennae are shorter, with the antennomere XI less than 3 times as long as wide. While the aedeagus of B. punctigera is similar to that of the New Guinean B. flagrans Löbl, 2002, the species differs strongly from the latter by its external characters., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on pages 108-110, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2002. On Baeocera (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea. Mitteilungen der Schweizerischen entomologischen Gesellschaft 75: 1 - 20."]}

Baeocera notata sp. nov. Figs 7-10 Materiel examined: Holotype; MHNG; male; MALUC: Aru Isls: Kola Isl.: Watan, 27.8.81, D. Agosti F91936. Paratype; MHNG; 1 female; MALUC: Aru Isl., Wokam, 22.8.81, D. Agosti F911970. Etymology: The species epithet is a Latin adjective referring to the pronotum. Diagnosis: The species is defined by the following characters in combination: Body-length about 1.35- 1.45 mm, body dark brown; pronotum coarsely and densely punctate, with many punctures larger than puncture intervals; lateral margins of pronotum and elytra separately arcuate; elytron entirely coarsely punctate, with sutural stria shortened, lacking basal stria; hypomeron and mesanepisternum distinctly punctate; metaventrite and ventrite I coarsely punctate; ventrite I without basal striae or wrinkles; aedeagus symmetrical, with apical process narrowed apically, bent near tip in lateral view, apical halves of parameres bent and bearing subapical membranous lobe (dorsal view), widened posterior of basal third in lateral view, internal sac with complex sclerites as in Fig. 9, lacking membranous scale or denticle-like structures. Description: Length 1.36-1.43 mm, width 0.90- 1.0 mm. Head and body dark brown, femora and tibiae lighter, reddish-brown, tarsi and antennae yellowish. Head with interocular distance slightly smaller than dorsoventral eye diameter. Antennomere II slightly longer than antennomere III. Length/width ratios of antennomeres as: III 28/7: IV 27/7: V 37/7: VI 35/7: VII 39/12: VIII 38/10: IX 39/12: X 37/14: XI 48/14. Pronotum and elytra separately arcuate. Pronotum not microsculptured, pronotal punctation conspicuously coarse and dense, distinct at 15 times magnification, consisting of sharply delimited punctures to part larger than puncture intervals; lateral carinae of pronotum hardly visible in dorsal view, bearing punctures very dense, larger than puncture intervals. Point of scutellum exposed. Elytra covering tip of abdomen, weakly narrowed apically, with lateral margins evenly, weakly rounded; adsutural area flat and narrow; sutural stria punctate, shortened, starting about 0.20 mm posterior of tip of scutellum; basal stria absent; lateral stria punctate. Elytral disc not microsculptured; punctation all over coarse and dense, punctures larger than those on pronotum but not as well delimited, puncture intervals mostly about as puncture diameters or slightly larger. Hind wings fully developed. Hypomeron not microsculptured, conspicuously punctate, with punctures about as large as pronotal punctures. Mesoventrite punctate, without median ridge, fused with metaventral process. Mesanepisternum conspicuously punctate, punctures similar to those on hypomeron. Mesepimeron impunctate, about four times as long as wide and three times as long as interval to coxa. Metaventrite flattened in middle, all over coarsely and densely punctate, smooth narrow apical band excepted; punctures on lateral part of metaventrite well delimited, to part as large or larger than puncture intervals. Submesocoxal lines slightly convex, with marginal punctures smaller than punctures on lateral area of metaventrite and somewhat extended laterally along margin of metaventrite. Submesocoxal areas about 0.03 mm long, as third of shortest interval to metacoxae. Metanepisternum flat, fused with metaventrite, its margin indicated by outer puncture row of metaventrite. Tibiae straight. Abdomen not microsculptured. Ventrite I densely and coarsely punctate, punctures smaller than those on lateral area of metaventrite, basal punctures not enlarged and not elongate, basal striae or wrinkles absent. Following ventrites finely punctate. Male characters: Protarsomeres I to III slightly widened. Aedeagus (Figs 7-10) 0.37 mm long, weakly sclerotized, symmetrical.Apical process gradually narrowed apically, with tip bent and acute in lateral view. Parameres straight in basal halves, bent in apical halves, each with weakly sclerotized subapical lobe (dorsal view), narrow near bases and nearly evenly broad in apical two thirds, with slightly concave ventral margins (lateral view). Internal sac with complex sclerites, as in Fig. 9, with flagellar guide-sclerite widened apically, narrow accessory lateral sclerite present, lacking membranous scale- or denticlelike structures. Habitat: Secondary/primary forest 5 km E of Mt. Sungai, shallow leaf litter, on limestone; understory with Pandanus. Distribution: Indonesia: Moluccas, Aru Islands. Remarks: The species is another member of the B. lenta group. It shares with the Moluccan B. kaibesara Löbl, 2014 and B. vicina Löbl, 2015, and the Papuan B. praedicta Löbl, 2002, elytra with shortened sutural striae. It may be readily distinguished from these three species by the hypomera and mesanepisterna conspicuously punctate. In addition, the shape of the parameres lobed subapically is diagnostic for this new species. Several species, such as the New Guinean B. fortepunctata Löbl, 2002 and the Philippine B. hypomeralis Löbl, 2012 also possess punctate hypomera but differs by the elytra having entire sutural striae and the impunctate or extremely finely punctate mesanepisterna. The Himalayan B. cribrata Löbl, 1992, shares the pronotum coarsely punctate, distinctly punctate hypomera and mesanepisterna, and elytra with shortened sutural striae. It may be easily distinguished by the punctation on lateral parts of the pronotum much coarser than that on pronotal centre., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on pages 106-107, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2014. On the Scaphidiinae Leach (Coleoptera, Staphylinidae) of the Moluccas. Mitteilungen der Schweizerischen entomologischen Gesellschaft 87: 49 - 60.","Lobl I. 2002. On Baeocera (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea. Mitteilungen der Schweizerischen entomologischen Gesellschaft 75: 1 - 20."]}

Scaphisoma sublimbatum Löbl, 1977 Scaphisoma sublimbatum Löbl, 1977: 57. Material examined: MHNG; 2 males; MALUK: Aru Isl., Wokam, 5 km E. Mt. Sungai, 22.VIII.1981, D. Agosti, F91936. Redescription: Length 1.53-1.75 mm, width 1.02- 1.22 mm. Head, thorax and most abdomen dark brown, apex of abdomen yellowish. Elytra on adsutural area, along basal and lateral margins, and on transverse stripe in middle as dark as pronotum, in apical sixth to fifth light brown, each with two large, irregular, transverse yellowish spots. Anterior spots extending to middle third of elytral length, posterior spot starting about at level of apical third of elytral length. Femora light brown, tibiae, tarsi and antennae yellowish. Length/ width ratios of antennomeres as: III 12/10: IV 27/8: V 50/9: VI 50/9: VII 57/16: VIII 38/10: IX 53/14: X 53/15: XI 60/14. Pronotum not microsculptured, with lateral margins strongly, evenly rounded, lateral margin carinae throughout visible in dorsal view, lateral margin carinae impunctate, discal punctation fine, sparse, most puncture intervals about 2 to 4 times as large as puncture diameters, punctures shallow, not well delimited, visible at 30 times magnification. Exposed point of scutellum minute. Elytra not microsculptured, with lateral margins evenly rounded, lateral margin carinae exposed in dorsal view, lateral margin striae impunctate, apical margins truncate, inner apical angles not prominent, situated in level with outer apical angles, marginal crenulation very fine, distinct, sutural striae deep, bent at bases and extending laterally to form basal striae reaching outer half of basal width, widely separated from lateral striae, sutural striae parallel, impunctate, adsutural areas flat, each with single, very fine and sparse puncture row, discal punctures irregular, fairly fine and dense, coarser than pronotal or adsutural punctures, not forming rows, puncture intervals mostly distinctly larger than puncture diameters. Hind wings fully developed. Hypomera not microsculptured, impunctate. Mesepimon about 4 times as long as wide and somewhat shorter than interval to mesocoxa. Metaventrite not microsculptured. Median part of metaventrite convex between mesocoxae, flattened apically, with three fovea-like apical impressions, middle impression larger and situated slightly anterior lateral impressions, often joint to short mesal stria; punctation indistinct, sparse and extremely fine on centre and on lateral areas of metaventrite. Lateral parts of metaventrite lacking antecoxal puncture rows. Submesocoxal area 0.02 mm long, about as sixth of shortest interval to metacoxa. Submesocoxal line parallel, densely and coarsely punctate, punctures extended laterally along anterior margin of metaventrite. Metanepisterna slightly convex, impressed along nearly straight suture, narrowed anteriad. Tibiae straight. Abdomen not microsculptured, with hardly visible punctation (at 100 times magnification). Ventrite I with submetacoxal area extremely narrow, about 0.01- 0.02 mm long, narrowed mesally and as fifteens of interval to apical margin of ventrite; marginal punctures coarse. Male characters: Protarsomeres I to III and mesotarsomeres I and II strongly widened, basal tarsomeres nearly as wide as apices of tibiae. Apical margin of ventrite VI trilobed, mesal lobe triangular, about 0.10 mm long, lateral lobes shorter and rounded. Habitat: Secondary/primary forest on limestone, leaf litter on rather dry soil, understory Pandanus. Distribution: Australia: Queensland; Indonesia: Moluccas, Aru Islands. Remarks: This species is a member of the S. unifasciatum group defined in Löbl, 1977. In addition to S. sublimbatum, the group includes four New Guinean species: S. audax Löbl, 1975, S. quadripunctatum (Pic, 1956), S. unifasciatum Pic, 1956, and S. unimaculatum Löbl, 1975. An additional species is described below. The original description of the external features S. sublimbatum was brief and limited to comparation to the similar S. audax. Therefore, it is supplement., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 112, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 1977. Beitrag zur Kenntnis der Scaphidiidae (Coleoptera) Australiens. Revue suisse de Zoologie 84: 3 - 69.","Lobl I. 1975. Beitrag zur Kenntnis der Scaphidiidae (Coleoptera) von Neuguinea. Revue suisse de Zoologie 82: 369 - 420."]}

Scaphoxium sp. Materiel examined: MHNG; 1 female; MOLUK: Kai Besar, Bombay, G. Dab, 3.IX.1981, D. Agosti, F911019. MHNG; 1 female; MOLUK: Kai Besar, G. Tukram, 300-500 m, 7.IX.1981, D. Agosti F911075. Remarks: These two specimens differ notably from S. lobatum by the very finely punctate lateral parts of the metaventrite. Habitat: Secondary forests on limestone; leaf litter and moss on fallen tree. Distribution: Indonesia: Moluccas, Kai Besar Island., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 117, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022

Baeocera incurva sp. nov. Figs 4-6 Materiel examined: Holotype; MHNG; male; MALUK: Kai Besar, Bombay, G. Dab, 3.IX.81, D. Agosti F911019. Paratypes; MHNG; 1 male, 2 ex. with the same data as the holotype. – MHNG; 2 males, 1 female; MALUK: Kai Besar, G. Tukrau nr. Bombay, 300 -400 m, 7.IX.81 D. Agosti, F91075. Etymology: The species epithet is a Latin adjective meaning bent. Diagnosis: The species is defined by the following characters in combination: Body-length about 1.25- 1.35 mm, body blackish with lighter apical abdominal segments; lateral margins of pronotum and elytra nearly continuously arcuate; pronotum finely punctate; elytron entirely coarsely punctate, lacking basal stria, sutural stria shortened, starting posterior of level of scutellum; hypomeron very finely punctate; mesepimeron, mesanepisterum, metaventrite and ventrite I distinctly punctate; ventrite I without basal wrinkles or striae; aedeagus symmetrical, with apical process in lateral view abruptly narrowed near apex, parameres sinuate, nearly evenly broad and bent dorsally in lateral view, internal sac with complex sclerites, as in Fig. 6, lacking membranous scale or denticle-like structures. Description: Length 1.25-1.33 mm, width 0.85- 0.88 mm. Head and most of body blackish, apical abdominal segments, femora and tibiae reddish-brown, tarsi and antennae yellowish. Head with interocular distance slightly larger than dorsoventral eye diameter. Antennomere II slightly longer than antennomere III. Length/width ratios of antennomeres as: III 24/6: IV 22/6: V 30/7: VI 30/7: VII 37/9: VIII 33/8: IX 41/12: X 41/14: XI 46/15. Pronotum and elytra nearly continuously arcuate. Pronotum not microsculptured, pronotal punctation hardly visible at 50 times magnification; lateral carinae of pronotum concealed in dorsal view. Point of scutellum exposed. Elytra nearly entirely covering abdomen. Elytron weakly narrowed apically, with lateral margin rounded; lateral margin carinae concealed in dorsal view, adsutural area flat and narrow; sutural stria punctate, shortened, starting well posterior of level of tip of scutellum; basal stria absent; lateral stria punctate. Elytral disc not microsculptured, with punctation well delimited, much coarser than on pronotum, nearly regular, puncture intervals mostly about as to three times puncture diameters. Hind wings fully developed. Impressed inferior part of hypomeron with few very fine punctures. Mesoventrite punctate, without median ridge, fused with metaventral process. Mesanepisternum pubescent, rather finely punctate. Mesepimeron punctate, about four times as long as wide and about three times as long as interval to coxa. Median part of metaventrite flattened, fairly coarsely, irregularly and densely punctate. Lateral area of metaventrite with punctation similar to that on elytron; punctures well delimited, clearly smaller than puncture intervals, evanescent near metacoxal margin. Submesocoxal lines slightly convex, with marginal punctures smaller than punctures on lateral area of metaventrite and not extended laterally along margin of metaventrite. Submesocoxal areas about 0.02-0.03 mm long, about as fifth of shortest interval to metacoxae. Metanepisternum flat, fused with metaventrite, its margin indicated by outer puncture row of metaventrite. Tibiae straight. Abdomen not microsculptured. Ventrite I about as coarsely and densely punctate as lateral area of metaventrite, basal punctures not enlarged and not elongate, basal wrinkles or striae absent. Following ventrites finely punctate. Male characters: Protarsomeres I to III slightly widened. Aedeagus (Figs 4-6) 0.35-0.36 mm long, weakly sclerotized, symmetrical. Apical process abruptly narrowed near apex (lateral view), gradually narrowed in dorsal view. Parameres conspicuously sinuate, nearly evenly broad and bent dorsally in lateral view, hardly sinuate in dorsal view. Internal sac with complex sclerites as in Fig. 6, with very short flagellar guide-sclerite, lacking membranous scale- or denticle-like structures. Habitat: Secondary forests on limestone, litter from dead log and rotting leaves. Distribution: Indonesia: Moluccas, Kai Besar Island. Remarks: The species is also member of the B. lenta group, as shown by its aedeagal characters. It falls in the key to the Moluccan species (Löbl, 2015a) to B. vicina Löbl, 2015. It may be easily distinguished from the latter as from other congeners by the shape of the parameres and by the internal sac possessing a short flagellum and reduced flagellar-guide sclerite. The New Guinean B. praedicta Löbl, 2002 possesses parameres rather similar to those of B. incurva. It differs notably in external characters, in particular by the not fused metanepisterna and the very finely punctate metaventrite., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on pages 104-106, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2015 a. Contribution to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas. Stuttgarter Beitrage zur Naturkunde A, Neue Serie 8: 165 - 187.","Lobl I. 2002. On Baeocera (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea. Mitteilungen der Schweizerischen entomologischen Gesellschaft 75: 1 - 20."]}

Scaphobaeocera sp. 2 Material examined: MHNG; 1 female; MALUK: Tanimbar Isl., Yamdena, 22 km N Samlaki via Aram, Desa Lorulun, 11.IX.1981, D. Agosti. Remarks: This specimen would fall in the key to the New Guinean Scaphobaeocera (Löbl, 2017) under the couplet 5, to S. antennalis Löbl, 1975 and S. remota Löbl, 1981, and it shares with S. remota the elytra darkened apically and the antennomeres XI more than twice as long as the antennomeres X. Habitat: Melaleuca open forest on limestone, leaf litter heavily infested by mycorrhiza. Distribution: Indonesia: Moluccas, Tanimbar Island., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 115, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2017. New species and records of Scaphobaeocera Csiki from New Guinea (Coleoptera: Staphylinidae: Scaphidiinae). Mitteilungen der Munchner entomologischen Gesellschaft 107: 33 - 41.","Lobl I. 1975. Beitrag zur Kenntnis der Scaphidiidae (Coleoptera) von Neuguinea. Revue suisse de Zoologie 82: 369 - 420.","Lobl I. 1981. Scaphidiidae. Insects of Micronesia 15 (2): 69 - 80."]}