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Increasing fire severity and warmer, drier postfire conditions are making forests in the western United States (West) vulnerable to ecological transformation. Yet, the relative importance of and interactions between these drivers of forest change remain unresolved, particularly over upcoming decades. Here, we assess how the interactive impacts of changing climate and wildfire activity influenced conifer regeneration after 334 wildfires, using a dataset of postfire conifer regeneration from 10,230 field plots. Our findings highlight declining regeneration capacity across the West over the past four decades for the eight dominant conifer species studied. Postfire regeneration is sensitive to high-severity fire, which limits seed availability, and postfire climate, which influences seedling establishment. In the near-term, projected differences in recruitment probability between low- and high-severity fire scenarios were larger than projected climate change impacts for most species, suggesting that reductions in fire severity, and resultant impacts on seed availability, could partially offset expected climate-driven declines in postfire regeneration. Across 40 to 42% of the study area, we project postfire conifer regeneration to be likely following low-severity but not high-severity fire under future climate scenarios (2031 to 2050). However, increasingly warm, dry climate conditions are projected to eventually outweigh the influence of fire severity and seed availability. The percent of the study area considered unlikely to experience conifer regeneration, regardless of fire severity, increased from 5% in 1981 to 2000 to 26 to 31% by mid-century, highlighting a limited time window over which management actions that reduce fire severity may effectively support postfire conifer regeneration.

Across the western United States, forests are changing rapidly, with uncertain impacts on snowmelt water resources. Snow partitioning is controlled by forest effects on interception, radiation, and sublimation. Yet, models often lack snow measurements with sufficiently high spatial and temporal resolution across gradients of forest structure to accurately represent these fine-scale processes. Here, we utilize four Snowtography stations in Arizona, in the lower Colorado River Basin, with daily measurements over 3-5 years at ~110 positions distributed across gradients of forest structure resulting from wildfires and mechanical thinning. We combine Snowtography with lidar snapshots of forest and snow to train a high-resolution snow model and run it for 6 years to quantify how forest structure regulates snowpack and snowmelt. These study sites represent a climate gradient from lower/warmer ephemeral snowpack (~2,100 m asl) to higher/colder seasonal snowpack (~2,800 m asl). Forest cover reduced snowpack and snowmelt through canopy sublimation. Forest advanced snowmelt timing at lower/warmer sites but delayed it at higher/colder sites. Within canopy gaps, shaded cool edges had the greatest peak snow water equivalent (SWE). Surprisingly, sunny/warm gap edges produced more snowmelt than cool edges, because high radiation melted snow quickly, reducing exposure to sublimation. Therefore, peak SWE is not an ideal proxy for snowmelt volume from ephemeral snowpacks, which are becoming more prevalent due to warming. The results imply that forest management can influence the amount and timing of snowmelt, and that there may be decision trade-offs between enhancing forest resilience through delayed snowmelt and maximizing snowmelt volumes for downstream water resources. [ABSTRACT FROM AUTHOR]

Increasing aridity is a challenge for forest managers and reducing stand density to minimize competition is a recognized strategy to mitigate drought impacts on growth. In many dry forests, the most widespread and common forest management programs currently being implemented focus on restoration of historical stand structures, primarily to minimize fire risk and enhance watershed function. The implications of these restoration projects for drought vulnerability are not well understood. Here, we examined how planned restoration treatments in the Four Forests Restoration Initiative, the largest forest restoration project in the United States, would alter landscape‐scale patterns of forest growth and drought vulnerability throughout the 21st century. Using drought–growth relationships developed within the landscape, we considered a suite of climate and treatment scenarios and estimated average forest growth and the proportion of years with extremely low growth as a measure of vulnerability to long‐term decline. Climatic shifts projected for this landscape include higher temperatures and shifting seasonal precipitation that promotes lower soil moisture availability in the early growing season and greater hot‐dry stress, conditions negatively associated with tree growth. However, drought severity and the magnitude of future growth declines were moderated by the thinning treatments. Compared to historical conditions, proportional growth in mid‐century declines by ~40% if thinning ceases or continues at the status quo pace. By comparison, proportional growth declines by only 20% if the Four Forest Restoration Initiative treatments are fully implemented, and <10% if stands are thinned even more intensively than currently planned. Furthermore, restoration treatments resulted in dramatically fewer years with extremely low growth in the future, a recognized precursor to forest decline and eventual tree mortality. Benefits from density reduction for mitigating drought‐induced growth declines are more apparent in mid‐century and under RCP4.5 than under RCP8.5 at the end of the century. Future climate is inherently uncertain, and our results only reflect the climate projections from the representative suite of models examined. Nevertheless, these results indicate that forest restoration projects designed for other objectives also have substantial benefits for minimizing future drought vulnerability in dry forests and provide additional incentive to accelerate the pace of restoration. [ABSTRACT FROM AUTHOR]

Recent streamflow declines in the Upper Colorado River Basin raise concerns about the sensitivity of water supply for 40 million people to rising temperatures. Yet, other studies in western US river basins present a paradox: streamflow has not consistently declined with warming and snow loss. A potential explanation for this lack of consistency is warming-induced production of winter runoff when potential evaporative losses are low. This mechanism is more likely in basins at lower elevations or latitudes with relatively warm winter temperatures and intermittent snowpacks. We test whether this accounts for streamflow patterns in nine gaged basins of the Salt River and its tributaries, which is a sub-basin in the Lower Colorado River Basin (LCRB). We develop a basin-scale model that separates snow and rainfall inputs and simulates snow accumulation and melt using temperature, precipitation, and relative humidity. Despite significant warming from 1968–2011 and snow loss in many of the basins, annual and seasonal streamflow did not decline. Between 25% and 50% of annual streamflow is generated in winter (NDJF) when runoff ratios are generally higher and potential evapotranspiration losses are one-third of potential losses in spring (MAMJ). Sub-annual streamflow responses to winter inputs were larger and more efficient than spring and summer responses and their frequencies and magnitudes increased in 1968–2011 compared to 1929–1967. In total, 75% of the largest winter events were associated with atmospheric rivers, which can produce large cool-season streamflow peaks. We conclude that temperature-induced snow loss in this LCRB sub-basin was moderated by enhanced winter hydrological inputs and streamflow production. [ABSTRACT FROM AUTHOR]