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The oldest oceanic basin (160–180 Ma) in the western Pacific is the birthplace of the Pacific Plate and is thus essential for understanding the formation and evolution of the oceanic plate. However, the upper mantle structure beneath the region has not been thoroughly investigated because of the remoteness and difficulties of long‐term in situ seismic measurements at the ocean bottom. From 2018 to 2019, the Oldest‐1 experiment on the oldest seafloor was conducted as part of the international Pacific Array initiative. We present the first three‐dimensional P‐wave velocity structure down to a depth of 350 km based on the relative travel time residuals of teleseismic earthquakes recorded by 11 broadband ocean‐bottom seismometers operated during the Oldest‐1 experiment. Our result shows a fast P‐wave velocity anomaly (VP perturbation of 2%–4% faster than average) at a depth of 95–185 km beneath the northeast of the study area. This structure is interpreted as evidence of dry, viscous, and rigid materials at depths below the lithosphere. Two slow anomalies (VP perturbation of 2%–4% slower than average) are seen beneath the southwestern and eastern (the oldest seafloor >170 Ma) parts of the array site. The low‐velocity zones are found at depths of 95–305 km. The observed velocity structures can be indicative of plume activities that affected the upper mantle as the Pacific Plate migrated over hotspots from the southeast. Alternatively, the observed velocity features may provide seismic evidence for small‐scale sublithospheric convection. Key Points: One‐year ocean‐bottom geophysical investigation on the oldest Pacific provides seismic mantle structure of the regionDetailed 3‐D mantle structure implies complex evolution process of Pacific PlateOur model implies thermochemical modification of the upper mantle by plume interaction or small‐scale convection [ABSTRACT FROM AUTHOR]

The mobility of the lithosphere over a weaker asthenosphere constitutes the essential element of plate tectonics, and thus the understanding of the processes at the [ithosphere-asthenosphere boundary (LAB) is fundamental to understand how our planet works. It is especially so for oceanic plates because their relatively simple creation and evolution should enable easy elucidation of the LAB. Data from borehole broadband ocean bottom seismometers show that the LAB beneath the Pacific and Philippine Sea plates is sharp and age-dependent. The observed large shear wave velocity reduction at the LAB requires a partially molten asthenosphere consisting of horizontal melt-rich layers embedded in melt-rich mantle, which accounts for the large viscosity contrast at the LAB that facilitates horizontal plate motions. [ABSTRACT FROM AUTHOR]