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Arthropod systematics & phylogeny 81, 475 - 498 (2023). doi:10.3897/asp.81.e101441, A new family of Diptera Acalyptratae, Christelenkidae Roháček fam. nov., is established for Christelenka multiplex Roháček gen. et sp. nov., an unusual extinct taxon described from a unique male specimen preserved in Baltic amber (Mid-late Eocene, ca 48–34 Ma). Apart from detailed examination by light microscopy and photography, the holotype of the new species has also been studied by means of X-ray synchrotron microtomography with the aim of obtaining additional morphological data for consideration of its relationships. Because of a very peculiar combination of morphological characters, the new family is tentatively considered a separate lineage of Acalyptratae having no apparent sister-group relationship with any of the known families. Its probable relationships to some families of Opomyzoidea and Ephydroidea are discussed., Published by Museum, Dresden

Materials and corrosion 74(7), 1116 - 1119 (2023). doi:10.1002/maco.202313751, Magnesium-based alloys are emerging as a capable alternative to traditional materials for bone implant applications. The implant's degradation rate is the main indicator of their performance; however, different formulas have been reported to determine it based on three-dimensional imaging. In this technical note, we are presenting the deviation in the degradation rate determined by different equations for two sets of data and the implications for the comparison of different studies., Published by Wiley-VCH, Weinheim [u.a.]

Developments in X-Ray Tomography XIV, San Diego, United States, 21 Aug 2022 - 26 Aug 2022; Proceedings of SPIE 12242, 1224213 (2022). doi:10.1117/12.2631184, Previously, we designed a new quantitative phase-contrast setup for the microtomography experiments at the hereon beamlines P05 and P07 at PETRA III, DESY. This setup is based on Talbot array illuminators (TAI) as high visibility wavefront markers and can reach a spatial resolution level comparable to propagation-based imaging. In this work, we focus on the progress of bringing this setup into user operation. We collaborated with INI-Research to investigate the vascular system of mouse lymph nodes, exploiting the spatial resolution capabilities and using the quantitative aspect of the data to compare different sample preparation methods. We could successfully visualize and trace the blood supply of the lymph nodes, even with fine capillaries, showing the stability and performance of the setup in user application., Published by SPIE, Bellingham, Wash.

Subfamily Ponerinae Lepeletier de Saint-Fargeau, 1835. Type genus. Ponera Latreille, 1804. Male diagnosis. Males of Ponerinae are best identified at the tribal level as there are as yet no clear male-based synapomorphies for the subfamily. Male Ponerinae share the following key diagnostic plesiomorphies: (1) wing venation complete or nearly complete, with at least four closed cells present; (2) petiole without tergosternal fusion; (3) abdominal segment III not petiolated; and (4) abdominal sternum IX without prongs or teeth. Remarks. The diagnosis provided for Ponerinae above and Platythyreini and Ponerini below collectively represent a revision of the global diagnosis for the subfamily of Boudinot [42]. Male Ponerinae have previously been diagnosed for the Malagasy region [47] and Japan [48, 49]. No single character has been discovered yet that uniquely identifies all male Ponerinae. Presence of posterolateral processes on the petiolar sternum which contact the outer margins of the helcial tergite, recognized as a ponerine synapomorphy for the female castes [46], are either poorly developed in males or obscured by the petiolar tergite, thus necessitating focused study. Notably, whereas female Ponerinae display a high degree of specialization with respect to mandibular and leg characters, these are universally lacking in the conspecific male., Published as part of Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej & Perrichot, Vincent, 2022, Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae), pp. 1-19 in Insects 73 (796) on page 7, DOI: 10.3390/insects13090796, http://zenodo.org/record/7046409, {"references":["42. Boudinot, B. E. Contributions to the knowledge of Formicidae (Hymenoptera, Aculeata): A new diagnosis of the family, the first global male-based key to subfamilies, and a treatment of early branching lineages. Eur. J. Taxon. 2015, 120, 1 - 62. [CrossRef]","47. Yoshimura, M.; Fisher, B. L. A revision of male ants of the Malagasy region (Hymenoptera: Formicidae): Key to subfamilies and treatment of the genera of Ponerinae. Zootaxa 2007, 1654, 21 - 40. [CrossRef]","48. Ogata, K. A generic synopsis of the poneroid complex of the family Formicidae in Japan (Hymenoptera). Part 1. Subfamilies Ponerinae and Cerapachyinae. Esakia 1987, 25, 97 - 132. [CrossRef]","49. Yoshimura, M.; Onoyama, K. Male-based keys to the subfamilies and genera of Japanese ants (Hymenoptera: Formicidae). Entomol. Sci. 2002, 5, 421 - 443.","46. Keller, R. A. A phylogenetic analysis of ant morphology (Hymenoptera: Formicidae) with special reference to the poneromorph subfamilies. Bull. Am. Mus. Nat. Hist. 2011, 355, 1 - 90. [CrossRef]"]}

Genus † Desyopone gen. nov. Boudinot and Perrichot Type species. † Desyopone hereon sp. nov., by present designation monotypy. ZooBank LSID: urn:lsid:zoobank.org:act: 7228E671-DF5E-44D1-ADE6-62FB6DB34220. Etymology. The genus name is a portmanteau of the traditional ponerine generic suffix, “-pone”, and the acronym for the Deutsches Elektronen-Synchrotron (DESY), whose storage ring and radiation beamline facilities were used to generate the phenomic data that were crucial for the correct identification of the new taxon. Diagnosis. † Desyopone has plesiomorphies 1–4 of Ponerinae and is identifiable as Ponerini at minimum due to the vestigial mandibles and infraaxial helcium. † Desyopone and Cryptopone are uniquely identified among all Ponerinae by: (1) subpetiolar process completely absent, with the poststernite low and nearly flat in profile. The new genus differs from the males of all known Cryptopone by the following: (2) head broader than long, excluding the compound eyes (vs. head narrower than long); (3) mandibles elongate (vs. short); (4) mandibles lobate (vs. spiniform); (5) mesospiracular sclerite evidently absent (vs. this sclerite present); (6) meso- and metatibiae with no spur and one spur, respectively (vs. two spurs each); and (7) petiolar peduncle long, about as long as node is tall (vs. peduncle short, considerable shorter than height of node). Remarks. The identity of Cryptopone is significantly clarified by the phenomic data from the new species and the phylogenomic revision of Branstetter and Longino [50]. Prior to this work, the diagnostic importance of the absent subpetiolar process was obscured by the inclusion of Wadeura guianensis in Cryptopone. Now it is clear that the absence of the subpetiolar process is a unique condition among extant Ponerinae that is shared between † Desyopone gen. nov. and Cryptopone, and thus constitutes a reasonable autapomorphy within the subfamily for the two genera. No known Cryptopone, however, matches the diagnostic character combination of † Desyopone gen. nov., with conditions 3–7 being apomorphic. Critically, the elongate peduncle of † D. hereon gen. et sp. nov. is nearly unique among Ponerinae; this condition is similarly derived in Harpegnathos and is approached by Dinoponera, some Odontomachus (e.g., O. chelifer, O. coquereli), and Platythyrea (although node at middle of segment rather than posterior).

† Desyopone hereon sp. nov. Boudinot and Perrichot (Figures 1–5) ZooBank LSID: urn: lsid:zoobank.org:act: 9E345965-6AA1-42D8-A468-187BABBB38D2. Etymology. The specific epithet gratefully recognizes the Helmholtz-Zentrum Hereon, the research center which funds and operates the Imaging Beamline (P05) at DESY, thus making the present work possible. Holotype. Male (m), MAIG 6016 (Figures 1B, 2A,B, 3 and 4), deposited in the Museum of Amber Inclusions at University of Gda´nsk, Poland. Paratypes. 12 males (m). Synincluded with holotype in amber piece MAIG 6016 (Figures 1A,C,D, 2C,D and 5). Type locality. Exact locality unknown in the Bashilo river gorge near Weldiya, Semien Wollo Zone, Amhara Region, Ethiopia. Type horizon. A fine siltstone/mudstone of Early Miocene age (16–23 Ma). Material. Holotype and paratypes 1–4 are nearly complete, preserved without apparent distortion but with integument entirely covered by a white, opaque (bacterial?) coat for paratypes 1, 2, 4 (Figures 1A,C and 2A). Paratypes 5–12 mostly complete but variously preserved, more or less distorted by apparent dehydration (e.g., Figure 1D). Diagnosis. † Desyopone hereon is uniquely identifiable among all Ponerinae by the distinctly elongate petiolar peduncle and the enlarged, lobate mandibles. Description. Measurements: Holotype (paratypes) (taken from paratypes 1,2,4)—BL 3.11 (3.00–3.50), HW 0.57 (0.50–0.60), HWE 0.70 (0.66–0.70), SL 0.12 (0.10–0.12), EL 0.23, OLL 0.08 (0.08), OIL 0.16 (0.16), WL 1.25 (1.25), ML 0.52 (0.50–0.56), MW 0.49, FWL (2.75), PH 0.26 (0.30), PL ~0.40 (0.40), GL 1.20 (1.00–1.50), GW 0.62 (0.65–0.80), HWI 81 (76–86), SI 21 (20), OCI 50 (50), PI 154 (133). Head (Figures 2–4): Head capsule ovoid-elliptical in lateral view; in full-face view, posterior head margin broadly and evenly convex to compound eyes; oral region of head, i.e., the malar areas, clypeus, and mouthparts, narrower than distance between compound eyes; postgenal bridge short, about 2/5 the length of the head in full-face view as measured from the postocciput to the hypostoma; malar areas distinctly developed; clypeus medially bulging, laterally depressed. Compound eyes situated almost entirely in anterior half of head; eyes subspherical and relatively small, with their length being about 1/3 head length. Ocelli distant from compound eyes, with the lateral ocelli separated from the compound eyes by slightly more than one compound eye length. Antennal toruli located at about head midlength in full-face view, distinctly posterad the posterior clypeal margin. Antennal scapes short, just barely longer than wide, with their length distinctly Mesosoma (Figures 2 and 3): Pronotum short and simple but with distinct muscular convexity as seen in lateral and dorsal views; posterad the anteromedian pronotal lobe (“nuchal lobe”), pronotum in the form of a simple arch, without distinct dorsal and ventral surfaces. Propleurae widely emarginate posteromedially, together forming a broad arch for the prosternum. Prosternum with basisternum apparently arcuate anteriorly; prosternal process developed. Mesoscutum somewhat narrow, with the anteroposterior length slightly greater than the lateromedial width. Notauli developed, and Y-shaped; meeting in the posterior half of the mesoscutum. Parapsides developed, although indistinct. Scutoscutellar sulcus broad, with at least five cross-costae. Mesoscutellum simple, convex, longer than broad in full-face view. Oblique mesopleural sulcus developed. Spiracular lobe (ventrad wing insertions) absent. Mesopleural area divided into dorsal and ventral regions; both regions dorsoventrally taller than anteroposteriorly long. Mesosternal and metasternal regions without processes. Propodeum convex, without armature or distinct sculpturation. Propodeal lobes developed, weak. Legs: Mesotibiae with no spur, metatibiae with a single, pectinate spur. Pretarsal claws narrow and simple, without additional teeth. Arolia well-developed but not very large. Fore wings (Figures 2C,D and 5): Costal vein (C) present, complete. Rsf1 diverging from Sc+ R +Rs well proximad pterostigma, with Sc+ R abscissa about 1/3 pterostigma length. Pterostigma well-developed, long, and narrow, with its length> 5 × its width. Rf distad pterostigma tubular. M+Cu tubular. Mf1 diverging from M+Cu at or slightly distad crossvein cu-a; this abscissa very weakly curved and meeting Rf1 at a distinct oblique angle. Rs+M tubular. Rsf2–3 diverging from Rs+M proximad 1m-cu. Crossvein 1r-rs absent. Crossvein 2r-rs anterior juncture at 2/3 length of pterostigma; this crossvein meeting Rsf proximad crossvein 2rs-m, which is tubular (paratype 4 has 2rs-m duplicated on the right wing, see Figure 2D). Rsf4+ tubular, meeting Rf distally). Mf2 (=abscissa between Rs+M and 1m-cu) short, shorter than 2r-rs and 2rs-m. Mf3 similar in length to but distinctly shorter than Rsf2–3. Mf4+ tubular proximally, becoming nebulous distally, with this occurring at a distance that is about 3 × the length of 2rs-m. Cuf3 (=abscissa of Cu after 1m-cu) joined to 1A posteriorly; 1A is tubular for its entire length. Submarginal cells 1 and 2 similar in size and shape, but with 2 distinctly smaller than 1. Marginal cell 1 long and narrow, with a length that is about 1.5 × pterostigma length. Discal cell 1 subrhomboidal, its length slightly less than 2 × its anteroposterior width. Subdiscal cell closed. Hind wings (Figures 2C and 5): Wing with eight distal hamuli. Jugal lobe absent. C not distinctly developed. R splitting from Sc+ R +Rs distad crossvein 1rs-m; Rf incomplete, not reaching anterior wing margin. Rsf tubular for a distance that is about 2 × the length of Sc+ R +Rs distad 1rs-m. M+Cu splitting well distad crossvein cu-a. Mf1 meeting rs-m at a broad, oblique angle; Mf developed as a stud distad this juncture. Cuf developed distad the split of M+Cu, but exact condition uncertain. Crossvein cu-a long, slightly longer than the length of Mf1; this crossvein situated proximad split of M+Cu by about twice its length. Anal vein (A) tubular past its juncture with cu-a. Anal cell relatively long; M+Cu proximad cu-a distinctly longer than M+Cu distad cu-a. Metasoma (Figures 2B and 3A,D,E): Petiole nodiform and distinctly pedunculate, albeit without a marked inflection between its anterior portion and the anterior surface of the petiolar node; peduncle about 2/5 petiole length; petiolar node height about 3/5 entire petiole length; node broad and convex; posterior collar well-developed; tergosternal fusion absent, laterotergites present; sternum low and very weakly sinuate in lateral view, without an anteroventral (=subpetiolar) process, nor with a posterior process; posterior sternal margin distinctly notched. Helcium infraaxial (=below abdominal segment III midheight); helcial tergite broad and overlapping sternite laterally. Prora not distinctly developed. Gastral segments homonomous in appearance and gradually decreasing in length from abdominal segment III to VIII. Abdominal segment IV without cinctus (=constriction). Abdominal spiracles IV–VIII obscured by preceding tergites. Abdominal tergum VIII apparently simple. Abdominal sternum IX lobate and somewhat narrow, with a length that is about twice its width. Cerci (=pygostyles) developed. Genitalia (Figures 2B and 3A): Cupula present (only visible in µ- CT cross-sections). Gonopods longer than tall. Gonostyli broad proximally, indistinctly set off from gonocoxa, and lobate in appearance, being narrowly rounded apically. Lateropenites (=digiti) thickened apically, downcurved. Penites (=penisvalvae) apparently longer than tall, curving strongly to their narrowly lobate apices. Setation: Body with two primary hair classes: (1) short pubescence, which covers the head and all segments of the antennae, mesosoma, legs, and metasoma; (2) long hairs, which are sparse on all body regions, but are denser on the metasoma., Published as part of Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej & Perrichot, Vincent, 2022, Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae), pp. 1-19 in Insects 73 (796) on pages 9-13, DOI: 10.3390/insects13090796, http://zenodo.org/record/7046409

Subfamily Aneuretinae Emery, 1913. Type genus. Aneuretus Emery, 1893. Diagnosis (all adult castes). All adults of Aneuretinae possess the following diagnostic plesiomorphies (1–6): (1) mandibles shovel-shaped (=“triangular”) [plesiomorphy of Poneroformicia, or at least Doryloformicia]; (2) meso- and metatibiae with one spur each [synapomorphy of clade Dolichoderomorpha]; (3) petiole with complete tergosternal fusion [synapomorphy of clade Dolichoderomorpha]; (4) petiole with an elongate anterior peduncle [possible synapomorphy of clade Myrmechoderines]; (5) helcium infraaxial [possible synapomorphy of clade Myrmechoderines]; and (6) abdominal segment IV unconstricted [synapomorphy of clade Dolichoderomorpha]. Males and queens of Aneuretinae share the following diagnostic traits (7–12): (7) near-complete fore wing venation, with only the subdiscal cell open; (8) crossvein 2rs-m furcal to post-furcal, i.e., 2r-sm meeting Rs at or distad 2r-rs; (9) well-prefurcal fore wing crossvein cu-a, i.e., cu-a meeting M+Cu proximad the split of M+Cu by more than one of its own lengths; (10) absence of free M in the hind wing after the juncture of rs-m and Mf1, i.e., the abscissa between Sc+ R +Rs and M+Cu linear, without a kink; (11) hind wing anal cell short, its length less than half that of the basal cell; and (12) absence of the hind wing jugal lobe. Males of Aneuretinae have the following diagnostic plesiomorphy (13): (13) genital gonocoxa and gonostylus not strongly differentiated in size, with the dorsal gonocoxal margin continuing more-or-less evenly to that of the gonostylus. Workers and queens of Aneuretinae share the following diagnostic plesiomorphies (14, 15): (14) mandible with biseriate dentition, i.e., with small teeth interspersed between large teeth [synapomorphy of Dolichoderomorpha]; (15) basal and masticatory margins of mandible not marked, i.e., these margins curving into one another, without a distinct angle [synapomorphy of Dolichoderomorpha]. Remarks. The operational paleontological definition of Aneuretinae has relied on character states 2, 4, 6, 7 (regardless of subdiscal cell state), and 12. With the explicit recognition of character state 3—which was previously indicated for the “formicomorph subfamilies” by Bolton [11] (p. 16)—it is possible to reject the placement of † Desyopone hereon gen. et sp. nov. from the Aneuretinae. The condition of helcial axiality is here reinterpreted from Bolton [11] (p. 18), who described the helcium of Aneuretinae as “high on [the] anterior face of abdominal segment III”, which is interpretable as supraaxial sensu Keller [46]. Although the helcial tergite of worker Aneuretus is dorsoventrally short, it can be seen that the helcium is at the ventralmost position of the sternum, which does not broaden. An axial helcium is confirmed for the Baltic amber taxa † Paraneuretus and † Protaneuretus as well. Wing venation was observable for Aneuretus and † Paraneuretus. Finally, we recognize the mandibular character states 14 and 15 as critical for the identification of Aneuretinae. The states of the mandibles have not been previously remarked upon, but along with the conformation of the clypeus (not defined here), they form the gestalt of the Aneuretinae and Dolichoderinae, which was likely used by Wheeler [7] to place † Paraneuretus and † Protaneuretus, although his justifications were not made explicit. Further refinement of the aneuretine diagnosis via comparative phenomics and traditional comparative morphology is highly desirable., Published as part of Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej & Perrichot, Vincent, 2022, Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae), pp. 1-19 in Insects 73 (796) on pages 6-7, DOI: 10.3390/insects13090796, http://zenodo.org/record/7046409, {"references":["11. Bolton, B. Synopsis and classification of Formicidae. Mem. Amer. Entomol. Inst. 2003, 71, 1 - 370.","46. Keller, R. A. A phylogenetic analysis of ant morphology (Hymenoptera: Formicidae) with special reference to the poneromorph subfamilies. Bull. Am. Mus. Nat. Hist. 2011, 355, 1 - 90. [CrossRef]","7. Wheeler, W. M. The ants of the Baltic Amber. Schrift. Physikal. Okonom. Gesell. Konigsberg 1915, 55, 1 - 142."]}

Tribe Platythyreini Emery, 1901. Type genus. Platythyrea Roger, 1863. Male diagnosis. In addition to the ponerine plesiomorphies, male Platythyreini are distinguished by the following: (1) mandibles worker-like, shovel-shaped (=“triangular”) [plesiomorphy]; (2) face between antennal toruli distinctly raised, such that the antennal toruli are directed relatively laterad, and are situated dorsad a depression which may receive the antenna [synapomorphy, homoplastic in Ponerini]; (3) antennal toruli usually close to or indenting the posterior margin of clypeus, toruli never distant from the posterior margin by more than half of one of their diameters [apomorphy]; (4) meso- and metatibiae with two spurs each [plesiomorphy]; (5) jugal lobe present [plesiomorphy]; (6) helcium not distinctly infraaxial, being axial to more-or-less axial, i.e., at about or just slightly below the midheight of abdominal segment III [plesiomorphy]; (7) abdominal segment IV with cinctus [plesiomorphy]; (8) cuticle pruinose [synapomorphy, homoplastic in Ponerini and some Proceratiinae]. Remarks. Because male Platythyreini have never been explicitly diagnosed, we found it necessary to provide a diagnosis in order to confirm the identification of the fossils in question. We observe that the mandibular form, tibial spur count, and cuticular sculpture of male Platythyreini are sufficient for identification at the global scale., Published as part of Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej & Perrichot, Vincent, 2022, Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae), pp. 1-19 in Insects 73 (796) on pages 7-8, DOI: 10.3390/insects13090796, http://zenodo.org/record/7046409

† CRETACEOZOMUS ANGUSTOCAUDATUS DE FRANCESCO MAGNUSSEN SP. NOV. FRANCESCO MAGNUSSEN SP. NOV. (FIGS 6–8) Zoobank registration: urn:lsid:zoobank.org:act: 7B08CF32-2C1F-4507-BA94-C87B97CFCCFB Type material: Holotype (F i g. 6A –D), a d u l t homeomorphic ♂, collection no. GPIH04896; ex collection Patrick Müller (no. BUB06). Diagnosis † Cretaceozomus angustocaudatus differs from † Cretaceozomus robustus by its thick pedipalps, femur of leg IV 3.1 times longer than deep and flagellum slender and oblong. Etymology The specific epithet is derived from the Latin words angustus, narrow, and cauda, tail, referring to the shape of the male flagellum. It is masculine in gender. Description Total length from anterior dorsal process to base of the flagellum ~ 3 mm. Colour (in amber): Dorsal side dark brown; patella, tibia and tarsus of the legs II–IV golden to light brown; ventral side golden brown; flagellum and pedipalps dark brown, original colouring equivocal. Prosoma (Fig. 7A, B): Length 1.36 mm; propeltidium length 1.12 mm, width 0.56 mm. Anterior process long and broad, length 0.18 mm; setation: two setae next to each other followed by a single median seta. Propeltidium setation: four pairs of setae, first pair aborted, tubercles visible, following three pairs left ones visible, right ones aborted. Corneate eyes present, clearly elevated above the surface of the propeltidium. Mesopeltidium width (left) 0.2 mm. Metapeltidium not divided (Fig. 8A, B), length 0.32 mm, width 0.64 mm. Anterior sternum with> 10 + 2 sternapophysial setae, length 0.31 mm, width 0.26 mm. Posterior sternum with seven setae, length 0.19 mm, width 0.25 mm. Chelicerae: Setation of the mesal side of chelicerae not visible, except for G6-1. Pedipalps: Robust, without armature except for a mesal spur on trochanter (Fig. 7C, D); trochanter produced distally, setation: one+ mesal, five ventral, two ventroectal, one+ ectal; femur: longer than trochanter, setation: eight+ dorsal, two+ dorsomesal, two+ ectal, three + mesal, one + ventral, two + dorsoectal, t w o+ v e n t r o e c t a l; p a t e l l a: l o n g e r t h a n f e m u r, setation: one+ mesal, four+ ectal, two+ dorsomesal, two+ dorsoectal, five dorsal; tibia: shorter than patella, setation: six dorsomesal, two+ ventromesal, four+ ectal, six dorsoectal, five+ mesal, six dorsal; tarsus: setation: two+ dorsoectal, two+ dorsomesal; claw not visible. Legs: Right leg I aborted at the end of the trochanter, left leg incomplete, tarsus missing, leg I ≥ 1.05 times longer than the body; anterodorsal margin of femur of leg IV produced at an angle of ~90°, femur of leg IV 3.1 times longer than deep, leg IV> 1.08 times longer than the total body length; leg formula presumed: 1423. Opisthosoma: Not entirely visible, length 1.60– 1.65 mm, width 0.9 mm; segments X–XII not elongated, segments IX–XII not visible owing to overlaying debris. Abdominal tergites setation: I–IV: Dm; V: not visible; VI: Dm, Dl1; VII: Dm, Dl1; VIII: Dm, Dl1; XI–XII not visible; setation pattern: 2:2:2:2:?:4:4:4:?:?:?:?. Flagellum: slim, dorsoventrally flattened with a slender and oblong shape (Fig. 8C, D), length 0.27 mm, width 0.18 mm; stalk length 0.09 mm; setation: Dm1 not visible; Dm4; Dl2 paired, sitting lateral; Dl3 paired, with the left one aborted; Vm1; Vm2 paired; Vm3 paired; Vm5; Vl2 paired. Pair Vl1 absent in this species. Further measurements: Pedipalp: trochanter 0.41 mm; femur 0.53 mm; patella 0.63 mm; tibia 0.49 mm; tarsus> 0.20 mm; claw not measurable; total length> 2.26 mm. Leg I: trochanter 0.20 mm; femur 1.09 mm; patella 1.11 mm; tibia> 0.75 mm; metatarsus not measurable; tarsus not measurable; total length> 3.15 mm. Leg II: trochanter 0.18 mm; femur 0.73 mm; patella 0.43 mm; tibia 0.51 mm; metatarsus 0.34 mm; tarsus (tarsus segments) 0.34 mm (0.12 mm; 0.08 mm; 0.14 mm); total length 2.53 mm. Leg III: trochanter 0.20 mm; femur 0.67 mm; patella 0.33 mm; tibia 0.38 mm; metatarsus 0.29 mm; tarsus (tarsus segments) 0.32 mm (0.14 mm; 0.08 mm; 0.10 mm); total length 2.19 mm. Leg IV: trochanter 0.22 mm; femur 1.09 mm; patella 0.36 mm; tibia 0.61 mm; metatarsus 0.56 mm; tarsus (tarsus segments) 0.39 mm (0.16 mm; 0.09 mm; 0.14 mm); total length 3.23 mm., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on pages 802-806, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082

† GROEHNIZOMUS RODRIGOI S.P. MÜLLER SP. NOV. (FIGS 20–23) Zoobank registration: urn:lsid:zoobank.org:act: 97BC6CF7-BF6A-48C1-968E-069699D99D4A Ty p e m a t e r i a l: Holotype (Fig. 2 0A–D), adult homeomorphic ♂, collection no. GPIH04891, ex collection Patrick Müller (no. BUB83). Diagnosis † Groehnizomus rodrigoi differs from † G. oculiferans by the state of the metapeltidium, which is divided in † G. oculiferans and entire in † G. rodrigoi. Although the state of the metapeltidium is no longer considered diagnostic at genus level (Lawrence, 1969), this character still shows great consistency within a given genus. Therefore, we consider † G. rodrigoi to be distinct from † G. oculiferans. Etymology The genus name is a patronym for Rodrigo MonjarazRuedas from the Universidad Nacional Autónoma de México and honours his outstanding contribution to modern schizomid taxonomy and phylogenetics. It is masculine in gender. Description Total length from anterior margin of the propeltidium to base of flagellum 2.72 mm. C o l o u r: L i g h t–d a r k b r o w n, o r i g i n a l c o l o u r i n g equivocal. Prosoma: Length 1.08 mm; propeltidium length 0.87 mm, width 0.38 mm. Anterior process with a single seta followed by a pair of setae. Further setation of propeltidium a single seta followed by four pairs of setae, all four pairs aborted (Fig. 21C, D). Corneate eyes present, clearly elevated above the surface of the propeltidium. Mesopeltidia length> 0.13 mm, width equivocal. Metapeltidium not divided (Fig. 21A, B), length 0.58 mm, width 0.28 mm. Anterior sternum with 12 + 2 sternapophysial setae. Posterior sternum with 5+ setae. Pedipalps: Stout and clearly shorter than the body length, without armature (Fig. 22A, B), mesal spur not visible; trochanter not produced anteriorly, setation: six strong, long ventral, six+ small ventromesal–ventroectal; femur setation: six+ dorsal, one dorsomesal, three ventral; patella slightly longer than femur, setation: three dorsal, three dorsomesal, one dorsoectal, two ventroectal, two ventral; tibia shorter than patella, setation: three+ dorsal, four dorsoectal, nine ventroectal–ventromesal, mesal setation equivocal; tarsus about two-thirds length of tibia, setation: three+ dorsoectal, three mesal, four ventromesal, dorsal setation equivocal; tarsal spurs equivocal; claw slightly shorter than half the length of tarsus. Chelicerae: Setation of the mesal side of chelicerae not visible, except for G6-1. 822 I. DE FRANCESCO MAGNUSSEN ET AL. Legs: Anterodorsal margin of femur of leg IV produced at an angle of ~90°; leg IV 1.37 times longer than the body, femur of leg IV slender, 3.27 times longer than deep; leg formula: 1423. Opisthosoma: Length 1.63 mm, width 0.60 mm; tergites X–XII not elongated. Abdominal tergites setation: I: Dm, five microsetae; II: Dm; III–XII: disrupted; setation pattern: 2 + 5:2:?:?:?:?:?:?:?:?:?:?. Flagellum: dorsoventrally flattened, rounded to subrhomboidal (Fig. 23A–D), length 0.30 mm, width 0.24 mm; stalk length 0.07; setation: Dm1 (lost), Dl2 paired, Dm4 (lost), Dl3 paired, Vm1, Vm2 paired, Vm3 paired, Vl1 paired, Vm5, Vl2 paired. Further measurements: Pedipalp: trochanter 0.36 mm; femur 0.49 mm; patella 0.55 mm; tibia 0.36 mm; tarsus 0.21 mm; total length 1.97 mm. Leg I: trochanter 0.27 mm; femur 1.20 mm; patella 1.52 mm; tibia 1.04 mm; metatarsus 0.34 mm; tarsus 0.41 mm; total length 4.78 mm. Leg II: trochanter 0.17 mm; femur 0.69 mm; patella 0.38 mm; tibia 0.52 mm; metatarsus 0.47 mm; tarsus 0.37 mm; total length 2.60 mm. Leg III: trochanter 0.16 mm; femur 0.66 mm; patella 0.26 mm; tibia 0.38 mm; metatarsus 0.43 mm; tarsus 0.39 mm; total length 2.28mm. Leg IV: trochanter 0.24 mm; femur 1.21 mm; patella 0.49 mm; tibia 0.72 mm; metatarsus 0.65 mm; tarsus 0.41 mm; total length 3.72 mm., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on pages 818-822, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082, {"references":["Lawrence RF. 1969. The Uropygi (Arachnida: Schizomidae) of the Ethiopian Region. Journal of Natural History 3: 217 - 260."]}

† ANNAZOMUS DE FRANCESCO MAGNUSSEN & S.P. MÜLLER GEN. NOV. (FIGS 3–5) Zoobank registration: urn:lsid:zoobank.org:act: 8FC98781-C14B-4E05-8E6F-A8EFD928E713 Type species: † Annazomus parvulus De Francesco Magnussen sp. nov. Remarks The tarsal spurs are not visible, but the specimen differs from species of Protoschizomidae by the lack of the pair of setae at the base of the anterior process and the presence of a distinct stalk at the male flagellum. It also differs from members of the Megaschizominae by having a smaller body size, the lack of eight or nine setae on its anterior process and the presence of only one pair of setae on the tergites II–VII. Therefore, it is clearly a member of subfamily Hubbardiinae. The specimen cannot be assigned to any extant genus because of its combination of characters. Diagnosis † Annazomus differs from all other hubbardiid genera by the following combination of characters: propeltidium with three pairs of setae; corneate eyes or eyespots absent; metapeltidium not divided; body without clavate setae; anterodorsal margin of femur of leg IV produced at an angle of ~90°; tergite II with one pair of setae, tergites X–XII not elongated; tergite XII with posterodorsal process; male flagellum dorsoventrally flattened. KEY TO THE FOSSIL GENERA FROM BURMESE AMBER 1. Propeltidium without single median seta …................................................................................................. 2– Propeltidium with single median seta…........................................................................................................ 4 2. Propeltidium with three pairs of setae, corneate eyes and eyespots absent, male flagellum shape rectangular–bulbous….................................................................................................. † Annazomus parvulus – Anterior process with a 2 + 1 setation, propeltidium with four pairs of setae, corneate eyes present …......................................................................................................................................................... 3 († Cretaceozomus) 3. Male flagellum shape long–bulbous …....................................................... † Cretaceozomus angustocaudatus – Male flagellum shape bulbous–subtriangular.........................................................† Cretaceozomus robustus 4. Segment XII without posterodorsal process …................................................................................................ 5– Segment XII with posterodorsal process …..................................................................................................... 6 5. Fixed finger of chelicerae with two small teeth between two larger outer teeth, femur IV length-to-width ratio Mesozomus groehni 6. Propeltidium with four pairs of setae, male flagellum shape rhomboidal–subtriangular …....................................................................................................................................................................... 7 († Groehnizomus) – Propeltidium with three pairs of setae, male flagellum shape well rounded to subtriangular …................................................................................................................................................................. 8 († Muellerizomus) 7. Metapeltidium entire, claw-to-tarsus ratio bigger than half the length of the tarsus......................................................................................................................................................................… † Groehnizomus rodrigoi – Metapeltidium divided, claw-to-tarsus ratio smaller than half the length of the tarsus …............................................................................................................................................................. † Groehnizomus oculiferans 8. Corneate eyes and eyespots absent, metapeltidium divided, female flagellum with four flagellomeres …....................................................................................................................................... † Muellerizomus amandae – Eyespots present, metapeltidium entire …........................................................ † Muellerizomus palicaudatus Etymology The genus name was chosen in honour of the first author’s grandmother Anna and is combined with the suffix -zomus, which is commonly used for schizomids. It is masculine in gender. Comparison † Annazomus is most similar to the new genus † Muellerizomus (described below), which has a single median seta followed by three pairs of setae on the propeltidium, whereas † Annazomus has three pairs of setae on the propeltidium. † Muellerizomus has a rounded and flat flagellum with an apical tip and a w-shaped depression on the dorsal side, but in † Annazomus the male flagellum is nearly rectangular, with a bulbous elevation on each dorsal side without any depressions. For comparison with Recent genera, see the Supporting Information (Table S1).

† ANNAZOMUS PARVULUS DE FRANCESCO MAGNUSSEN SP. NOV. (FIGS 3–5) Zoobank registration: urn:lsid:zoobank.org:act: 9221DD2F-0C17-48AC-8904-C18E18A05F96 Type material: Holotype (Fig. 3A –D), adult homeomorphic ♂, collection no. GPIH 04893; ex collection Patrick Müller (no. BUB556). Diagnosis Pedipalps short; flagellum shape nearly rectangular, with a bulbous elevation on the left and right dorsal side; small body size of 1.95 mm. Etymology The specific epithet is Latin parvulus, small, referring to the small body size of the specimen. It is masculine in gender. Description Total length from the anterior process to the base of the flagellum 1.95 mm. Colour (in amber): Light brown and golden; prosoma, opisthosoma, flagellum, femur IV and pedipalps slightly darker brown golden; leg I golden and translucent, original colouring equivocal. Prosoma: Length 0.83 mm; propeltidium length 0.68 mm, width 0.32 mm. Anterior process sharply bent down, length not measurable; setation: one seta, sitting on the left side of the anterior process, no further setation visible; small round holes in the proximal part of the propeltidium prevent reconstruction of further setation, but there might have been a pair of setae with the right one now missing, definite setation remains unclear. Propeltidium setation prosoma: three pairs, first pair of setae still in place, second pair left one aborted, third pair both aborted with only left tubercle visible, right one not visible owing to a crack in the amber covering parts of the propeltidium. Corneate eyes and eyespots absent. Mesopeltidia width (right): ~ 0.11 mm, left one not visible owing to an air bubble in the amber. Metapeltidium not divided with a thin suture (Fig. 4A, B), length 0.2 mm, width 0.42 mm. Anterior sternum with 10 + 2 sternapophysial setae, length 0.24 mm, width 0.22 mm. Posterior sternum with six setae, length 0.12 mm, width 0.11 mm. Chelicerae: Setation of the mesal side of chelicerae not visible, except for G6-1. Pedipalps: Short and slim, without armature (Fig. 4C, D), mesal spur on trochanter not visible; trochanter setation: five ventral strong setae, two ectal, four mesal setae; femur: longer than trochanter, setation: five dorsal, three ectal, two ventroectal, three mesal; patella: shorter than femur, setation: four mesal, two ectal, four dorsomesal, two dorsoectal, three dorsal; tibia: shorter than patella, setation: three dorsomesal, three ectal, two ventroectal, two dorsoectal, five mesal; tarsus: with tarsal spurs, spur length: 0.02 mm, setation: three ectal, four dorsoectal, four ventroectal, two dorsal, three dorsomesal, three+ ventromesal, five mesal, two+ ventral; claw: straight, slightly longer than half the length of tarsus. Legs: Right leg I aborted at the trochanter, left leg well preserved and complete, tarsus with six segments, leg I ~1.37 times longer than the body; anterodorsal margin of femur of leg IV produced at an angle of ~90°, femur of leg IV 2.65 times longer than deep; leg IV> 1.04 times longer than the total body length; leg formula: 1423. Opisthosoma: Length 1.12 mm, width 0.42 mm; tergites X–XII not elongated, tergite XII with a large, rectangular posterodorsal process. Abdominal tergites setation: I: Dm; II: Dm; III: Dm, left missing; IV: Dm, both missing; V: Dm, both missing; VI: Dm; VII: Dm, both missing; VIII: Dm, left missing, Dl1; IX: Dl1, Dl2; X: Dl1; XI: Dl1; XII: Dm, Dl1, setation pattern: 2:2:2:2:2:2:2:4:4:2:2:4. Flagellum: dorsoventrally flattened, rectangular with an apical tip at the posterior end, on the dorsal side with a bulbous elevation each left and right near the posterior end (Fig. 5A–G), length 0.17 mm, width 0.22 mm; stalk length ~ 0.09 mm; setation: Dm1, Dl2 paired, Dm4, Dl3 paired and nearly at the posterior end of the flagellum, Vm1, Vm2 paired, Vm3 paired, Vl1 paired, Vm5, Vl2 paired. Further measurements: Pedipalp: trochanter 0.16 mm; femur 0.26 mm; patella 0.23 mm; tibia 0.20 mm; tarsus 0.13 mm; claw 0.07 mm; total length 1.05 mm. Leg I: trochanter 0.19 mm; femur 0.61 mm; patella 0.71 mm; tibia 0.58 mm; metatarsus 0.24 mm; tarsus (tarsus segments) 0.34 mm (0.04 mm; 0.04 mm; 0.04 mm; 0.04 mm; 0.05 mm; 0.13 mm); total length 2.67 mm. Leg II: t r o c h a n t e r 0.1 2 m m; f e m u r 0.4 9 m m; p a t e l l a 0.22 mm; tibia 0.25 mm; metatarsus 0.29 mm; tarsus (tarsus segments) 0.26 mm (0.10 mm; 0.06 mm; 0.10 mm); total length 1.63 mm. Leg III: trochanter 0.13 mm; femur 0.48 mm; patella 0.17 mm; tibia 0.19 mm; metatarsus 0.25 mm; tarsus (tarsus segments) 0.26 mm (0.12 mm; 0.06 mm; 0.08 mm); total length 1.48 mm. Leg IV: trochanter 0.19 mm; femur 0.70 mm; patella> 0.22 mm; tibia> 0.29 mm; metatarsus 0.35 mm; tarsus (tarsus segments) 0.2 8 m m (0.1 1 m m; 0.0 6 m m; 0.1 1 m m); t o t a l length> 2.03 mm., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on pages 797-801, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082

† MUELLERIZOMUS AMANDAE DE FRANCESCO MAGNUSSEN & S.P. MÜLLER SP. NOV. MAGNUSSEN & S.P. MÜLLER SP. NOV. (FIGS 27–31) Zoobank registration: urn:lsid:zoobank.org:act: 63A0355B-C647-414E-8C0F-2C5BD807495E Type material: Holotype (Fig. 27 A – D), adult homeomorphic ♂, collection no. GPIH04895, ex collection Patrick Müller (no. BUB3553); paratype (Fig. 29A–D), adult ♀, GPIH04892, ex collection Patrick Müller (no. BUB557). Remarks The female specimen is matched with the male holotype of † Mu. amandae because both share the following characters: propeltidium with a single median seta followed by three pairs of setae; corneate eyes or eyespots absent; body without clavate setae; anterodorsal margin of femur of leg IV produced at an angle of ~90°; tergite II with one pair of setae; tergites X–XII not elongated. Diagnosis † Muellerizomus amandae differs from its congener † Mu. palicaudatus by the lack of eyespots and a more rounded male flagellum shape. Etymology The specific epithet name is a patronym for Dr Amanda Eva Bauzá Iraola, former directorial assistant of the Center of Natural History in Hamburg, for her outstanding commitment to staff and collectionbased science during the past years. It is feminine in gender. Description Total length from anterior dorsal process to base of the flagellum 1.83 mm. Colour (in amber): Golden brown; prosoma and pedipalps with a slightly darker touch of brown than the opisthosoma; femur and patella of leg IV light brown; all the other legs and flagellum golden yellowish and translucent. Prosoma: Length 0.80 mm; propeltidium length 0.47 mm, width 0.27 mm. Anterior process much rounded, without tip, length 0.03 mm; setation: one seta on the right side of the anterior process, which implies an aborted seta on the left, but definite setation remains unclear. Further propeltidium setation: a single median seta followed by three pairs of setae (paratype Fig. 30A, B). Neither corneate eyes nor eyespots present. Mesopeltidia and metapeltidium compressed owing to taphonomic processes to the point where the dimensions cannot be measured. Ventral side of the prosoma pressed inwards into the propeltidium. Anterior sternum: not entirely visible, with more than 3 + 2 sternapophysial setae, length and width not measurable owing to the compression of the ventral side of the prosoma. Posterior sternum not visible. Chelicerae: Setation of the mesal side of chelicerae not visible, except for G6-1. Pedipalps: Without armature, mesal spur not visible; trochanter hardly visible, setation: three+ strong setae ventral; femur: setation: two+ dorsal, two+ dorsoectal, two+ dorsomesal; patella: setation: four ventromesal, two mesal, four dorsomesal, three ectal, three dorsal; tibia: shorter than patella, setation: five dorsal, three ventromesal, four dorsoectal, three ventral; tarsus: with small, thin, asymmetrical tarsal spurs, spur length 0.01 mm, setation: two+ dorsal, four dorsomesal, four mesal; claw: bent downwards, slightly longer than half the length of tarsus. Legs: Legs I well preserved and complete on each side, tarsus with six segments, leg I ~1.42 times longer than the body; anterodorsal margin of femur of leg IV produced at an angle of ~90°, femur of leg IV 2.71 times longer than deep, leg IV 1.19 times longer than the total body length, leg formula: 1423. Opisthosoma: Length 1.03 mm, width 0.51 mm; segments XI – XII partly retracted into the opisthosoma, XII with a posterodorsal process, slightly bent upwards. Abdominal tergites setation: I–VIII: Dm; IX: Dl1; X: not visible; XI: Dl1, XII: Dm, Dl1, setation pattern: 2:2:2:2:2:2:2:2:2:?:2:4. Flagellum of male: dorsoventrally flattened, nearly round and flat, with an apical tip at the posterior end, two small depressions near the posterior end on the dorsal side of the flagellum (Fig. 28A–D), length 0.22 mm, width 0.21 mm; stalk length 0.06 mm; setation: Dm1, Dl2 paired, Dm4, Dl3 paired, Vm1, Vm2 paired, Vm3 paired, Vl1 paired, Vm5, Vl2 paired. Paratype female: Metapeltidium barely divided (Fig. 30C, D); pedipalp trochanter with mesal spur, claw slightly longer than half the length of the tarsus; body size 2.35 mm; leg IV 0.89 times longer than the total body length, femur of leg IV 2.76 times longer than deep; flagellum with three annuli, flagellomeres I, II, III, V + VI (Fig. 31A, B), total length 0.27 mm, lengths of segments: 0.05 mm, 0.05 mm, 0.03 mm, 0.12 mm; setation: Dm1, Dm4, Vm1 missing, Vm2 paired, left one probably lost, Vm3 paired, Vm5, Dl1 paired, right one lost, Dl2 paired, Dl3 paired, Dl4 absent, Vl1 paired, Vl2 paired. Holotype further measurements: Pedipalp: trochanter not measurable; femur not measurable; patella 0.28 mm; tibia 0.17 mm; tarsus 0.12 mm; claw 0.07 mm; total length not measurable. Leg I: trochanter 0.14 mm; femur 0.68 mm; patella 0.55 mm; tibia 0.7 mm; metatarsus 0.21 mm; tarsus (tarsus segments) 0.31 mm (0.03 mm; 0.04 mm; 0.04 mm; 0.03 mm; 0.05 mm; 0.12 mm); total length 2.59 mm. Leg II: trochanter> 0.05 mm; femur 0.41 mm; patella 0.26 mm; tibia 0.26 mm; metatarsus 0.25 mm; tarsus (tarsus segments) 0.22 mm (0.09 mm; 0.06 mm; 0.07 mm); total length> 1.45 mm. Leg III: trochanter 0.13 mm; femur 0.34 mm; patella 0.21 mm; tibia 0.22 mm; metatarsus 0.24 mm; tarsus (tarsus segments) 0.22 mm (not measurable); total length 1.36 mm. Leg IV: trochanter 0.16 mm; femur 0.65 mm; patella 0.24 mm; tibia 0.45 mm; metatarsus 0.39 mm; tarsus (tarsus segments) 0.29 mm (0.12 mm; 0.06 mm; 0.11 mm); total length 2.18 mm. Paratypefurthermeasurements: P edipalp:trochanterand femur not measurable; patella 0.25 mm; tibia 0.34 mm; tarsus 0.17 mm; total length not measurable. Leg I: trochanter 0.18 mm; femur 0.59 mm; patella 0.72 mm; tibia 0.53 mm; metatarsus 0.20 mm; tarsus 0.33 mm; total length 2.55 mm. Leg II: trochanter 0.13 mm; femur 0.41 mm; patella 0.27 mm; tibia 0.32 mm; metatarsus 0.24 mm; tarsus 0.26 mm; total length 1.63 mm. Leg III: trochanter 0.16 mm; femur 0.43 mm; patella 0.10 mm; tibia 0.19 mm; metatarsus 0.22 mm; tarsus 0.17 mm; total length 1.27 mm. Leg IV: trochanter 0.25 mm; femur 0.72 mm; patella 0.19 mm; tibia 0.34 mm; metatarsus 0.32 mm; tarsus 0.28 mm; total length 2.10 mm., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on pages 824-829, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082, {"references":["Muller SP, Dunlop JA, Kotthoff U, Hammel JU, Harms D. 2019. The oldest short-tailed whipscorpion (Schizomida): a new genus and species from the Upper Cretaceous amber of northern Myanmar. Cretaceous Research 106: 104227."]}

† CRETACEOZOMUS ROBUSTUS DE FRANCESCO MAGNUSSEN SP. NOV. MAGNUSSEN SP. NOV. (FIGS 9–11) Zoobank registration: urn:lsid:zoobank.org:act: 5CCD94A5-55B5-44A2-929C-F441016866F9 Type material: Holotype (F i g. 9A –D), a d u l t homeomorphic ♂, collection no. GPIH05048; ex collection Carsten Gröhn (no. 11206). Diagnosis † Cretaceozomus robustus can be characterized by the following features: femur of leg IV 2.76 times longer than wide; flagellum shape semi-oval but thicker near the stalk of the flagellum. Etymology The specific epithet is the Latin, robustus, stout, referring to the robust body shape of the specimen. It is masculine in gender. Description Total length from anterior dorsal process to base of the flagellum ~ 2.86 mm. Colour (in amber): Pedipalps dark brown reddish; prosoma dark brown to reddish; metapeltidial region lighter brown yellowish; femora dark brown-reddish, other leg segments golden-ochre; chelicerae dark brown; opisthosoma dark brown; flagellum golden brown; original colouring equivocal. Prosoma: Length 1.25 mm; propeltidium length 1.07 mm, width 0.66 mm. Anterior process bent downwards with an apical tip, length 0.07 mm; setation: two setae next to each other followed by a single median seta. Propeltidium setation: four pairs of setae, first and second pair complete, third pair missing the left and fourth pair missing the right one owing to preservation (Fig. 10C, D). Corneate eyes present, clearly elevated above the surface of the propeltidium. Mesopeltidium width 0.2 mm. Metapeltidium not divided (Fig. 10A, B), length 0.31 mm, width 0.73 mm. Anterior sternum not entirely visible, with> 4 + 2 sternapophysial setae, length 0.31 mm, width not measurable owing to taphonomic processes. Posterior sternum covered by a syninclusion and not visible. Chelicerae: Setation on the mesal side of chelicerae not visible, except for G6-1 and some setae of G1 and G2. Pedipalps: Without armature except for a mesal spur on trochanter; trochanter setation: six ventral strong setae, three ventroectal, three mesal setae in tubercles; femur: longer than trochanter, setation: seven dorsal, two ectal, two+ dorsomesal, two+ mesal; patella: shorter than femur, setation: three mesal, seven ectal, four dorsomesal, four dorsoectal, three dorsal; tibia: shorter than patella, setation: five dorsomesal, four ventromesal, five ectal, three dorsoectal, nine mesal, two dorsal; tarsus: with asymmetrical tarsal spurs, spur length: 0.03 mm, setation: five ectal, four dorsoectal, three dorsal, four dorsomesal, five mesal; claw: long and straight, slightly longer than half the length of tarsus. Legs: Right leg I aborted at the end of the coxa, left leg well preserved and complete, tarsus with six segments, leg I ~1.29 times longer than the body; anterodorsal margin of femur of leg IV produced at an angle of ~90°, leg IV 2.76 times longer than deep; leg IV 1.15 times longer than the total body length; leg formula: 1423. Opisthosoma: Length 1.61 mm, width 0.93 mm (opisthosoma compressed owing to preservation). Tergites X–XII not elongated, tergite XII with a small posterodorsal process. Abdominal tergites setation: I– VII: Dm; VIII: Dm, Dl1, left one missing; IX: Dl1, Dl2; X: not visible; XI: Dl1, Dl2; XII: Dm, Dl1; setation pattern: 2:2:2:2:2:2:2:4:4:?:4:4. Flagellum: dorsoventrally flattened, semioval shape, broader near the stalk, slightly more slender at the posterior end, tip rounded (Fig. 11A–D), length 0.34 mm, width 0.33 mm; stalk length 0.14 mm; setation: Dm1 not visible; Dm4; Dl2 paired, sitting lateral; Dl3 paired with the left one aborted; Vm1; Vm2 paired; Vm3 paired; Vm5; Vl2 paired with right one aborted; between Dl2 and Dl3 four microsetae on each side. Pair Vl1 absent in this species. Further measurements: Pedipalp: trochanter 0.39 mm; femur 0.63 mm; patella 0.56 mm; tibia 0.49 mm; tarsus 0.24 mm; claw 0.14 mm; total length 2.45 mm. Leg I: trochanter 0.23 mm; femur 0.84 mm; patella 0.94 mm; tibia 0.95 mm; metatarsus 0.35 mm; tarsus (tarsus segments) 0.39 mm (0.06 mm; 0.06 mm; 0.06 mm; 0.05 mm; 0.05 mm; 0.11 mm); total length 3.70 mm. Leg II: trochanter 0.16 mm; femur 0.73 mm; patella 0.45 mm; tibia 0.35 mm; metatarsus 0.42 mm; tarsus (tarsus segments) 0.36 mm (0.19 mm; 0.07 mm; 0.10 mm); total length 2.47 mm. Leg III: trochanter 0.20 mm; femur 0.72 mm; patella 0.30 mm; tibia 0.32mm; metatarsus 0.44 mm; tarsus (tarsus segments) 0.31 mm (0.13 mm; 0.06 mm; 0.12 mm); total length 2.29 mm. Leg IV: trochanter 0.21 mm; femur 1.16 mm; patella 0.39 mm; tibia 0.60 mm; metatarsus 0.55 mm; tarsus (tarsus segments) 0.39 mm (0.20 mm; 0.06 mm; 0.13 mm); total length 3.30 mm.

† MESOZOMUS S.P. MÜLLER, DUNLOP, KOTTHOFF, HAMMEL & HARMS, 2019 (FIGS 24–26) Type species: † Mesozomus groehni S.P. Müller, Dunlop, Kotthoff, Hammel & Harms, 2019, by original designation., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on page 822, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082, {"references":["Muller SP, Dunlop JA, Kotthoff U, Hammel JU, Harms D. 2019. The oldest short-tailed whipscorpion (Schizomida): a new genus and species from the Upper Cretaceous amber of northern Myanmar. Cretaceous Research 106: 104227."]}

† MUELLERIZOMUS PALICAUDATUS DE FRANCESCO MAGNUSSEN SP. NOV. (FIGS 32–34) Zoobank registration: urn:lsid:zoobank.org:act: FAEA5DC9-66A7-4500-822C-D37A86820FB3 Type material: Holotype Holotype (Fig. 32A–D), adult homeomorphic ♂, collection no. GPIH04894, ex collection Patrick Müller (no. BUB558). Diagnosis † Muellerizomus palicaudatus differs from other genera by a combination of characters: pedipalps short and delicate; flagellum spatulate and semicircular at the base but narrowed and triangular posteriorly; depression on the dorsal side of the flagellum shaped like a dumbell. Etymology The specific epithet is based on the Latin words pala, shovel, and cauda, tail, referring to the spatulate male flagellum. It is masculine in gender. Description Total length from the anterior process to the base of the flagellum 2.15 mm. C o l o u r (i n a m b e r): O c h r e – g o l d e n; p r o s o m a, opisthosoma, flagellum and pedipalps light brown– golden; legs greenish golden, original colouring equivocal. Prosoma (Fig. 33A, B): Length 0.95 mm; propeltidium length ~ 0.83 mm (owing to the anterior process being missing), width 0.25 mm. Anterior process ripped apart, no setation visible. Propeltidium setation: a single median seta followed by three pairs of setae, left one missing in second pair, only left tubercle visible in third pair. Eye spots present. Mesopeltidia width (left) 0.17 mm, right one not visible owing to an air bubble in the amber. Metapeltidium not divided, length 0.25 mm, width 0.48 mm. No sternum visible owing to air bubbles in the amber. Chelicerae: Setation of the mesal side of chelicerae not visible, except for G6-1. Pedipalps: Short, without armature except for a mesal spur on trochanter (Fig. 33C, D); trochanter setation: ventral five strong setae in bulbous tubercles, three mesal setae in tubercles; femur: longer than trochanter, setation: six dorsal, two dorsomesal, two mesal; patella: shorter than femur, setation: four mesal, four ectal, five dorsomesal, three dorsoectal, three dorsal, two ventral; tibia: shorter than patella, setation: four ventral, one ventroectal, five dorsal, four ectal, four dorsoectal, three mesal; tarsus: with asymmetrical tarsal spurs, spur length 0.02 mm, setation: three ectal, three dorsoectal, three dorsal, four dorsomesal, five mesal, four ventral; claw: long, about two-thirds as long as the length of tarsus. Legs: Left leg I aborted at the trochanter, right leg I well preserved and complete, tarsus with six segments, leg I ~1.58 times as long as the body; anterodorsal margin of femur of leg IV produced at an angle of ~90°, femur of leg IV 3.12 times longer than deep, leg IV 1.19 times longer than the total body length; leg formula: 1423. Opisthosoma: Length 1.2 mm, width 0.49 mm; segments X–XII not elongated, XII with a rounded posterodorsal process pointing upwards at an angle of ~45°. Abdominal tergites setation: I–VIII: Dm; IX: Dl1, Dl2; X: not visible; XI: Dl1; XII: Dm, Dl1; setation pattern: 2:2:2:2:2:2:2:2:4:?:2:4. Flagellum: dorsoventrally flattened, semicircular shape at the base that narrows down towards the posterior end into a triangular shape with a rounded tip, depression on the dorsal side of the flagellum shaped like a dumbbell (Fig. 34A–D), length 0.26 mm, width 0.24 mm; stalk length 0.08 mm; setation: Dm1, Dl2 paired, Dm4, Dl3 paired, Vm1, Vm2 paired, Vm3 paired, Vl1 paired, Vm5, Vl2 paired. Further measurements: Pedipalp: trochanter 0.19 mm; femur 0.29 mm; patella 0.25 mm; tibia 0.17 mm; tarsus 0.15 mm; claw 0.10 mm; total length 1.15 mm. Leg I: trochanter 0.19 mm; femur 0.79 mm; patella 0.99 mm; tibia 0.69 mm; metatarsus 0.28 mm; tarsus (tarsus segments) 0.46 mm (0.05 mm; 0.06 mm; 0.05 mm; 0.06 mm; 0.06 mm; 0.18 mm); total length 3.40 mm. Leg II: trochanter 0.14 mm; femur 0.59 mm; patella 0.29 mm; tibia 0.31 mm; metatarsus 0.30 mm; tarsus (tarsus segments) 0.30 mm (0.09 mm; 0.10 mm; 0.11 mm); total length 1.93 mm. Leg III: trochanter 0.15 mm; femur 0.52 mm; patella 0.19 mm; tibia 0.26 mm; metatarsus 0.32 mm; tarsus (tarsus segments) 0.27 mm (0.11 mm; 0.06 mm; 0.10 mm); total length 1.71 mm. Leg IV: trochanter 0.18 mm; femur 0.78 mm; patella 0.35 mm; tibia 0.52 mm; metatarsus 0.42 mm; tarsus (tarsus segments) 0.30 mm (0.10 mm; 0.08 mm; 0.12 mm); total length 2.55 mm., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on pages 832-833, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082

† MESOZOMUS GROEHNI S.P. MÜLLER, DUNLOP, KOTTHOFF, HAMMEL & HARMS, 2019 (FIGS 24–26) Type material: Holotype, collection no. GPIH04986. Remarks The original description of † Me. groehni states that a mesal spur on the pedipalp trochanter is present, but this character could not be verified; thereby the presence or absence of a mesal spur in † Me. groehni remains unclear. For a full description, see Müller et al. (2019)., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on page 822, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082, {"references":["Muller SP, Dunlop JA, Kotthoff U, Hammel JU, Harms D. 2019. The oldest short-tailed whipscorpion (Schizomida): a new genus and species from the Upper Cretaceous amber of northern Myanmar. Cretaceous Research 106: 104227."]}

SUBFAMILY HUBBARDIINAE COOK, 1899 Remarks There are three schizomid families: Protoschizomidae, Hubbardiidae and the extinct Calcitronidae. The status of Calcitronidae as a valid family is still contradictory because the tarsomere formula of 7:5:4:4 (Pierce, 1951) deviates from the general pattern of 7:3:3:3 shared by Protoschizomidae, Hubbardiidae and Thelyphonida (Cokendolpher & Reddell, 1992). The presence of six tarsomeres on legI is a rare occurrence in Old and New World schizomids alike and has been interpreted as a randomly occurring malformation caused by regeneration of a limb after loss or damage (Rowland, 1973). The two Recent families for schizomids can be distinguished by 29 morphological characters (Cokendolpher & Reddell, 1992; Monjaraz-Ruedas et al., 2017). Out of these characters, only the symmetry of the tarsal spurs (symmetrical in Protoschizomidae, asymmetrical in Hubbardiidae), body size (usually between 6 and 9 mm in Protoschizomidae and ~ 3–6 mm in Hubbardiidae), male and female flagellum morphology and the setation of the base of the anterior process (present in Protoschizomidae, absent in Hubbardiidae) can be assessed in the amber fossils owing to fossil preservation, but all of the described fossil specimens align well with the diagnosis for Hubbardiidae. Within this family, the Megaschizominae (endemic to southern Africa) have a body size of ~ 8 mm, a row of eight or nine setae along the frontal margin of the propeltidium, and their tergites II–VII on the opisthosoma have two submarginal rows of setae, whereas Hubbardiinae are smaller and have only one row. All fossil species align with Hubbardiinae rather than with Megaschizominae. The key provides basic diagnoses for all fossil genera described from Burmese amber. As the fossil schizomids are often altered and deformed by taphonomical processes, a schizomid body plan with labelling of body parts is given in Figure 2 for a better understanding of the morphology., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on page 795, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082, {"references":["Cokendolpher JC, Reddell JR. 1992. Revision of the Protoschizomidae (Arachnida: Schizomida) with notes on the phylogeny of the order. Texas Memorial Museum, Speleological Monographs 3: 31 - 74.","Rowland JM. 1973. A new genus and several new species of Mexican schizomids (Arachnida: Schizomida). Occasional Papers Museum of Texas Tech University 11: 1 - 24.","Monjaraz-Ruedas R, Francke OF, SantibanezLopez CE. 2017. The morphological phylogeny of the family Protoschizomidae revisited (Arachnida: Schizomida): setal characters, fossil and paraphyletic genera. Journal of Arachnology 45: 99 - 111."]}

† CRETACEOZOMUS DE FRANCESCO MAGNUSSEN & S.P. MÜLLER GEN. NOV. (FIGS 6–11) Zoobank registration: urn:lsid:zoobank.org:act: 36AEDFB3-9F9B-4A46-9330-C272B75F352C Type species: † Cretaceozomus robustus De Francesco Magnussen sp. nov. Remarks Specimens of this genus show asymmetrical tarsal spurs (equivocal in † Cretaceozomus angustocaudatus) and also differ from species of Protoschizomidae by lack of the pair of setae at the base of the anterior process and by the presence of a distinct stalk at the male flagellum. They also differ from members of the Megaschizominae by having a smaller body size, lack of eight or nine setae on their anterior process and the presence of only one pair of setae on the tergites II– VII. Therefore, they are clearly members of subfamily Hubbardiinae. The specimens cannot be assigned to any extant genus because of their combination of characters and the particular setation of the male flagellum. Diagnosis † Cretaceozomus differs from all other hubbardiid genera by the following combination of characters: anterior process with two setae next to each other followed by a single median seta (2 + 1), propeltidium with four pairs of setae; corneate eyes present; metapeltidium not divided; body without clavate setae; pedipalp trochanter with a mesal spur; anterodorsal margin of femur of leg IV produced at an angle of ~90°; tergite II with one pair of setae; tergites X–XII not elongated; tergite XII with posterodorsal process (equivocal in † Cretaceozomus angustocaudatus); male flagellum dorsoventrally flattened and without the pair Vl1. Note Although the presence of a posterodorsal process in † Cretaceozomus angustocaudatus cannot be verified, the species can still be discriminated genuinely from any other Recent fauna (see Supporting Information, Table S1). Etymology The genus name stems from the Cretaceous Era, which is the geological period of the formation of Burmese amber, and the suffix -zomus, the last letters of Schizomus. It is masculine in gender. Comparisons † Cretaceozomus is similar to Zomus Reddell & Cokendolpher, 1995, an extant genus widespread in SoutheastAsiaandOceania(Reddell&Cokendolpher,1995; Harvey, 2001b; Manzanilla, 2010). A secure assignment of † Cretaceozomus to the genus Zomus is not possible owing to the fact that males of Zomus are still unknown, and all specimens of † Cretaceozomus are males. Therefore, important diagnostic characters (e.g. spermathecae, male flagellum shape) cannot be compared. Until additional material becomes available, it seems premature to merge the genera Zomus and † Cretaceozomus. † Cretaceozomus is also similar to Ovozomus Harvey, 2001b found from the Comoros to the Cook Islands, but Ovozomus has eye spots, whereas † Cretaceozomus bears corneate eyes. The male flagellum is spade shaped in Ovozomus and bulbous in † Cretaceozomus. Also, the body size range of Ovozomus is larger at 4.00– 5.51 mm compared with 2.86–3.00 mm in † Cretaceozomus. † Cretaceozomus shares characters with † Annazomus, which has three pairs of setae on the propeltidium and lacks eyes (four pairs of setae and eyes present in † Cretaceozomus). In † Annazomus, the male flagellum shape is nearly rectangular with a bulbous elevation, whereas the flagellum of † Cretaceozomus has a semi-oval shape. † Cretaceozomus shares characters with † Muellerizomus, which has three pairs of setae on the propeltidium and eye spots (four pairs and corneate eyes in † Cretaceozomus). Both genera differ also in flagellum shape. † Cretaceozomus shares characters with † Muellerizomus, but the latter has a single median seta followed by three pairs of setae on the propeltidium (four pairs in † Cretaceozomus). † Muellerizomus also lacks eyes and differs in flagellum shape. For comparison with Recent genera, see the Supporting Information (Table S1)., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on pages 801-802, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082, {"references":["Reddell JR, Cokendolpher JC. 1995. Catalogue, bibliography, and generic revision of the order Schizomida (Arachnida). Texas Memorial Museum, Speleological Monographs 4: 1 - 170.","Harvey MS. 2001 b. The Schizomida (Arachnida) of the Seychelle Islands. Invertebrate Systematics 15: 681 - 693.","Manzanilla OV. 2010. The first record of the genus Zomus Reddel & Cokendolpher, 1995 (Schizomida: Hubbardiidae) from Samoa. Arthropoda Selecta 19: 81 - 83."]}

† GROEHNIZOMUS DE FRANCESCO MAGNUSSEN & S.P. MÜLLER GEN. NOV. S.P. MÜLLER GEN. NOV. (FIGS 12–23) Zoobank registration: urn:lsid:zoobank.org:act: 64D36CD9-3AEE-4899-8DFA-5F6BAB6415F7 Type species: † Groehnizomus oculiferans De Francesco Magnussen & S.P. Müller sp. nov. Remarks Specimens of this genus show asymmetrical tarsal spurs and also differ from species of the Protoschizomidae by lack of the pair of setae at the base of the anterior process and the presence of a distinct stalk at the male flagellum. They also differ from members of Megaschizominae by having a smaller body size, lack of eight or nine setae on their anterior process and the presence of only one pair of setae on the tergites II– VII. Therefore, they are clearly members of subfamily Hubbardiinae. The specimens cannot be assigned to any extant genus because of their combination of characters. Diagnosis † Groehnizomus differs from all other hubbardiid genera through a combination of characters: anterior process with one seta followed by a pair of setae; propeltidium with a single seta followed by four pairs of setae (equivocal in † G. oculiferans); corneate eyes present; metapeltidium divided; body without clavate setae; pedipalp trochanter with a mesal spur; anterodorsal margin of femur of leg IV produced at an angle of ~90°; tergite II with one pair of setae; tergites X–XII not elongated; tergite XII with posterodorsal process; male flagellum dorsoventrally flattened. Etymology The genus name is a patronym for Carsten Gröhn, who donated the specimen, combined with the suffix -zomus, which is commonly used for schizomids. It is masculine in gender. Comparisons † Groehnizomus has similarities to Oculozomus Reddell & Cokendolpher, 1995 from Sumatra, but Oculozomus has a larger body size range of 3.5–6.0 mm. Both genera also differ in the shape of the male flagellum (round-bulbous in Oculozomus; subrhomboidal in † Groehnizomus). In Oculozomus, the femur of leg IV is Groehnizomus. There are also similarities to † Mesozomus, but this genus lacks a posterodorsal process, has a single median seta followed by only three pairs of setae on the propeltidium and has an oblong-oval male flagellum. For comparison with Recent genera, see the Supporting Information (Table S1). Note In † G. oculiferans, the setation of the propeltidium is equivocal. Therefore, this character is not included in the comparison., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on page 810, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082, {"references":["Reddell JR, Cokendolpher JC. 1995. Catalogue, bibliography, and generic revision of the order Schizomida (Arachnida). Texas Memorial Museum, Speleological Monographs 4: 1 - 170."]}

† MUELLERIZOMUS DE FRANCESCO MAGNUSSEN & S.P. MÜLLER GEN. NOV. S.P. MÜLLER GEN. NOV. (FIGS 27–34) Z o o b a n k r e g i s t r a t i o n: u r n: l s i d: z o o b a n k. o r g: act: 1E280AB6-17C2-4F05-A4C4-E021DF9F2722 Type species: † Muellerizomus palicaudatus De Francesco Magnussen sp. nov. Remarks Specimens of this genus show asymmetrical tarsal spurs and also differ from species of Protoschizomidae by the lack of a pair of setae at the base of the anterior process and the presence of a distinct stalk at the male flagellum. They also differ from members of Megaschizominae by having a smaller body size, lack of eight or nine setae on their anterior process and the presence of only one pair of setae on the tergites II -VII. Therefore, they are clearly members of subfamily Hubbardiinae. The specimens cannot be assigned to any extant genus because of their combination of characters. Diagnosis † Muellerizomus differs from all other hubbardiid genera by the following combination of characters: propeltidium with a single median seta followed by three pairs of setae; eye spots present or absent; metapeltidium not divided; body without clavate setae; pedipalp trochanter with a mesal spur (equivocal in † Mu. amandae); anterodorsal margin of femur of leg IV produced at an angle of ~90°; tergite II with one pair of setae; tergites X–XII not elongated; tergite XII with posterodorsal process; male flagellum dorsoventrally flattened; female flagellum with three annuli. Etymology The genus name is a patronym for private collector Patrick Müller, who provided the specimen and numerous others for examination, with the common suffix -zomus, from the last letters of Schizomus. It is masculine in gender. Comparison † M u e l l e r i z o m u s s h a r e s c h a r a c t e r s w i t h † Groehnizomus, but the latter has corneate eyes and a divided metapeltidium (eye spots and metapeltidium not divided in † Muellerizomus). Secozomus Harvey, 2001b from the Seychelles is also similar, but it lacks a posterodorsal process, and the anterodorsal margin of femur IV is angled at> 90°. For comparison with Recent genera, see the Supporting Information (Table S1)., Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on page 824, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082, {"references":["Harvey MS. 2001 b. The Schizomida (Arachnida) of the Seychelle Islands. Invertebrate Systematics 15: 681 - 693."]}