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Exhaustive similarities in terminal Ediacaran–Ordovician cover sequences between tectonic inliers in the Appalachian–Caledonian (A–C) orogen define the Avalonian terrane from coastal northeast North America through southern Britain to Belgium. However, Barr et al. effectively block inclusion of the Mira belt (Mb) in Avalonia by masking a long-documented trans-Avalonian succession under locally defined lithostratigraphic names that prove to be confusing homonyms of themselves. Earlier revisions of Mira belt stratigraphy allow assignment of the Mira belt to a precise location in the Avalonian strike-slip regime—on the marginal–inner platform transition just as the northern Antigonish Highlands and southeast Burin Peninsula. The Mira belt is a tectonic inlier not a "terrane" in the A–C orogen. [ABSTRACT FROM AUTHOR]

The Avalonia microcontinent has diagnostic terminal Ediacaran–Ordovician lithostratigraphy, depositional sequence architecture, and igneous activity that extends for 2000+ km reflecting epeirogeny related to the Avalonian transform fault. Avalonia records an abrupt early Middle Cambrian (late Wuliuan) change from green, purple, or light grey to overlying black, dark grey, and brown facies in platform and off-platform areas (Meguma, North Wales). This change within one trilobite zone marks onset of ca. 26 m.y. of shallow-marine anoxia/strong dysoxia lasting into the Ordovician with Hatch Hill oxygenated mid-water zone (OMZ) onlap onto the shelf. A Bakken model (new, based on the middle Paleozoic Bakken Formation) is applied to shallow-shelf–shoreline organic-rich mud deposition. Erosion of greenish Avalonian depositional sequence (Ads) 7 was followed by Ads 8 tilting, volcanism, debris flows, and bentonite deposition on a cryptic unconformity in SE Newfoundland. The early Middle Cambrian age of the Ads 7–8 boundary is obscured by referring the lower Manuels River Formation and Cristallinium cambriense Zone to the younger Drumian Stage. Ads 8 has thin ashes in coterminous British and North American Avalonia where erosion and subaerial exposure with caliche development preceded onlap of upper Middle or Upper Cambrian Ads 9 black muds and sands. The green–black change emphasizes Avalonian unity; it precludes multiple Avalonian "micro-terranes" or assigning parts of Avalonia to West Gondwana or "Ganderia" (the Little River, Brookville, and Bras d'Or "terranes" are part of the Avalonian marginal platform). Coeval green–black transitions and similar later Cambrian faunas show comparable paleoenvironments in Avalonia and Baltica. [ABSTRACT FROM AUTHOR]

Episodic low oxygenated conditions on the sea-floor are likely responsible for exceptional preservation of animal remains in the upper Amouslek Formation (lower Cambrian, Stage 3) on the northern slope of the western Anti-Atlas, Morocco. This stratigraphic interval has yielded trilobite, brachiopod, and hyolith fossils with preserved soft parts, including some of the oldest known trilobite guts. The "Souss fossil lagerstätte" (newly proposed designation) represents the first Cambrian fossil lagerstätte in Cambrian strata known from Africa and is one of the oldest trilobite-bearing fossil lagerstätten on Earth. Inter-regional correlation of the Souss fossil lagerstätte in West Gondwana suggests its development during an interval of high eustatic levels recorded by dark shales that occur in informal upper Cambrian Series 2 in Siberia, South China, and East Gondwana. [ABSTRACT FROM AUTHOR]

Episodic low oxygenated conditions on the sea-floor are likely responsible for exceptional preservation of animal remains in the upper Amouslek Formation (lower Cambrian, Stage 3) on the northern slope of the western Anti-Atlas, Morocco. This stratigraphic interval has yielded trilobite, brachiopod, and hyolith fossils with preserved soft parts, including some of the oldest known trilobite guts. The "Souss fossil lagerstätte" (newly proposed designation) represents the first Cambrian fossil lagerstätte in Cambrian strata known from Africa and is one of the oldest trilobite-bearing fossil lagerstätten on Earth. Inter-regional correlation of the Souss fossil lagerstätte in West Gondwana suggests its development during an interval of high eustatic levels recorded by dark shales that occur in informal upper Cambrian Series 2 in Siberia, South China, and East Gondwana. [ABSTRACT FROM AUTHOR]

Two species of the Middle Cambrian trilobite Eodiscus Hartt, in Walcott, 1884, E. punctatus (Salter, 1864) and E. scanicus (Linnarsson, 1883), have been reported from several paleocontinents. However, in their respective type areas of Avalonian Britain and Baltica (Sweden), both species are poorly documented from moulds preserved in siliciclastic mudstone that are variably compacted and distorted. Moreover, variation in such characters as surface sculpture between putative occurrences suggests that widespread use of these names may mask species differentiation within and between paleocontinents. Detailed examination of Eodiscus sclerites that are exquisitely preserved in full relief in limestone from the Manuels River Formation of Avalonian Newfoundland and the Shallow Bay Formation of Laurentian Newfoundland demonstrates the presence of multiple species that are distinct from both E. punctatus and E. scanicus. We interpret them as a group of pseudocryptic species that are comparable to groups that are now identified routinely among modern invertebrates. New species are E.confossus, E.tuberculus, and E.coloholcus. At the current state of knowledge, E. punctatus and E. scanicus are best restricted to their respective types. [ABSTRACT FROM AUTHOR]

The Chesley Drive Group, an Upper Cambrian-Lower Ordovician mudstone-dominated unit, is part of the Ediacaran-Ordovician cover sequence on the North American part of the Avalon microcontinent. The upper Chesley Drive Group on McLeod Brook, Cape Breton Island (previously "McLeod Brook Formation"), has two lithofacies-specific Tremadocian biotas. An older low-diversity benthic assemblage (shallow burrowers, Bathysiphon, phosphatic brachiopods, asaphid trilobites) is in lower upper Tremadocian green-gray mudstone. This wave-influenced, slightly dysoxic facies has Bathysiphon-brachiopod shell lags in ripple troughs. The upper fauna (ca. 483 +/- 1 Ma) is in dysoxic-anoxic (d-a), unburrowed, dark gray-black, upper upper (but not uppermost) Tremadocian mudstone with a "mass kill" of the olenid Peltocare rotundifrons (Matthew)—a provincial trilobite in Avalonian North America that likely tolerated low oxygen bottom waters. Scandodus avalonensis Landing n. sp. and Lagenochitina aff. conifundus (Poumot), probable nektic elements and the first upper Tremadocian conodont and chitinozoan reported from Avalon, occur in diagenetic calcareous nodules in the dark gray-black mudstone. An upper Tremadocian transition from lower greenish to upper black mudstone is not exposed on McLeod Brook, but is comparable to a coeval green-black mudstone transition in Avalonian England. The successions suggest that late late Tremadocian (probable Baltic Hunnebergian Age) sea level was higher in Avalon than is suggested from successions on other paleocontinents. The Tremadocian sea-level history of Avalon was a shoaling-deepening-shoaling sequence from d-a black mudstone (lower Tremadocian), to dysoxic green mudstone (lower upper Tremadocian), and back to black mudstone (upper upper Tremadocian). Scandodus Lindström is emended, with the early species S. avalonensis Landing n. sp. assigned to the emended Family Protopanderodontidae. Triangulodus Van Wamel is considered a junior synonym of Scandodus. Peltocare rotundifrons is emended on the basis of complete specimens. [ABSTRACT FROM AUTHOR]

Exquisite Pywackia baileyi Landing n. gen. and sp. specimens from the lower Tiñu Formation, southern Mexico, extend the bryozoan record into the Upper Cambrian. They are ∼8 m.y. older than the purported oldest bryozoans from South China, and show that all skeletalized metazoan phyla appeared in the Cambrian. The new form differs from similar, twig-like cryptostomes by its shallow autozooecia and an elongate axial zooid, which may be homologous to the stolon in nonmineralized ctenostomes. It may morphologically resemble mineralized stem group bryozoans that retained a stolon-like individual, although an ability to bud was acquired by the feeding individuals (autozooids). The latest Cambrian origin of bryozoans, several mollusk classes (polyplacophorans, cephalopods), and euconodonts was a major evolutionary development and can be considered the onset of the Ordovician radiation of more complex marine communities [ABSTRACT FROM AUTHOR]

The dramatic late Early Ordovician radiation of cephalopods on tropical paleocontinents is illustrated by the diverse fauna (21 genera, 30 species) of the Fort Cassin Formation (Floian and lower Blackhillsian Stage) in northeast Laurentia. Cephalopods occur through the thin (ca. 30-65 m) depositional sequence of the Fort Cassin but are most common and diverse in mollusk-rich, trilobite-poor parts of the formation that characterize the thrombolite-bearing intervals in the shoaling part of the highstand systems tract. This lithofacies-biofacies linkage persists from the Tribes Hill and Rochdale Formations (lower and lower upper Tremadocian, and upper Skullrockian and Stairsian Stages, respectively), and suggests that the Early Ordovician radiations of cephalopods took place in shallow-marine, thrombolite reef facies of tropical carbonate platforms. These habitats differed strongly from the near-shore, peritidal habitats of the older Cambrian evolutionary radiation. Genus-level diversity and absolute abundance changed little through the Skullrockian-Blackhillsian, but morphologic diversity and body size increased dramatically by the late Early Ordovician. The morphological diversification suggests cephalopods diversified into a wider variety of macropredators and more complex late Early Ordovician ecosystems. Anrangeroceras whitehallense n. gen. and n. sp. is proposed. The following are emended: the Protocycloceratidae, Centrotcirphyceras and C. seelyi, Protocycloceras and P. Iamarcki, and Rudolfoceras cornuoryx. The following are indeterminate and abandoned: Baltoceras? pusillum Ruedemann, 1906; Carneroceras annuliferum Flower, 1941; Cyptendoceras whitfieldi Ulrich et al., 1944; Endoceras? champlainense Ruedemann, 1906; Wolungoceras valcourense Flower, 1964. Beekmanoceras Ulrich and Foerste, 1936 is a gastropod. [ABSTRACT FROM AUTHOR]

The article presents a study that examines the oldest cephalopods from East Laurentia. It describes a newly recognized fauna called Rathbunville School Limestone Cephalopods from Washington County, New York. It shows that these earliest Ordovician limestone cephalopods are Upper Cambrian. Therefore, these cephalopods comprise only the third Upper Cambrian assemblage known in Laurentia.

The Cambrian inlier at Beaver Harbour, southern New Brunswick, is now confidently referred to the marginal platform of the late Proterozoic-Early Paleozoic Avalon microcontinent. The sub-trilobitic Lower Cambrian Chapel Island and Random Formations are unconformably overlain by the mafic volcanic-dominated Wade's Lane Formation (new). Late Early Cambrian trilobites and small shelly taxa in the lowest Wade's Lane demonstrate a long Random-Wade's Lane hiatus (middle Terreneuvian-early Branchian). Latest Early-middle Middle Cambrian pyroclastic volcanism produced a volcanic edifice at Beaver Harbour that is one of three known volcanic centers that extended 550 km along the northwest margin of Avalon. Middle Middle Cambrian sea-level rise, probably in the Paradoxides eteminicus Chron, mantled the extinct volcanics with gray-green mudstone and limestone of the Fossil Brook Member. Black, dysoxic mudstone of the upper Manuels River Formation (upper Middle Cambrian, P. davidis Zone) is the youngest Cambrian unit in the Beaver Harbour inlier. Lapworthella cornu (Wiman, 1903) emend., a senior synonym of the genotype L. nigra (Cobbold, 1921), Hyolithellus sinuosus Cobbold, 1921, and probably Acrothyra sera Matthew, 1902a, range through the ca. 8 m.y. of the trilobite-bearing upper Lower Cambrian, and H. sinuosus and A. sera persist into the middle Middle Cambrian. Lapworthella cornu and H. sinuosus replaced the tropical taxa L. schodackensis (Lochman, 1956) and H. micans Billings, 1872, in cool-water Avalon. [ABSTRACT FROM AUTHOR]

The Tribes Hill Formation (upper Skullrockian) of New York records the earliest Ordovician diversification of cephalopods, in particular ellesmerocerids, on the east Laurentian, shallow carbonate platform. Revision of this cephalopod fauna on the basis of approximately 430 specimens collected across eastern New York has led to new information on inter- and intraspecific variation of the taxa and extensive synonymization of species-level taxa. The Ellesmeroceratidae and Protocycloceratidae, Ellesmeroceras, and Eremoceras are emended, Eorudolfoceras n. gen. and Dakeoceras champlainense n. sp. are erected. A rank abundance plot of the 342 specimens at a locality in the Lake Champlain lowlands provides information on the community structure of the nautiloid fauna, where small orthoconic taxa arc shown to dominate strongly. The small orthocone Ectenolites was the most common genus in terms of total occurrences, was the most paleogeographically widespread genus, and was the only genus to cross the Cambrian-Ordovician boundary. [ABSTRACT FROM AUTHOR]

The Lower Ordovician middle Beekmantown Group is a very thin carbonate platform succession on the northern New York Promontory that thickens north into the Ottawa aulacogen near Montréal. The Tribes Hill Formation (Rossodus manitouensis Zone) records the earliest Ordovician (late Skullrockian, late early Tremadocian) eustatic high that submerged Laurentia, and produced the lowest Ordovician sequence along the New York Promontory. These dolostones are succeeded in the Beekmantown, New York, area by late Tulean?-Blackhillsian transgressive systems tract quartz arenites of the lower Fort Cassin Formation (Ward Member). The "Fort Ann Formation" (middle Stairsian, upper Tremadocian) of the southern Lake Champlain lowlands (=Theresa Formation sandstones in the Ottawa graben) is absent at Beekmantown, and moderate Stairsian (late Tremadocian) eustatic rise apparently did not inundate the Beekmantown area after Skullrockian-Stairsian boundary interval offlap. Highstand carbonates of the upper Fort Cassin Formation [Sciota Member = "Spellman Formation" and "Ogdensburg Member" of the "Beauharnois Formation" in the Montréal area; designations abandoned] at Beekmantown yield diverse conodonts seemingly characteristic of the Oepikodus communis-Fahraeu-sodus marathonensis Zone (new). However, associated trilobites, particularly Carolinites tasrnanensis (Etheridge, 19 19), show a cor- relation with the upper Trigonocerca lypica (trilobite) Zone of the Utah and the overlying Reutterodus andinus (conodont) Zone. This abrupt early Blackhillsian lithofacies change features the appearance of chitinozoans and conodonts known from marginal successions, and records the Laignet Point highstand (new). This highstand is recognized across Laurentia on the west Newfoundland and southern Midcontinent platforms. It is recorded on the east Laurentian continental slope by lower Oepikodus evae Zone dysoxic black mudstone in the Taconian allochthons. Taxonomic re-evaluations include Utrichodina Branson and MehI, 1933, with its genotype species U. abnormalis (Branson and MehI, 1933) emend., as the senior synonym of Colaptoconus Kennedy, 1994; Eucharodus Kennedy, 1980; and Glyptoconus Kennedy, 1980. Paraserratognathus An in An et at., 1983, emend. is the senior synonym of Wandelia Smith, 1991 and Stultodontus Ji and Barnes, 1994. Tropodus Kennedy, 1980 is the senior synonym of Chionoconus Smith, 1991. The trilobite fauna of the Sciota Member includes species of Isoteloides, Benthamaspis, Acidiphorus and Carolinites, of which I. fisheri is new. [ABSTRACT FROM AUTHOR]

Fossiliferous clasts occur in Carboniferous conglomerate in the Horton Group on western Isle Madame and in the Mabou Group on eastern Isle Madame. Most of the clasts (21 of 23 examined) are calcareous siltstone and sandstone that contain Silurian – Lower Devonian faunas comparable to those in the Arisaig area, northern mainland Nova Scotia, although the lithologies are coarser grained and less calcareous than those of the Arisaig section. These middle Paleozoic faunas are well constrained to the Silurian (uppermost Llandovery through Pridoli) and lowest Devonian and are characteristic of those known from shallow siliciclastic-dominated platforms of the Avalon microcontinent in Wales and England. The remaining two clasts have abundant inarticulate brachiopod shells that indicate provenance from Middle Cambrian proximal marine facies on the Avalonian marginal platform. No clasts were found that are likely to have been derived from the Torbrook Formation, and thus from the Meguma terrane in southwestern Nova Scotia, as has been previously reported. The association of relatively large, reworked fossiliferous clasts in Carboniferous conglomerate on Isle Madame suggests local derivation from lower and middle Paleozoic units not presently exposed, although probably present as subcrop under the Carboniferous units, in southwestern Cape Breton Island and adjacent mainland Nova Scotia. [ABSTRACT FROM AUTHOR]

The only pillow basalt in synorogenic sedimentary rocks at the exterior margin of the Taconic orogen in eastern North America is at Stark's Knob in eastern New York. Earlier reported as extrusive into allochthonous Ordovician slope and rise facies, this small lens (ca. 125+ m long, 39 m thick) is a fault-bounded block in Upper Ordovician melange under the Taconian frontal thrust. Its N-MORB (normal mid-ocean ridge basalt) basalt geochemistry and spinel composition are characteristic of oceanic ridge settings at a water depth of 2 km or more. Abundant limestone lenses on pillows and lava shelves within pillows yielded a middle Late Ordovician gastropod. The limestones are reconciled with this extrusion depth and with limited early Paleozoic pelagic carbonate production by lime mud transport from the Laurentian platform or abiotic carbonate precipitation with sea-water heating during basalt extrusion. A genetic relationship between the parautochthonous Stark's Knob basalts and the allochthonous Jonestown volcanics in slope and rise facies of the Hamburg klippe, eastern Pennsylvania, is likely. Both are Ordovician MORB basalts that reflect volcanism on the subducting outer trench slope prior to the Taconic arc – Laurentia collision. Taconic orogenesis may have led to basalt production on the subducting plate by (1) the setting up of orogen-parallel, predominantly strike-slip motion on the subducting slab with MORB basalt generated at offsets in a setting analogous to the Gulf of California or (2) development of faults in a flexure-induced extensional regime. By either process, mafic volcanism appears to be a rare but tectonically significant process on outer trench slopes as continental margins or oceanic plates enter subduction zones. [ABSTRACT FROM AUTHOR]

Deals with a study which documented the development of comparable shallow-marine facies in the terminal Arenig of Avalon from eastern Newfoundland to southern New England. Limestone lithology; Faunas; Discussion; Systematic paleontology; Discussion.

The Cambrian-Ordovician boundary is a type 1 depositional sequence boundary with dramatic local erosional incision in restricted marine facies on the easternmost New York Promontory. The systemic boundary is bracketed below by Late Cambrian, upper Cordylodus proavus Zone (s.s). conodonts from carbonates of the upper Little Falls Formation (=Whitehall Formation, abandoned). Presumed Lower Ordovician ellesmeraceratoid cephalopods from the upper Little Falls are uppermost Cambrian and among the oldest known in North America. The overlying deepening-shoaling cycle of the Tribes Hill Formation (=Cutting and Great Meadows Formations, abandoned) is the local expression of a lowermost Ordovician (Rossodus manitouensis Zone) depositional sequence recognizable across Laurentia. Complete replacement of conodonts takes place in the late Tremadocian or Tremadocian-Arenigian boundary interval with onlap of the "Fort Ann Formation" across the paleokarst cap of the Tribes Hill. The trilobites Hystricurus sp. and Symphysurina myopia Westrop new species occur in less restricted, thrombolitic facies of the middle Tribes Hill that have the highest conodont diversity. Ulrichodina Furnish, 1938, emend, is regarded as the senior synonym of the conodont Colaptoconus Kennedy, 1994 (=Glyptoconus Kennedy, 1980). [ABSTRACT FROM AUTHOR]

Investigates the shelly fossils Trilobites and Hatch Hill dysaerobic interval in Quebec. Association of sea-level change with Hawke Bay regression; Correlation between Trilobites and Bicella bicensis assemblage; Denitrification of the Olenellus Chron.

Examines Middle Cambrian tribolites of the Braintree Member in eastern Massachusetts. Species categorized as tribolites; Correlation of Braintree tribolites with those found in other Cambrian paleocontinents; Stratigraphy of Braintree tribolites; Effects of faunal exchanges between continents.

Examines the stratigraphy and correlation of Middle Cambrian trilobites in Massachusetts. Impact of taxonomical re-examination on the specimen; Effect of provincial barrier breakdown on faunal distribution; Classification of polymeroid species.

Reports on the species of trilobites located in the Hanford Brook Formation, a Cambrian unit of the Avalonian North America. Complexity of the vertical succession of the faunas; Breakdown of biogeographic barriers in the late Early Cambrian; Basis for the correlation of the Middle and Lower Cambrian interval.

The Cambrian olenid trilobite PelturaMilne-Edwards, 1840 is widespread in the Furongian strata of the Cambrian continents of Baltica and Avalonia, and has played a significant role in biostratigraphic zonations for both regions. Despite this importance, the genus is poorly known in Avalonian Canada. New collections from the Chelsey Drive Group on Cape Breton Island, Nova Scotia, yielded numerous sclerites of a new species, Peltura hutchinsoni sp. nov., that may also occur in the Parabolina lobata Zone of Baltica. Archival specimens from Sweden and Norway allow the revision of Peltura scarabaeoides (Wahlenberg, 1821) and Peltura acutidensBrögger, 1882, respectively. [ABSTRACT FROM AUTHOR]

Focuses on the discovery of a section in Taconic allochthon that has the most diverse and one of the lowest Elliptocephala asaphoides assemblage in eastern New York. Oldest body fauna; Diversity, abundance and excellent preservation; Standard for taxonomic and biostratigraphic comparison of the oldest Cambrian body fossils in eastern Laurentia.

Studies conodonts, stratigraphy and relative sea-level changes of the Tribes Hill Formation in the Lower Ordovician in east-central New York. Depositional environments and stratigraphy indicating that Tribes Hill was deposited on a wave; Deepening-shoaling sequence of the Tribes Hill; Shoaling half-cycle of the Tribes hill.

Reports on the stratigraphy of the Precambrian-Cambrian global boundary stratotype. Biostratigraphic significance; Importance of trace-fossil assemblages; Discussion on global correlation; Views on stratigraphic continuity.

Comments on the study on comparison of C isotope trends in Early Cambrian history by Andrew Knoll, et al. Sub-Tommotian unconformity in Siberia; Biostratigraphic and geochronologic ties; Reply of authors of the study.

The article presents a commentary on the study "Tropical weathering of the Taconic orogeny as a driver for Ordovician cooling: Geology," by N. L. Swanson-Hysell and F. A. Macdonald. Topics discussed include effect of carbon dioxide CO2 sequestration on global temperatures, record of Ordovician temperature, and interpretation of the partial pressure of carbon dioxide (pCO2).

Comments on the article `Precambrian-Cambrian boundary global stratotype ratified and new perspective of Cambrian time,' by Ed Landing published in `Geology,' Volume 22, 1994. Misconceptions on small shelly fossils (SSF); Gap between the Manykay and Tommotian stages; Justification of combined section for Siberia and Karatau; Landing's reply.

Assemblages of mineralized skeletal fossils are described from limestone rocks of the lower Cambrian Nemakit-Daldyn, Medvezhya, Kugda-Yuryakh, Manykay, and lower Emyaksin formations exposed on the western and eastern flanks of the Anabar Uplift of the northern Siberian Platform. The skeletal fossil assemblages consist mainly of anabaritids, molluscs, and hyoliths, and also contain other taxa such as Blastulospongia, Chancelloria, Fomitchella, Hyolithellus, Platysolenites, Protohertzina, and Tianzhushanella. The first tianzhushanellids from Siberia, including Tianzhushanella tolli sp. nov., are described. The morphological variation of Protohertzina anabarica and Anabarites trisulcatus from their type locality is documented. Prominent longitudinal keels in the anabaritid Selindeochrea tripartita are demonstrated. Among the earliest molluscs from the Nemakit-Daldyn Formation, Purella and Yunnanopleura are interpreted as shelly parts of the same species. Fibrous microstructure of the outer layer and a wrinkled inner layer of mineralised cuticle in the organophosphatic sclerites of Fomitchella are reported. A siliceous composition of the globular fossil Blastulospongia is reported herein and a possible protistan affinity similar to Platysolenites is discussed. New carbon isotope data facilitate correlation both across the Anabar Uplift and with the Terreneuvian Series of the IUGS chronostratigraphical scheme for the Cambrian System. The base of Cambrian Stage 2 is provisionally placed herein within the Fortunian Cambrian Stage 2 transitional interval bracketed by the lowest appearance of Watsonella crosbyi and by a slightly higher horizon at the peak of carbon isotope excursion I' from western flank of the Anabar Uplift. Correlation across the Siberian Platform of the fossiliferous Medvezhya and lower Emyaksin formations showing δ13Ccarb excursion I' with the upper Sukharikha Formation containing excursion 5p and upper Ust'-Yudoma Formation containing excursion I is supported herein. [ABSTRACT FROM AUTHOR]

Whereas cosmopolitan distribution patterns are established for many Late Devonian vertebrates (e.g., placoderms, onychodontiforms), few palaeobiogeographic studies have considered chondrichthyans. Recent discoveries of shark material demonstrate that some chondrichthyans were cosmopolitan by the Middle Devonian. Abundant Givetian microremains have been recovered from the Cairo quarry in eastern New York State, USA. These include teeth of two shark species with Gondwanan affinities, the omalodontid Portalodus mannoliniae sp. nov. and the antarctilamnid Wellerodus priscus. Abundant teeth of P. mannoliniae sp. nov. are characterized by a smooth diplodont crown, polarized cusps, and a labially oriented base. The teeth demonstrate monognathic heterodonty. The juvenile morph is distinguished from the adult by smaller size, slender cusps, and variation in the shape of the base. W. priscus is represented by rare juvenile teeth. Two groups of scales that show affinity to material from northern (Spain) and East Gondwana (Antarctica) are tentatively attributed to the two described species. Antarctilamnid distribution suggests a north Gondwanan origin and a colonization of the margin of the landmass before dispersing to Laurentia by the Middle Devonian. This material further indicates that vertebrate global dispersal was initiated by the Middle Devonian, and emphasizes earlier palaeogeographic interpretations that the Middle Devonian "Hamilton fauna" of North American Laurussia originated in the Early Devonian in South American Gondwana. [ABSTRACT FROM AUTHOR]

Eighteen taxa of linguliform brachiopods, mainly represented by acrotretoids, are reported from the Upper Cambrian (Furongian, Stage 10) and Lower Ordovician (Tremadocian) Tiñu Formation of Oaxaca State, Mexico. At the time of deposition, this area was part of Oaxaquia, which was either a microcontinent or an integral part of the Gondwanan margin. Whereas certain trilobites seem to indicate a Gondwanan affinity, the Tiñu brachiopod faunas show a less definite paleogeographic relationship. Some taxa have previously only been reported from Laurentia (Eurytreta cf. fillimorensis, Eurytreta cf. campaniformis), and one taxon is best known from the Avalon microcontinent (Eurytreta cf. sabrinae). However, the relatively high percentage of new and potentially endemic taxa (Oaxaquiatreta labrifera n. gen. n. sp., Tapuritreta reclinata n. sp., Oaxaquiatreta sp., Eurytreta? n. sp., Acrotretidae n. gen. n. sp., Obolinae gen. and sp. indet.) and the lack of other typical Laurentian, Gondwanan, or Avalonian taxa suggest either a certain degree of insularity of Oaxaquia or reflects a more temperate, unrestricted marine environment during the Early Paleozoic. Other taxa reported from the Tiñu Formation include Semitreta sp., Lingulella? spp., Obolinae gen. and sp. indet., Eoscaphelasma? sp., Ottenbyella? sp. A and sp. B, and Acrotretidae gen. and sp. indet. A, B, and C. Eurytreta and Semitreta are critically reviewed and several taxa previously assigned to them have been excluded. An emended diagnosis for the genus Eurytreta is presented. The presence of delthyrium and notothyrium-like structures in the siphonotretid Oaxaquiatreta n. gen. further strengthens the previously proposed relationship between the Siphonotretida and Paterinida. [ABSTRACT FROM AUTHOR]

The Early and Middle Ordovician Orthocerida and Lituitida of Precordilleran Argentina are described, and their systematics and paleogeographic significance are revised. These cephalopods show a strong affinity to coeval faunas of North China, suggesting a location of the Precordillera at middle latitudes in the Southern Hemisphere east of the North China block and relatively close to the Gondwanan margin during the early Middle Ordovician. The descriptive terminology of characters of the septal necks, the position and shape of the siphuncule, and the shape of the connecting ring is improved. The distribution of these characters support an emendation of the Baltoceratidae, Sactorth- oceratidae, and Proteoceratidae. Braulioceras n. gen. (Sactorthoceratidae) and Palorthoceras n. gen. (Orthoceratidae) are erected. The new species Braulioceras sanjuanense, Eosomichelinoceras baldisii, Gangshanoceras villicumense, and Rhynchorthoceras minor are proposed. Palorthoceras n. gen. from the Lower Ordovician Oepikodus evae Zone represents the earliest known orthocerid. [ABSTRACT FROM AUTHOR]

The Fossil Brook Member of the upper Chamberlain's Brook Formation is a thin (up to 14 m) but distinctive, unconformitybound depositional sequence recognizable from Rhode Island to eastern Newfoundland in Avalonian North America. Its diverse trilobite fauna was first described more than century ago from the limestone-rich facies of the member in southern New Brunswick. However, the systematics, stratigraphic context, and biostratigraphic significance of these trilobites have remained poorly known. A revision of the conocoryphid and paradoxidid trilobites has been completed, and the taxa set into their stratigraphic context within the middle Middle Cambrian. The faunas of the Fossil Brook are assigned to the Eccaparadoxides eteminicus Zone of Avalon. Although biogeographic barriers between Avalon and Gondwana remained strong in the Middle Cambrian and few shared trilobite species are present, a generalized correlation of the E. eteminicus Zone into Gondwana is with the Badulesia tenera Zone of the Toushamian Stage in Morocco and the Badulesia Zone of the Caesaraugustian Stage in Spain. [ABSTRACT FROM AUTHOR]

The snowball Earth hypothesis predicts that low-latitude glaciation lasted millions of years while CO2 built up to critical levels to culminate in catastrophic deglaciation in a supergreenhouse climate. The Gaskiers Formation of eastern Newfoundland (Canada) has been attributed to a snowball glaciation event, but the lack of robust paleomagnetic data and precise geochronological constraints has precluded tests of the hypothesis. Here we present high-precision U-Pb zircon geochronology (chemical abrasion-isotope dilution-thermal ionization mass spectrometry) from eight tuffs from multiple distant stratigraphic sections that bracket glacial diamictites and the first appearance of large Ediacaran fossils. Including internal error, deposition of the Gaskiers diamictite on the Avalon Peninsula is constrained to have been between 580.90 ± 0.40 and 579.88 ± 0.44 Ma, and the Trinity diamictite on Bonavista Peninsula was deposited between 579.63 ± 0.15 and 579.24 ± 0.17 Ma. Assuming approximately synchronous deglaciation, these results imply a maximum duration for deposition of the Trinity diamictite of ≥ 340 k.y.; this is inconsistent with the multimillion year duration predicted by the snowball Earth hypothesis. Our geochronologic data also constrain the first appearance datum of Ediacaran fossils to <9.5 m.y. after the Gaskiers glaciation. Thus, despite existing paleomagnetic constraints that indicate that marine ice sheets extended to low to middle latitudes, it appears that Earth narrowly escaped a third Neoproterozoic snowball glaciation just prior to the late Ediacaran expansion of metazoan ecosystems. [ABSTRACT FROM AUTHOR]

The evolution of trees of modern size growing together in forests fundamentally changed terrestrial ecosystems. The oldest trees are often thought to be of latest Devonian age (about 380–360 Myr old) as indicated by the widespread occurrence of Archaeopteris (Progymnospermopsida). Late Middle Devonian fossil tree stumps, rooted and still in life position, discovered in the 1870s from Gilboa, New York, and later named Eospermatopteris, are widely cited as evidence of the Earth’s ‘oldest forest’. However, their affinities and significance have proved to be elusive because the aerial portion of the plant has been unknown until now. Here we report spectacular specimens from Schoharie County, New York, showing an intact crown belonging to the cladoxylopsid Wattieza (Pseudosporochnales) and its attachment to Eospermatopteris trunk and base. This evidence allows the reconstruction of a tall (at least 8 m), tree-fern-like plant with a trunk bearing large branches in longitudinal ranks. The branches were probably abscised as frond-like modules. Lower portions of the trunk show longitudinal carbonaceous strands typical of Eospermatopteris, and a flat bottom with many small anchoring roots. These specimens provide new insight into Earth’s earliest trees and forest ecosystems. The tree-fern-like morphology described here is the oldest example so far of an evolutionarily recurrent arborescent body plan within vascular plants. Given their modular construction, these plants probably produced abundant litter, indicating the potential for significant terrestrial carbon accumulation and a detritus-based arthropod fauna by the Middle Devonian period. [ABSTRACT FROM AUTHOR]

Investigates the systematics of the Ordovician trilobites Ischyrotoma and Dimeropygiella with species from the type Ibexian area in western Utah. Reinvestigation of faunas; Reassessment of the dimeropygid genera Ischyrotoma and Dimeropygiella; Availability of morphological information for most part of the exoskeleton.

The origin of trees and forests in the Mid Devonian (393–383 Ma) was a turning point in Earth history, marking permanent changes to terrestrial ecology, geochemical cycles, atmospheric CO 2 levels, and climate. However, how all these factors interrelate remains largely unknown. From a fossil soil (palaeosol) in the Catskill region near Cairo NY, USA, we report evidence of the oldest forest (mid Givetian) yet identified worldwide. Similar to the famous site at Gilboa, NY, we find treefern-like Eospermatopteris (Cladoxylopsida). However, the environment at Cairo appears to have been periodically drier. Along with a single enigmatic root system potentially belonging to a very early rhizomorphic lycopsid, we see spectacularly extensive root systems here assigned to the lignophyte group containing the genus Archaeopteris. This group appears pivotal to the subsequent evolutionary history of forests due to possession of multiple advanced features and likely relationship to subsequently dominant seed plants. Here we show that Archaeopteris had a highly advanced root system essentially comparable to modern seed plants. This suggests a unique ecological role for the group involving greatly expanded energy and resource utilization, with consequent influence on global processes much greater than expected from tree size or rooting depth alone. • The earliest fossil forest to date is recovered from the Devonian of New York • Three types of trees are identified from fossil soil evidence in plan view • Early lignophyte relatives of seed plants have surprisingly modern root systems • Advanced energetics in this group suggests a unique role in changing Earth history Using data from a Middle Devonian fossil soil, Stein et al. report root systems from the earliest intact forest to date, including cladoxylopsids, possibly stigmarians and Archaeopteris. Striking seed plant-like features of the latter indicate a special role for this clade in the profound changes in Earth global systems that took place at that time. [ABSTRACT FROM AUTHOR]

An introduction is presented in which the authors discuss various reports within the issue on topics including bioturbation, sediments, and fossilization.