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Private forest owners are the main forest ownership group within Europe, and important conservation values have been found on their land. Yet, small plot sizes, societal heterogeneity, and structural changes impede developing and implementing effective conservation programs in private forests. We present a systematic literature review focusing on small-scale private forest owners and their perspectives on nature conservation by synthesizing research approaches, social-ecological drivers, and policy recommendations. Conservation perspectives were positively related to female gender, higher levels of education, formalized forest management, an active relation to the forest, and ecological values of the property. In contrast, high age, rural orientation, economic forest management factors, large parcel size, and economic and sentimental property values negatively influenced conservation perspectives. Applying a natural resource conflict management framework, we synthesized recommendations covering three dimensions: substance, procedure, relationship. Considering perspectives of small-scale private forest owners in current forestry decision-making has great potential to strengthen sustainable forest management that integrates nature conservation and resource use. [ABSTRACT FROM AUTHOR]

The protection and sustainable management of habitat trees is an integral part of modern forest nature conservation concepts such as retention forestry. Bats, cavity‐nesting birds, arboreal marsupials, and many different saproxylic species depend on habitat trees and their great variety of microhabitats and old‐growth characteristics. With a focus on insights from temperate forests, we traced the development of habitat‐tree protection over 200 years. The idea was first conceptualized by foresters and natural scientists in the early 19th century. At that time, utilitarian conservation aimed to protect cavity trees that provided roosts and nesting holes for insectivorous bats and birds. By the second half of the 19th century, habitat‐tree protection was well known to foresters and was occasionally implemented. Knowledge of the protection of large old trees, a special kind of habitat tree, for sociocultural and aesthetic reasons developed similarly. But, many foresters of that time and in the following decades fundamentally rejected protection of habitat trees for economic reasons. Beginning in the 1970s, forest conservation and integrative forest management became increasingly important issues worldwide. Since then, the protection of habitat trees has been implemented on a large scale. Long‐term views on the development of conservation concepts are important to inform the implementation of conservation today. In particular, historical analyses of conservation concepts allow the testing of long‐term conservation outcomes and make it possible to study the resilience of conservation approaches to changing social or ecological conditions. We encourage all conservation ecologists to assess the practical and conceptual impact of the initial ideas that led to modern conservation concepts in terms of long‐term biodiversity conservation. Article Impact Statement: It took 200 years to implement the habitat‐tree protection ideas of far‐sighted, trend‐setting foresters, and natural scientists. [ABSTRACT FROM AUTHOR]

Three questions have been addressed. Firstly, where in the pre-nineteenth-century landscape did farmers hold strips, camps, meadows and shares in the commons? Secondly, did farmers each own strips and camps or were some specialised strip and others exclusively 'camp' farmers? Finally, can we corroborate or reject one of the alternative hypotheses: strip-field-first versus camp-first.The area of interest is the current cadastral district cum medieval parish Epe at today's German-Dutch border as pars pro toto for the surrounding area of about 100 km diameter in the North-West European cover sand belt. Our key data source was the 1827 cadastre complemented by the historical topographic map and geological, soil and elevation maps. For population estimates, we used six tax registers from 1499–1750. All parcels per farmstead were identified in the cadastral registry, farms located on parcel maps and hamlet territories delineated as the aggregate of its farms. The following farm features were extracted from the cadastre and averaged per settlement: number of strips and strip-fields, parcel type, farm size, tenure, number of meadows, oak camps, crop camps, and pasture camps. Next, the following landscape features were identified from the map set per settlement: farmstead pattern, type of settlement, commons, strip-field, soil, and watercourse. In Part 1, we provided the introduction, the materials & methods section, and the historical and landscape context (Gils & Mölder 2019a). In this Part 2, we present our findings, a discussion, a hypothetical narrative consistent with our findings, and answers to our research questions. [ABSTRACT FROM AUTHOR]

Tree saplings are exposed to a competitive growth environment in which resources are limited and the ability to adapt determines general vitality and specific growth performance. In this study we analyzed the aboveground spatial neighborhood of oak [ Quercus petraea (Matt.) Liebl.] and beech (Fagus sylvatica L.) saplings growing in Germany, by using hemispherical photography and terrestrial laser scanning as proxy for the competitive pressure saplings were exposed to. The hemispherical images were used to analyze the light availability and the three-dimensional (3D) point clouds from the laser scanning were used to assess the space and forest structure around the saplings. The aim was to increase the precision with which the biomass allocation, growth, and morphology of the saplings could be predicted by including more detailed information of their environment. The predictive strength of the models was especially increased through direct neighborhood variables (e.g., relative space filling), next to the light availability being the most important predictor variable. The biomass allocation patterns within the more light demanding oak were strongly driven by the space availability around the saplings. Diameter and height growth variables of both species reacted significantly to changes in light availability, and partly also to the neighborhood variables. The leaf morphology [as leaf-area ratio (LAR)] was also driven by light availability and decreased with increasing light availability. However, the branch morphology (as mean branch weight) could not be explained for oak and the model outcome for beech was hard to interpret. The results could show that individuals of the same species perform differently under constant light conditions but differing neighborhoods. Assessing the neighborhood of trees with highly precise measurement devices, like terrestrial laser scanners, proved to be useful. However, the primary response to a dense neighborhood seemed to be coping with a reduction of the lateral light availability aboveground, rather than responding to an increase of competition belowground. The results suggest continuing efforts to increase the precision with which plant environments can be described through innovative and efficient methods, like terrestrial laser scanning. [ABSTRACT FROM AUTHOR]

Three questions have been addressed. Firstly, where in the pre-nineteenth-century landscape did farmers hold strips, camps, meadows and shares in the commons? Secondly, did farmers each own strips and camps or were some specialised strip and others exclusively 'camp' farmers? Finally, can we corroborate or reject one of the alternative hypotheses: strip-field-first versus camp-first. The area of interest is the current cadastral district cum medieval parish Epe at today's German-Dutch border as pars pro toto for the surrounding area of about 100 km diameter in the North-West European cover sand belt. Our key data source was the 1827 cadastre complemented by the historical topographic map and geological, soil and elevation maps. For population estimates, we used six tax registers from 1499–1750. All parcels per farmstead were identified in the cadastral registry, farms located on parcel maps and hamlet territories delineated as the aggregate of its farms. The following farm features were extracted from the cadastre and averaged per settlement: number of strips and strip-fields, parcel type, farm size, tenure, number of meadows, oak camps, crop camps, and pasture camps. Next, the following landscape features were identified from the map set per settlement: farmstead pattern, type of settlement, commons, strip-field, soil, and watercourse. We present the introduction, the materials and methods section, the historical and landscape context in Part 1, followed in Part 2 by our findings, a discussion, a hypothetical narrative consistent with our findings, and answers to our research questions. [ABSTRACT FROM AUTHOR]

Graphical abstract Highlights • Oak forests are of great economic and conservation importance. • We identify integrative management options for forestry and nature conservation. • Light availability is a strong link between forestry and conservation demands. • There is an urgent need for integrative oak forest planning approaches. • Ecological continuity should be secured in structural "sustainability units". Abstract Central European temperate oak woodlands are highly valued for their rich biodiversity. They are also of great economic importance and forest management aims to produce high quality timber, which demands high investments. The aim of this literature review is to identify management options for forestry and nature conservation that sustain both the ecological value of oak forests and the economic viability of oak silviculture. We addressed three main questions: (a) Oaks and close-to-nature forestry – what are the key silvicultural challenges and options?, (b) What is the particular significance of ecological continuity and which structural features are of importance for biodiversity conservation in oak forests?, (c) What are the key elements and possible strategies of forest management that sustain the ecological values in oak forests in combination with viable forestry? Light availability appeared to be a conspicuous link connecting the conservation and the silvicultural aspects of multifunctional oak forest management: Both young oak trees and multiple oak woodland specialist species are characterized by their need for increased sunlight exposure. This common denominator provides a sound basis for integrative management practices for forestry and nature conservation. The concept of retention forestry offers purposeful approaches. So the harvest of valuable timber oaks or the creation of canopy gaps for oak regeneration can be used to release the crowns and trunks of habitat oaks from shading and competition. When looking at the management of oak woodland biodiversity hotspots, the re-establishment of (modified) historical forest management techniques, which increase stand openness and create transitional habitats that provide suitable oak regeneration niches, seems to be necessary. Not only the continuity of oak woodland cover and natural site conditions, but also the uninterrupted temporal continuity and availability of wood-related structural features turned out to be of particular importance for oak woodland specialist species. We identified an urgent need for systematic forest planning approaches that secure the long-term availability of these structural features within areas or "sustainability units" that are large enough to maintain viable populations of oak woodland specialist species. In particular, conservation-oriented forestry measures should mainly be implemented in those areas, where the greatest effectiveness is to be expected. In the sustainability units, oak regeneration measures ought to take place either in close vicinity to old oak stands or directly in these stands. The choice of one of these options should be based on a careful consideration of the needs and possibilities of both silvicultural and nature conservation management. [ABSTRACT FROM AUTHOR]

Abstract Forests are under pressure from accelerating global change. To cope with the multiple challenges related to global change but also to further improve forest management we need a better understanding of (1) the linkages between drivers of ecosystem change and the state and management of forest ecosystems as well as their capacity to adapt to ongoing global environmental changes, and (2) the interrelationships within and between the components of forest ecosystems. To address the resulting challenges for the state of forest ecosystems in Central Europe, we suggest 45 questions for future ecological research. We define forest ecology as studies on the abiotic and biotic components of forest ecosystems and their interactions on varying spatial and temporal scales. Our questions cover five thematic fields and correspond to the criteria selected for describing the state of Europe's forests by policy makers, i.e. biogeochemical cycling, mortality and disturbances, productivity, biodiversity and biotic interactions, and regulation and protection. We conclude that an improved mechanistic understanding of forest ecosystems is essential for the further development of ecosystem-oriented multifunctional forest management in the face of accelerating global change. [ABSTRACT FROM AUTHOR]

This study underlines the often under-estimated importance of forest ownership and land tenure in European forest biodiversity studies which are crucial for the management, structure, and tree species composition of woodland. In particular it is assumed that, in regions with both state-owned forests and smaller private forests, the latter contain more relict habitats shaped by historical woodland management practices. A government decree of 1721, a political compromise, was crucial to the present-day woodland ownership pattern and distribution of woodland habitats in the Osnabrück region (northwest Germany). It resulted in the privatization of woodlands held in common for centuries and created a huge number of small, private forest parcels in the 18th century. These developments are discussed in relation to Europe-wide processes in forest affairs. Mainly due to the low economic importance of these forest parcels, as well as the individualism of the forest owners, coppice structures providing valuable habitats have persisted until today. For instance, over-aged coppice stands provide important habitat conditions for saproxylic species and unique herbaceous layers. These valuable habitats must be protected while creating new coppice stands to eventually take their place in future decades. Management plans for Natura 2000 sites in the Osnabrück region should address this problem while reconciling any conflict of interests between private owners and nature conservation organizations. Researchers are encouraged to give more consideration to the important relationship between current woodland biodiversity and the history of forest ownership patterns. [ABSTRACT FROM AUTHOR]

Ancient woodlands, with their long ecological continuity, frequently harbor a high number of typical, rare and threatened species, and are therefore of particular importance for nature conservation. To pinpoint these habitats, a common application is the use of plants as “ancient woodland indicators”. The occurrence of these particular species allows for evaluating the continuity of woodland cover in time. While lists of ancient woodland vascular plants have been derived for many regions, the identification and use of bryophytes as ancient woodland indicators has been widely neglected. This is a bit surprising because certain woodland bryophytes are very sensitive to varying environmental conditions or changes in land management. It therefore appeared promising to compile an ecologically grounded list of ancient woodland indicator bryophytes for practical use. In this study, we present a set of ancient woodland indicator bryophytes based on the analysis of datasets from the North German federal state of Schleswig-Holstein. To compile this list, we systematically evaluated the bryophyte distribution data from floristic surveys in relation to ancient woodland cover data from state-wide inventories. In this way, we were able to determine ancient woodland bryophytes using consistent and repeatable statistical methods. The presented list of 31 ancient woodland indicator bryophytes is ecologically sound and corresponds well with data from the sparse literature. We could distinguish two groups of ancient woodland indicator bryophytes. The first group is linked to base-rich, semi-natural deciduous woodlands with high soil and air humidity. The second group comprises acidophilic bryophytes that occur not only in acidic beech and oak woods, but also in acidic mixed or coniferous forests on ancient woodland sites. Apart from the ancient woodland indicator bryophytes, we could identify one group of recent woodland bryophytes and four groups of bryophytes that are more or less indifferent with respect to woodland continuity. Finally, we provide recommendations for the application of ancient woodland indicator bryophytes in nature conservation practice. Management suggestions for the conservation of the typical bryophyte diversity of ancient semi-natural woodlands are also given. [ABSTRACT FROM AUTHOR]

Ancient woodlands that have been in continuous existence for hundreds of years have a floristic composition which greatly differs from younger afforestations. The occurrence of certain associated vascular plant species, termed “ancient woodland indicator plants“, can be used to recognise the continuity of woodland cover. Ancient woodland habitats frequently contain a typical and rich forest biodiversity and can often be regarded as “biodiversity hotspots”. To pinpoint these habitats for nature conservation, there is a need to compile ancient woodland indicator lists with a widespread validity. In this study, we introduce a new methodical approach that enables the compilation of such lists from the readily available resources of plant species monitoring programs, archive records, and land cover data. Using northwest Germany as a model region, we have developed an ecologically grounded list of 67 ancient woodland indicator plants for this area. In this context, we consider the “ancient woodland indicator plants” as a subset of the larger group of “ancient woodland plants”. The widely applicable ancient woodland indicator plants list presented here may be a useful tool for future forest nature conservation. Potential applications include: (a) the identification of ancient woodlands in areas where historical maps are lacking, (b) the identification of biodiversity hotspots of ancient woodland indicator plants, and (c) locating ancient semi-natural woodlands. Finally, we highlight the importance of effective conservation management, which should seek to promote the typical plant diversity of ancient semi-natural woodlands. In doing so, conservation management should promote the preservation of remaining ancient deciduous woodlands and inhibit the conversion of ancient woodlands to coniferous or mixed forests. Additionally, conservation management should strengthen the connections between recent and ancient woodlands through habitat corridors. Furthermore, careful forest management of deciduous ancient woodland sites with high typical woodland plant diversity has to be ensured to avoid soil damage. [ABSTRACT FROM AUTHOR]

Highlights: [•] European beech is considered to spread out in strictly protected European deciduous forests prospectively. [•] We analyse the effects of increasing beech proportion on herb-layer diversity and productivity. [•] In dark unmanaged stands, increasing beech proportion reduces herb-layer diversity and productivity. [•] In managed stands, light availability is significant for herb-layer diversity and productivity. [•] Forest management cessation may result in lower herb-layer diversity and productivity in many regions. [Copyright &y& Elsevier]

Abstract: How tree species diversity affects ecosystem functioning is a topic of intensive research. This study compares monospecific and species-rich broad-leaved forests under similar bedrock and climate conditions for the size and composition of their seed bank. We tested the hypotheses that (i) the actual herb-layer vegetation has an only weak influence on the composition of the seed bank, (ii) the species diversity of the seed bank increases with tree-layer diversity, and (iii) tree species forming a more persistent litter layer reduce the number of germinating seeds. The number of seeds and their species composition were investigated in soil cores taken from three soil depths (0–5, 5–10 and 10–20cm; n =4, with each 6 sub-samples) in 9 study plots differing in tree species diversity (3 monospecific Fagus plots, 3 plots with Fagus, Fraxinus and Tilia, and 3 plots with Fagus, Fraxinus, Tilia, Carpinus and Acer). Tree species diversity had a much stronger influence on the size and composition of the seed bank than herb-layer diversity or composition, the latter revealing only a low similarity to the corresponding seed bank. The number and species diversity of emerging seedlings decreased significantly with the amount of acidifying Fagus litter, but increased with litter mass of Tilia and other trees with nutrient-rich, rapidly decomposing litter. We conclude that tree species diversity does not influence the seed bank through effects on herb-layer composition, but mostly through differential disturbance histories of the stands and litter quality effects on germination and soil chemistry. From the contrasting effects of Fagus and Tilia leaf litter, it appears that effects of tree species identity are more relevant than influences of tree species diversity itself. [Copyright &y& Elsevier]

Abstract: Where natural production capacity permits, modern silvicultural management in Central Europe frequently aims at the development of mixed broadleaved stands, instead of pure European beech (Fagus sylvatica) stands. It is crucial to study the effects of these tree-layer diversity variations on herb-layer vegetation, since herb-layer vegetation contributes significantly to ecosystem functioning in forests. In Hainich National Park (Thuringia, Germany), we conducted observational research in deciduous stands to investigate whether herb-layer diversity was related to canopy-layer diversity, and to ascertain possible causal mechanisms. We found that herb-layer vegetation of deciduous forest stands rich in canopy species appeared to be more diverse than herb-layer vegetation of beech-dominated stands. We surmise that herbaceous understorey diversity was indirectly influenced by canopy tree species through the medium of the altered environmental factors soil pH and litter layer thickness. Apparently, lower beech proportion had a more profound effect than the number of secondary tree species. There were no correlations between herb-layer diversity and light transmissibility of the canopy layer, indicating that the light factor was not crucial for herb-layer diversity. At least for the Hainich research sites, our results indicated that small-scale light and soil heterogeneity is insignificant for herb-layer diversity. We found several herb-layer species whose occurrence was particularly correlated with tree-layer diversity and environmental factors. Remarkably, all species positively correlated with soil pH were important for the phytosociological classification of the research sites. Beech-dominated research sites showed high tree-layer volumes, whereas research sites with high tree-layer diversity tended to feature lower tree-layer volumes. These findings could be the result of differing former silvicultural systems and varying soil clay contents affecting tree species composition. In contrast, herb-layer biomass was positively correlated with tree-layer diversity. Herb-layer productivity might be promoted in more diverse research sites by increased nutrient supply and base saturation. It is also possible that greater beech proportion interfered with herb-layer productivity. However, herb-layer biomass was also positively correlated with herb-layer diversity. Hence, our study hints that positive diversity-functioning relationships might occur in the herb-layer of the deciduous forest under investigation. [Copyright &y& Elsevier]