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The future of dry forests around the world is uncertain given predictions that rising temperatures and enhanced aridity will increase drought‐induced tree mortality. Using forest management and ecological restoration to reduce density and competition for water offers one of the few pathways that forests managers can potentially minimize drought‐induced tree mortality. Competition for water during drought leads to elevated tree mortality in dense stands, although the influence of density on heat‐induced stress and the durations of hot or dry conditions that most impact mortality remain unclear.Understanding how competition interacts with hot‐drought stress is essential to recognize how, where and how much reducing density can help sustain dry forests in a rapidly changing world. Here, we integrated repeat measurements of 28,881 ponderosa pine trees across the western US (2000–2017) with soil moisture estimates from a water balance model to examine how annual mortality responds to competition, temperature and soil moisture conditions.Tree mortality responded most strongly to basal area, and was elevated in places with high mean temperatures, unusually hot 7‐year high temperature anomalies, and unusually dry 8‐year low soil moisture anomalies. Mortality was also lower in places that experienced unusually wet 3‐year soil moisture anomalies between measurements. Importantly, we found that basal area interacts with temperature and soil moisture, exacerbating mortality during times of stress imposed by high temperature or low moisture.Synthesis and applications. Our results imply that a 50% reduction in forest basal area could reduce drought‐driven tree mortality by 20%–80%. The largest impacts of density reduction are seen in areas with high current basal area and places that experience high temperatures and/or severe multiyear droughts. These interactions between competition and drought are critical to understand past and future patterns of tree mortality in the context of climate change, and provide information for resource managers seeking to enhance dry forest drought resistance. [ABSTRACT FROM AUTHOR]

Increasing aridity is a challenge for forest managers and reducing stand density to minimize competition is a recognized strategy to mitigate drought impacts on growth. In many dry forests, the most widespread and common forest management programs currently being implemented focus on restoration of historical stand structures, primarily to minimize fire risk and enhance watershed function. The implications of these restoration projects for drought vulnerability are not well understood. Here, we examined how planned restoration treatments in the Four Forests Restoration Initiative, the largest forest restoration project in the United States, would alter landscape‐scale patterns of forest growth and drought vulnerability throughout the 21st century. Using drought–growth relationships developed within the landscape, we considered a suite of climate and treatment scenarios and estimated average forest growth and the proportion of years with extremely low growth as a measure of vulnerability to long‐term decline. Climatic shifts projected for this landscape include higher temperatures and shifting seasonal precipitation that promotes lower soil moisture availability in the early growing season and greater hot‐dry stress, conditions negatively associated with tree growth. However, drought severity and the magnitude of future growth declines were moderated by the thinning treatments. Compared to historical conditions, proportional growth in mid‐century declines by ~40% if thinning ceases or continues at the status quo pace. By comparison, proportional growth declines by only 20% if the Four Forest Restoration Initiative treatments are fully implemented, and <10% if stands are thinned even more intensively than currently planned. Furthermore, restoration treatments resulted in dramatically fewer years with extremely low growth in the future, a recognized precursor to forest decline and eventual tree mortality. Benefits from density reduction for mitigating drought‐induced growth declines are more apparent in mid‐century and under RCP4.5 than under RCP8.5 at the end of the century. Future climate is inherently uncertain, and our results only reflect the climate projections from the representative suite of models examined. Nevertheless, these results indicate that forest restoration projects designed for other objectives also have substantial benefits for minimizing future drought vulnerability in dry forests and provide additional incentive to accelerate the pace of restoration. [ABSTRACT FROM AUTHOR]

Inter-annual variability (IAV) of forest Net Primary Productivity (NPP) is a function of both extrinsic (e.g., climate) and intrinsic (e.g., stand dynamics) drivers. As estimates of NPP in forests are scaled from trees to stands to the landscape, an understanding of the relative effects of these factors on spatial and temporal behavior of NPP is important. Although a high degree of spatial coherence (i.e., the degree of spatial synchrony over time) is often assumed, this inherent behavior is rarely examined. Quantifying this term may improve future predictions as site-level estimates are scaled up spatially. We quantified the spatial coherence of bole biomass production (BBP) within and between trees, and bole-related NPP (NPP B ) between sites of varying age, elevation, moisture, and species composition across a forested landscape in the western Cascade Range of Oregon. Within sites, individual trees with lower than average BBP were the most coherent. IAV of BBP increased as average tree BBP increased and spatial coherence was reduced. Among sites, NPP B was the most spatially coherent ( r = 0.92) between young sites, while older sites and comparisons between age classes revealed a much larger range in spatial coherence ( r = −0.18 to 0.85). Our findings indicate climate variability may be of greater importance for spatial coherence between young sites, and that intrinsic factors could be decreasing spatial coherence between older sites or sites not in close proximity. The wide range in spatial coherence between sites found in this study, coupled with the complex land use history patterns across forested landscapes, has significant implications for modeling and scaling of NPP. [ABSTRACT FROM AUTHOR]

Lodgepole pine (Pinus contorta) forests are widely distributed throughout North America and are subject to mountain pine beetle (Dendroctonus ponderosae) epidemics, which have caused mortality over millions of hectares of mature trees in recent decades. Mountain pine beetle is known to influence stand structure, and has the ability to impact many forest processes. Dwarf mistletoe (Arceuthobium americanum) also influences stand structure and occurs frequently in post-mountain pine beetle epidemic lodgepole pine forests. Few studies have incorporated both disturbances simultaneously although they co-occur frequently on the landscape. The aim of this study is to investigate the stand structure of lodgepole pine forests 21–28 years after a mountain pine beetle epidemic with varying levels of dwarf mistletoe infection in the Deschutes National Forest in central Oregon. We compared stand density, stand basal area, canopy volume, proportion of the stand in dominant/codominant, intermediate, and suppressed cohorts, average height and average diameter of each cohort, across the range of dwarf mistletoe ratings to address differences in stand structure. We found strong evidence of a decrease in canopy volume, suppressed cohort height, and dominant/codominant cohort diameter with increasing stand-level dwarf mistletoe rating. There was strong evidence that as dwarf mistletoe rating increases, proportion of the stand in the dominant/codominant cohort decreases while proportion of the stand in the suppressed cohort increases. Structural differences associated with variable dwarf mistletoe severity create heterogeneity in this forest type and may have a significant influence on stand productivity and the resistance and resilience of these stands to future biotic and abiotic disturbances. Our findings show that it is imperative to incorporate dwarf mistletoe when studying stand productivity and ecosystem recovery processes in lodgepole pine forests because of its potential to influence stand structure. [ABSTRACT FROM AUTHOR]

• Dry western US forests are at risk of large-scale tree die-offs due to hot-drought. • We modeled forest restoration effects on future ponderosa pine drought mortality. • Without thinning, mortality will increase 45–57% over current rates by mid-century. • With thinning, mid-century mortality rates remain near or below contemporary rates. • Lower tree density can mitigate the effects of climate change on drought mortality. Drought-induced tree mortality is predicted to increase in dry forests across the globe as future projections show hotter, drier climates. This could potentially result in large-scale tree die-offs, changes in species composition, and loss of forest ecosystem services, including carbon storage. While some studies have found that forest stands with greater basal areas (BA) have higher drought mortality, many have not evaluated the extent to which forests restored to lower densities via restoration activities affect drought mortality. The southwestern USA is particularly susceptible to tree mortality due to the predicted increases in temperature, drier soils, and forests with high density. Our objective was to evaluate how ponderosa pine mortality is expected to be influenced by the Four Forests Restoration Initiative (4FRI), a large-scale forest restoration effort ongoing in northern Arizona, USA, that will reduce stand BA by approximately 40%. Specifically, we modeled drought mortality in three time periods, one contemporary (1970–2010), and two future (2020–2059 and 2060–2099) under three restoration scenarios: no thinning, 4FRI thinning, and a BA reduction beyond the 4FRI plan (4FRI-intensive). We estimated mortality using 11 climate models under two emissions scenarios. Without thinning, our model predicted that by mid-century (2020–2059), changes in climate could increase annual ponderosa pine mortality rates by 45–57% over contemporary rates. However, with thinning, mid-century mortality was predicted to remain near or below contemporary rates and these rates are 31–35% (4FRI) and 46–51% (4FRI-intensive) less than the mid-century scenarios without thinning. Our study shows that while climate change is likely to increase tree mortality rates, large-scale forest restoration projects, such as 4FRI, have the potential to ameliorate the effects of climate change and keep mortality rates near contemporary levels for decades. [ABSTRACT FROM AUTHOR]

Abstract: Two common sampling methodologies coupled with a simple statistical model were evaluated to determine the accuracy and precision of annual bole biomass production (BBP) and inter-annual variability estimates using this type of approach. We performed an uncertainty analysis using Monte Carlo methods in conjunction with radial growth core data from trees in three Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) dominated sites (young, mature, and old-growth) in the western Cascades of Oregon. A model based on the mean and standard deviation of annual radial growth from sampled trees was used with and without stratification by tree size to predict radial growth for non-sampled trees. Sample sizes of 64–128 trees per stand were required to achieve accuracy and precision within ±10%. Without stratification the model underestimated annual BBP (Mgha−1 year−1) in all three age classes by up to 28%, and inter-annual variability by as much as 26%. Applying stratification increased accuracy of estimates at least twofold, and precision of estimates improved by 3–10%, resulting in decreased sample size requirements. The coefficient of variation of error of estimates was half that of inter-annual variability over the study period. Thus, this approach can be used to examine patterns of inter-annual variability of BBP in response to changing climate and land use patterns. [Copyright &y& Elsevier]