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Highlights • Plant functional traits (PFTs) significantly contributed to tree species classification. • Combining PFTs with spectral features and LiDAR metrics yielded the highest classification accuracy. • Equivalent water thickness was the most robust PFTs for the classification. Abstract Plant functional traits have been extensively used to describe, rank and discriminate species according to their variability between species in classical plant taxonomy. However, the utility of plant functional traits for tree species classification from remote sensing data in natural forests has not been clearly established. In this study, we integrated three selected plant functional traits (i.e. equivalent water thickness (Cw), leaf mass per area (Cm) and leaf chlorophyll (Cab)) retrieved from hyperspectral data with hyperspectral derived spectral features and airborne LiDAR derived metrics for mapping five tree species in a natural forest in Germany. Our results showed that when plant functional traits were combined with spectral features and LiDAR metrics, an overall accuracy of 83.7% was obtained, which was statistically significantly higher than using LiDAR (65.1%) or hyperspectral (69.3%) data alone. The results of our study demonstrate that plant functional traits retrieved from hyperspectral data using radiative transfer models can be used in conjunction with hyperspectral features and LiDAR metrics to further improve individual tree species classification in a mixed temperate forest. [ABSTRACT FROM AUTHOR]

Numerous airborne LiDAR-derived metrics have been proposed for classifying tree species. Yet an in-depth ecological and biological understanding of the significance of these metrics for tree species mapping remains largely unexplored. In this paper, we evaluated the performance of 37 frequently used LiDAR metrics derived under leaf-on and leaf-off conditions, respectively, for discriminating six different tree species in a natural forest in Germany. We firstly assessed the correlation between these metrics. Then we applied a Random Forest algorithm to classify the tree species and evaluated the importance of the LiDAR metrics. Finally, we identified the most important LiDAR metrics and tested their robustness and transferability. Our results indicated that about 60% of LiDAR metrics were highly correlated to each other (| r | > 0.7). There was no statistically significant difference in tree species mapping accuracy between the use of leaf-on and leaf-off LiDAR metrics. However, combining leaf-on and leaf-off LiDAR metrics significantly increased the overall accuracy from 58.2% (leaf-on) and 62.0% (leaf-off) to 66.5% as well as the kappa coefficient from 0.47 (leaf-on) and 0.51 (leaf-off) to 0.58. Radiometric features, especially intensity related metrics, provided more consistent and significant contributions than geometric features for tree species discrimination. Specifically, the mean intensity of first-or-single returns as well as the mean value of echo width were identified as the most robust LiDAR metrics for tree species discrimination. These results indicate that metrics derived from airborne LiDAR data, especially radiometric metrics, can aid in discriminating tree species in a mixed temperate forest, and represent candidate metrics for tree species classification and monitoring in Central Europe. [ABSTRACT FROM AUTHOR]

• Combining multi-temporal aerial photographs with LiDAR significantly improved the accuracy of individual tree species mapping. • Texture features captured by multi-temporal digital CIR orthophotos with different view-illuminations were species-specific. • The average gray-level and the dissimilarity of the NIR band contributed most (among texture features) to the classification. Digital colour-infrared (CIR) aerial photographs, which have been collected routinely in many parts of the world, are an invaluable data source for the monitoring and assessment of forest resources. Yet, the potential of these data for automated individual tree species mapping remains largely unexplored. One way to maximize the usefulness of digital CIR aerial photographs for individual tree species mapping is to integrate them with modern and complementary remote sensing technologies such as the light detection and ranging (LiDAR) system and 3D segmentation algorithms. In this study, we examined whether multi-temporal digital CIR orthophotos could be used to further increase the accuracy of airborne LiDAR-based individual tree species mapping for a temperate mixed forest in eastern Germany. Our results showed that the texture features captured by multi-temporal digital CIR orthophotos under different view-illumination conditions were species-specific. As a consequence, combining these texture features with LiDAR metrics significantly improved tree species mapping accuracy (overall accuracy: 77.4%, kappa: 0.68) compared to using LiDAR data alone (overall accuracy: 69.3%, kappa: 0.58). Among various texture features, the average gray level in the near-infrared band was found to contribute most to the classification. Our results suggest that the synergic use of multi-temporal digital aerial photographs and airborne LiDAR data has the potential to accurately classify individual tree species in Central European mixed forests. [ABSTRACT FROM AUTHOR]

Gap fraction ( P gap ) and vertical gap fraction profile (vertical P gap profile) are important forest structural metrics. Accurate estimation of P gap and vertical P gap profile is therefore critical for many ecological applications, including leaf area index (LAI) mapping, LAI profile estimation and wildlife habitat modelling. Although many studies estimated P gap and vertical P gap profile from airborne LiDAR data, the scan angle was often overlooked and a nadir view assumed. However, the scan angle can be off-nadir and highly variable in the same flight strip or across different flight strips. In this research, the impact of off-nadir scan angle on P gap and vertical P gap profile was evaluated, for several forest types. Airborne LiDAR data from nadir (0°∼7°), small off-nadir (7°∼23°), and large off-nadir (23°∼38°) directions were used to calculate both P gap and vertical P gap profile. Digital hemispherical photographs (DHP) acquired during fieldwork were used as references for validation. Our results show that angular P gap from airborne LiDAR correlates well with angular P gap from DHP ( R 2  = 0.74, 0.87, and 0.67 for nadir, small off-nadir and large off-nadir direction). But underestimation of P gap from LiDAR amplifies at large off-nadir scan angle. By comparing P gap and vertical P gap profiles retrieved from different directions, it is shown that scan angle impact on P gap and vertical P gap profile differs amongst different forest types. The difference is likely to be caused by different leaf angle distribution and canopy architecture in these forest types. Statistical results demonstrate that the scan angle impact is more severe for plots with discontinuous or sparse canopies. These include coniferous plots, and deciduous or mixed plots with between-crown gaps. In these discontinuous plots, P gap and vertical P gap profiles are maximum when observed from nadir direction, and then rapidly decrease with increasing scan angle. The results of this research have many important practical implications. First, it is suggested that large off-nadir scan angle of airborne LiDAR should be avoided to ensure a more accurate P gap and LAI estimation. Second, the angular dependence of vertical P gap profiles observed from airborne LiDAR should be accounted for, in order to improve the retrieval of LAI profiles, and other quantitative canopy structural metrics. This is especially necessary when using multi-temporal datasets in discontinuous forest types. Third, the anisotropy of P gap and vertical P gap profile observed by airborne LiDAR, can potentially help to resolve the anisotropic behavior of canopy reflectance, and refine the inversion of biophysical and biochemical properties from passive multispectral or hyperspectral data. [ABSTRACT FROM AUTHOR]