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Gryllobencain n. gen. urn:lsid:zoobank.org:act: BF978806-B42C-489E-BAE2-306DAC00B8B5 TYPE SPECIES. — Gryllobencain patrickmuelleri n. sp. DERIVATIO NOMINIS. — ‘Gryllo’forcricket;‘ben’isHebrewfor‘sonof’and provides connection to the next part of the name;‘cain’ for the biblical figure. According to some legends,the children of Cain became the vampires.The name hence means ‘vampire cricket’ in reference to the prominent sickleshaped proximal parts of the maxilla,distantly reminding of vampire teeth. DIAGNOSIS. — As for the species. REMARK The current version of the International Code of Zoological Nomenclature demands that a newly described species is assigned to a genus. Due to the uncertainty of relationship and the apparent unique combination of characters of the new fossils we cannot assign them to any known genus. We therefore must erect a new one. While it is in principle possible to avoid creating this factually unnecessary additional category and we generally support the idea to not erect a new genus in such a case, we here provide a classical approach for the convenience of readers used to this approach. We still want to point out that the uncertainty concerning relationship could much better be expressed with the combination of the next higher group (in this case ‘ Ensifera patrickmuelleri ’, see for this approach discussion in Haug & Haug 2016 and references therein; Wagner et al. 2019).

Gryllobencain n. gen. urn:lsid:zoobank.org:act: BF978806-B42C-489E-BAE2-306DAC00B8B5 TYPE SPECIES. ��� Gryllobencain patrickmuelleri n. sp. DERIVATIO NOMINIS. ��� ���Gryllo���forcricket;���ben���isHebrewfor���sonof���and provides connection to the next part of the name;���cain��� for the biblical figure. According to some legends,the children of Cain became the vampires.The name hence means ���vampire cricket��� in reference to the prominent sickleshaped proximal parts of the maxilla,distantly reminding of vampire teeth. DIAGNOSIS. ��� As for the species. REMARK The current version of the International Code of Zoological Nomenclature demands that a newly described species is assigned to a genus. Due to the uncertainty of relationship and the apparent unique combination of characters of the new fossils we cannot assign them to any known genus. We therefore must erect a new one. While it is in principle possible to avoid creating this factually unnecessary additional category and we generally support the idea to not erect a new genus in such a case, we here provide a classical approach for the convenience of readers used to this approach. We still want to point out that the uncertainty concerning relationship could much better be expressed with the combination of the next higher group (in this case ��� Ensifera patrickmuelleri ���, see for this approach discussion in Haug & Haug 2016 and references therein; Wagner et al. 2019)., Published as part of Haug, Joachim T., H��rnig, Marie K., Kiesm��ller, Christine, Pazinato, Paula G., Baranov, Viktor & Haug, Carolin, 2022, A 100 - million-year-old ensiferan with unusual mouthparts and comments on the evolution of raptorial appendages within Polyneoptera, pp. 57-73 in Geodiversitas 44 (4) on pages 59-60, DOI: 10.5252/geodiversitas2022v44a4, http://zenodo.org/record/6000901, {"references":["HAUG J. T. & HAUG C. 2016. - \" Intermetamorphic \" developmental stages in 150 million-year-old achelatan lobsters - The case of the species tenera Oppel, 1862. Arthropod Structure & Development 45 (2): 108 - 121. https: // doi. org / 10.1016 / j. asd. 2015","WAGNER P., HAUG J. T. & HAUG C. 2019. - A new calmanostracan crustacean species from the Cretaceous Yixian Formation and a simple approach for differentiating fossil tadpole shrimps and their relatives. Zoological Letters 5 (1): art. 20. https: // doi. org / 10.1186 / s 40851 - 019 - 0136 - 0"]}

Gryllobencain patrickmuelleri n. gen., n. sp. urn:lsid:zoobank.org:act: 80C207C0-1B2E-44F9-90F8-A53E9229461B HOLOTYPE. ��� PED 0147. PARATYPE. ��� PED 0178. ADDITIONAL MATERIAL. ��� BUB 3072, 3073, from the collection of Patrick M��ller. DERIVATIO NOMINIS. ��� In honour of Patrick M��ller and his effort for research, especially on Myanmar amber. DIAGNOSIS. ��� Specimen with prominent maxillae [maxillulae]; medio-proximal part prominent sickle-shaped; palp prominent with long and strong setae; foreleg and mid leg [thorax appendages 1 and 2] with many long and strong setae mostly medially, on femur and tibia. Tibia additionally with strong disto-median spur-like spine. Cerci prominent with numerous very long and strong setae. DESCRIPTION General Body organised into head and trunk, the latter subdivided into thorax and abdomen (Figs 1A, B; 4A, C; 5A, C; 6A, C), presumably with 20 segments, ocular segment plus 19 post-ocular segments. Head Ocular segment and post-ocular segments1-5forming distinct capsule (head capsule) (Figs1A; 4A; 5A; 6A).Head capsule oval in dorsal(or ventral) view; about 1.6�� wider than long. Surface of head capsule dorsallywithprominentY-shapedmoultingsuture(epicranialsuture) (Fig. 5A). Lateral edges of head capsule rounded with few setae. Ocular segment recognisable by prominent compound eyes laterally projecting from the head capsule; additionally, three ocelli present [median eyes] (Fig. 5B). Compound eyes with numerous facets; difficult to count, at least 40 rows of facets (Fig. 7A, B). Ocelli arranged in a triangle; central ocellus dorsal to lateral ones. Appendage derivative of ocular segment (clypeo-labrum complex) [hypostome-labrum complex] well developed (Fig. 7D). Clypeus [hypostome] rectangular, about 2�� wider than long. Labrum lobe-like, about as large as clypeus, anterior edge gently rounded. Post-ocular segment 1 recognisable by its appendage, antenna [antennula]. Antenna arising from head capsule antero-dorsally; lateral to the clypeus; very long, longer than main body, with numerous elements (antennomeres), more than 40 (Figs 1A, B; 5A, C); some elements are longer and possess a distinct constriction (indication of future subdivision?); length of elements strongly varying; proximal elements with few, but prominent setae. More distal elements with numerous, but shorter setae. Post-ocular segment 2 (intercalary segment) not recognisable externally. Post-ocular segment 3 recognisable by its appendage, mandible (Fig. 7 A-D). Mandibles prominent, curved, bearing teeth on inner surface. Post-ocular segment 4 recognisable by its appendage, maxilla [maxillula] (Fig. 7 A-F). Proximal part of maxilla (cardo) [basipod?] medially drawn out; cardo distally carrying palp [endopod]. Drawn out part (cardo? stipes? both?) prominent, sickle-shaped; laterally with at least three setae. Palp with five elements (Figs 1C; 6D; 7F). Element 1 (most proximal one) short, as long as wide (diameter). Element 2 sub-similar to element 1. Element 3 elongate, at least 4.5�� longer than wide (diameter); with at least two strong spine-like setae and numerous smaller setae of varying sizes. Element 4 shorter than preceding element, elongate, at least 2.8�� longer than wide (diameter); with at least two strong spine-like setae and numerous smaller setae of varying sizes. Element 5 (most distal one) elongate, about 4�� longer than wide (diameter); with at least six strong spine-like setae and numerous smaller setae of varying sizes. Post-ocular segment 5 recognisable by its appendage, labium [maxilla] (Fig. 7E, F); only distal parts, palp [endopod], apparent. Palp mostly concealed, with numerous setae. Thorax [anterior trunk] Post-ocular segments 6-8 differentiated from posterior ones, forming a distinct unit, thorax (Fig. 5A). Tergite of postocular segment 6 (tergite of thorax segment 1; pronotum) about 0.55�� narrower than head; about 1.6�� wider than long; rectangular-shaped in dorsal view; with six smaller setae medio-anteriorly. Post-ocular segment 6 (thorax segment 1; prothorax) ventrally with a pair of prominent appendages (foreleg) [thoracopod 1] (Figs1D; 2A; 4B, D; 5D); basically cylindrical, composed of five elements. Element 1 (coxa) [basipod?] about as long (proximal-distal axis) as wide (diameter); with at least three smaller, thin setae posteriorly. Element 2 (trochanter) [endopod element 1?] slightly longer than wide, slightly longer than coxa; with more than nine small, thin setae medially. Element 3 (femur) [endopod element 2?] very elongate; about 10�� longer than coxa, similar width at proximal part, tapering towards distally to about 0.48�� of maximum width; with more than ten small, thin setae proximo-medially and disto-laterally; also with more than ten strong spine-like setae arranged in two rows (Fig. 4D), the majority of which is located proximally, with one exception disto-laterally. Element 4 (tibia) [endopod element 3?] elongate, about 0.75�� the length of the femur; about as wide as the femur, though slightly less proximally; covered with smaller setae, more distally than proximally; with three strong spine-like setae disto-medially of varying size and one strong spine-like seta laterally, more towards proximal than distal; with one very strong spine very medio-distally. Element 5 (tarsus) [endopod element 4?] slightly shorter than preceding element, more than 0.8�� of the length and about half of the width; with numerous smaller, thin setae throughout its surface; distally with a pair of claws, possible attachment structure between claws not apparent; tarsus subdivided into three sub-elements (tarsomeres) which are of mostly similar length (0.3�� of total length) and similar width, most proximal element has three stronger spine-like setae distally. Post-ocular segment 7 (thorax segment 2; mesothorax) about half as long as prothorax and about as wide as the latter, but slightly tapering posteriorly; trapezoid-shaped; with one small seta on each side respectively (antero-laterally), close to the coxa. Ventrally with a pair of prominent appendages (mid leg) [thoracopod 2] (Fig. 2B); basically cylindrical, composed of five elements; as a whole slightly longer than foreleg [thoracopod 1]. Element 1 (coxa) [basipod?] about as long (proximal-distal axis) as wide (diameter); with two smaller setae medially. Element 2 (trochanter) [endopod element 1?] slightly longer than wide, but as wide as the coxa; with one small, thin seta and two stronger, spine-like setae medio-distally. Element 3 (femur) [endopod element 2?] very elongate; about 10�� longer than coxa, almost similar width at proximal part, then widening to 1.5�� width at about 30 % of total femur length, then tapering towards distally to about 0.67�� of maximum width; with more than ten small, thin setae laterally throughout its surface; also with more than ten strong spine-like setae with varying sizes, the majority of which located medially, with one exception laterally. Element 4 (tibia) [endopod element 3?] elongate, slightly shorter than femur; about as wide as femur, though slightly less so proximally; covered with smaller setae, more distally than proximally; with two strong spine-like setae and more than five smaller spine-like setae disto-medially; also with one strong spine-like seta laterally, more towards proximal than distal, and one very strong spine very medio-distally. Element 5 (tarsus) [endopod element 4?] slightly shorter than tibia, more than 0.8�� of the length and about half of the width; with numerous smaller, thin setae throughout; distally with a pair of claws, possible attachment structure between claws not apparent; tarsus subdivided into three sub-elements which are of mostly similar length (0.3�� of total length) and similar width. Post-ocular segment 8 (thorax segment 3; metathorax) about 1.4�� longer than preceding segment and about as wide as that as well; rectangular shaped. Ventrally with a pair of prominent appendages (hind leg) [thoracopod 3] (Figs 2C; 3); basically cylindrical, composed of supposedly five elements; slightly longer than mid leg [thoracopod 2]. Element 1 (coxa) [basipod?] not differentiable from next element, i.e. element 2 (trochanter) [endopod element 1?]. Proximal region (coxa + trochanter?) slightly longer than wide. Element 3 (femur) [endopod element 2?] very elongate; about 8�� longer than proximal region, similar width as preceding element, but tapering slightly distally; with four strong spine-like setae laterally. Element 4 (tibia) [endopod element 3?] elongate, but about 0.8�� shorter than femur; about 0.67�� the width of the femur, tapering slightly distally; covered with smaller setae and stronger spine-like setae; no very strong spine very medio-distally, but with one strong spine-like seta on a similar position as on the tibia of fore and mid legs, but shorter than in the latter. Element 5 (tarsus) [endopod element 4?] as long as tibia and slightly less wide than that as well; with numerous smaller, thin setae throughout; distally with a pair of claws, possible attachment structure between claws not apparent; tarsus subdivided into three sub-elements which are of similar width, but with varying length (from proximal to distal: 0.52��, 0.22�� and 0.26�� of total length). Abdomen [posterior trunk] Post-ocular segments 9-19 differentiated from further anterior ones, forming distinct unit (abdomen; not corresponding to abdomen in other crustacean groups) (Figs 1A, B; 5A, C). Only eight distinct units differentiable, first seven probably corresponding to abdominal segments 1 to 7, rectangularshaped. Last unit most likely corresponding to several abdomen segments (8-11?), triangular-shaped. Post-ocular segment 9 (abdomen segment 1) about 1.4�� longer than metathorax; but less wide, about 0.67�� of metathorax width; one spine-like seta postero-laterally on each side. Post-ocular segment 10 (abdomen segment 2) similar to preceding segment, but very slightly shorter and slightly wider posteriorly (widening posteriorly); one spine-like seta postero-laterally on each side. Post-ocular segment 11 (abdomen segment 3) about half as long, but as wide as the preceding segment; one spine-like seta postero-laterally respectively. Post-ocular segment 12 (abdomen segment 4) slightly longer and wider than preceding segment; two spine-like setae postero-laterally respectively, one slightly longer than the other. Post-ocular segment 13 (abdomen segment 5) similar to preceding segment, but slightly longer and very slightly tapering posteriorly; two spine-like setae postero-laterally on each side, one slightly longer than the other. Post-ocular segment 14 (abdomen segment 6) slightly shorter and less wide than preceding segment, tapering slightly posteriorly; one spine-like seta postero-laterally on each side. Post-ocular segment 15 (abdomen segment 7) similar to preceding segment, but less wide and tapering posteriorly; one spine-like seta postero-laterally on each side. Trunk end (post-ocular segments 16-19?; abdomen segments 8-11?) as long as first abdomen segment; anteriorly as wide as preceding segment, but tapering drastically posteriorly. Ventrally with a pair of prominent appendages (cerci) (Figs 1A, B; 3; 5A, C; 6B); basically cylindrical, not subdivided; about 0.8�� the length of the antenna; as wide as the tibia of the hind leg proximally and tapering slightly until about 70 % of its length, then strongly tapering towards tip; bearing numerous spine-like setae of varying sizes, mostly strong and very long (compared to similar structures elsewhere); also with a few smaller, shorter setae throughout. Size of specimens While the specimens are very similar in overall appearance, they differ in size. Though it remains difficult to measure, it appears that the four specimens fall into three size classes (Fig. 8). It seems likely that these represent three successive stages. It remains unclear whether the largest size class corresponds to the adult stage., Published as part of Haug, Joachim T., H��rnig, Marie K., Kiesm��ller, Christine, Pazinato, Paula G., Baranov, Viktor & Haug, Carolin, 2022, A 100 - million-year-old ensiferan with unusual mouthparts and comments on the evolution of raptorial appendages within Polyneoptera, pp. 57-73 in Geodiversitas 44 (4) on pages 60-67, DOI: 10.5252/geodiversitas2022v44a4, http://zenodo.org/record/6000901

Studies on aquatic Diptera in the Plitvice Lakes National Park (Croatia) conducted in the last 50 years have produced 157 species and 7 taxa of aquatic Diptera placed in 13 families. Samples were collected at 25 sampling sites representing the four main types of karst aquatic habitats: spring, stream, tufa barriers and lakes. All records of all the aquatic families of Diptera in Plitvice Lakes NP are summarized, including previously unpublished data. Twelve species new for Plitvice Lakes NP are recorded for the first time, belonging to the families: Chironomidae – Labrundinia longipalpis (Goetghebuer, 1921), Nilothauma brayi (Goetghebuer, 1921), Potthastia longimanus Kieffer, 1922, Polypedilum (Polypedilum) nubeculosum (Meigen, 1804), Tanytarsus brundini Lindeberg, 1963; Dixidae – Dixella autumnalis (Meigen, 1838); Scathophagidae – Acanthocnema latipennis Becker, 1894 and Stratiomyidae – Oxycera pardalina Meigen, 1822, Oxycera limbata Loew, 1862, Oxycera turcica Ustuner & Hasbenli, 2004, Nemotelus pantherinus (Linnaeus, 1758), Oplodontha viridula (Fabricius, 1775). The most species-rich family was the Chironomidae with 62 species (and an additional seven taxa), followed by the Empididae with 22 species and Limoniidae with 19 species. The highest number of species was recorded in springs. The relatively low number of species in certain families and the complete absence of some aquatic families shows that further research into the aquatic Diptera in Plitvice Lakes NP is needed.

Hydrobaenus korneyevi (Baranov, 2011), comb. nov. (Figs 10–20, 25–26, 29) Parorthocladius korneyevi Baranov, 2011a: 406. Type material examined. HOLOTYPE: ♁ (SIZK), ʻ UKRAINE: CRIMEA: leg. V. Baranov; Simferopol, Salgir River, 29.XI.2010, handnetʼ. PARATYPES: 2 ♁♁ (JBMF), same locality as for holotype. Diagnosis. The description of male adult of H. korneyevi, as provided in the original paper (BARANOV 2011a), can be completed and supplemented with the following additional characters: lobes of antepronotum distinctly gaping; acrostichals reduced, composed of 4–5 setae starting some distance from antepronotum; sensilla chaetica present on tibia and tarsomeres ta1–ta5 of PI, PII and PIII; tergite IX broadly triangular and narrowing distally, bearing medially a distinct elevated truncate hump; distal part of tergite IX and anal point densely covered with about 60 setae which are placed mainly on sides of dorsal hump; anal point triangular to drop-shaped, elongate, nearly parallel-sided to weakly narrowing distally, ending with a short finger-like apex; virga consisting of 6–8 long subequal thin spines; inferior volsella composed of 2 distinct lobes, proximal one semi-spherical and much larger; gonostylus massively bulbous medially, with blackish long and short setae covering posterior and lateral sides; crista dorsalis consisting of strong preapical triangular tooth, orally projecting and placed close to megaseta, which is markedly bent inwards. Redescription. Male adult (n = 3; Figs 10–20, 25–26, 29). Large sized Hydrobaenus species. Total length 4.40–4.60 mm; wing length 2.60–2.70 mm. General colouration blackish with contrasting brown to blackish thorax and legs. Mesonotal stripes distinctly blackish; legs dark brown to blackish; tergites I–VIII brown to dark brown; anal segment blackish, crista dorsalis distinctly dark brown. Head. Eyes bare. Temporals consisting of 12–13 setae including 7–8 inner and 5 outer verticals. Antenna 810– 830 μm long, 13-segmented, length (μm) of segments: 1, 50; 2–12, 30–35 (nearly sub-equal), last flagellomere 420–430; distal part of last flagellomere (Figs 23–24) moderately clubbed ending with forked rounded apex; antennal groove beginning on segment 1 and reaching ultimate flagellomere; AR 1.10. Clypeus with 8 setae. Palp 5-segmented, not reduced, palpomere 3 (Fig. 10) with 4–5 sensilla clavata. Thorax. Lobes of antepronotum (Fig. 29) distinctly gaping, lateral antepronotals 6–7; acrostichals (Fig. 11) reduced, composed of 4–5 setae in 1 row, starting some distance from antepronotum; dorsocentrals 15–17 in 2 rows; prealars 4–5 in 1 row. Scutellum with 13–14 setae in 2 rows. Wing. Brachiolum with 1 seta. Distribution of setae on veins: R, 8–9; R1, 0; R2+3, 1–2; remaining veins bare. Squama with 10–14 setae in 1–2 rows. Legs. Sensilla chaetica present on tibia and tarsomeres ta1–ta5 of PI, PII and PIII. Length (μm) and proportions of legs as in Table l. Hypopygium in dorsal and ventral view (Figs 15–16). Tergite IX broadly triangular, narrowing distally, with elevated truncate hump medially (Figs 12–14) which is clearly visible in lateral view, distal part of tergite IX and anal point densely covered with setae (about 60). Anal point (Figs 12–14) about 90–100 μm long, 60–70 μm maximum width at base and 20–25 μm at apex, long triangle to parallel-sided, slightly narrowing distally, entirely covered with setae (less at base than at apex), apex ending with small finger-like apex (Figs 12 a–12b). Virga (Figs 15–16, 18) consisting of 6–8 subequal fine spines about 55–60 μm long and fused at base. Gonocoxite 250–260 μm long, maximum width 125–130 μm, with rounded apex; inferior volsella (dorsal, Figs 15, 17) composed of 2 large lobes, proximal lobe distinctly larger and semi-circular. Gonostylus (Figs 19–20) 120–125 μm long, maximum width 60–70 μm, massively bulb-shaped; posterior and posterolateral margins densely covered with characteristic short and stout blackish setae, posterior and lateral side with 9–10 long stout characteristic setae; crista dorsalis consistently chitinous, triangular with pointed apex, orally directed and placed pre-apically close to megaseta, clearly visible in both dorsal and lateral view (Figs 19–20); megaseta about 20 μm long, markedly bent inwards. Taxonomic position. The male adult of H. korneyevi resembles that of H. dentistylus and H. lugubris in having a similar shape of inferior volsella and a high number of setae on tergite IX and anal point. Nevertheless, the massive structure of gonostylus and the strong triangular crista dorsalis look more similar to those of H. dentistylus. However, H. korneyevi can be separated from other related Hydrobaenus species by the following characters: presence of a distinct truncate elevated hump on median part of tergite IX (clearly visible in lateral view); anal point long triangle to long drop-shaped, often ending in a finger-like apex (Figs 12 a–12b); virga with 6–8 long thin spines; lobes of inferior volsella differently shaped, proximal one rounded and much larger; gonostylus densely covered with blackish short and long setae, crista dorsalis strong tooth-like, triangular with pointed apex, placed pre-apically close to megaseta. Ecology. Male adults of H. korneyevi are collected only during winter time between December and January: dates of emergence recorded in the Salgir River basin are: 29.12.2010 and 21.1.2013. Adults were recorded swarming, flying and moving on the vegetation and on the snow at low temperatures (Distribution. Hydrobaenus korneyevi is reported from the Central part of the Crimean Peninsula, including its type locality (the Salgir river in Gagarin Park in the centre of Simferopol) as well as from Botanical Garden of Taurida State University and from Western Bulganak river, next to Pozharske village (BARANOV 2011a,b)., Published as part of Moubayed-Breil, Joel & Baranov, Viktor, 2018, Taxonomic notes on the genus Hydrobaenus with description of H. simferopolus sp. nov. from Crimea (Diptera: Chironomidae), pp. 347-355 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 58 (2) on pages 349-351, DOI: 10.2478/aemnp-2018-0029, http://zenodo.org/record/4504794, {"references":["BARANOV V. 2011 a: New and rare species of Orthocladiinae (Diptera, Chironomidae) from the Crimea, Ukraine. Vestnik Zoologii 45: 405 - 410."]}

Hydrobaenus dentistylus Moubayed, 1985 (Figs 1–9, 23–24) Type material examined. PARATYPES: LEBANON: Litany River, Bekaa plain, 800–850 m a.s.l., 29.iii.1982, 5 ♁♁, J. Moubayed-Breil lgt. (2 ♁♁ IBNC, 2 ♁♁ JBMF, 1 ♁ ZSMC). Diagnostic characters. The differentiating characters provided in the original paper (MOUBAYED 1985) can be summarized and supplemented by the following main distinguishing characters found in the male adult: antenna, palp and legs not reduced; lobes of antepronotum gaping; acrostichals 4–5 in total, reduced, starting some distance from antepronotum; anal point drop-like, tergite IX and anal point densely covered with about 60–70 setae reaching apex of anal point; virga consisting of 3 curved unequal teeth (median one shorter); inferior volsella consisting of 2 unequal large lobes with proximal lobe broader; gonostylus bulb-like, globular and bearing a distinct pre-apical pointed tooth placed close to the megaseta. Redescription. Male adult (n = 5; Figs 1–9, 23–24). Small to medium sized Hydrobaenus species. Total length 2.85–3.00 mm. Wing length 2.20–2.40 mm. General colouration contrasting brown to blackish. Head dark brown, antennae pale; thorax contrasting brown to blackish, mesonotal stripes distinctly blackish; wing pale; legs dark brown to blackish. Tergites I–VIII brown; anal segment brown to dark brown, crista dorsalis dark brown. Head. Eyes bare. Temporals consist of 9–11 setae including 5–6 inner and 4–5 outer verticals.Antenna and palp are not reduced. Antenna 810–830 μm long, 13-segmented, length (μm) of segments: 1, 50; 2–12, 30–35 (nearly sub-equal), last flagellomere 420–430; distal part of last flagellomere (Figs 35–36) moderately clubbed ending with forked rounded apex; antennal groove beginning on segment 1 and reaching ultimate flagellomere; AR 1.10. Clypeus with 8 setae. Palp 5-segmented, palpomere 3 (Fig. 1) with 3–4 sensilla clavata. Thorax. Acrostichals reduced, consisting of 4–5 uniserial setae beginning some distance from antepronotum; antepronotum with 6–7 lateral antepronotals, lobes well gaping as in H. simferopolus sp. nov. (Fig. 28); prealars 4–5 in 1 row. Scutellum with 8 uniserial setae. Wing. Brachiolum with 1 seta. Distribution of setae on veins: R, 8–12; R1, 25–7; remaining veins bare. Squama with 9 uniserial setae. Legs. Sensilla chaetica present on tibia and tarsomeres ta 1 –ta 5 of PI, PII and PIII. Hypopygium in dorsal view with gonostylus removed (Fig. 2). Tergite IX (Figs 2–3) broadly rectangular, narrowing distally, slightly sinuous in lateral view (Fig. 3), 60–70 setae present on both tergite IX and anal point. Anal point (dorsal, Figs 2, 4; lateral, Fig. 3) about 40–45 μm long, 30 μm maximum width at base, drop-like and densely covered with setae. Virga (Figs 2, 5) consisting of 3 long spines about 45 μm long, fused at base, median one shorter. Gonocoxite 250–260 μm long, maximum width 120–125 μm, with rounded apex; inferior volsella (dorsal, Figs 2, 7; lateral, Fig. 6) 95–105 μm long, 30–35 μm maximum width, consisting of 2 large lobes; proximal lobe distinctly larger. Gonostylus (Figs 8–9) 115 μm long, maximum width 65–70 μm, massively spherical to bulb-shaped medially and distally; hyaline area present on posterolateral side; anterior margin swollen and bearing 5–6 setae placed on proximal part; crista dorsalis consisting of 1 triangular characteristic and strong pre-apical teeth, orally directed and placed close to megaseta, clearly visible in both dorsal and lateral view (Figs 8–9); megaseta dark brown, conspicuous and slightly bent inwards. Ecology. Large material composed of adults and pupae of H. dentistylus is recorded from shallow flowing water with rich aquatic vegetation. Emergence reported in early springtime: from March to April. Chironomid species encountered in the type locality of H. dentistylus include: Potthastia gaedii (Meigen, 1838), P. longimanus Kieffer, 1922, Sympotthastia zavreli Pagast, 1947, Smittia durandae Moubayed, 1989, Polypedilum anjarum Moubayed, 1989, P. aegyptium Kieffer, 1925, P. longisetum Moubayed, 1992, P. yammounei Moubayed, 1992, Micropsectra lindrothi Goetghebuer, 1931, M. sofiae Stur & Ekrem, 2006, Rheotanytarsus curtistylus (Goetghebuer, 1921), R. rhenanus Klink, 1983 and R. ringei Lehman, 1970. Distribution. Hydrobaenus dentistylus is considered a typical biogeographic representative of the Levantine Province. This species is only known from its type locality: middle basin of the Litany River, Bekaa plain, Lebanon, alt. 800– 850 m., Published as part of Moubayed-Breil, Joel & Baranov, Viktor, 2018, Taxonomic notes on the genus Hydrobaenus with description of H. simferopolus sp. nov. from Crimea (Diptera: Chironomidae), pp. 347-355 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 58 (2) on pages 348-349, DOI: 10.2478/aemnp-2018-0029, http://zenodo.org/record/4504794, {"references":["MOUBAYED Z. 1985: Les Chironomidae (Diptera) du Liban. III. Hydrobaenus dentistylus n. sp. (Orthocladiinae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 121: 73 - 76."]}

Hydrobaenus lugubris Fries, 1830 (Figs 21, 22, 27, 30–34) Material examined. FRANCE: Vosges Region, Mairy and Valtin areas, 400–600 m a.s.l., 24.ii.2014 & 11.iii.2014, 47 ♁♁ 6♀♀, Gennaro Coppa lgt.(JBMF); Natural Reserve of Lake Remoray, 27.xi.2017, 2♀♀ (pupal exuviae), Bruno Tissot lgt. (JBMF). Diagnosis. Male adult of H. lugubris resembles that of H dentistylus and H. korneyevi in having: a similar shape of inferior volsella; high number of dorsal setae on tergite IX and anal point. Its description, as provided in SAETHER (1976), can be supplemented by the following additional relevant characters: antenna, palp and legs reduced; palpomere 3 without 3 sensilla clavata; acrostichals not reduced, composed of 4–6 curved setae in 1 row; sensilla chaetica present on tibia and tarsomeres ta1–ta5 of PI–PIII, tarsomere ta4 of PI–PII distinctly shorter than ta5, BR of all legs lower than 1 (0.60–0.95); tergite IX with distinct elevated hump clearly visible in lateral view), anterior part with blackish transverse band, about 130–140 setae present on tergite I and anal point; anal point drop-shaped; virga present, consisting of 4–5 long unequal thin spines; inferior volsella composed of 2 subequal well separated lobes; gonostylus without crista dorsalis. Redescription. Male adult (n = 5; Figs 21–22, 27, 31–34, 38). Medium sized Hydrobaenus species. Total length 3.60–3.80 mm; wing length 1.75–1.85 mm. General colouration contrasting brownish to yellowish with contrasting brown to dark brown thorax, legs and abdomen. Head brownish; legs dark brown; abdomen with contrasting brownish to yellowish tergites and sternites, anal segment brown to dark brown. Head. Eyes bare. Temporals consisting of 21–23 setae including 13–14 inner and 8–9 outer verticals.Antenna and palp reduced. Antenna 410–430 μm long, 13-segmented; last flagellomere (Fig. 21, with 2 preceding segments) 180–190 μm long, entirely divided with forked rounded apex (Fig. 21); antennal groove beginning on segment 1 and reaching ultimate flagellomere;AR 0.53–0.61. Clypeus with about 40 setae placed in 6–7 rows. Palp (Fig. 22) reduced, 5-segmented, palpomeres 1–4 fused, palpomere 3 without sensilla clavata. Thorax. Acrostichals (Fig. 38) not reduced, consisting of 4–6 markedly curved setae 30–35 μm long, starting some distance from antepronotum and placed in 1 row; antepronotum (Fig. 29) with 17–21 lateral antepronotals; prealars 5– 5 in 1 row. Scutellum with 30–35 setae in 5–6 rows. Wing. Brachiolum with 5–6 setae. Distribution of setae on veins: R, 17–19; R1, 4–5; R2+3, 5–6; R4+5, 2–3, remaining veins bare. Squama with 10–13 uniserial setae. Legs. Sensilla chaetica present on tibia and tarsomeres ta1–ta5 of PI–PIII, tarsomere ta4 of PI–PII distinctly shorter than ta5, BR of all legs lower than 1 (0.60–0.95). Length (μm) and proportions of legs as in Table 2. Hypopygium in dorsal view (Fig. 31). Tergite IX semicircular, narrowing distally, distinctly truncate medially (clearly visible in lateral view, Fig. 30), about 130–140 setae present on tergite IX and anal point. Anal point drop-like; virga consisting of 4–5 long unequal thin spines; inferior volsella composed of 2 subequal well separated large lobes. Gonostylus (Fig. 34, lateral) elongated and narrowing distally, anterior area with 9–10 stout setae which are orally directed, crista dorsalis as low elongated ridge or absent; megaseta conspicuous and slightly bent inwards. Ecology. Up to 50 male and female adults of H. lugubris have been examined based on large material collected in cold limnocrenes and pristine peat bogs located in NE-France. Emergence of chironomid fauna reported from February to March revealed that H. lugubris largely dominates the other associated species which are mainly encountered during the first seasonal inundations of floodplains of some rivers and streams located in both the Vosges Region and upper basin of the River Doubs. Species encountered in the same localities with H. lugubris include: Bryophaenocladius aestivus (Brundin, 1947), B. nidorum (Edwards, 1929), Chaetocladius dentiforceps (Edwards, 1929), C. melaleucus (Meigen, 1818), C. perennis (Meigen, 1830), Limnophyes pentaplastus (Kieffer, 1921), L. difficilis Brundin, 1947, Metriocnemus albolineatus Meigen, 1818, M. eurynotus (Holmgren, 1883), M. fuscipes (Holmgren, 1818), M. hirticollis (Staeger, 1839). Distribution. Hydrobaenus lugubris is actually widespread in Europe and the Palaearctic Region (FRIES 1830, ZETTERSTEDT 1850, GIARD 1904, SAETHER 1976, SERRA- TOSIO & LAVILLE 1991, BARANOV 2011b). In France, this species was reported for the first time from North Western areas (GIARD 1904). Records cited here from NE-France, after MOUBAYED- BREIL (2017), are coming from Vosges Region and the upper basin of the Doubs River (altitude 400–600 m). This is considered to be the second record from France., Published as part of Moubayed-Breil, Joel & Baranov, Viktor, 2018, Taxonomic notes on the genus Hydrobaenus with description of H. simferopolus sp. nov. from Crimea (Diptera: Chironomidae), pp. 347-355 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 58 (2) on pages 351-352, DOI: 10.2478/aemnp-2018-0029, http://zenodo.org/record/4504794, {"references":["FRIES B. F. 1830: Beskrifnig ofver et nytt slagte, Hydrobaenus, horande till Tipulariae. Kungliga Svenska Vetenskaps-Akademiens Handlingar 1829: 176 - 187.","SAETHER O. A. 1976: Revision of Hydrobaenus, Trissocladius, Zalutschia, Paratrissocladius, and some related genera (Diptera, Chironomidae). Bulletin of the Fisheries Research Board of Canada 195: 1 - 287.","ZETTERSTEDT J. W. 1850: Diptera Scandinaviae. Disposita et descripta. Vol. 9. Officina Lundbergiana, Lund, pp. 3367 - 3710.","GIARD A. 1904: Quelques mots sur l'Hydrobaenus lugubris Fries (( Dipt.) Bulletin de la Societe Entomologique de France 9: 164 - 165.","BARANOV V. 2011 b: A preliminary annotated checklist of non-biting midges (Diptera, Chironomidae) of Ukraine. Ukrainska Entomofaunistyka 2: 7 - 24."]}

Hydrobaenus simferopolus sp. nov. (Figs 29, 35–37, 39–47) Type material. HOLOTYPE: ♁ (SIZK), ʻ UKRAINE: CRIMEA: leg. V. Baranov; Simferopol, Salgir River, 29.XI.2010, handnetʼ. PARATYPE: ♁ (JBMF) (mounted on the same slide), same locality as for holotype. Diagnosis. Based on some typical and specific characters found in the hypopygium of H. simferopolus sp. nov., in particular the singular shape of its phallapodeme, this new species apparently belongs to a separate group. This new species can be easily distinguished from all members of the genus by having: acrostichals not reduced, consisting of 5–6 long markedly curved setae; tergite IX without setae, dorsal margin sinuous; laterosternite IX with 7 setae on each side; anal point large, drop-shaped, proximal half with 12 setae, distal half bare and ending with a thumb-like apex; virga composed of 3 strong equal teeth; phallapodeme characteristic, terminating in 2 unusual and characteristic spirals; inferior volsella consisting of a large plain lobe ending with a hyaline and bare nose-like apex which is projecting downwards; gonocoxite with a swollen ventral lobe placed distally; gonostylus linearly elongated and slender, posterior part hyaline and bare; crista dorsalis large tooth-like, smooth apically and nearly hyaline, orally projecting and placed pre-apically close to the megaseta. Description. Male adult (n = 2; Figs 28, 35–37, 39–47). Medium sized Hydrobaenus species. Total length 3.50– 3.60 mm; wing length 2.45–2.50 mm. General colouration contrasting brown to dark brown. Thorax dark brown with blackish mesonotal stripes. Legs dark brown. Tergites I– VIII and anal segment dark brown; crista dorsalis distinctly hyaline to yellowish. Head. Eyes bare. Temporals consisting of 10–11 setae including 6–7 inner and 4 outer verticals. Palp 5-segmented, not reduced; length (in μm) of segments: 35, 45, 63, 136, 225; palpomere 3 (Fig. 35) with 3 sensilla clavata, palpomere 4 (Fig. 36) with 2 circular area of microtrichia placed proximally and distally. Clypeus with 6 setae in 2 rows. Thorax. Antepronotum with gaping lobes (Fig. 28), lateral antepronotals 3–4; acrostichals (Fig. 37) not reduced, composed of 6–7 distinctly pin-like setae starting some distance from antepronotum; dorsocentrals 10 in 1 row; prealars 5–7 in 1 row. Scutellum with 10 uniserial setae. Wing. Brachiolum with 1 seta. Distribution of setae on veins: R 4–5 placed on proximal part; remaining veins bare. Squama with 12–13 setae in 1–2 rows. Legs. Tarsomeres ta2–ta5 of PI are missing; sensilla chaetica present on tibia of PI and tarsomeres ta1–ta5 of PII–PIII. Length (μm) and proportions of legs as in Table 3. Hypopygium in dorsal, ventral and lateral view as in Figs 39, 40, and 47. Tergite IX broadly semi-circular, narrowing distally, dorsal margin sinuous (clearly visible in lateral view, Fig. 41); posterior area (Fig. 39) with 12–14 dorsal setae placed near the base of anal point (6–7 setae on each side). Laterosternite IX with 7 setae on each side. Anal point (Figs 39, 41–42) 75–85 μm long, 60 μm maximum width in median part; broad drop-shaped ending with thumb-like apex; distal part hyaline and lacking microtrichia. Virga (Figs 39, 44) 45–50 μm long, consisting of 3–4 equal pointed teeth, entirely fused except for distal part. Phallapodeme unusually shaped, characteristic, terminating in 2 characteristic spirals (Fig. 40). Gonocoxite 285–300 μm long, with rounded apex; ventral margin with large swollen lobe. Inferior volsella (dorsal, Figs 39, 43; lateral, Fig. 47) 100–105 μm long, consisting of 2 unequal elongated lobes, proximal one nearly plain larger and covered with setae, distal lobe nose-like, hyaline and bare. Gonostylus (Figs 45–46) 120–125 μm long, maximum width 50–55 μm, linearly elongated and slender, bearing distinct hyaline and bare posterior area; crista dorsalis strong tooth-like, hyaline with smooth and rounded apex, orally projecting and placed pre-apically close to megaseta. Taxonomic position. Male adult of H. simferopolus sp. nov. can be keyed near that of H. dentistylus based on the following resembling characters: lobes of antepronotum not in contact (Fig. 28); acrostichals not reduced; virga composed of 3 strong pointed teeth; inferior volsella bilobed; crista dorsalis large tooth-like, orally projecting and placed pre-apically. However, the newly described species can be easily distinguished by the following combination of characters: acrostichals consisting of 6–7 long pin-like setae; tergite IX without setae while densely covered with setae in H. dentistylus (Fig.2; MOUBAYED 1985: Fig. 1b); anal point large, drop-shaped, with a thumb-like apex (Figs 39, 42), while densely covered with setae in H. dentistylus (Figs 2, 4; Fig. 1b, MOUBAYED 1985) and differently figured in lateral view (Fig. 41 for H. simferopolus sp. nov., Fig. 3 for H. dentistylus); phallapodeme (Fig. 40) unusually shaped and terminating in 2 characteristic spirals; basal lobe of inferior volsella (Figs 39, 47) ending with a noselike lobe which is hyaline, bare and projecting downwards; gonostylus (Figs 45–46) linearly elongated and slender, while bulb-like and spherical in H. dentistylus (Figs 8–9; MOUBAYED 1985: Figs 1b, 2); crista dorsalis large toothlike, hyaline and smooth (Figs 45–46) while triangular with pointed apex and consistently chitinous in H. dentistylus. Etymology. The new species is named ‘ simferopolus ’ after the Ukrainian city of Simferopol where the type material was collected; adjective. Ecology. A typical rheophilic and oxybiontic species occurring in pristine section of streams and rivers. Species is active in winter, observed in a few swarms over water and on the grass at the near zero to sub-zero temperatures (from 5°C to -1°C). Specimens were active before the sunset in December and January. Distribution. Only known from the type locality which is situated in the Crimean Peninsula (Crimea, Ukraine), Salgir River, Gagarin City Park, Simferopol City., Published as part of Moubayed-Breil, Joel & Baranov, Viktor, 2018, Taxonomic notes on the genus Hydrobaenus with description of H. simferopolus sp. nov. from Crimea (Diptera: Chironomidae), pp. 347-355 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 58 (2) on pages 352-354, DOI: 10.2478/aemnp-2018-0029, http://zenodo.org/record/4504794, {"references":["MOUBAYED Z. 1985: Les Chironomidae (Diptera) du Liban. III. Hydrobaenus dentistylus n. sp. (Orthocladiinae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 121: 73 - 76."]}

Cricotopus sp Locality: Spain, Mallorca, Puigpunyant, between Ma-1041-11 and 12, 39.614133 oN, 2.553593 oE, 1♀, 12 February 2015, waterfall and stream edges near bridge., Published as part of Baranov, Viktor & Kvifte, Gunnar Mikalsen, 2016, NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract, pp. 11-14 in CHIRONOMUS Journal of Chironomidae Research 29 on page 12, DOI: 10.5324/cjcr.v0i29.1990, http://zenodo.org/record/7991027

Orthocladius (O.) maius Goetghebuer, 1942 Locality: Spain, Mallorca, Esporles, Torrent de San Vic, 39.670459 oN, 2.569193 oE, 2♂♂, 10 February 2015, stream edges. Distribution: Austria, Czech Republic, Germany, Great Britain, Italy, Spain, Sweden (Ashe and O’Connor 2012, Spies and Saether 2013). New to the Balearic Islands., Published as part of Baranov, Viktor & Kvifte, Gunnar Mikalsen, 2016, NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract, pp. 11-14 in CHIRONOMUS Journal of Chironomidae Research 29 on page 12, DOI: 10.5324/cjcr.v0i29.1990, http://zenodo.org/record/7991027, {"references":["Ashe, P. and O'Connor, J. P. 2012. A world catalogue of Chironomidae (Diptera). Part 2. Orthocladiinae. Irish Biogeographical Society, Dublin. xvi + 968 p.","Spies, M. and Saether, O. A. 2013. Fauna Europaea: Chironomidae. In Beuk, P. and Pape, T. (Eds.) Fauna Europaea: Diptera Nematocera. Fauna Europaea version 2.6, http: // www. fauna-eu. org [Accessed 25 March 2016]"]}

Paracladius conversus (Walker, 1856) Locality: Spain, Mallorca, Deía, loc. 2, 39.750554 oN, 2.642722 oE, 1♂, 08 February 2015, rock surface with sparse vegetation and some moisture. Distribution: Widespread in the Holarctic region (Ashe and O’Connor 2012, Spies and Saether 2013, Soriano et al 1997). New to the Balearic Islands., Published as part of Baranov, Viktor & Kvifte, Gunnar Mikalsen, 2016, NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract, pp. 11-14 in CHIRONOMUS Journal of Chironomidae Research 29 on page 12, DOI: 10.5324/cjcr.v0i29.1990, http://zenodo.org/record/7991027, {"references":["Ashe, P. and O'Connor, J. P. 2012. A world catalogue of Chironomidae (Diptera). Part 2. Orthocladiinae. Irish Biogeographical Society, Dublin. xvi + 968 p.","Spies, M. and Saether, O. A. 2013. Fauna Europaea: Chironomidae. In Beuk, P. and Pape, T. (Eds.) Fauna Europaea: Diptera Nematocera. Fauna Europaea version 2.6, http: // www. fauna-eu. org [Accessed 25 March 2016]","Soriano, O., Cobo, F., Rieradevall, M. Y. Prat, N. 1997. Lista faunistica y bibliografica de los quironomidos (Diptera, Chironomidae) de la Peninsula Iberica e Islas Baleares. Asociacion Espanola de Limnologia, Madrid. 210 p."]}

Bryophaenocladius sp. cf. scanicus (Brundin, 1947) sensu Langton and Pinder 2007 Locality: Spain, Mallorca, Puigpunyant, between Ma-1041-11 and 12, 39.614133 oN, 2.553593 oE, 1♂, 12 February 2015, waterfall and stream edges near bridge. The species recorded here is the same as the one illustrated by Langton and Pinder (2007), which appears to be quite different from the B. scanicus sensu stricto redescribed and illustrated in Du et al. (2011). Bryophaenocladius scanicus sensu stricto is listed as occurring in Spain (Soriano et al. 1997, Ashe and O’Connor 2012), but the species must be considered as new to the fauna of the Balearic Islands. However, further studies of Bryophaenocladius sp. cf. scanicus sensu Langton and Pinder 2007 is required to clarify its status and distribution., Published as part of Baranov, Viktor & Kvifte, Gunnar Mikalsen, 2016, NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract, pp. 11-14 in CHIRONOMUS Journal of Chironomidae Research 29 on page 12, DOI: 10.5324/cjcr.v0i29.1990, http://zenodo.org/record/7991027, {"references":["Brundin, L. 1947. Zur kenntnis der schwedischen Chironomiden. Almquist et Wiksells boktryckeri. 95 p.","Langton, P. and Pinder, L. C. V. 2007. Keys to the adult male Chironomidae of Britain and Ireland. - Freshwater Biological Association, Scientific Publication 64 (1 - 2), 239 + 168 p.","Du, J., Wang, X. H. and Saether, O. A. 2011. Redescriptions of species of Bryophaenocladius Thienemann, 1934 (Diptera: Chironomidae) described by Brundin (1947). - Zootaxa 2743: 40 - 48. DOI: http: // dx. doi. org / 10.11646 / % 25 x","Soriano, O., Cobo, F., Rieradevall, M. Y. Prat, N. 1997. Lista faunistica y bibliografica de los quironomidos (Diptera, Chironomidae) de la Peninsula Iberica e Islas Baleares. Asociacion Espanola de Limnologia, Madrid. 210 p.","Ashe, P. and O'Connor, J. P. 2012. A world catalogue of Chironomidae (Diptera). Part 2. Orthocladiinae. Irish Biogeographical Society, Dublin. xvi + 968 p."]}

Bryophaenocladius nidorum (Edwards, 1929) Localities: Spain, Mallorca, Campos, 1 km from city centre, 39.425695 oN, 2.957022 oE, 7♂♂, 06 February 2015, garden; Esporles, Torrent de San Vic, 39.670459 oN, 2.569193 oE, 1♂, 10 February 2015, stream edges; Esporles, Son Tria, 39.663051 oN, 2.573698 oE, 1♂, 09 February 2015, recreational area; Banyalbufar, 39.690612 oN, 2.525107 oE, 1♂, 09 February 2015, shaded ditch between Banyalbufar and Esporles. Distribution: Austria, Finland, France (Incl. Corsica), Germany, Great Britain, Mongolia, Norway, Novaya Zemlya (Russia), Netherlands, Romania, Russia (Nothern European part), Sweden, Switzerland (Ashe and O’Connor 2012, Spies and Saether 2013). New to Spain inclusive the Balearic Islands., Published as part of Baranov, Viktor & Kvifte, Gunnar Mikalsen, 2016, NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract, pp. 11-14 in CHIRONOMUS Journal of Chironomidae Research 29 on pages 11-12, DOI: 10.5324/cjcr.v0i29.1990, http://zenodo.org/record/7991027, {"references":["Ashe, P. and O'Connor, J. P. 2012. A world catalogue of Chironomidae (Diptera). Part 2. Orthocladiinae. Irish Biogeographical Society, Dublin. xvi + 968 p.","Spies, M. and Saether, O. A. 2013. Fauna Europaea: Chironomidae. In Beuk, P. and Pape, T. (Eds.) Fauna Europaea: Diptera Nematocera. Fauna Europaea version 2.6, http: // www. fauna-eu. org [Accessed 25 March 2016]"]}

Paraphaenocladius impensus (Walker, 1856) Locality: Spain, Mallorca, Puigpunyant, between Ma-1041-11 and 12, 39.614133 oN, 2.553593 oE, 1♂, 12 February 2015, waterfall and stream edges near bridge. Distribution: Widespread in the Holarctic region (Ashe and O’Connor 2012, Spies and Saether 2013, Soriano et al. 1997). New to the Balearic Islands., Published as part of Baranov, Viktor & Kvifte, Gunnar Mikalsen, 2016, NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract, pp. 11-14 in CHIRONOMUS Journal of Chironomidae Research 29 on page 12, DOI: 10.5324/cjcr.v0i29.1990, http://zenodo.org/record/7991027, {"references":["Ashe, P. and O'Connor, J. P. 2012. A world catalogue of Chironomidae (Diptera). Part 2. Orthocladiinae. Irish Biogeographical Society, Dublin. xvi + 968 p.","Spies, M. and Saether, O. A. 2013. Fauna Europaea: Chironomidae. In Beuk, P. and Pape, T. (Eds.) Fauna Europaea: Diptera Nematocera. Fauna Europaea version 2.6, http: // www. fauna-eu. org [Accessed 25 March 2016]","Soriano, O., Cobo, F., Rieradevall, M. Y. Prat, N. 1997. Lista faunistica y bibliografica de los quironomidos (Diptera, Chironomidae) de la Peninsula Iberica e Islas Baleares. Asociacion Espanola de Limnologia, Madrid. 210 p."]}

Smittia pratorum (Goetghebuer, 1927) Locality: Spain, Mallorca, Esporles, Torrent de San Vic, 39.670459 oN, 2.569193 oE, 1♂, 10 February 2015, stream edges; Cala Figuera, 39.330871 oN, 3.165476 oE, 2♂♂, 1♀, 10 February 2015, forest glade in front of hotel; Banyalbufar, 39.690612 oN, 2.525107 oE, 1♂, 09 February 2015, shaded ditch between Banyalbufar and Esporles. Distribution: Widely distributed in the Holarctic region; occurring in Oriental China, and possibly also in Argentina in the Neotropical region. In Spain, the species was previously recorded in the Sierra Nevada (Ashe and O’Connor 2012, Spies and Saether 2013). New to Balearic Islands (Ashe and O’Connor 2012, Spies and Saether 2013, Soriano et al. 1997). Remarks: The male collected at Banyalbufar exhibits a curious abnormality in the development of the anal point as the apex of the anal point is bifurcate and Y-shaped (Fig. 2). We have not found similar examples in the literature, but a wide variety of other developmental abnormalities resulting from parasites or gene or chromosomal mutations have been thoroughly documented (see Rempel 1940; Martin and Lee 2000)., Published as part of Baranov, Viktor & Kvifte, Gunnar Mikalsen, 2016, NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract, pp. 11-14 in CHIRONOMUS Journal of Chironomidae Research 29 on pages 12-13, DOI: 10.5324/cjcr.v0i29.1990, http://zenodo.org/record/7991027, {"references":["Ashe, P. and O'Connor, J. P. 2012. A world catalogue of Chironomidae (Diptera). Part 2. Orthocladiinae. Irish Biogeographical Society, Dublin. xvi + 968 p.","Spies, M. and Saether, O. A. 2013. Fauna Europaea: Chironomidae. In Beuk, P. and Pape, T. (Eds.) Fauna Europaea: Diptera Nematocera. Fauna Europaea version 2.6, http: // www. fauna-eu. org [Accessed 25 March 2016]","Soriano, O., Cobo, F., Rieradevall, M. Y. Prat, N. 1997. Lista faunistica y bibliografica de los quironomidos (Diptera, Chironomidae) de la Peninsula Iberica e Islas Baleares. Asociacion Espanola de Limnologia, Madrid. 210 p.","Rempel, J. G. 1940. Intersexuality in Chironomidae induced by nematode parasitism. - Journal of Experimental Zoology 84: 261 - 289.","Martin, J. and Lee, B. T. O. 2000. Sex determination in Chironomus and the Drosophila paradigm. In Hoffrichter, O. (Ed.) Late 20 th Century Research on Chironomidae. Shaker Verlag, Aachen, Germany, pp. 177 - 181."]}

Limnophyes minimus (Meigen, 1818) Localities: Spain, Mallorca, Puigpunyant, between Ma-1041-11 and 12, 39.614133 oN, 2.553593 oE, 10♂♂, 10♀♀, 12 February 2015, waterfall and stream edges near bridge; Esporles, Torrent de San Vic, 39.670459 oN, 2.569193 oE, 3♂♂, 1♀, 10 February 2015, stream edges; Banyalbufar, 39.690612 oN, 2.525107 oE, 3♂♂, 09 February 2015, shaded ditch between Banyalbufar and Esporles; Deía, loc. 1, 39.748072 oN, 2.643385 oE, 9♂♂, 08 February 2015, water-through and stream near hygropetric surface; Deía, loc. 2, 39.750554 oN, 2.642722 oE, 1♂, 08 February 2015, rock surface with sparse vegetation and some moisture; Deía, loc. 3, 39.752634 oN, 2.642415 oE, 1♂, 08 February 2015, half-dry streambed with rocks and bryophytes. Distribution: Cosmopolitan; previously recorded from Balearic isles (Ashe and O’Connor 2012, Malo and Garcìa-Avilès 1999, Soriano et al. 1997)., Published as part of Baranov, Viktor & Kvifte, Gunnar Mikalsen, 2016, NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract, pp. 11-14 in CHIRONOMUS Journal of Chironomidae Research 29 on page 12, DOI: 10.5324/cjcr.v0i29.1990, http://zenodo.org/record/7991027, {"references":["Ashe, P. and O'Connor, J. P. 2012. A world catalogue of Chironomidae (Diptera). Part 2. Orthocladiinae. Irish Biogeographical Society, Dublin. xvi + 968 p.","Soriano, O., Cobo, F., Rieradevall, M. Y. Prat, N. 1997. Lista faunistica y bibliografica de los quironomidos (Diptera, Chironomidae) de la Peninsula Iberica e Islas Baleares. Asociacion Espanola de Limnologia, Madrid. 210 p."]}

Baranov, Viktor, Kvifte, Gunnar Mikalsen (2016): NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract. CHIRONOMUS Journal of Chironomidae Research 29: 11-14, DOI: 10.5324/cjcr.v0i29.1990

Bryophaenocladius inconstans (Brundin, 1947) Locality: Spain, Mallorca, Banyalbufar, 39.690612 oN, 2.525107 oE, 2♂♂, 09 February 2015, shaded ditch between Banyalbufar and Esporles. Distribution: Finland, Germany, Great Britain, Italy, Norway, Romania, Russia (East Siberia), Sweden, Spain (Sierra Nevada) (Ashe and O’Connor 2012, Spies and Saether 2013, Casas et al., 2013). New to Balearic Islands., Published as part of Baranov, Viktor & Kvifte, Gunnar Mikalsen, 2016, NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract, pp. 11-14 in CHIRONOMUS Journal of Chironomidae Research 29 on page 12, DOI: 10.5324/cjcr.v0i29.1990, http://zenodo.org/record/7991027, {"references":["Brundin, L. 1947. Zur kenntnis der schwedischen Chironomiden. Almquist et Wiksells boktryckeri. 95 p.","Ashe, P. and O'Connor, J. P. 2012. A world catalogue of Chironomidae (Diptera). Part 2. Orthocladiinae. Irish Biogeographical Society, Dublin. xvi + 968 p.","Spies, M. and Saether, O. A. 2013. Fauna Europaea: Chironomidae. In Beuk, P. and Pape, T. (Eds.) Fauna Europaea: Diptera Nematocera. Fauna Europaea version 2.6, http: // www. fauna-eu. org [Accessed 25 March 2016]","The present study is based on a small sample of chironomids collected as by-catch in a study targeted at moth flies (Diptera, Psychodidae) and em- phasis was thus on madicolous habitats (Kvifte et al. 2016). Chironomidae from such habitats have received a lot less attention than those of larger aquatic habitats such as lakes and rivers (Przhibroro and Baranov, 2014). The records of six new species records for the Balearic Islands from such a small sample highlights that madicolous habitats in the Mediterranean accommodate a large and still comparatively poorly understood diversity of Chironomidae. Casas, J. J., Langton, P. H. and Fuentes-Rodriguez, F. 2013. Los Quironomidos (Diptera, Chironomidae). In Ruano, F., Tierno de Figueroa, M. y Tinaut, A. (Eds). Los insectos de Sierra Nevada. 200 anos de historia. Vol 2. Universidad de Granada & Consejeria de Medio Ambiente, pp. 194 - 215."]}

Corynoneura sp. Locality: Spain, Mallorca, Puigpunyant, between Ma-1041-11 and 12, 39.614133 oN, 2.553593 oE, 1♂, 12 February 2015, waterfall and stream edges near bridge., Published as part of Baranov, Viktor & Kvifte, Gunnar Mikalsen, 2016, NEW RECORDS OF NON-BITING MIDGES (DIPTERA: CHIRONOMIDAE, ORTHOCLADIINAE) FROM MALLORCA, SPAIN Abstract, pp. 11-14 in CHIRONOMUS Journal of Chironomidae Research 29 on page 12, DOI: 10.5324/cjcr.v0i29.1990, http://zenodo.org/record/7991027

Palaeoboreochlus inornatus Baranov et Andersen, sp. n. (Figs 1���6) Type material. Holotype male, Ukraine: Pugach open-pit mine, Klesov, Rovno amber, Late Eocene (SIZK: K- 27535). Etymology: From Latin, inornatus, meaning unadorned, referring to the lack of megaseta or strong spines on the gonostylus. Diagnosis. See generic diagnosis. Description. Male (n = 1). Total length 1.88 mm. Wing length 1.51 mm. Total length / wing length 1.24. Wing length / length of profemur 4.30. Head (Fig. 2). Eye without dorsomedial prolongation. At least 2 strong orbitals, frontals not visible. Length / width of palpomeres 2���5 (in ��m): 68/20, 92/26, 80/24, 100/18. Antenna (Fig. 4). AR = 0.66, length of flagellomeres (in ��m): 44, 36, 36, 36, 36, 40, 40, 40, 40, 40, 40, 40, 216, 92. Thorax. With at least 14 strong dorsocentrals. Wing (Fig. 5). VR = 1.13. Costal extension 132 ��m long. Distance between arculus and crossvein MCu 192 ��m. Legs. Spur of fore tibia about 28 ��m long; spurs of mid tibia about 44 ��m long; spurs of hind tibia about 56 ��m long. Width at apex of fore tibia 28 ��m, of mid tibia 30 ��m, of hind tibia 36 ��m. Comb of hind tibia composed of several setae, longest about 30 ��m long. Length and proportion of leg segments as in Table 1. Hypopygium (Figs 3, 6). Tergite IX with rounded posterior margin, without anal point. Gonocoxite about 110 ��m long. Gonostylus 80 ��m long, 36 ��m wide at base, apically conical, tapering to blunt apex., Published as part of Baranov, Viktor, Andersen, Trond & Perkovsky, Evgeny, 2014, A new genus of Podonominae (Diptera: Chironomidae) in Late Eocene Rovno amber from Ukraine, pp. 581-586 in Zootaxa 3794 (4) on pages 583-584, DOI: 10.11646/zootaxa.3794.4.9, http://zenodo.org/record/4915023

Palaeoboreochlus Baranov et Andersen, gen. n. Type species. Palaeoboreochlus inornatus Baranov et Andersen, sp. n. (present designation by monotypy). Etymology. Named after the related genus Boreochlus, using the Greek palaiois, ancient, as a prefix. Diagnostic characters. The new genus groups with Boreochlus in having a simple gonostylus; flagellum with 14 flagellomeres; a long costal extension reaching the wing tip; no pulvilli; no anal point; crossvein MCu strongly retracted; and reniform, naked eyes. It can be separated from Boreochlus by the comparatively long flagellomere 14, slightly shorter than half the length of flagellomere 13; palpus with palpomere five about 5 times as long as wide; slender, smoothly rounded gonocoxite lacking volsella; and gonostylus with weak, rounded basal heel with few setae on low pedestals, and conical, apical portion without megaseta. Generic diagnosis. Male. Small species, wing length about 1.5 mm. Antenna. Flagellum with 14 flagellomeres. Flagellomere 14 slightly less than half the length of flagellomere 13, flagellomere 13 slightly longer than the length of fagellomeres 7���11 together. Plume well developed. Apex of flagellomere 14 apparently with several sensilla. Head. Eye reniform, naked. Orbitals present. Palp with five palpomeres, length ratio between palpomeres 2���5 as: 0.20: 0.27: 0.24: 0.29. Thorax. Scutal setation well developed, details poorly observable. Wing. Membrane with coarse punctuation, setae not observed. Costa extended, ending near wing tip. MCu weak, strongly retracted, situated at about 1/3 the distance between arculus and RM. FCu slightly distal to RM. Legs. LR 1 0.68. Tibial spurs with basal denticles. Tibial comb of several setae. Pulvilli absent. Hypopygium. Abdominal segment IX well developed; without anal point. Gonocoxite slender, smoothly rounded, with setae along inner margin. Gonostylus simple, with weak, rounded basal heel with few setae on low pedestals, and conical apical portion, with distinct y-seta at base, without megaseta., Published as part of Baranov, Viktor, Andersen, Trond & Perkovsky, Evgeny, 2014, A new genus of Podonominae (Diptera: Chironomidae) in Late Eocene Rovno amber from Ukraine, pp. 581-586 in Zootaxa 3794 (4) on pages 582-583, DOI: 10.11646/zootaxa.3794.4.9, http://zenodo.org/record/4915023

The genus Palaeoboreochlus Baranov et Andersen, n. gen. is erected based on P. inornatus Baranov et Andersen, n. sp. described from a male found in Late Eocene Rovno amber. The new genus groups with Boreochlus Edwards in the tribe "Boreochlini". publishedVersion

Micropsectra uva sp. nov. Type material. Holotype, male (in DIZP): CROATIA, Plitvice Lakes National Park, springs of Bijela Rijeka (44 �� 50 ���N 15 �� 33 ���E, 720 m a.s.l.), 25 July 2007, pyramid emergence trap, leg. M. Ivković. Paratypes: 6 males (4 in DIZP, 2 in MSIZ), same data as holotype. Derivation of the name. From Latin, meaning 'bunch of fruits', in reference to the shape of the hypopygial median volsella; noun in apposition. Diagnosis. Stem of median volsella short, straight or slightly curved at tip posterolaterally turned, blunt apex reaching midpoint of superior volsella and half length of inferior volsella at most; spoon-shaped lamellae arranged in 3���4 rows, placed on distal half of stem and posteromedially directed; stem/spoon-shaped lamellae length ratio c. 2.5. Inferior volsella slightly curved at base, narrowest in mid length, with swollen and posteromedially turned distal half, transverse protrusion very slight if present. Description. Adult male (n = 7, unless otherwise stated) Colouration. Eyes and tibial combs black, other parts of the body yellowish green, with antenna, scutal stripes and sternum slightly darker. Head. Antenna with 13 flagellomeres, plume well developed, AR 0.60���0.70 (0.64, n = 3). Frontal tubercles absent. Length of palpomeres 2���5 (��m): 48���64 (56, n = 3), 155���181 (164, n = 3), 128���155 (137, n = 3), 227���271 (244, n = 3). Clypeus with 10���18 (13) setae. Thoracic chaetotaxy. Ac 20���25, reaching antepronotum; Dc 8���10; Pa 2���3; Scts 7���8 in row. Wing. Slightly variable in shape as shown in Figure 1. Length (arculus���tip) 1.99���2.15 mm (2.07 mm). Sc, R 2 + 3, M, short proximal part of M 1 + 2, RM, 1 / 3 proximal section of Cu bare, remaining veins with macrotrichia; membrane except basal part covered with macrotrichia. R 2 + 3 weak. FCu placed under RM-R connection. VR Cu 1.09���1.14 (1.12, n = 2). Veins ending as follows (in order from base to tip): An (under FCu), Sc, Cu 1, R 1, R 2 + 3, M 3 + 4, R 4 + 5, M 1 + 2. Anal lobe reduced. Legs. Fore tibia with straight, c. 10���20 ��m long spur. Combs of mid and hind tibiae contiguous, placed on 50��� 65 % circumference of tibial apices, each comb with single somewhat longer spur-like tooth. Basitarsus of mid leg bearing 1���3 hook-shaped sensilla chaetica. For lengths of leg segments and leg ratios see Table 1. Hypopygium (Fig. 2, 3 D). Anal tergite with bands of V-type, fused at elevated hump at base of anal point, bearing 3���4 median tergite setae, lateral teeth and basilateral processes well developed. Anal point wide at base, with long crests separated distally by large knob, apex blunt, subrectangular, microtrichia-free area surrounding base of anal point extensive. Strong Micropsectra -seta sensu S��wedal (1982) placed on prominent conical tubercle at base of superior volsella. Superior volsella round, bearing 2���3 setae on anteromedian margin (proximal seta weaker) and 6���9 dorsal setae. Digitus moderately long, almost reaching margin of superior volsella, apically blunt or acute. Stem of median volsella short, c. 45���50 ��m, straight or slightly curved at tip posterolaterally turned, blunt apex reaching midpoint of superior volsella and half length of inferior volsella at most; c. 20 spoon-shaped lamellae arranged in 3���4 rows, placed on distal half of stem, posteromedially directed; stem/spoon-shaped lamellae length ratio c. 2.5. Inferior volsella with slight knee-like curve at base, narrowest in mid length, with swollen and posteromedially turned distal half, transverse protrusion usually absent or very slight. Gonostylus c. 145���170 ��m long, broadest near mid-length, tapering to rounded apex. Micropsectra atrofasciata group; compiled from Stur & Ekrem (2006), Rossaro et al. (2009), Rossaro (pers. comm.), material presently examined. character/species M. pallidula M. schrankelae M. sofiae M. uva Fig. 3 A Fig. 3 B Fig. 3 C Fig. 3 D Discussion. We compared Micropsectra uva with three species of the Micropsectra atrofasciata group: M. pallidula (Meigen), M. schrankelae Stur et Ekrem and M. sofiae Stur et Ekrem, which have so far been recognized as close relatives (Stur & Ekrem 2006, Rossaro et al. 2009). With respect to the variability known from the species compared, only a few differences in metric/meristic characters are sufficiently distinct and diagnostically significant. Among these M. pallidula has a relatively long wing, and high AR, LR ratio values, whereas M. uva is the smallest species, but has a relatively stout and long gonostylus. In contrast to rather low usefulness of most metric characters, the shape and arrangement of hypopygial volsellae are the essential characters in diagnostics of the four species compared. We present them in Figure 3 and Table 2. These four species dwell in similar habitats. The adult males of Micropsectra uva were collected in the spring (rheocrene) of the Bijela Rijeka. The stream, together with the Crna Rijeka, forms a common watercourse, the Matica river, a water-replenishment for the Plitvice Lakes. Water temperature in the spring of the Bijela Rijeka at an altitude of 720 m varies between 7.3���7.8 ��C, has a slightly acid to weakly alkaline pH (6.9���7.8) (Ivković et al. 2012). The two closest species compared, Micropsectra schrankelae and M. sofiae, are known from similar habitats, being associated with persistent low temperature in cold-water habitats at high elevations, mainly in springs (cf. Stur & Ekrem 2006, Rossaro et al. 2009); Micropsectra pallidula, well-known throughout Europe, inhabits springs and streams, as well as cold-water lakes and small standing water bodies. We presume that female specimens found in the sample with males of Micropsectra uva belong to the same species. Unfortunately, due to lack of pupae and/or pharate specimens, association of the adults would be uncertain, thus the female could not be described., Published as part of Gi��ka, Wojciech, Zakrzewska, Marta, Baranov, Viktor A. & Dominiak, Patrycja, 2013, Diagnostic clues for identification of selected species of the Micropsectra atrofasciata group, with description of M. uva sp. nov. from Croatia (Diptera: Chironomidae: Tanytarsini), pp. 288-294 in Zootaxa 3702 (3) on pages 289-292, DOI: 10.11646/zootaxa.3702.3.6, http://zenodo.org/record/224007, {"references":["Sawedal, L. (1982) Taxonomy, morphology, phylogenetic relationships and distribution of Micropsectra Kieffer, 1909 (Diptera: Chironomidae). Entomologica scandinavica, 13, 371 - 400. http: // dx. doi. org / 10.1163 / 187631282 x 00228","Stur, E. & Ekrem, T. (2006) A revision of West Palaearctic species of the Micropsectra atrofasciata species group (Diptera: Chironomidae). Zoological Journal of the Linnean Society, 146, 165 - 225. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2006.00198. x","Rossaro, B., Lencioni, V. & Marziali, L. (2009) Micropsectra sofiae Stur & Ekrem, 2006 (Diptera, Chironomidae) redescribed. Bollettino di Zoologia agraria e di Bachicoltura, Ser. II, 41, 129 - 139.","Ivkovic, M., Micetic Stankovic, V. & Mihaljevic, Z. (2012) Emergence patterns and microhabitat preference of aquatic dance flies (Empididae; Clinocerinae and Hemerodromiinae) on a longitudinal gradient of barrage lake system. Limnologica, 42, 43 - 49. http: // dx. doi. org / 10.1016 / j. limno. 2011.07.003"]}

Micropsectra uva sp. nov. Type material. Holotype, male (in DIZP): CROATIA, Plitvice Lakes National Park, springs of Bijela Rijeka (44 º 50 ’N 15 º 33 ’E, 720 m a.s.l.), 25 July 2007, pyramid emergence trap, leg. M. Ivković. Paratypes: 6 males (4 in DIZP, 2 in MSIZ), same data as holotype. Derivation of the name. From Latin, meaning 'bunch of fruits', in reference to the shape of the hypopygial median volsella; noun in apposition. Diagnosis. Stem of median volsella short, straight or slightly curved at tip posterolaterally turned, blunt apex reaching midpoint of superior volsella and half length of inferior volsella at most; spoon-shaped lamellae arranged in 3–4 rows, placed on distal half of stem and posteromedially directed; stem/spoon-shaped lamellae length ratio c. 2.5. Inferior volsella slightly curved at base, narrowest in mid length, with swollen and posteromedially turned distal half, transverse protrusion very slight if present. Description. Adult male (n = 7, unless otherwise stated) Colouration. Eyes and tibial combs black, other parts of the body yellowish green, with antenna, scutal stripes and sternum slightly darker. Head. Antenna with 13 flagellomeres, plume well developed, AR 0.60–0.70 (0.64, n = 3). Frontal tubercles absent. Length of palpomeres 2–5 (Μm): 48–64 (56, n = 3), 155–181 (164, n = 3), 128–155 (137, n = 3), 227–271 (244, n = 3). Clypeus with 10–18 (13) setae. Thoracic chaetotaxy. Ac 20–25, reaching antepronotum; Dc 8–10; Pa 2–3; Scts 7–8 in row. Wing. Slightly variable in shape as shown in Figure 1. Length (arculus–tip) 1.99–2.15 mm (2.07 mm). Sc, R 2 + 3, M, short proximal part of M 1 + 2, RM, 1 / 3 proximal section of Cu bare, remaining veins with macrotrichia; membrane except basal part covered with macrotrichia. R 2 + 3 weak. FCu placed under RM-R connection. VR Cu 1.09–1.14 (1.12, n = 2). Veins ending as follows (in order from base to tip): An (under FCu), Sc, Cu 1, R 1, R 2 + 3, M 3 + 4, R 4 + 5, M 1 + 2. Anal lobe reduced. Legs. Fore tibia with straight, c. 10–20 Μm long spur. Combs of mid and hind tibiae contiguous, placed on 50– 65 % circumference of tibial apices, each comb with single somewhat longer spur-like tooth. Basitarsus of mid leg bearing 1–3 hook-shaped sensilla chaetica. For lengths of leg segments and leg ratios see Table 1. Hypopygium (Fig. 2, 3 D). Anal tergite with bands of V-type, fused at elevated hump at base of anal point, bearing 3–4 median tergite setae, lateral teeth and basilateral processes well developed. Anal point wide at base, with long crests separated distally by large knob, apex blunt, subrectangular, microtrichia-free area surrounding base of anal point extensive. Strong Micropsectra -seta sensu Säwedal (1982) placed on prominent conical tubercle at base of superior volsella. Superior volsella round, bearing 2–3 setae on anteromedian margin (proximal seta weaker) and 6–9 dorsal setae. Digitus moderately long, almost reaching margin of superior volsella, apically blunt or acute. Stem of median volsella short, c. 45–50 Μm, straight or slightly curved at tip posterolaterally turned, blunt apex reaching midpoint of superior volsella and half length of inferior volsella at most; c. 20 spoon-shaped lamellae arranged in 3–4 rows, placed on distal half of stem, posteromedially directed; stem/spoon-shaped lamellae length ratio c. 2.5. Inferior volsella with slight knee-like curve at base, narrowest in mid length, with swollen and posteromedially turned distal half, transverse protrusion usually absent or very slight. Gonostylus c. 145–170 Μm long, broadest near mid-length, tapering to rounded apex. Micropsectra atrofasciata group; compiled from Stur & Ekrem (2006), Rossaro et al. (2009), Rossaro (pers. comm.), material presently examined. character/species M. pallidula M. schrankelae M. sofiae M. uva Fig. 3 A Fig. 3 B Fig. 3 C Fig. 3 D Discussion. We compared Micropsectra uva with three species of the Micropsectra atrofasciata group: M. pallidula (Meigen), M. schrankelae Stur et Ekrem and M. sofiae Stur et Ekrem, which have so far been recognized as close relatives (Stur & Ekrem 2006, Rossaro et al. 2009). With respect to the variability known from the species compared, only a few differences in metric/meristic characters are sufficiently distinct and diagnostically significant. Among these M. pallidula has a relatively long wing, and high AR, LR ratio values, whereas M. uva is the smallest species, but has a relatively stout and long gonostylus. In contrast to rather low usefulness of most metric characters, the shape and arrangement of hypopygial volsellae are the essential characters in diagnostics of the four species compared. We present them in Figure 3 and Table 2. These four species dwell in similar habitats. The adult males of Micropsectra uva were collected in the spring (rheocrene) of the Bijela Rijeka. The stream, together with the Crna Rijeka, forms a common watercourse, the Matica river, a water-replenishment for the Plitvice Lakes. Water temperature in the spring of the Bijela Rijeka at an altitude of 720 m varies between 7.3–7.8 ºC, has a slightly acid to weakly alkaline pH (6.9–7.8) (Ivković et al. 2012). The two closest species compared, Micropsectra schrankelae and M. sofiae, are known from similar habitats, being associated with persistent low temperature in cold-water habitats at high elevations, mainly in springs (cf. Stur & Ekrem 2006, Rossaro et al. 2009); Micropsectra pallidula, well-known throughout Europe, inhabits springs and streams, as well as cold-water lakes and small standing water bodies. We presume that female specimens found in the sample with males of Micropsectra uva belong to the same species. Unfortunately, due to lack of pupae and/or pharate specimens, association of the adults would be uncertain, thus the female could not be described.

Giłka, Wojciech, Zakrzewska, Marta, Baranov, Viktor A., Dominiak, Patrycja (2013): Diagnostic clues for identification of selected species of the Micropsectra atrofasciata group, with description of M. uva sp. nov. from Croatia (Diptera: Chironomidae: Tanytarsini). Zootaxa 3702 (3): 288-294, DOI: http://dx.doi.org/10.11646/zootaxa.3702.3.6