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Leucobaeta smithi sp. n. (Figs. 4���6, 18, 24) Holotype. ♂, ��� Mozambique, 630m, Manica Province, Chimanimani National Reserve, Moribane Forest, Ndzou Camp, (Moist Forest), 19��44���01.4���S, 33��20���15.1���E, 3���5.VIII.2018, Actinic Light Trap, Laszlo, G., Miles, W., Vetina, A. leg., ANHRT:2018.30���, unique number: ANHRTUK 00047856, gen. slide No.: LGNA 961 (ANHRT). Paratypes (11 ♂, 5 ♀ in total). Mozambique . 1 ♂, Maputo Special Reserve, West Gate (Sand Forest), 22m, 26��30���14.2���S, 32��42���59.6���E, 13���15.ii.2018, Actinic Light Trap, L��szl��, G., Mulvaney, J., Smith, L. leg., ANHRT:2018.2, unique number: ANHRTUK 00027539, gen. slide No.: LGNA 962 (ANHRT). Ethiopia. 1 ♂, Abyssinia, Harar, 18.vii.1939, R. E. Ellison, R.E. Ellison coll., B.M. 1960-550, QR code label with unique number: NHMUK 014046237, gen. slide No.: NHMUK 010316556 (NHMUK). Tanzania. 1 ♂, Empakani Crater, Ngorongoro Highlands, 2278m, 02��54���55���S, 35��51���23���E, 26���28.vii.2012, Light Trap, leg. Smith, R. & Takano, H., ANHRT:2017.6, unique number: ANHRTUK 00073440, gen. slide No.: LGNA 963 (ANHRT); 1 ♀, Malaria Institute Amani, P. 604, 2/1963, G. Pringle Coll., B.M. 1966-281, QR code label with unique number: NHMUK 014046236, gen. slide No.: NHMUK 010316555; 1 ♀, same site, 3/1964, QR code label with unique number: NHMUK 010917505 (NHMUK). Kenya. 1 ♂, Central Province, Thika District, 8 km SW Thika, Karamaini Estate, 1550m, 01��02���7.46���S, 36��59���4.72���E, 25.ix.2012, leg. A.J. Kingston (coll. A.J. Kingston, Albrighton, UK). Zimbabwe. 1 ♂, Khami, nr. Bulawayo, S. Rhodesia, III.1956, Nat. Mus. S. Rhodesia, Brit. Mus. 1962-143, QR code label with unique number: NHMUK 014046235, gen. slide No.: NHMUK 010316554 (NHMUK); 1 ♂, Manicaland, Chipinge Highlands, Chirinda Forest, 1170m, 20��24���6.04���S, 32��41���9.62���E, 19���20.xi.2017, leg. A. Kingston, K. Larsen & A. Cipolla (coll. A.J. Kingston, Albrighton, UK). South Africa. 1 ♂, Natal, Weenen, IX���X.1925, H.P. Thomasset, Pres. By Imp. Bur. Ent. Brit. Mus. 1927-387., QR code label with unique number: BMNH(E) 1403484, gen. slide No.: Arctiidae 553; 1 ♂, Durban, Natal, Bred 8.viii.1919, E.E. Platt, 1919-89, QR code label with unique number: BMNH(E) 1403484, gen. slide No.: Arctiidae 554 (NHMUK); 2 ♂, KwaZulu Natal, Karkloof, Rockwood Forest Lodge, 26���29.ii.2020, 1290m, leg. A.J. Kingston, K. Larsen & A. Cipolla (coll. A.J. Kingston, Albrighton, UK); 1 ♂, 2 ♀, Mpumalanga, Barberton Nature Reserve, Berghuisi, 2.5 km S Barberton, 900���1200m, 25��48���S, 31��02���E, 7���9.xi.2015, leg. Fiebig, Schellhorn & Stadie (coll. R. Fiebig, Rossleben, Germany); 1 ♀, same site, date and collectors, 1100m (coll. D. Stadie, Lutherstadt-Eisleben, Germany). Diagnosis. Leucobaeta smithi sp. n. is externally reminiscent of its sibling L. semialba, but easily distinguished by its considerably larger size (forewing length of the new species is 8.5���10.0 mm, that of L. semialba is 5.5���8.0 mm), the more elongate forewing and the larger subapical patch surrounded by a more extensive whitish area. Despite the fairly similar external habitus, the male genitalia of the two species show substantial differences expressed by the distally much narrower uncus, the considerably shorter and broader tegumen, and the narrower valva of the new species. The most salient difference in the male genitalia of the two species is expressed by the configuration of the costal process which is an excessively enlarged, arrowhead-shaped lobe with its apex pointing caudad in L. smithi, while the costal process is much smaller, finger-like, apically rounded, directing ventrad in L. semialba. Further remarkable differences between the two species are the poorly developed, membranous transtillae, the much smaller fultura inferior (juxta) and the markedly shorter vinculum of L. smithi compared to those of its West African congener. The phallus of the two species are also very different: it is gently S-shaped without a carina process in the new species, while it is straight with a long, robust, claw-like carina process in L. semialba; in addition, the vesica of L. smithi bears a large, spike-like cornutus associated with an elongate sclerotized area, whereas that of L. semialba lacks cornuti and other sclerotization. Compared to those of L. semialba, the female genitalia of the new species are markedly larger, with a somewhat shorter and thicker apophyses posteriores, a considerably longer, thicker, apically club-shaped apophyses anteriores (those of L. semialba are pointed apically) and a shorter 8 th tergite with parallel proximal and distal margins (it is medially constricted in the allied species). In addition, the new species has a shorter, less sclerotized antrum, an approximately 20% longer ductus bursae with larger lateral protrusion, and an unmodified cervix bursae, compared to those of L. semialba. The corpus bursae of L. smithi is markedly larger than that of its West African congener with 20% longer tubular distal section and somewhat smaller signum bursae, lacking an appendix bursae. Description. Adult. (Figs. 4���6). Forewing length 8.5���10.0 mm. Antenna fasciculate in males, filiform in females. Head relatively large, labial palp length 1.2 times the diameter of eye, moderately thick, porrect, dark brown; frons pale brown posteriorly, off-white anteriorly, vertex whitish-grey; compound eyes relatively large, globular. Thorax, collar, tegulae and abdomen bright white. Intraspecific variability and sexual dimorphism negligible, expressed only by the slightly darker hindwing of females. Forewing moderately elongate, rather quadrangular, apically rounded. Basal and antemedial area bright white with two, pale grey costal dashes, the proximal one larger than the distal one. Medial area dark brownish grey, inner margin straight, oblique, sharply separated from white basal area. Postmedial and tornal areas brownish grey, fading to pale grey distally, dorsal half of postmedial and terminal area creamy-white, apically with an extensive, dark ovoidal blotch consisting of a diffuse ochreous patch encompassing a small black amorphous area anteriorly, and a dark brown round macule adjoining apex posteriorly. Transverse lines deleted except for the diffuse, very narrow, evenly arcuate, whitish postmedial line, and the very narrow, interrupted pale grey terminal line. Cilia long, darkening from pale grey to dark brownish-grey towards tornus. Hindwing of male off-white in basal three-quarters, pale greyish near termen, that of female dark grey in distal three-quarters with off-white basal area; cilia off white. Underside of forewing dark grey in the dorsal two-thirds, pale greyish-white in the ventral third; hindwing underside pale grey, somewhat darker near termen; pattern undetectable. Male genitalia. (Fig. 18). Uncus medium long, broad at base, tapered in basal third, slightly dilated medially, then abruptly tapered, distal third very narrow, apically pointed, spike-like. Tegumen short with relatively wide arms fused at middle of tegumen. Valva relatively short, narrow, apically evenly rounded, membranous. Valva costa straight, subbasally with a large, heavily sclerotized arrowhead-shaped lobe pointing caudad, conjunct with sacculus bearing a short, rounded subventral process pointing distally; distal two-thirds of costa weakly sclerotized. Sacculus half as long as valva, rather broad, heavily sclerotized, fused with costal process by a narrow, sclerotized ventral band. Transtillae poorly developed, represented by membranous ribbon without medial plate. Fultura inferior (juxta) very small, sclerotized, broad V-shaped. Vinculum conspicuously short, rounded triangular. Phallus tubular, short and thin, gently S-shaped, medially moderately dilated, coecum penis short and rounded, heavily sclerotized, apex straight, without carina process; vesica with a robust, spike-like cornutus and an elongate, moderately sclerotized area. Female genitalia. (Fig. 24). Ovipositor short, conical, papillae anales trapezoidal, apically with a short protracted process, covered in long, dense setae, apophyses posteriores short, basally broad, apically pointed, apophyses anteriores ca. twice as long as posterior ones, narrow, slightly curved, apically club-shaped. Eighth tergite heavily sclerotized, short, distal and proximal margins straight, parallel. Ostium bursae narrow, slit-like, antrum very short, poorly sclerotized, somewhat tapered anteriorly. Ductus bursae membranous, moderately long, five times as long as antrum, medially dilated forming a lateral broad, rounded triangular protrusion on right side. Cervix bursae unmodified, membranous. Distal third of corpus bursae tubular, as long as ductus bursae, slightly dilated proximally, proximal two-thirds globular, without appendix bursae. Signum bursae represented by a relatively small, round finely scobinated area. Genetic divergence. Pairwise distance between L. smithi and its Madagascan sibling L. malagassa sp. n. is 3.5��� 4.1%, with 1.7% intraspecific divergence between the southern and central Mozambican populations. In comparison to the West African L. semialba, the pairwise distance is 7.6���7.7%, the significant genetic divergence concurs with the high level of morphological differences. Distribution (Fig. 26). The new species is widely distributed in Eastern and Southern Africa with records from Ethiopia, Kenya, Tanzania, Mozambique, Zimbabwe and South Africa. Etymology. The new species is dedicated to Mr Richard Smith, founder and director of the African Natural History Research Trust acknowledging his prominent support of entomological research in Sub-Saharan Africa., Published as part of L��szl��, Gyula M., 2022, On the taxonomy of Negeta semialba Hampson, 1918 with descriptions of two new Leucobaeta L��szl��, Ronkay & Witt, 2010 species (Lepidoptera, Nolidae, Nolinae), pp. 56-68 in Zootaxa 5091 (1) on pages 63-65, DOI: 10.11646/zootaxa.5091.1.2, http://zenodo.org/record/5840401

Leucobaeta malagassa sp. n. (Figs. 7���9, 19, 25) Holotype. ♂, ��� Madagascar N, Montagne d���Ambre NP, 3 km SO of Joffreville, S 12��30���50/ E 49��11���02,05./ 06.11.2018 LF, 970 m, leg. R. Fiebig & D. Stadie ���, gen. slide No.: RF 699.2021 (coll. R. Fiebig, to be deposited later in ZSM). Paratypes (6 ♂, 15 ♀ in total). Madagascar . 2 ♂, 4 ♀, Montagne d���Ambre, 4 km NW of Joffreville, 12��27���50���S, 49��13���29���E, 480m, 4.xi.2018, Light Trap, leg. R. Fiebig & D. Stadie; 2 ♂, 3 ♀, Ankarana NP, Ankarana Lodge, 12��57���40���S, 49��08���58���E, 500m, 9���11.xi.2018, Light Trap, leg. R. Fiebig & D. Stadie, gen. slide Nos.: RF 697.2021 (♀), RF 698.2021 (♀); 1 ♂, 4 ♀, Ranomafana NP, Setam, Lodge, 21��14���58.5������S, 47��25���36.9������E, 913m, 9���13.iv.2018, Light Trap, leg. D. Stadie & R. Fiebig (coll. R. Fiebig); 2 ♀, 3km N Vorondolo / Ranamofana, 21��13.319������S, 47��22.069������E, 10.iv.2018, LT, 1218m, leg. D. Stadie & R. Fiebig; 1 ♂, Montagne d`Ambre, NW of Joffreville, 12��27���50������S, 49��13���29������E, 04.xi.2018, LT, 480m, leg. D. Stadie & R. Fiebig (coll. D. Stadie); 1 ♀, Fianarantsoa pr., 20 km NE Ranohira, 26���29.i.2014, Sakamaningy vill. Env., 828m, 22��30.457���S, 45��29.761���E, M. Tryzna leg., ANHRT:2018.4, unique number: ANHRTUK 00073441, gen. slide No.: LGNA 964; 1 ♀, Mahafaly Plateau, Antanambao, near Bezaha village, 14.i.2013, M. Tryzna leg., ANHRT:2018.4, unique number: ANHRTUK 00214293 (ANHRT). Diagnosis. The external appearance of Leucobaeta malagassa sp. n. is nearly identical to that of L. smithi, but distinguishable by its noticeably narrower, oblique, dark medial fascia which is paler, more diffuse, less contrasting in the females and somewhat darker in the males, the hindwing of which is also considerably darker than in L. smithi. In the male genitalia, compared to L. smithi, the new species has an approximately 30% smaller genital capsule, a considerably shorter uncus with a much broader basal part, a broader, dorso-apically more angular valva, and a markedly shorter, distally curved, more or less trapezoidal costal lobe (it is straight distally, arrowhead-shaped in L. smithi), with slightly longer rounded subventral process arising closer to the ventral margin than in its congener. The phallus of the new species is approximately 20% shorter and narrower than that of L. smithi, bearing slightly shorter cornutus of the vesica. The configuration of the female genitalia of the two species are nearly identical, the only noticeable differences are the less swollen distal section of the ductus bursae with a considerably shorter lateral protrusion, the approximately 50% shorter and narrower tubular distal part of the corpus bursae and the slightly smaller signum bursae of L. malagassa compared to those of its continental congener. Description. Adult. (Figs. 7���9). Forewing length 7.5���10.0 mm. Antenna fasciculate in males, filiform in females. Head relatively large, labial palp length 1.2 times the diameter of eye, moderately thick, porrect, dark brown; frons pale brown posteriorly, off-white anteriorly, vertex whitish grey; compound eyes relatively large, globular. Thorax, collar, tegulae and abdomen bright white. Intraspecific variability and sexual dimorphism negligible, expressed only by the somewhat larger size and slightly paler, more diffuse forewing medial fascia of females. Forewing elongate, rather quadrangular, apically rounded. Basal and antemedial area bright white with two, pale grey costal dashes, proximal one of which larger than distal one. Medial area dark brownish-grey, inner margin straight, oblique, sharply separated from white basal area. Postmedial and tornal areas dark brownish-grey, fading to paler grey distally, dorsal half of postmedial and terminal area creamy-white, subapically with a large, dark ovoidal blotch consisting of a dark diffuse-ochreous anterior patch followed by a narrow black area fading to grey distally, and a dark brown round posterior macule adjoining apex. Transverse lines deleted except for the narrow, diffuse, interrupted pale grey terminal line. Cilia long, grey, slightly darkened towards tornus. Hindwing of male pale grey, somewhat lighter in basal quarter, that of female dark grey with a small off-white basal area; cilia off-white. Underside of forewing dark grey in the dorsal twothirds, pale greyish-white in the ventral third; hindwing underside grey, somewhat darker near the termen; pattern undetectable. Male genitalia. (Fig. 19). Uncus relatively short, very broad at base, abruptly tapered in basal third forming a trapezoidal base, gradually tapered in medial third, distal third very narrow, spine-like, apically pointed. Tegumen short with wide arms fused in distal three-quarter. Valva short, narrow, dorso-apically right-angled, ventrally evenly rounded, membranous. Valva costa straight, basally with a large, heavily sclerotized more or less trapezoidal lobe with short, curved pointed apex projecting caudad, costal lobe conjunct with sacculus by narrow sclerotized band, bearing a short, rounded subventral process pointing caudad; distal two-thirds of costa weakly sclerotized, sparsely setose. Sacculus approximately half as long as valva, rather broad, heavily sclerotized, fused with costal process by a narrow ventral band. Transtillae reduced to a membranous ribbon without medial plate. Fultura inferior very small, sclerotized, somewhat U-shaped. Vinculum conspicuously short, medially with a short, rounded anterior projection. Phallus tubular, short and thin, straight, medially slightly dilated, coecum penis short and rounded, heavily sclerotized, apex straight, without carina process; vesica with robust, spike-like cornutus and an elongate moderately sclerotized area. Female genitalia. (Fig. 25). Ovipositor short, conical, papillae anales trapezoidal, apically with a short protracted process, covered in long, dense setae. Apophyses posteriores short, basally broad, apically pointed, apophyses anteriores ca. 20% longer than posterior ones, narrow, gently curved, apically slightly dilated. Eighth tergite heavily sclerotized, short, with straight, parallel distal and proximal margins. Ostium bursae narrow, slitlike, antrum very short, poorly sclerotized, somewhat tapered anteriorly. Ductus bursae short, five times as long as antrum, membranous, medial section slightly dilated with a short lateral, moderately sclerotized protrusion on right side. Cervix bursae unmodified, membranous. Distal quarter of corpus bursae tubular, narrow, gradually dilated proximally, proximal three-quarters ovoidal, without appendix bursae. Signum bursae represented by small, ovoidal scobination. Genetic divergence. Pairwise distances between L. malagassa and L. smithi is 3.5���4.1% and between L. malagassa and L. semialba, is 7.1%. As only a single specimen sequenced successfully, the intraspecific divergence of L. malagassa could not be calculated. Distribution (Fig. 26). Leucobaeta malagassa is a species endemic to Madagascar with sporadic records from the island. Etymology. The specific name is dedicated to the Malagasy people of Madagascar, referring to the distribution of the new species., Published as part of L��szl��, Gyula M., 2022, On the taxonomy of Negeta semialba Hampson, 1918 with descriptions of two new Leucobaeta L��szl��, Ronkay & Witt, 2010 species (Lepidoptera, Nolidae, Nolinae), pp. 56-68 in Zootaxa 5091 (1) on pages 65-66, DOI: 10.11646/zootaxa.5091.1.2, http://zenodo.org/record/5840401

Leucobaeta semialba (Hampson, 1918) comb. n. (Figs. 1���3, 17, 23) Negeta semialba Hampson, 1918, Novitates Zoologicae 25(1): 204. Type locality: [Ghana] Gold Coast, Bibianaha. Holotype, ♀ (NHMUK). Type material examined. Holotype, ♀, red ring label ���Type H.T.��� / ��� Gold Coast. Bibianaha. 700 ft. XII.1911. H.G.F. Spurrell. 1912-275��� / with handwritten: ��� Negeta semialba type ♀. Hmpsn.���, QR code label with unique number: NHMUK 010917504 (NHMUK). (Fig. 1) Additional material examined. Liberia. 1 ♂, Nimba Mts., ENNR, Nimba county (Cellcom road), 1000��� 1100m, 7��32���45.88���N, 8��31���21.04���W, 16���28.xii.2018, Cold Cathode UV Light Trap (8W), S��fi��n, Sz., Simonics, G. leg., ANHRT:2018.43, unique number: ANHRTUK 00073977, gen. slide No.: LGNA 955; 1 ♂, same site, 2���14.xii.2017, Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33, unique number: ANHRTUK 00142092; 3 ♂, 2♀, Mount Nimba, Cellcom Road, ENNR, Nimba County, 750���1100m, 7��32���35.89���N, 8��31���20.63���W, 14���28.xii.2017, Cold Cathode Light Bucket Trap, S��fi��n, Sz., Simonics, G. leg., ANHRT:2017.33, unique numbers: ANHRTUK 00194493-00194496, gen. slide No.: LGNA 1293 (♀); 8 ♂, 1 ♀, Nimba County, Nimba Mts, ENNR, Cellcom road, 750m, 7��33���3.78���N, 8��31���46.49���W, 16���28.xii.2018, Cold Cathode UV Light Trap (8 W), S��fi��n, Sz., Simonics, G. leg., ANHRT:2018.43, unique numbers: ANHRTUK 00073958, 00142173, 00142169-00142172, 00219576-00219578 (ANHRT). Nigeria. 1 ♂, Oyo State, Gambari Forest, 6.viii.1977, M.A. Cornes, QR code label with unique number: NHMUK 014046238, gen. slide No.: NHMUK 010316557 (NHMUK). Diagnosis. In comparison with the Oriental Leucobaeta hemiphaea, L. semialba is considerably smaller in size (forewing length 5.5���8.0 mm, that of hemiphaea is 8.5���9.5 mm), the border of the whitish basal-antemedial area of the forewing is straight, while it is slightly curved in its ventral quarter in the Oriental species. The postmedial fascia consists of a very narrow cream line in L. semialba, while it is marked by a row of three to four black dots anteromedially in L. hemiphaea. In addition, the apical area of the forewing has a large amorphous dark brown patch in the Afrotropical species, whereas it is greyish white, without a distinct dark marking in L. hemiphaea. In the male genitalia, compared to the type species of the genus, L. semialba has a somewhat longer, basally narrower uncus, and wider, reverse U-shaped medial recess of the tegumen (which is reverse V-shaped in L. hemiphaea). The valva is considerably narrower and longer with a broadly rounded apex in L. semialba, while it is rectangular and distally truncate in the Oriental species. The transtillae are medially fused with a shield-like medial plate in L. semialba, whereas they are connected by a membranous ribbon in L. hemiphaea. The costal process of L. semialba is markedly wider at the base and considerably longer than in L. hemiphaea forming a finger-like process outreaching the ventral margin of the valva, lacking a short, elongate medial crest which is present in the Oriental species. The phallus of L. semialba is armed with a large, robust, curved, claw-like carina, whereas L. hemiphaea lacks a carinal process, but its vesica bears a pair of large, thumbtack-shaped cornuti, which is absent in the Afrotropical species. Comparing the female genitalia of the Afrotropical and Oriental species, the former has a conical ovipositor, with apically protracted papillae anales, while it is quadrangular and apically rounded, respectively, in L. hemiphaea. The posterior apophyses are longer, while the anterior ones are shorter in L. semialba than in its Oriental congener. The eighth tergite is considerably longer and medially more constricted and the anterior margin of the eighth sternite is evenly curved in both sides in the Afrotropical species, while the eighth sternite bears a pair of short and narrow, slightly conical, apically rounded anterior processes in L. hemiphaea. Further differences are expressed by the markedly shorter and less sclerotized ductus bursae, the much smaller appendix bursae and the presence of signum bursae in L. semialba. Description. Adult. (Figs. 1���3). Forewing length 5.5���8.0 mm. Antenna fasciculate in male, filiform in female. Head relatively large, labial palp medium long, rather thick, porrect, pale brown; frons pale brown posteriorly, off-white anteriorly, vertex niveous; compound eye relatively large, globular. Thorax, collar, tegulae and abdomen bright white. Intraspecific variability limited. Sexual dimorphism negligible, expressed only by the somewhat darker hindwing of females. Forewing relatively short and narrow, rather quadrangular, apically rounded. Basal and antemedial area bright white with a pair of elongate, pale grey costal dashes. Medial area dark brownish-grey, isolated abruptly from basal area by straight, oblique border, postmedial and tornal areas brownish grey, fading somewhat to pale grey towards tornus; dorsal half of postmedial area off-white, with a large round, dark brownish-grey apical patch. Transverse lines poorly visible except for the fine, evenly arcuate, postmedial line well-defined by creamy scales and the very narrow, continuous pale brownish-grey terminal line. Cilia long, darkening from pale grey to dark brown towards tornus. Hindwing of male off-white in basal half, gradually darkened distally, pale greyish-brown near termen, that of female pale greyish-brown with a small off-white basal area; cilia pale yellowish-brown. Forewing underside dark graphite-grey in dorsal two-thirds, pale greyish-white in ventral third; hindwing underside pale grey, somewhat darker near termen; pattern undetectable. Male genitalia. (Fig. 17). Uncus medium long, as long as vinculum, broad at base, abruptly tapered medially, gradually tapered distally, pointed apically. Tegumen medium long with narrow arms, medially widely open, arms forming a reverse U-shape. Valva moderately long and broad, as long as tegumen, apically evenly rounded, membranous. Valva costa almost straight, in basal half with a proximally smooth, distally sparsely setose triangular plate serving as a base for well-developed costal process; distal half of costa with narrow marginal sclerotization. Costal process long, medially arched, finger-like with rounded apex outreaching the ventral margin of valva. Sacculus short, broad at base, tapered distally, weakly sclerotized, sparsely setose, conjunct with costal process. Transtillae broad at base, tapered medially and strongly dilated apically, arms fused medially, forming a moderately large, proximally concave, distally rounded, shield-like plate. Fultura inferior (juxta) relatively small, heavily sclerotized, narrow V-shaped. Vinculum rather long, broad at base, abruptly tapered medially into conspicuously narrow, apically rounded, finger-like process (saccus). Phallus tubular, short and thin, straight, medially slightly dilated; coecum penis short and rounded, heavily sclerotized, apex armed with a conspicuously long, robust, medially curved, apically pointed claw-shaped carina process; vesica without cornuti or scobination. Female genitalia. (Fig. 23). Ovipositor short, conical, papillae anales triangular, apically with a short protracted process, covered in fine setae, apophyses short and thin, apically pointed, posterior ones twice as long as anterior ones. Eighth tergite weakly sclerotized, short, medially constricted. Ostium bursae narrow, more or less round, antrum heavily sclerotized, relatively short, distal margin deeply concave, lateral margins gently arched, proximal margin rounded with a moderately sclerotized, short conical antero-lateral protrusion on right side. Ductus bursae short, membranous, twice as long as sclerotized antrum, medially slightly dilated. Cervix bursae with a short, rounded, sclerotized protuberance on right side. Distal third of corpus bursae conical, proximal part ovoidal, with semi-globular appendix bursae fused with corpus bursae antero-laterally. Signum bursae represented by a small, rounded, fine scobination. Genetic divergence. Pairwise distance between L. semialba and the Madagascan endemic L. malagassa sp. n. is 7.1% and between L. semialba and the southeast African L. smithi sp. n. is 7.6���7.7%. Distribution (Fig. 26). This rare species was described from a single female specimen collected in southern Ghana. An additional male specimen was found in the NHMUK collection from Gambari forest, South Nigeria. A short series of specimens was collected on recent ANHRT expeditions at a medium-high elevation in the Nimba Mountains, Liberia., Published as part of L��szl��, Gyula M., 2022, On the taxonomy of Negeta semialba Hampson, 1918 with descriptions of two new Leucobaeta L��szl��, Ronkay & Witt, 2010 species (Lepidoptera, Nolidae, Nolinae), pp. 56-68 in Zootaxa 5091 (1) on pages 58-63, DOI: 10.11646/zootaxa.5091.1.2, http://zenodo.org/record/5840401, {"references":["Hampson, G. F. (1918) Descriptions of new genera and species of Amatidae, Lithosidae and Noctuidae. Novitates Zoologicae, 25 (1), 93 - 217."]}

Angustopila Jochum, Slapnik & P��ll-Gergely, 2014; Jochum et al., 2014: 410: 26. Type species: Systenostoma tamlod Panha & Burch, 1999, by original designation., Published as part of P��ll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, Andr��s, ��rstan, Aydin, Szab��, ��bel, D��nyi, L��szl�� & Schilthuizen, Menno, 2022, The world's tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae), pp. 62-78 in Contributions To Zoology 91 on page 66, DOI: 10.1163/18759866-BjA10025, http://zenodo.org/record/6147303, {"references":["Jochum, A., Slapnik, R., Kampschulte, M., Martels, G., Heneka, M. & Pall-Gergely, B. (2014) A review of the microgastropod genus Systenostoma Bavay & Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae). ZooKeys, 410, 23 - 40."]}

Angustopila psammion P��ll-Gergely, Anker & Vermeulen, n. sp. urn:lsid:zoobank.org:act:EAD52AC1-985A-48E7-8AF9-4C2629C416BF Type material: Vietnam, Quang Ninh Province, Ha Long Bay, Cap La Cave, 20��51.793���N, 107��13.541���E, soil deposit fallen through roof in pristine cave, vegetation outside cave tall and woody, leg. J.J. Vermeulen & K. Anker, 07 March 2018, holotype (HNHM 104886); CUMZ 7436 / 2 paratypes; SMF 365052 / 2 paratypes; VNMN / 2 paratypes; ZRC. MOL.23218/ 2 paratypes, jjV 17633/ 409 paratypes. Diagnosis: An Angustopila species with a depressed-globular shell with dome-shaped spire, thick spiral striae, kidney-shaped aperture with single parietal denticle not reaching parietal callus. Description: Shell tiny, off-white, depressed-globular form with domed spire; body whorl widest in standard apertural view; protoconch with 1.5 whorls and a minute and very uneven pattern of polygonal, rather sharp-crested ridges locally forming minute, conical peaks where ridges split (the structure also reminiscent of cross-sectional trabecular bone), no spiral striation discernible; teleoconch ornamented by some weak radial growth lines and much stronger, ropelike, equidistantly-arranged spiral striae (ca. 14���15 on body whorl from standard apertural view); the 3.25���3.5 whorls are separated by a deep suture; whorls slightly shouldered; aperture oblique to shell axis in lateral view; umbilicus relatively wide, less than one third of shell width; aperture kidney-shaped with strongly concave parietal side; sinulus wide, weakly separated due to weak parietal tooth; peristome expanded, not reflected; parietal callus strongly protruding, but in line with curvature of penultimate whorl and beyond aperture edge (profile) in lateral view (fig. 3C), detached from penultimate whorl; parietal tooth weak, low, short, rounded, does not reach peristome. Soft anatomy unknown. Measurements (in mm): SW = 0.6���0.68, SH = 0.46���0.57 (n = 24). Differential diagnosis: Angustopila pallgergelyi Dumrongrojwattana, Chuenit & Wongkamhaeng, 2021 is similar in shell and aperture shape, but is larger, has a prominent palatal tooth (absent in Angustopila psammion n. sp.)and a stronger parietal tooth. Angustopila coprologos n. sp. is slightly larger, has a rough shell surface, and has an additional subcolumellar tooth and two palatal teeth. Etymology: The specific epithet (��������������) means a grain of sand in Greek and is used as a noun in apposition. Distribution: This new species is known only from the type locality, Cap La Cave, Ha Long Bay, Quang Ninh Province, Vietnam. Habitat: Angustopila psammion n. sp. was found in large numbers in small sediment deposits along the walls of a dry cave (Cap La Cave), in complete darkness. We assume that the sediment had fallen in through crevices in the rock, because it contains a species-rich assemblage of bleached, opaque shells of surface-dwelling terrestrial gastropods. Shells of Angustopila psammion n. sp., however, look fresh and are probably autochthonous., Published as part of P��ll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, Andr��s, ��rstan, Aydin, Szab��, ��bel, D��nyi, L��szl�� & Schilthuizen, Menno, 2022, The world's tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae), pp. 62-78 in Contributions To Zoology 91 on pages 69-71, DOI: 10.1163/18759866-BjA10025, http://zenodo.org/record/6147303, {"references":["Dumrongrojwattana, P., Chuenit, S. & Wongkamhaeng, K. (2021) A new species of the world's smallest cave snail of the genus Angustopila Jochum, Slapnik & Pall- Gergely in Jochum, et al., 2014 (Gastropoda: Hypselostomatidae) from eastern Thailand. Raffles Bulletin of Zoology, 69, 102 - 108."]}

Meganola perfusca (Hampson, 1911) Nola perfusca Hampson, 1911, Annals and Magazine of Natural History (8) 8: 398. Type locality: S. Nigeria, Lagos. Syntypes, ♂, ♀ (NHMUK). Remark. The taxonomy of this species has not been clarified in Hacker et al. (2012) as none of the syntypes were examined and dissected. The author of the present paper was also unable to examine the genitalia of the species and thus the morphology of the male genitalia of M. perfusca remains unknown. The female genitalia of a specimen from Ivory Coast illustrated in Hacker et al. (2012) is inadequate for identification. Distribution. Nigeria, Ghana, Burkina Faso and Ivory Coast (Hacker et al. 2012)., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 180, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hampson, G. F. (1911) Descriptions of new genera and species of Syntomidae, Arctiadae, Agaristidae and Noctuidae. Annals and Magazine of Natural History, Series 8, 8 (46), 393 - 445. https: // doi. org / 10.1080 / 00222931108693053","Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Meganola smithi sp. n. (Figs 1���3, 57, 88) Holotype. ♂, Ivory Coast, 1171m, Mt. Tonkoui Peak, 07��27���15.2���N, 07��38���12.5���W, 1-8.XI.2015, Light Trap, Aristophanous, M., Moretto, P., Ruzzier, E. leg., ANHRT:2017.16, unique number: ANHRTUK 00009093, slide No.: LGNA 242 ♂ (ANHRT). Paratypes. Ivory Coast. 1 ♂, with the same data as the holotype, unique number: ANHRTUK 00009094. Liberia. 4 ♂, 1 ♀, 1165m, Nimba Mts camp, ENNR, Nimba county, 7��31���45���N, 8��31���37���W, 03-13.xii.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33, unique numbers: ANHRTUK 00019910 - 00019913, 00145314, slide Nos.: LGNA 496 ♂, LGNA 497 ♂, LGNA 498 ♀; 1 ♂, 1 ♀, 1000���1100m, Nimba Mts., ENNR, Nimba county (Cellcom road), 7��32���45.9���N, 8��31���21���W, 27.iii��� 04.iv.2017, Light Trap (250W blended bulb) & Cold Cathode UV light bucket trap (8W), S��fi��n, Sz., Simonics, G. leg., ANHRT:2017.36, unique numbers: ANHRTUK 00056451, 00056444, slide No.: LGNA 525 ♂; 1 ♂, same locality and collectors, but collected at 12���16.iii.2017, unique number: ANHRTUK 00022987; 6 ♂, 750m, Nimba Mts., ENNR, Cellcom road, 7��33���3.8���N, 8��31���46.5���W, 10���12.iii.2017, Light Trap (250W blended bulb) & Cold Cathode UV light bucket trap (8W), S��fi��n, Sz., Simonics, G. leg., ANHRT:2017.36, unique numbers: ANHRTUK 00058565, 00058566, 00058571, 00058572, 00058567, 00058568, slide No.: LGNA 798 ♂; 1 ♂, 1♀, 1000���1100m, Nimba Mts., ENNR, Nimba county (Cellcom road), 7��32���45.88���N, 8��31���21.04���W, 02���14.xii. 2017, Cold Cathode Light Bucket, Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33, unique numbers: ANHRTUK 00019947, 00019948, slide No.: LGNA 499 ♀; 1 ♀, 865m, Lofa County, 8��07���10���N, 9��57���11���W, 24���29.xi.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G. & Smith, L. leg. ANHRT:2017.33, unique number: ANHRTUK 00034351, slide No.: LGNA 523 ♀. Guinea . 4 ♂, 1500m, Nimba Mts, SMFG concession area (Societ�� des Mines de Fer de Guin��e) 600 Forest, 7��39���49.93���N, 8��22���22.19���W, 21��� 30.viii.2017, Cold Cathode UV Light Trap (8W), S��fi��n, Sz., Simonics, G. leg., ANHRT:2017.36, unique numbers: ANHRTUK00059223, 00058585 - 00058587, slide Nos: LGNA 796, LGNA 797 (all ♂); 1 ♂, 1400���1752m, Guine�� Foresti��re Monts Nimba UNESCO World Heritage Site, Mont Richard Molard camp and ridge (High-Altitude Grassland & Forest) 07��36���19���N, 08��25���30���W, 1���7.VII.2019, 250W Blended Light Trap, Derozi��r, V., Miles, W., S��fi��n, S. leg., ANHRT: 2019.11, unique number: ANHRTUK 00143347 (ANHRT)., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 152, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542

Meganola endoscota (Hampson, 1914) (Figs 4���7, 58, 89) Nola endoscota Hampson, 1914, Catalogue of the Lepidoptera Phalaenae in the British Museum, Supplement 1: 407. Type locality: [Ghana] Gold Coast, Bibianaha. Holotype, ♀ (NHMUK). = Meganola endoscota undosaria Hacker, 2012, Esperiana 17: 256, syn. n. = Meganola kaduna Hacker, 2012, Esperiana 17: 290, syn. n. = Meganola peksi Hacker, 2012, Esperiana 17: 295 (Hacker 2014) = Meganola mabiriaria Hacker, 2012, Esperiana 17: 291, syn. n. = Meganola simplicifacta Hacker, 2012, Esperiana 17: 294, syn. n. Type material examined. Holotype. ♀, red ring ���Type H. T.��� label; ��� Gold Coast. Bibianaha. 24���26.X.1911. H.G.F. Spurrell. 1911-413���, with handwritten: ��� Nola endoscota type ♀ Hmpsn.���, unique number: NHMUK010598804, Slide No.: NHMUK010315151 (prep. by Gy.M. L��szl��) (NHMUK). Additional material examined. Cameroon. 1 ♂, 612m, Central Region, Nkoteng, On Sanaga River, Nkoteng Forest, 04��33���34.2���N, 11��59���37.6���E, 24���28.x.2018, LepiLED Light Trap, S��fi��n, Sz., Simonics, G. leg. ANHRT:2018.36, slide No.: LGNA 888♂, 1 ♂, same data, but collected by Cold Cathode UV Light Trap, slide No.: LGNA 972♂. Guinea . 1 ♂, 435m, Geipa Camp, Foret de Diecke, 7��26���7.06���N, 8��50���47.87���W, 05���14.iv.2019, Light Trap, Blended Bulb (250W), S��fi��n, Sz., Koivogui, S. leg. ANHRT:2019.7, slide No.: LGNA 969♂. Liberia . 2 ♂, 750m, Nimba County, Nimba Mts., ENNR, Cellcom road, 7��33���3.78���N, 8��31���46.49���W, 16���28.xii.2018, Cold Cathode UV Light Trap (8 W), S��fi��n, Sz., Simonics, G. leg. ANHRT:2018.43, slide Nos: LGNA 970♂, LGNA 971♂ (ANHRT) Ghana. 1 ♂, 1 ♀, Bunso Arboretum, 6��15���58.03���N, 0�� 27���45.72���W, 13���14.IX.2010, leg. Sz. S��fi��n (HNHM). Remark. The taxonomy of M. endoscota (Hampson, 1914) has not been satisfactorily clarified by Hacker et al. (2012), having failed to illustrate the copulatory organ of the female holotype and figuring instead, an externally similar female specimen from Ivory Coast. Despite not examining the genital morphology of the primary type of the species, Hacker described a new subspecies of M. endoscota from Kivu Province, D.R. Congo (M. endoscota undosaria Hacker, 2012). The author of this present paper had the opportunity to dissect the female holotype of M. endoscota housed in the NHMUK, the genitalia of which is illustrated here for the first time (Fig. 89). As there are no considerable differences in the external habitus or the female genitalia of the two subspecies, M. endoscota undosaria is considered here as a mere synonym of M. endoscota. In the same work, Hacker described three new species based exclusively on male specimens (M. peksi, M. simplicifacta and M. mabiriaria Hacker, 2012) despite their undoubted similarities with M. endoscota, namely their markedly short, brownish grey forewings with the conspicuously dark brown median area and the oblique, slightly dentate postmedial line. A fourth species, M. kaduna Hacker, 2012 also reminiscent of M. endoscota, shares the same, rather characteristic male genital morphology with the above-mentioned three taxa. A female paratype of M. kaduna illustrated by Hacker et al. (2012: 290) has proved to be a misidentification of Meganola cirrhographa Hacker & Hoppe, 2012 (cf. p. 288 in Hacker et al. (2012)). Subsequently, Hacker (2014: 143) figured a male specimen identified as M. endoscota undosaria from Nyungwe Forest (Rwanda), without illustrating its genitalia, unfortunately rendering this identification doubtful. Hacker (2014) himself synonymised his M. peksi with M. kaduna, admitting the lack of distinctive features to separate the two species. A series of male specimens in the ANHRT collection reminiscent of M. endoscota recently collected in Cameroon, Guinea and Liberia display identical male genitalia characters (fig. 58) to Hacker���s taxa discussed above (cf. Hacker et al. 2012: 290, 291, 294, 295), namely the conspicuously narrow, medially constricted valva, the long and robust, curved harpe and the short but conspicuous, heavily sclerotized crest-like ventral carinal process of the aedeagus. From the comparative study of genital morphology based on a dozen specimens from various West African localities, it has been possible to conclude that M. kaduna, M. mabiriaria, M. peksi and M. simplicifacta are conspecific and represent the hitherto unknown male of M. endoscota; the four former species are here synonymised with the latter. It is also worth noting that this species has a much wider distribution than had been proposed earlier (Hacker et al. 2012; Hacker 2014). Distribution. Ghana, Ivory Coast, D.R. Congo, Uganda (Hacker et al. 2012) and Rwanda (Hacker 2014). The specimens from Cameroon, Guinea and Liberia represent new country records., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on pages 157-158, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hampson, G. F. (1914) Catalogue of the Amatidae and Arctiadae (Nolinae and Lithosianae) in the Collection of the British Museum. Catalogue of the Lepidoptera Phalaenae in the British Museum, London, Supplement 1, 1 - 858.","Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612.","Hacker, H. H. (2014) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae) Supplement 1. Esperiana, 19, 121 - 179."]}

Meganola xantholeuca Hacker, 2012 (Figs 29���30, 97) Meganola xantholeuca Hacker, 2012, Esperiana 17: 498. Type locality: Ivory Coast, Danane. Holotype, ♂ (ZSM). Material examined. Ivory Coast. 2 ♀, 1171m, Mt. Tonkoui Peak, 07��27���15.2���N, 07��38���12.5���W, 1���8.XI.2015, Light Trap, Aristophanous, M., Moretto, P. & Ruzzier, E. leg., ANHRT:2017.16, slide Nos: LGNA 86 ♀, LGNA 253 ♀. Liberia . 1 ♀, 883m, Lofa County, Wologizi Mts, Ridge Camp 2, 8��07���20.79���N, 9��56���50.75���W, 22��� 31.xi.2018, Cold Cathode UV Light Trap (8W), S��fi��n, Sz., Simonics, G. leg. ANHRT:2018.43, slide No.: LGNA 977 (ANHRT). Remark. M. xantholeuca was considered to be a sister species of M. cretacea (Hampson, 1914) by Hacker et al. (2012). The taxonomy of M. cretacea has been clarified by L��szl�� (2020) and the species has been transferred to the genus Ezishnola L��szl��, Ronkay & Witt, 2010. The former presumption namely that M. xantholeuca is related to E. cretacea was due to a misidentification of the latter taxon by Hacker et al. (2012) (L��szl�� 2020). Distribution. Ivory Coast and Nigeria (Hacker et al. 2012). New record for Liberia., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 167, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612.","Hampson, G. F. (1914) Catalogue of the Amatidae and Arctiadae (Nolinae and Lithosianae) in the Collection of the British Museum. Catalogue of the Lepidoptera Phalaenae in the British Museum, London, Supplement 1, 1 - 858.","Laszlo, Gy. M. (2020) On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae). Ecologica Montenegrina, 30, 82 - 92. https: // doi. org / 10.37828 / em. 2020.30.8","Laszlo, Gy. M., Ronkay, G. & Witt, T. (2010) Contribution to the Nolinae (Lepidoptera, Noctuidae) fauna of North Thailand. Esperiana, 15, 7 - 125."]}

Meganola allardi Hacker, 2012 Meganola allardi Hacker, 2012, Esperiana 17: 328. Type locality: Ivory Coast, Abidjan. Holotype, ♂ (RMCA). Remark. The species is known only from the male holotype specimen. Distribution. Ivory Coast (Hacker et al. 2012)., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 158, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Meganola canocolorata Hacker, 2012 (Figs 43, 102) Meganola canocolorata Hacker, 2012, Esperiana 17: 440. Type locality: Ivory Coast, Katiola. Holotype, ♀ (ZSM). Material examined: Ivory Coast. 1 ♀, 174m Tai NP. Tai Research Station (SRET), 05��50���00���N, 07��20���32.0W, 27.III.���16.IV.2017, Actinic Light Trap, Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P. leg. ANHRT:2017.25, slide No.: LGNA 981 ♀. Liberia. 1 ♀, 140m, Krahn-Bassa Reserve, Sinoe County, Juboe River, 7.5km South West Pellokon Town, 5��39���4���N, 8��39���4���W, 14���21.i.2018, Cold Cathode UV Bucket Light Trap, Geiser, M., S��fi��n, Sz., Simonics, G. leg. ANHRT:2017.33, slide No.: LGNA 985 ♀. Distribution. The species was known only from its holotype collected at Katiola, Ivory Coast (Hacker et al. 2012). The specimen from Liberia represents a new country record., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 173, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Meganola subchionea sp. n. (Figs 34, 76) Holotype. ♂, Ivory Coast, 174m, Tai NP, Tai Research Station, 05��49���59.8���N, 07��20���32.0���W, 14���23.XI.2015, Light Trap, Aristophanous, M., Moretto, P., Ruzzier, E. leg., ANHRT:2017.16, unique number: ANHRTUK 00009099, slide No.: LGNA 139 ♂ (ANHRT). Diagnosis. The new species belongs to the M. chionea species-group, being most closely related to M. chionea (Hampson, 1911) with its pale greyish white forewing ground colour and the broad, almost straight, obtuse pale grey medial band of the forewing. As the species-group comprises several externally indistinguishable species, examination of the genitalia is required for reliable identification. In the male genitalia, M. subchionea differs clearly from M. chionea by its somewhat broader uncus and considerably narrower harpe and the vesica armed with a pair of short but heavily sclerotized, thorn-like cornuti (the vesica of M. chionea bears a peculiar elongate, crest-like, dentate cornutus). The female is unknown., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 168, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hampson, G. F. (1911) Descriptions of new genera and species of Syntomidae, Arctiadae, Agaristidae and Noctuidae. Annals and Magazine of Natural History, Series 8, 8 (46), 393 - 445. https: // doi. org / 10.1080 / 00222931108693053"]}

Meganola rhyssomorpha Hacker, 2012 (Figs 31���32, 74, 98) Meganola rhyssomorpha Hacker, 2012, Esperiana 17: 390. Type locality: Ivory Coast, San Pedro. Holotype, ♀ (ZSM). Material examined. Ivory Coast. 3 ♂, 174m, Tai NP., Tai Research Station, 05��49���59.8���N, 07��20���32���W, 14��� 23.XI.2015, Light Trap, Aristophanous, M., Moretto, P., Ruzzier, E. leg., ANHRT:2017.16, slide Nos.: LGNA 140 ♂, LGNA 210 ♂, LGNA 233 ♂; 2 ♀, from the same site, but collected at 25.III.���17.IV.2017 by Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P., ANHRT:2017.25, slide Nos: LGNA 399 ♀, LGNA 984 ♀. Sierra Leone . 5 ♂, 1 ♀, 120m, Tiwai Island, Moa River, N07��33���00���, W11��21���09���, 17���22.vi.2016, Light Trap, leg. Takano, Miles & Goff, ANHRT:2017.18, slide Nos: LGNA 406 ♂, LGNA 407 ♂, LGNA 410 ♂, LGNA 467 ♂, LGNA 408 ♀. Liberia . 1 ♂, 530m, Lofa County, Foya Proposed Protected Area, 7��56���36���N, 10��16���36���W, 10���19.xi.2017, MV Light Trap (125W), Aristophanous, M., S��fi��n, Sz., Simonics, G. & Smith, L. leg., ANHRT:2017.33, slide No.: LGNA 550 ♂ (ANHRT). Remark. The species was described based on a single female specimen. A series of males of this rather distinctive species have recently been collected in West Africa and the genitalia are illustrated here for the first time. Distribution. Ivory Coast (Hacker et al. 2012). The specimens from Sierra Leone and Liberia represent new country records., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on pages 167-168, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Meganola cinereoparva sp. n. (Figs 35���36, 77���78) Holotype. ♂, Ivory Coast, 174m, Tai NP., Tai Research Station (SRET), 05��50���00���N, 07��20���32.0���W, 25.III��� 17.IV.2017, MV Light, Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P. leg., ANHRT:2017.25, unique number: ANHRTUK 00001457, slide No.: LGNA 397 ♂ (ANHRT). Paratype. ♂, Ivory Coast, 174m, Tai NP., Tai Research Station, 05��49���59.8���N, 07��20���32.0���W, 14���23.XI.2015, Light Trap, Aristophanous, M., Moretto, P., Ruzzier, E. leg., ANHRT:2017.16, unique number: ANHRTUK 00009100, slide No.: LGNA 137 ♂ (ANHRT). Diagnosis. The new species is similar externally to M. spermophaga Fletcher, 1962, but distinguished by its characteristically long, wedge-shaped costal patch of the forewing, which is much shorter and dash-like in M. spermophaga. In the male genitalia the most conspicuous differences between the two species are the much narrower valva, the more slender and shorter harpe, and the significantly shorter aedeagus of the new species compared to those of M. spermophaga; in addition a short, triangular carinal process of the aedeagus is present in M. spermophaga (that character is absent in the new species) and a pair of short, thorn-like cornuti is present on the vesica in the new species (the cornutus of the vesica is a relatively larger plate with a single thorn-like process in M. spermophaga). The genital morphology suggests M. cinereoparva is closely related to M. subchionea, in particular the similar ground plan of their clasping apparatus and the presence of a characteristic pair of short, thorn-like cornuti of the vesica situated near the posterior end of the aedeagus when the vesica is not everted. The main morphological differences between M. cinereoparva and M. subchionea are as follows: the forewing ground colour of the new species is dark greyish, while that of M. subchionea is off white; in addition the medial area of the forewing of the new species is not filled in with dark grey scales, only the costal half of the antemedial line is connected to a wedgeshaped dark grey costal patch, while in M. subchionea the medial area of forewing is completely filled in with darker scales, forming a wide medial band. In the male genitalia, M. cinereoparva and M. subchionea differ in the following characters: the uncus of the new species is significantly shorter, the apical part of its valva is considerably broader and its harpe is much more slender, longer and more curved than those of the related species. The aedeagus of M. cinereoparva is similar to that of M. subchionea, but considerably shorter and the pair of small cornuti of the vesica are somewhat longer than those of the related species. The female is unknown., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 170, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Fletcher, D. S. (1962) Nolinae from Tanganyika collected by Dr. Christa Lindemann and Nina Pavlitzki. Veroffentlichungen der zoologischen Staatssammlung Munchen, 7, 1 - 8."]}

Meganola dananae Hacker, 2012 (Figs 19���20, 66, 94) Meganola dananae Hacker, 2012, Esperiana 17: 382. Type locality: Ivory Coast, Danane. Holotype, ♀ (coll. Hacker /ZSM). Material examined. Guinea. 1 ♂, 435m, Geipa Camp, Foret de Diecke, 7��26���7.06���N, 8��50���47.87���W, 05���14.iv.2019, Cold Cathode UV Light Trap (8W), S��fi��n, Sz., Koivogui, S. leg. ANHRT: 2019.7, slide No.: LGNA 975 ♂; 1 ♀, same data but collected by light trap equipped with 250W blended bulb, slide No.: LGNA 988 ♀ (ANHRT). Remark. The species was described from a single female specimen. The external habitus as well as the genital morphology of the species suggest a close relationship to M. bispermutata Hacker, 2012. The male is illustrated here for the first time (Figs 19, 66). Distribution. Ivory Coast (Hacker et al. 2012). New country record for Guinea., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 162, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Meganola hackeri sp. n. (Figs 48���49, 85) Holotype. ♂, Ivory Coast, 1171m, Mt. Tonkoui Peak, 07��27���15.2���N, 07��38���12.5���W, 1���8.XI.2015, Light Trap, Aristophanous, M., Moretto, P., Ruzzier, E. leg., ANHRT:2017.16, unique number: ANHRTUK 00009101, slide No.: LGNA 239 ♂ (ANHRT). Paratypes. Ivory Coast. 1 ♂, 39���48m Abidjan, Banco Forest (Parc-National du Banco), 05��23���03.8���N, 04��03���11.2W, 21���30.IV.2017, MV light Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P. leg. ANHRT:2017.25, unique number: ANHRTUK 00052754, slide No.: LGNA 982 ♂. Sierra Leone . 1 ♂, 80m, Kalainkay nr. Kamabai, Northern Prov., 3���6.XI.2015, N09��10���52���, W11��56���44���, Light Trap, R. Goff coll., leg. Smith, R. & Takano, H., ANHRT:2018.20, unique number: ANHRTUK 00009102, slide No.: LGNA 286 ♂. Liberia . 1 ♂, 1165m, Nimba Mts camp, ENNR, Nimba county, 7��31���45���N, 8��31���37���W, 03���13.xii.2017 Light Trap (blended bulb 250W) Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg. ANHRT:2017.33, unique number: ANHRTUK 00019908, slide No.: LGNA 488 ♂; 1 ♂, 865m, Lofa County, Wologizi Mts, Ridge Camp, 8��07���10���N, 9��57���11���W, 24���29.xi.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G. & Smith, L. leg. ANHRT:2017.33, unique number: ANHRTUK 00145396, slide No.: LGNA 1152 ♂ (ANHRT). Diagnosis. The new species is related to M. conspicillaris (Fletcher, 1962), but easily distinguishable by its larger size (wingspan of M. conspicillaris is 14���16 mm, that of M. hackeri is 17���21 mm) and the much narrower, dash-like medio-costal patch, which is much broader, longer and rather quadrangular in M. conspicillaris. In addition, both the fore- and hindwing ground colour of the new species is considerably paler than that of M. conspicillaris. It is worth mentioning the extraordinarily long rami of the male bipectinate antennae of M. hackeri, although it is not possible to make any comparisons with the holotype of M. conspicillaris as the antennae are missing. The configuration of the male genitalia of M. hackeri suggests a close relationship to M. conspicillaris, both taxa sharing a rather unique, long, asymmetrical basal processes of the valva. However, the new species has a somewhat longer, apically protracted uncus (which is evenly tapering in the related species), an apically more dilated valva, considerably shorter and pointed asymmetrical basal processes of the valva (these are apically dilated and rounded in M. conspicillaris), a much shorter and thicker, almost straight harpe (which is slightly S-shaped in the related species) and a somewhat longer vinculum compared to M. conspicillaris. The aedeagus of the new species is similar in shape to that of the sister species, but considerably shorter and slightly thicker. The female is unknown., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 175, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Fletcher, D. S. (1962) Nolinae from Tanganyika collected by Dr. Christa Lindemann and Nina Pavlitzki. Veroffentlichungen der zoologischen Staatssammlung Munchen, 7, 1 - 8."]}

Meganola taiana sp. n. (Figs 26���28, 71���72, 95) Holotype. ♂, Ivory Coast, 174m, Tai NP., Tai Research Station, 05��49���59.8���N, 07��20���32.0���W, 14���23.XI.2015, Light Trap, Aristophanous, M., Moretto, P., Ruzzier, E. leg., ANHRT:2017.16, unique number: ANHRTUK 00009096, slide No. LGNA 142 ♂ (ANHRT). Paratypes. Ivory Coast. 2 ♂, with the same data as the holotype, unique numbers: ANHRTUK 00009097, 00009098, slide Nos: LGNA 147 ♂, LGNA 209 ♂; 1 ♂, 1 ♀, same site, but collected at 25.III.���17.IV.2017 by Aristophanous, A., Aristophanous, M., Geiser, M., Moretto P., ANHRT:2017.25, unique numbers: ANHRTUK 00001412, ANHRTUK 00049668, slide Nos: LGNA 395 ♂, LGNA 983 ♀. Sierra Leone . 3 ♂, 120m, Tiwai Island, Moa River, N07��33���00���, W11��21���09���, 17���22.vi.2016, Light Trap, leg. Takano, Miles & Goff, ANHRT:1017.18, unique numbers: ANHRTUK 00016243, 00026228, 00026230, slide Nos: LGNA 409 ♂, LGNA 515 ♂, LGNA 564 ♂. Liberia . 1 ♂, 530m, Lofa County, Foya Proposed Protected Area, 7��56���36���N, 10��16���36���W, 10���19.xi.2017, MV Light Trap (125W), Aristophanous, M., S��fi��n, Sz., Simonics, G. & Smith, L. leg., ANHRT:2017.33, unique number: ANHRTUK 00039105, slide No.: LGNA 551 ♂; 1 ♂, 1165m, Nimba Mts camp, ENNR, Nimba county, 7��31���45���N, 8��31���37���W, 03���13.xii.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33, unique number: ANHRTUK 00019898, slide No.: LGNA 487 ♂ (ANHRT)., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 164, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542

Meganola pyrrhomorpha Hacker, 2012 (Figs 41���42, 81, 101) Meganola pyrrhomorpha Hacker, 2012, Esperiana 17: 398. Type locality: Ethiopia, Oromia Prov., Bonga. Holotype, ♂ (coll. Hacker /ZSM). = Meganola tabbertiella Hacker & Hoppe, 2012, Esperiana 17: 414, syn. n. = Meganola amaniella Hacker, 2012, Esperiana 17: 409, syn. n. = Meganola fuscostriata Hacker, 2012, Esperiana 17: 411, syn. n. Material examined. Ivory Coast. 1 ♂, 174m, Tai NP., Tai Research Station, 05��49���59.8���N, 07��20���32.0���W, 14��� 23.XI.2015, Light Trap, Aristophanous, M., Moretto, P., Ruzzier, E. leg., ANHRT:2017.16, slide No.: LGNA 138 ♂; 1 ♂, 3 ♀, 1171 m, Mt Tonkoui Peak, N07��27���15,2���, W07��38���12,5��� 1���8.XI.2015, Light Trap, leg. Aristophanous, M., Moretto, P. & Ruzzier, E., ANHRT:2017.16, slide Nos: LGNA 232 ♂, LGNA 48 ♀, LGNA 49 ♀, 145 ♀; 1 ♂, same site, but collected at 9���16.IV.2016 by Aristophanous, M. and Moretto, P., ANHRT:2017.17, slide No.: LGNA 565 ♂. Sierra Leone . 1 ♀, Baoma, Goderich, 8��25���41���N, 13��15���47���W, leg. R.W. Goff, 18.III.2015, ANHRT:2019.1, slide No.: LGNA 171 ♀. Liberia . 2 ♂, 530m, Lofa County, Foya Proposed Protected Area, 7��56���36���N, 10��16���36���W, 10���19.xi.2017, MV Light Trap (125W), Aristophanous, M., S��fi��n, Sz., Simonics, G. & Smith, L. leg., ANHRT:2017.33, slide Nos: LGNA 548 ♂, LGNA 552 ♂; 1 ♀, 1165m, Nimba Mts camp, ENNR, Nimba county, 7��31���45���N, 8��31���37���W, 03���13.xii.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33, slide No.: LGNA 553 ♀; 1 ♀, 1000���1100m, Nimba Mts., ENNR, Nimba county (Cellcom road), 7��32���45.9���N, 8��31���21���W, 27.iii���04.iv.2017, Light Trap (250W blended bulb) & Cold Cathode UV light bucket trap (8W), S��fi��n, Sz., Simonics, G. leg., ANHRT:2017.36, slide No.: LGNA 546 ♀ (ANHRT). Remarks. M. pyrrhomorpha Hacker, 2012 is a rather characterless small moth, rendering it difficult to identify. It resembles other highly similar species (e.g. M. spermophaga, M. illaudata, M. antennata) requiring the examination of the genitalia for reliable identification. The most important distinctive features in the male of the species are the conspicuously curved valva with a heavily sclerotized, broad costal margin, the long, distally slightly curved, apically dilated and broadly rounded harpe, the short aedeagus bearing a short, curved carinal process and the single, short and thin spike-like cornutus of the vesica. The most distinctive features in the female are the short, narrow, distally asymmetrical antrum, the short, rounded lateral protrusion at the distal end of the very long and membranous ductus bursae and the elongate ovoid scobinated plate of signum bursae. As several species described in Hacker et al. (2012) share these characters without any detectable distinctive features, the following species are considered to be mere synonyms of M. pyrrhomorpha: M. tabbertiella Hacker & Hoppe, 2012 syn. n., M. amaniella Hacker, 2012 syn. n. and M. fuscostriata Hacker, 2012 syn. n. The validity of several further, externally very similar species is supported by minor, but stable differences in the genitalia. These species are as follows: M. spermophaga (Fletcher, 1962), M. stictographa Hacker, 2012, M. amphigrapha Hacker, 2012, M. foviferoides (Poole, 1989), M. pedata (Fletcher, 1962) and M. obscuritata Hacker, 2012. It is worth noting that the externally strikingly different M. loxoleuca (Fletcher, 1958) and M. loxodentata Hacker & Hoppe, 2012 belong to the same species-group based on the configuration of their genitalia. Distribution. Ethiopia, Tanzania, Uganda, D.R. Congo, Kenya, Ghana (Hacker et al. 2012), Sierra Leone and Ivory Coast (Hacker 2014). The specimens from Liberia represent new country record., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on pages 172-173, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612.","Fletcher, D. S. (1962) Nolinae from Tanganyika collected by Dr. Christa Lindemann and Nina Pavlitzki. Veroffentlichungen der zoologischen Staatssammlung Munchen, 7, 1 - 8.","Poole, R. W. (1989) Noctuidae, Part 2. In: Heppner, J. B. (Ed.), Lepidopterorum Catalogus. New Series. Fascicle 118. Flora and Fauna Publications. E. J. Brill, New York, pp. 635 - 639.","Fletcher, D. S. (1958) Ruwenzori Expedition 1952. Arctiidae, Nolidae. Ruwenzori Expedition 1952, 1 (4), 55 - 67.","Hacker, H. H. (2014) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae) Supplement 1. Esperiana, 19, 121 - 179."]}

Meganola florianhackeri Hacker, 2012 Meganola florianhackeri Hacker, 2012, Esperiana 17: 482. Type locality: Kenya, Mt. Kenya, Thego. Holotype, ♂ (ZSM). Remark. Hacker et al. (2012) refer to a single male specimen from Abidjan, Ivory Coast deposited in RMCA. Distribution. Kenya, Tanzania, Ethiopia, Rwanda and Ivory Coast (Hacker et al. 2012)., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 172, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Meganola despicillaris Hacker, 2012 Meganola despicillaris Hacker, 2012, Esperiana 17: 455. Type locality: Ivory Coast, Gouedie. Holotype, ♀ (MNVD). Distribution. Ivory Coast (Hacker et al. 2012). F FIGURES 104���108: female genitalia (all in coll. ANHRT). 104, Meganola leucosigna, Ivory Coast, LGNA 264; 105, M. pyrrhosoma, Ivory Coast, LGNA 226; 106, M. illaudata, Liberia, LGNA 501; 107, M. monofascia, Ivory Coast, LGNA 256; 108, M. spherographa, Liberia, LGNA 973., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on pages 178-179, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Meganola illaudata (Fletcher, 1958) (Figs 54���56, 87, 106) Vandamia illaudata Fletcher, 1958, Ruwenzori Expedition 1952 1 (4): 56, figs 22���24, 37. Type locality: Uganda, Ruwenzori, Ibanda. Holotype, ♂ (NHMUK). = Meganola eburneana Hacker, 2012, Esperiana 17: 416, syn. n. Material examined. Ivory Coast. 1 male, 1171m, Mt. Tonkoui Peak, 07��27���15.2���N; 07��38���12.5���W, 1���8.XI.2015, Light Trap, Aristophanous, M., Moretto, P., Ruzzier, E. leg., ANHRT:2017.16, slide No.: LGNA 224 ♂. Liberia . 4 ♂, 6 ♀, 1165m, Nimba Mts camp, ENNR, Nimba county, 7��31���45���N, 8��31���37���W, 03���13.xii.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33, slide Nos: LGNA 486 ♂, LGNA 500 ♂, LGNA 555 ♂, LGNA 556 ♂, LGNA 489 ♀, LGNA 501 ♀, LGNA 557 ♀, LGNA 558 ♀; 1 ♂, 1 ♀, 1000���1100m, Nimba Mts., ENNR, Nimba county (Cellcom road), 7��32���45.88���N, 8��31���21.04���W, 02���14.xii.2017, Cold Cathode Light Bucket, Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33, slide No.: LGNA 493 ♂; 1 ♀, 611m, Lofa County, Wologizi Mts, base camp forest, 8��07���17���N, 9��57���42���W, 20.xi.���01.xii.2017, MV Light Trap (125W), Aristophanous, M., S��fi��n, Sz., Simonics, G. & Smith, L. leg., ANHRT:2017.33; 1 ♀, 865m, Lofa County, Wologizi Mts, Ridge Camp, 8��07���10���N, 9��57���11���W, 24���29.xi.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G. & Smith, L. leg., ANHRT:2017.33 (ANHRT). Remark. Meganola illaudata has unique male genitalia bearing a bifid uncus (Fig. 87), allowing for easy identification of an otherwise rather featureless moth. The species was described from the Ruwenzori Mts, Uganda. The genitalia of the male holotype and a female paratype are illustrated by Hacker et al. (2012) and despite his knowledge of both sexes of M. illaudata, Hacker described M. eburneana Hacker, 2012 based on a single female specimen collected in Ivory Coast showing no distinctive characters distinguishing it from M. illaudata. A long series of externally well matching males and females of M. illaudata from Liberia in the ANHRT collection confirms the conspecificity of the two species, therefore M. eburneana is synonymised here with M. illaudata. It is worth noting that the species is also figured by Hacker et al. (2012) as a female paratype specimen of M. antennata Hacker, 2012 from Pwani, Tanzania which is undoubtedly as a result of a misidentification. Distribution. Uganda, Ethiopia, Tanzania, D.R. Congo, Equatorial Guinea (Bioko Island) and Ivory Coast (Hacker et al. 2012). The specimens from Liberia are new country records representing the westernmost occurrence of the species., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on pages 180-181, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Fletcher, D. S. (1958) Ruwenzori Expedition 1952. Arctiidae, Nolidae. Ruwenzori Expedition 1952, 1 (4), 55 - 67.","Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Meganola spermophaga (Fletcher, 1962) (Figs 39���40, 80, 100) Nola spermophaga Fletcher, 1962, Ver��ffentlichungen der zoologischen Staatssammlung M��nchen 7: 5. Type locality: Uganda, Kampala. Holotype, ♂ (NHMUK). = Meganola obscuritata Hacker, 2012, Esperiana 17: 413, syn. n. Material examined. Sierra Leone. 2 ♀, 80m, Kalainkay nr. Kamabai, Northern Prov., 3���6.XI.2015, N09��10���52���, W11��56���44��� Light Trap, R. Goff coll., leg. Smith, R. & Takano, H., ANHRT:2018.20, slide Nos: LGNA 291, LGNA 462 (all ♀). Cameroon. 1 ♂, 1246m, Adamawa Region, Adamawa Plateau, 7.3km West of Bazanga, Chute de Tello, 07��13���50.6���N, 13��56���29.2���E, 24���30.ix.2018, MV Light Trap, S��fi��n, Sz., Simonics, G. leg., ANHRT:2018.36, slide No.: LGNA 575 ♂. Liberia . 1 ♂, 530m, Lofa County, Foya Proposed Protected Area, 7��56���36���N, 10��16���36���W, 10��� 19.xi.2017, MV Light Trap (125W), Aristophanous, M., S��fi��n, Sz., Simonics, G. & Smith, L. leg., ANHRT:2017.33, slide No.: LGNA 547 ♂. Zambia . 1 ♀, 1030m, Kundabwika Falls, Kalungwishi River, S 09��13���00���, E29��18���17���, 5���7. V.2013, Light Trap, leg. Smith, Takano & Oram, ANHRT:2017.9, slide No.: LGNA 43 ♀ (ANHRT). Remark. The male holotype of M. obscuritata Hacker, 2012 as illustrated by Hacker et al. (2012: 413) does not show any distinctive features in genitalia and external habitus compared to M. spermophaga and thus the former species is synonymised here with the latter. It is worth noting however, that the female specimens illustrated in Hacker et al. (2012) as M. obscuritata are apparently not conspecific with the male holotype, judging from their considerably different coloration and wing pattern. Distribution. Uganda, D.R. Congo, Republic of Congo, Burundi, Kenya, Tanzania, Ethiopia, Nigeria, Burkina Faso and Ivory Coast (Hacker et al. 2012). Specimens from Sierra Leone, Cameroon, Liberia and Zambia represent new country records., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 172, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Fletcher, D. S. (1962) Nolinae from Tanganyika collected by Dr. Christa Lindemann and Nina Pavlitzki. Veroffentlichungen der zoologischen Staatssammlung Munchen, 7, 1 - 8.","Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Meganola palaeographa Hacker, 2012 (Figs 14���15, 62, 93) Meganola palaeographa Hacker, 2012, Esperiana 17: 417. Type locality: Tanzania, Iringa. Holotype, ♂ (coll. Hacker /ZSM). Material examined. Ivory Coast. 7 ♂, 10 ♀, 1171 m, Mt Tonkoui Peak, N07��27���15,2���,W07��38���12,5���, 1���8.XI.2015, Light Trap, leg. Aristophanous, M., Moretto, P. & Ruzzier, E., ANHRT:2017.16, slide Nos LGNA 24 ♀, LGNA 39 ♂, LGNA 40 ♀, LGNA 41 ♂, LGNA 87 ♀, LGNA 148 ♂, LGNA 150 ♀, LGNA 165 ♀, LGNA 166 ♂, LGNA 167 ♀, LGNA 168 ♀, LGNA 235 ♂, LGNA 236 ♂, LGNA 238 ♂, LGNA 240 ♀, LGNA 252 ♀, LGNA 254 ♀; 2 ♂, 174m, Tai NP., Tai Research Station, 05��49���59.8���N, 07��20���32���W, 14���23.XI.2015, Light Trap, Aristophanous, M., Moretto, P., Ruzzier, E. leg., ANHRT:2017.16, slide Nos: LGNA 227 ♂, LGNA 234 ♂; 2 ♂, from the same site, but collected at 25.III.���17.IV.2017 by Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P., ANHRT:2017.25, slide Nos: LGNA 394 ♂, LGNA 398 ♂. Liberia . 5 ♂, 1 ♀, 1165m, Nimba Mts camp, ENNR, Nimba county, 7��31���45���N, 8��31���37���W, 03���13.xii.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33, slide Nos LGNA 478 ♂, LGNA 544 ♂; 2 ♂, 1000���1100m, Nimba Mts, ENNR, Nimba county (Cellcom road), 7��32���45.88���N, 8��31���21.04���W, 02���14.xii.2017, Cold Cathode Light Bucket, Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33; 1 ♂, 1 ♀, 1327m, Nimba Mts main ridge (montane forest), ENNR, Nimba county, 7��31���1.3���N, 8��31���1.0���W, 9.xii.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg. ANHRT:2017.33, slide No.: LGNA 538 ♀; 1 ♀, 230m, Lofa county, Zuwulor Village School, 7��54���52���N, 9��31���08���W, 8.xi.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33, slide No.: LGNA 543 ♀ (ANHRT). Remark. The species was known only from two specimens: the male holotype specimen from Udzungwa Mts, Tanzania (Hacker et al. 2012) and an additional record from the same area published later (Hacker 2014). Surprisingly, the specimens recently collected in West Africa hardly show any morphological differences to the ones from Tanzania, despite the large distance between these localities. The female and its copulatory organ are illustrated here for the first time. Distribution. Tanzania (Hacker et al. 2012, Hacker 2014). The specimens from Ivory Coast and Liberia represent new country records. The species is likely to be distributed throughout Equatorial Africa., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on page 161, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612.","Hacker, H. H. (2014) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae) Supplement 1. Esperiana, 19, 121 - 179."]}

Meganola monofascia (van Son, 1933) (Figs 16���18, 63���65, 107) Nola major var. monofascia van Son, 1933, Annals of the Transvaal Museum 15: 214. Type locality: [RSA] Transvaal, Pietersburg. Holotype, ♂ (TMSA). = Meganola septima Hacker, 2012, Esperiana 17: 375, syn. n. = Meganola octava Hacker, 2012, Esperiana 17: 377, syn. n. = Meganola heteromorpha Hacker, 2012, Esperiana 17: 383, syn. n. = Meganola dissoluta Hacker, 2012, Esperiana 17: 384, syn. n. = Meganola brachyvalva Hacker, 2012, Esperiana 17:385, syn. n. = Meganola pseudofuscata Hacker, 2012, Esperiana 17: 468, syn. n. Material examined. Ivory Coast. 1 ♂, 1171 m, Mt Tonkoui Peak, N07��27���15���, W07��38���13���, 12���18.VII.2015, Light Trap, leg. Aristophanous, M., Moretto, P. & Ruzzier, E., ANHRT:2017.14, slide No.: LGNA 26 ♂; 3 ♂, same site, but collected at 1���8.XI.2015, ANHRT:2017.16, slide Nos: LGNA 237 ♂, LGNA 245 ♂, LGNA 246 ♂. Liberia . 1 ♂, 1327m, Nimba county, ENNR, Nimba Mts, main ridge (montane forest), 7��31���1.3���N, 8��31���1.0���W, 9.xii.2017, Light Trap (blended bulb 250W), Aristophanous, M., S��fi��n, Sz., Simonics, G., Smith, L. leg., ANHRT:2017.33, slide No.: LGNA 583 ♂. Cameroon . 2 ♂, 1246m, Adamawa Region, Adamawa Plateau, 7.3km West of Bazanga, Chute de Tello, 07��13���50.6���N, 13��56���29.2���E, 24���30.ix.2018, MV Light Trap, S��fi��n, Sz., Simonics, G. leg., ANHRT:2018.36, slide Nos: LGNA 568 ♂, LGNA 570 ♂. Sierra Leone . 1 ♂, 80m, Kalainkay nr. Kamabai, Northern Prov., 3���6.XI.2015, N09��10���52���, W11��56���44���, Light Trap, R. Goff coll., leg. Smith, R. & Takano, H., ANHRT:2018.20, slide No.: LGNA 289 ♂; 1 ♂, 420m, Mansonia village at the foothills of Loma Mts, N09��07���47���, W11��05���06���, 6.vi.2016, Light Trap, leg. Takano, Miles & Goff, ANHRT:2017.18, slide No.: LGNA 471 ♂. Zambia. 1 ♂, 1400m, Hillwood, Ikelenge, S11��16���02���, E24��18���59���, 30.IV.���11. V.2014, Light Trap, leg. Smith, R., Takano, H., Chmurova, L. & Smith, L., ANHRT:2017.11, slide No.: LGNA 103 ♂; 1 ♂, same locality, but collected at 25���27.xi.2014 by Smith, R. & Takano, H., ANHRT:2017.12, slide No.: LGNA 393 ♂. Mozambique . 1 ♂, Maputo Special Reserve, 22m, West Gate, Sand Forest, 26��30���14.2���S, 32��42���59.6���E, 3���13.XII.2016, Light Trap, Aristophanous, M., Cristovao, J., L��szl��, G., Miles, W. leg., ANHRT:2017.22, slide No.: LGNA 354♂; 2 ♂, same locality and collectors, but collected at 21���30.XII.2016, slide Nos: LGNA 517 ♂, LGNA 518 ♂; 1 ♂, same site, but collected at 21���22.II.2018, by L��szl��, G., Mulvaney, J. & Smith, L., ANHRT:2018.2, slide No.: LGNA 519 ♂; 1 ♂, Ponta Milibangalala, Dune Grassland, 26��26���58.6���S, 32��55���29.8���E, 25���30. V.2017, MV Light Trap, Aristophanous, M., L��szl��, G., Miles, W., Vetina, A. leg., ANHRT:2017.26 (ANHRT). Remark. The externally variable but internally (genital morphology) fairly constant and distinctive M. monofascia has been found to be surprisingly widely distributed throughout Sub-Saharan Africa. As the validity of the numerous related species described by Hacker (Hacker et al. 2012) in the M. monofascia species-group is not supported by any firm distinctive morphological characters (cf. Hacker et al. 2012: 366, 375, 377, 383, 384, 385, 468), M. septima, octava, heteromorpha, dissoluta, brachyvalva and pseudofuscata are synonymized here with M. monofascia. The closest relative of M. monofascia is the Madagascan taxon M. praefica (Saalm��ller, 1884). Meganola monofascia is hardly distinguishable externally from the Southern African M. bispermutata Hacker, 2012 despite the considerable differences in their male genitalia (L��szl�� & Vetina 2019). It is also worth noting that the three specimens identified as M. furvitincta (Hampson, 1914) from Nigeria and Ivory Coast figured by Hacker et al. (2012: 380) belong in fact to M. monofascia. Distribution. South Africa, Zimbabwe, Tanzania, Uganda, Ethiopia, Tanzania, Kenya, D.R. Congo, Republic of Congo, Burundi, Nigeria, in addition, Ivory Coast (Hacker et al. 2012), Senegal (Hacker 2014) and Mozambique (L��szl�� & Vetina 2019). The specimens from Sierra Leone, Liberia, Cameroon and Zambia represent new country records., Published as part of L��szl��, Gyula M., 2020, Review of the genus Meganola Dyar, 1898 of Ivory Coast and adjacent areas with descriptions of 5 new species and several taxonomic updates (Lepidoptera Nolidae, Nolinae) - Taxonomic studies on West African Nolinae I., pp. 151-182 in Zootaxa 4853 (2) on pages 161-162, DOI: 10.11646/zootaxa.4853.2.1, http://zenodo.org/record/4410542, {"references":["Son, V. G. (1933) A revision of the South African moths of the tribe Nolini. Annals of the Transvaal Museum, 15 (2), 181 - 232.","Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612.","Saalmuller, M. (1884) Lepidopteren von Madagascar. Erste Abtheilung. Rhopalocera, Heterocera: Sphinges et Bombyces. Im Selbstverlas, der Gesellscliaft, Frankfurt am Main, 246 pp. [in German]","Hampson, G. F. (1914) Catalogue of the Amatidae and Arctiadae (Nolinae and Lithosianae) in the Collection of the British Museum. Catalogue of the Lepidoptera Phalaenae in the British Museum, London, Supplement 1, 1 - 858.","Hacker, H. H. (2014) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae) Supplement 1. Esperiana, 19, 121 - 179."]}

Cyana (Cyabarda) nambi sp. nov. (Figs 1, 2, 16, 24) urn:lsid:zoobank.org:act: 9A954F72-F42F-4697-8BD3-BDC6D4CF9B9E Type material. Holotype (Figs 1, 16): ♂, ��� Uganda, Kibale National Park, Biol. Field Station, 19.���24.XI.2014, LF, leg. W. Mey ���, slide AV4250 ♂ Volynkin (Coll. ZMB). Paratypes: 3 ♂, 1 ♀, with the same data as the holotype, slide Nos: AV4251 ♀, AV4849 ♂ (Coll. ZMB); 2 ♂, Uganda, Mpigi, Mpanga Forest, 25���30.XI.2014, LF, leg. W. Mey (Coll. ZMB); 1 ♀, Uganda, Kibale Forest, 1510m, Biological Field Station, Nordrand, ca. 14 km SO Fort Portal, Rand gest��rter Regenwald, 0��33���44.6������N, 30��21���24.7������O, 19.X.2014, LF, leg. K.-R. Beck / Gen. Pr��p. 3309 pr��p. KARISCH 2015 (Coll. CKDT). Diagnosis. Cyana nambi (Figs 1, 2) is a sister species of C. torrida (Figs 3, 4), from which it can be distinguished by its larger size, darker, spot-like posteriorly broadened postmedial line, subapically more strongly broadened subterminal line, and the presence of greyish brown irroration on both wings (which is absent in C. torrida). The male genitalia of C. nambi (Fig. 16) are similar to those of C. torrida (Fig. 17), but in C. nambi the uncus is narrower, the lateral apical processes of juxta are slightly larger, the saccus is rectangular apically (that is more rounded in C. torrida), the ampulla is narrower and hook-like curved (that is club-shaped in C. torrida), the distal saccular process is longer and slightly curved dorsad (that is straight in C. torrida), the aedeagus is much narrower distally and has shorter transverse dentate carinal crest, the cornuti of the subbasal patch are more robust, the sclerotised area on the lateral diverticulum is broader, and the distal diverticulum is slightly smaller. The female genitalia of C. nambi (Fig. 24) differ from those of C. torrida (Figs 25, 26) by its longer apophyses posteriores, the slightly broader and more rectangular antrum and the slightly larger signum bursae. Description. External morphology of adults. Male (Fig. 1). Forewing length 9���9.5. Antenna ciliate. Head white; thorax white with four greyish brown spots; abdomen white with admixture of greyish hairs. Forewing ground color off-white with pale greyish irroration, pattern greyish brown. Subbasal dot large. Costa at base with short longitudinal dash. Antemedial line S-curved, interrupted into four spots: one small at costa, one (the smallest) in the cell, one (the largest) postero-medially and one small at anal margin. Three discal spots present at the end of the cell. Postmedial line interrupted, represented by a large, broad spot at anal margin and a small diffuse spot anteriorly of it. Subterminal line interrupted, represented by a relatively large spot at apex and a diffuse small spot posteriorly of it situated near the outer margin. Cilia creamy-white. Hindwing off-white with intense greyish brown irroration at apex and along the outer margin. Female (Fig. 2). Forewing length 12���13 mm. Antenna weakly ciliate, with shorter ciliae than in male. Forewing ground colour pale grayish white, wing pattern similar to that of male but greyish brown suffusion on hindwing much more intense than in male, and cell with two discal spots only: a medial and a distal one. Male genitalia (Fig. 16). Uncus short, narrowly trigonal, dorso-ventrally flattened, fused with tuba analis basally. Tuba analis narrow, subscaphium setose. Tegumen short, relatively narrow. Juxta broadly trapezoidal, with a pair of short club-like latero-apical processes. Vinculum broad, robust, rectangular with rounded corners, saccus well developed, quadrangular, heavily sclerotised. Valva broad at base, slightly tapered distally. Costa concave; transtilla large, robust, broadly trigonal with rounded apex and convex dorsal margin. Ampulla short, curved, hook-like, densely setose. Cucullus short, narrow, weakly setose, apically rounded. Sacculus moderately broad, heavily sclerotised. Distal saccular process robust, elongated, laterally flattened, slightly curved dorsad, with small pointed denticle on apex. Aedeagus tubular, robust, tapered medially, slightly broadened proximally with small trigonal coecum, considerably broadened distally. Distal section of aedeagus with one long and robust weakly dentate transverse crest ventrally and one small and weakly dentate crest dorsally. Vesica short, with short subbasal dorsal diverticulum bearing three short but robust dent-like cornuti at base; lateral globular diverticulum with granulation and sclerotised area latero-ventrally; small granulated diverticulum laterally; and broad globular, heavily granulated distal diverticulum. Female genitalia (Fig. 24). Ovipositor short, broad; papillae anales broadly rectangular with rounded corners. Apophyses anteriores short, relatively broad, flattened, pointed apically. Apophyses posteriores thin, approximately 3 times longer than apophyses anteriores. Ostium bursae very broad. Antrum conspicuously broad, rectangular with rounded anterior corners, heavily sclerotised, rugose. Corpus bursae sack-like, its posterior two thirds sclerotised, strongly rugose anteriorly; anterior third of corpus bursae membranous, with small rounded signum. Appendix bursae small, narrowly conical, membranous, situated ventro-posteriorly, directed laterally. Distribution. Uganda (Western and Central Regions). Etymology. Nambi is the daughter of Ggulu, the creator deity in the Ugandan mythology., Published as part of Volynkin, Anton V. & L��szl��, Gyula M., 2019, On the taxonomy of the subgenera Cyabarda Karisch, 2013, Idiovulpecula Karisch, 2013 and Paravulpeculella Karisch, 2013 of the genus Cyana Walker, 1854, with descriptions of three new species from Africa (Lepidoptera, Erebidae, Arctiinae, Lithosiini), pp. 377-389 in Zootaxa 4664 (3) on pages 379-382, DOI: 10.11646/zootaxa.4664.3.5, http://zenodo.org/record/3385571

Cyana (Idiovulpecula) foya sp. nov. (Figs 11, 12, 22, 30) urn:lsid:zoobank.org:act: 0EE1ECB0-4A80-4A05-A506-E527746FC642 Type material. Holotype (Figs 11, 22): ♂, ��� Liberia, 530m, Lofa County, Foya Proposed Protected Area, 7��56���36������N 10��16���36������W, 10���19.XI.2017, MV Light Trap (125w), Aristophanous, M., S��fi��n, Sz., Simonics, G. & Smith, L., leg., ANHRT:2017.33���, ANHRT unique number ANHRTUK 00020066, slide No. AV 4594 ♂ (Coll. ANHRT). Paratypes: 3 ♂, 1 ♀, with same data as the holotype, ANHRT unique numbers ANHRTUK 00026066, 00020067, 00020076 and 00020520, slide Nos AV4627 ♂ and AV4628 ♀ (Coll. ANHRT). Diagnosis. Cyana foya (Figs 11, 12) resembles externally most of other members of the subgenus Idiovulpecula, but differs by its amber yellow transverse lines, whereas the transverse lines are crimson or absent in the related species (Figs 13���15). The ground plane of male genitalia of C. foya (Fig. 22) is similar to those of C. maculata (Fig. 20) and C. lowa (Fig. 21), but can be distinguished by its conspicuously shorter and broader uncus, more elongated juxta, broader vinculum with short but broad medial concavity, narrower medial protrusion of valva costa, narrower cucullus, and shorter but thicker distal saccular process. The aedeagus of C. foya is more elongated compared to the genital capsule than that of C. maculata, has longer and unilobate coecum and larger, broadly trigonal carinal process (that is smaller, finger-like in C. maculata); in comparison to that of C. lowa, the aedeagus of C. foya is narrower, straight distally (that is curved distally in C. lowa), with longer coecum and narrower carinal process directed distally (that is broader and directed dorsally in C. lowa). The vesica structure of C. foya differs clearly from that of C. maculata by the presence of the large, sack-like subbasal diverticulum with a conical tip (that is very small, tubercle-like in C. maculata), the absence of numerous small cornuti medially and the presence of the subapical field of short, but thick spinules (absent in C. maculata), and is most similar to that of C. lowa. However, the vesica of C. foya is curved subbasally around the aedeagus axis, and differs from that of C. lowa by its larger, unilobate subbasal diverticulum directed laterally (that is bilobate and directed distally in C. lowa), the presence of a sclerotised area at vesica base (absent in C. lowa), its larger lateral diverticulum and shorter subapical field of shorter, thicker and less numerous spinules. The female genitalia of C. foya (Fig. 30) are most similar to those of C. magnitrigutta (Fig. 31) by their antrum shape, but the antrum of C. foya is longer and broader than that of C. magnitrigutta; in addition, the anterior section of corpus bursae of C. foya is broader with smaller signum, and the posterior sclerotised section of corpus bursae is broader and less rugose compared to those of C. magnitrigutta. It is impossible to recognize the appendix bursae on the holotype slide of C. magnitrigutta, probably it was damaged or cut off accidentally during the preparation. Description. External morphology of adults (Figs 11, 12). Forewing length 11.5 mm in males and 13 mm in female. Sexual dimorphism barely expressed, wing pattern very similar in both sexes. Head milk white. Antennae ciliate with slightly longer cilia in males. Thorax, patagia and tegulae milk white, latter with one small amber spot. Abdomen whitish with admixture of pale ochreous scales. Forewing ground color milk white. Subbasal dot small, blackish. Cell with two rounded black spots medially and distally. Transverse lines amber. Subbasal line reduced to a short dash at costa. Antemedial line almost straight, oblique outwards posteriorly. Postmedial line evenly curved outwards. Terminal line diffuse, slightly broadened at costa. Cilia white. Hindwing pure milk white without pattern. Male genitalia (Fig. 22). Uncus short, broad, arrowhead-shaped, dorso-ventrally flattened, fused with tuba analis basally. Tuba analis broad, subscaphium weakly setose. Tegumen short and narrow. Juxta broad, inverse shieldshaped, medio-posteriorly shortly and narrowly incised, with a well-developed, elongated ventro-medial crest. Vinculum robust, nearly rectangular, anterior margin slightly concave, with well developed, trapezoidal saccus. Valva narrow at base, broadened medially. Transtillae broad, band-like. Costa with large trigonal dorsal protrusion bearing short subapical inner crest. Cucullus narrow, gently tapering, apically rounded, weakly setose. Sacculus moderately broad, evenly dilated medially, heavily sclerotised, with a robust, strongly curved dorsad, apically pointed distal process. Aedeagus relatively narrow, nearly straight, slightly broadened basally; coecum long, narrowly conical with rounded apex; carina with robust, broadly trigonal process directed dorsally. Vesica twisted around aedeagus axis, vesica ejaculatorius directed proximally. Subbasal section of vesica with a sclerotised plate dorsally. Lateral diverticulum large, membranous, sack-like with conical apex, directed ventrally. Subapical diverticulum small, globular, granulated. Subapical field of spinules narrow, elongated, consists of numerous short but relatively thick spinules of various lengths. Female genitalia (Fig. 30). Ovipositor short and broad, nearly rectangular. Papillae anales broadly trapezoidal with rounded corners, weakly setose. Apophyses thin medium long; apophyses anteriores slightly narrower and shorter than apophyses posteriores. Ostium bursae moderately broad. Antrum caliciform, heavily sclerotised. Ductus bursae narrow, short, sclerotised, rugose. Corpus bursae ovoid, its posterior section sclerotised and rugose at the connection with ductus bursae. Anterior section of corpus bursae membranous, with a small rounded signum. Appendix bursae approximately twice longer than corpus bursae, its basal section rounded, swollen and sclerotised, apical section large, gradually dilated, drop-shaped, membranous. Distribution. The species is known so far only from Liberia. Etymology. The specific epithet refers to its type locality in the Foya District of Lofa County, northwestern Liberia., Published as part of Volynkin, Anton V. & L��szl��, Gyula M., 2019, On the taxonomy of the subgenera Cyabarda Karisch, 2013, Idiovulpecula Karisch, 2013 and Paravulpeculella Karisch, 2013 of the genus Cyana Walker, 1854, with descriptions of three new species from Africa (Lepidoptera, Erebidae, Arctiinae, Lithosiini), pp. 377-389 in Zootaxa 4664 (3) on pages 384-387, DOI: 10.11646/zootaxa.4664.3.5, http://zenodo.org/record/3385571

Deltote turcica sp. n. (Figs 1 a���d, 2 c���d, 3 a) Type material. Holotype (Fig. 1 b): Male, Ankara-Kayseri, 26.7.1965, 1000 m, Central Turkey, Asia minor, leg. M. u. W. Glaser, slide No. RL 12402m (coll. Vartian, NHMW). Paratypes. Turkey. 2 males (Fig. 1 a and 1 c), with the same data as holotype, slide Nos RL 12401m, RL 12403m (coll. NHMW). Turkmenistan. 1 male, 1 female, Yol-Dere, 15 km NE Kara-Kala, 12. VI.1953, leg. V. Kuznetsov, slide Nos RL 11679m, RL11687f (coll. HNHM); 2 males, Kopet-Dagh Mts, Sayvana valley, 1000 m, 56��50���E, 38��17���N, L60, 28. VI.1992, leg. Gy. F��bi��n, B. Herczig, A. Podluss��ny & Z. Varga (coll. G. Ronkay). Description. Adult (Figs 1 a���d): Small noctuid moth with slender body, wingspan 13���14 mm, forewing length 6���7 mm. Head ���Antenna filiform in both sexes, dorsum covered with milky white scales; scape short and only ca twice as wide as basal segments of antenna. Eye large, globular, dark grey. Labial palp brown, short, upturned, third segment very short, pointed. Proboscis long, light brown. Head scales pale off white to light brown. Thorax ��� Ground colour similar to head, tegulae slender, metathorax somewhat more greyish. Legs gracile, brown, tibiae with longer milky white hair-scales, tarsi brown. Forewing: Ground colour and fringe light brown; basal and postmedial areas broad, white or brownish off-white; all lines wavy, terminal line beige to dark brown; orbicular stigma minute, rounded, dark brown with paler centre, reniform stigma fine, blackish, streak-like with fine white definition. Hindwing uniform beige, fringe brownish beige, terminal line narrow, black; discal spot, median line, and subterminal line dark, indistinct. Underside of forewing brown with dark subterminal line; that of hindwing pale brown, lacking lines and spots. Abdomen ���Slender, cylindrical, pale brown, basal 3���4 segments covered with smooth beige-whitish scales, dorsal crest absent, anal tuft of male weak, ochreous-white. Male genitalia (Figs 2 c���d): Genital capsule very small, weakly sclerotised. Uncus thick, medium-long, thickened towards acutely pointed and hooked. Tegumen weakly sclerotised, penicular lobes reduced; juxta large, flask-shaped; vinculum short, broadly U-shaped. Valva rather short (only slightly longer than tegumen), tapering towards apex; cucullus apically rounded, sparsely setose, apical thorn strong but short, finely arched; corona absent; ampulla short, mushroom-shaped. Aedeagus as long as valva, thick, subapically broadest, laterally and ventrally extended; carina less sclerotized; vesica (uneverted) membranous, with large L-shaped (boot-shaped) plate flattened, heavily sclerotized, positioned in distal half of aedeagus. Female genitalia (Fig. 3 a): Papillae anales nearly as long as wide, densely setose with short setae; both pairs of gonapophyses short (not or only slightly longer than papillae anales). Ostium bursae broad; antrum membranous, broad, funnel-like, with sclerotised ring at junction to ductus bursae; ductus bursae long, broadened towards corpus bursae, with sclerotised quadrangular plate at anterior section; appendix bursae small, subconical, membranous; corpus bursae elliptical-ovoid, with large scobinate area medially; signum sclerotised, narrow, Y-shaped. Differential diagnosis. The two species of this group are similar externally, with subtle differences in the forewing shape and pattern. Deltote turcica (Figs 1 a���d) has, comparing with D. delicatula (Figs 1 e���h) slightly narrower and more acutely pointed forewings and the postmedial line is running more obliquely to inner margin. The more recently collected specimens are remarkably darker and greyer than the older (50���100 years old) moths. In the male genitalia, the valvae of D. turcica (Figs 2 c���d) are somewhat shorter, more slender and tapered distally than those of D. delicatula (Figs 2 a���b), and the apical thorn of the cucullus is shorter and finer in D. turcica than in D. delicatula. In the female genitalia, the ductus bursae of D. turcica (Fig. 3 a) is broader than in D. delicatula (Fig. 3 b), having a more sclerotised posterior ring and anterior plate. In addition, the signum of the corpus bursae of D. turcica is thicker and continuous, not separated into two parts as in D. delicatula. Distribution. Deltote turcica is known from two rather remote areas in central Turkey (Ankara-Kayseri region; the records of D. delicatula published by Baisch et al. [1998] from the G��reme valley and ��rg��p, and by, Ko��ak and Kemal (2018) from Nevşehir and Kırıkkale probably refer to this species as well) and from the Kopet- Dagh area in Turkmenistan. Biology and Habitat. The populations of D. turcica occurring in the Kopet-Dagh Mts. inhabit dry rocky slopes with open steppe vegetation and trees limited to small shrubby patches. The Turkish populations live in similarly xerothermic rocky sites. Specimens were collected in June and July. Etymology. The epitheton name turcica refers to Turkey and Turkmenistan, the states, where the new species is discovered., Published as part of Seven, Erdem, Ronkay, L��szl�� & Ronkay, G��bor, 2019, Description of a new species of Deltote Reichenbach, Leipzig, 1817 (Lepidoptera, Noctuidae, Eustrotiinae) from Turkey and Turkmenistan, pp. 171-178 in Zootaxa 4576 (1) on pages 173-177, DOI: 10.11646/zootaxa.4576.1.10, http://zenodo.org/record/2624600, {"references":["Baisch, G., Beshkov, S. V., Gelbrecht, J., Hacker, H., Huber, K., Kallies, A., Kautt, P., Lobel, H., Lehmann, L. & Petersen, M. (1998) Bausteine zur Fauna der Noctuoidea der Turkei Mit einer Ubersicht aller in der Turkei bisher beobachteten Noctuoidea (Lepidoptera). Esperiana, 6, 213 - 373.","Kocak, A. O. & Kemal, M. (2018) A synonymous and distributional list of the species of the Lepidoptera of Turkey. '' Central entomological Studies Memoirs, 8, 1 - 487."]}

Genus Evonima Walker, 1865 Evonima Walker, 1865, List of the specimens of lepidopterous insects in the collection of the British Museum 32: 505. Type species: Evonima aperta Walker, 1865, Published as part of L��szl��, Gyula M. & Vetina, Alvaro A., 2019, Contribution to the knowledge of the Nolini of the Maputo Special Reserve in South Mozambique with description of two new species and several taxonomic updates (Lepidoptera, Nolidae, Nolinae), pp. 225-246 in Zootaxa 4571 (2) on page 244, DOI: 10.11646/zootaxa.4571.2.3, http://zenodo.org/record/2608276, {"references":["Walker, F. (1865) List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Part XXXII, Supplement 2, 505."]}

Nola politzari Hacker, 2012 (Figs 6, 7, 47, 68) Nola politzari Hacker, 2012, Esperiana 17: 75. Type locality: [Burkina Faso], Folonzo am Fluss, Comoe. Holotype male, in coll. ZSM. Material examined: Mozambique, Maputo Special Reserve. 2 ♂, 9m, Mangrove Camp, Mangrove-Woodland Mosaic, 26��19'35.9"S, 32��42'35.7"E, 7���9.XII.2016, Light Trap, Aristophanous, M., Cristovao, J., Laszlo, G., Miles, W. leg., slide No.: LGNA 329 ♂; 1 ♂, 15m, Ponta Milibangalala, Dune Grassland, 26��26���58.6���S, 32��55���29.8���E, 25���30.V.2017, MV Light Trap, Aristophanous, M., Laszlo, G., Miles, W., Vetina, A. leg.; 2 ♀, 22m, West Gate, Sand Forest, 26��30'14.2"S, 32��42'59.6"E, 30.V.���9. VI.2017, MV Light Trap, Aristophanous, M., Laszlo, G., Miles, W., Vetina, A. leg., slide Nos: LGNA 509 ♀, LGNA 510 ♀ (coll. ANHRT, MHNM). Remark. The species is very similar to Nola squalida Staudinger, 1870, and Nola afrotaeniata Hacker, 2012, the differences between the three species, however, are well expressed in the female genitalia (Hacker et al 2012). The concept of Hacker on the distribution of N. squalida seems to be correct considering the species being widely distributed from the West Mediterranean, throughout the Arabian peninsula towards Central Asia and the westernmost chains of the Himalayas. In the Subsaharan Africa N. squalida is replaced by its sister species N. afrotaeniata and N. politzari which are two evidently different species due to their female genitalia. The males are, however not distinguishable displaying in all three species the typical squalida -like genitalia configuration. Thus, it can be declared, that only the female specimens of the species group can satisfactorily be identified. The female specimens collected in the Maputo Special Reserve are identical in genitalia with the holotype of N. politzari described from Burkina Faso, thus the squalida -like species from South Mozambique is identified here as N. politzari, new country record for Mozambique representing the southernmost occurrence of the species known to date. Nevertheless it cannot be excluded, that N. afrotaeniata also occurs in South Mozambique., Published as part of L��szl��, Gyula M. & Vetina, Alvaro A., 2019, Contribution to the knowledge of the Nolini of the Maputo Special Reserve in South Mozambique with description of two new species and several taxonomic updates (Lepidoptera, Nolidae, Nolinae), pp. 225-246 in Zootaxa 4571 (2) on page 228, DOI: 10.11646/zootaxa.4571.2.3, http://zenodo.org/record/2608276, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612.","Staudinger, O. (1870) Beschreibung neuer Lepidopteren des europaischen Faunengebiets. Berliner entomologische Zeitschrift, 14, 102."]}

Mecothrix mineti Hacker, 2012 (Figs 13, 14, 51, 71) Nola (Mecothrix) mineti Hacker, 2012, Esperiana 17: 186. Type locality: Madagascar, Tananarive. Holotype male, in coll. MNHN. Material examined: Mozambique, Maputo Special Reserve. 1 ♂, 1 ♀, 18m, Hygrophilous Grass-Sand Forest Ecotone, 26��28'32.6"S, 32��45'7.7"E, 10.XII.2016, Light Trap, Aristophanous, M., Cristovao, J., Laszlo, G., Miles, W. leg., slide No.: LGNA 298 ♂, LGNA 317 ♀; 1 ♀, 22m, West Gate, Sand Forest, 26��30'14.2"S, 32��42'59.6"E, 3��� 13.XII.2016, Light Trap, Aristophanous, M., Cristovao, J., Laszlo, G., Miles, W. leg., slide No.: LGNA 299 ♀; 2 ♂, 1 ♀, 22m, West Gate, Sand Thicket, 26��30'14.2"S, 32��42'59.6"E, 21���30.XI.2016, Light Trap, Aristophanous, M., Cristovao, J., Laszlo, G., Miles, W. leg., slide Nos: LGNA 315 ♂, LGNA 316 ♂, LGNA 300 ♀; 1 ♀, 9m, Mangrove Camp, Mangrove-Woodland Mosaic, 26��19'35.9"S, 32��42'35.7"E, 7���9.XII.2016, Light Trap, Aristophanous, M., Cristovao, J., Laszlo, G., Miles, W. leg., slide No.: LGNA 304 ♀; 1 ♀, 17m, Futi Corridor (Sand Forest Woodland Mosaic), 26��32'10.1"S, 32��43'09.7"E, 23���24.ii.2018, MV Light Trap, Laszlo, G., Mulvaney, J., Smith, L. leg., slide No.: LGNA 432 ♀; 1 ♀, 22m, West Gate, Sand Forest, 26��30'14.2"S, 32��42'59.6"E, 21���22.ii.2018, Actinic Light Trap, Laszlo, G., Mulvaney, J., Smith, L. leg., slide No.: LGNA 434 ♀ (coll. ANHRT, MHNM). Remark. The species was described from Madagascar and considered to be endemic to the island. Its occurrence in South Mozambique is rather surprising, although it is worth noting that not all species which were found first in Madagascar are necessarily endemic to the island. Despite of a thorough study of the external and genital morphology of the specimens collected in the Maputo Special Reserve no significant difference was found in comparison with the Madagascan specimens. The most important diagnostic character in the species group is the configuration of the robust cornutus of vesica which was found to be completely identical in all Mozambican and Madagascan specimens. New country record for Mozambique., Published as part of L��szl��, Gyula M. & Vetina, Alvaro A., 2019, Contribution to the knowledge of the Nolini of the Maputo Special Reserve in South Mozambique with description of two new species and several taxonomic updates (Lepidoptera, Nolidae, Nolinae), pp. 225-246 in Zootaxa 4571 (2) on page 230, DOI: 10.11646/zootaxa.4571.2.3, http://zenodo.org/record/2608276, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Genus Nolidia van Son, 1933 Nolidia van Son, 1933, Annals of the Transvaal Museum 15: 229. Type species: Nolidia unipuncta van Son, 1933., Published as part of L��szl��, Gyula M. & Vetina, Alvaro A., 2019, Contribution to the knowledge of the Nolini of the Maputo Special Reserve in South Mozambique with description of two new species and several taxonomic updates (Lepidoptera, Nolidae, Nolinae), pp. 225-246 in Zootaxa 4571 (2) on page 228, DOI: 10.11646/zootaxa.4571.2.3, http://zenodo.org/record/2608276, {"references":["Son, van G. (1933) A revision of the South African moths of the tribe Nolini. Annals of the Transvaal Museum, 15 (2), 181 - 232."]}

Mecothrix aegyptiaca cana Hacker, 2012 stat. n. (Figs 11, 12, 50, 70) Nola (Mecothrix) cana Hacker, 2012, Esperiana 17: 158. Type locality: Mozambique, Magude. Holotype female in coll. ZSM. Material examined: Mozambique, Maputo Special Reserve. 1 ♂, 2 ♀, 9m, Mangrove Camp, Mangrove- Woodland Mosaic, 26��19'35.9"S, 32��42'35.7"E, 7���9.XII.2016, Light Trap, Aristophanous, M., Cristovao, J., Laszlo, G., Miles, W. leg., slide Nos: LGNA 322 ♂, LGNA 308 ♀, LGNA 321 ♀; 1 ♀, 22m, West Gate, Sand Forest, 26��30'14.2"S, 32��42'59.6"E, 3���13.XII.2016, Light Trap, Aristophanous, M., Cristovao, J., Laszlo, G., Miles, W. leg., slide No.: LGNA 318 ♀; 1 ♂, 22m, West Gate, Sand Thicket, 26��30'14.2"S, 32��42'59.6"E, 3���13.XII.2016, Light Trap, Aristophanous, M., Cristovao, J., Laszlo, G., Miles, W. leg., slide No.: LGNA 423 ♂; 1 ♀, 22m, West Gate, Sand Thicket, 26��30'14.2"S, 32��42'59.6"E, 21���30.XI.2016, Light Trap, Aristophanous, M., Cristovao, J., Laszlo, G., Miles, W. leg., slide No.: LGNA 319 ♀; 2 ♂, 2 ♀, 22m, West Gate, Sand Forest, 26��30'14.2"S, 32��42'59.6"E, 21���22.ii.2018, Actinic Light Trap, Laszlo, G., Mulvaney, J., Smith, L. leg., slide Nos: LGNA 435 ♂, LGNA 436 ♂, LGNA 424 ♀, LGNA 431 ♀; 1 ♂, 1 ♀, 22m, West Gate, Sand Forest, 26��30'14.2"S, 32��42'59.6"E, 13���15.ii.2018, MV Light Trap, Laszlo, G., Mulvaney, J., Smith, L. leg., slide Nos: LGNA 425 ♂, LGNA 426 ♀; 3 ♂, 1 ♀, 17m, Futi Corridor (Sand Forest Woodland Mosaic), 26��32'10.1"S, 32��43'09.7"E, 23���24.ii.2018, Actinic Light Trap, Laszlo, G., Mulvaney, J., Smith, L. leg., slide Nos: LGNA 427 ♂, LGNA 428 ♂, LGNA 429 ♂, LGNA 430 ♀; 1 ♀, 17m, Futi Corridor (Sand Forest Woodland Mosaic) 26��32'10.1"S, 32��43'09.7"E, 23���24.ii.2018, MV Light Trap, Laszlo, G., Mulvaney, J., Smith, L. leg., slide No.: 437♀; 1 ♂, 1 ♀, 22m, West Gate, Sand Thicket, 26��30'14.2"S, 32��42'59.6"E, 24���25.ii.2018, Actinic Light Trap, Laszlo, G., Mulvaney, J., Smith, L. leg., slide Nos: LGNA 433 ♀, LGNA 438 ♂; 1 ♂, 22m, Maputo Special Reserve, West Gate, Sand Thicket, 26��30'14.2"S, 32��42'59.6"E, 9���17.ii.2018, Actinic Light Trap, Laszlo, G., Mulvaney, J., Smith, L. leg., slide No.: LGNA 439 ♂ (coll. ANHRT, MHNM). Remark. M. cana was originally described as a distinct species. The detailed comparison of specimens collected in South Mozambique and Egypt revealed no significant differences in the configuration of genitalia, however the Southern African specimens are uniformly paler, having graphite greyish forewing ground colour, while the North African ones are much darker, with dark brownish forewing ground colour. The species might have had in the past a continuous distribution in Eastern Africa along the coast of Red Sea and Indian Ocean, later it had probably been splitted into morphologically slightly different populations (M. aegyptiaca sahelica Hacker, 2012 in the Sahel and Tanzania, and M. aegyptiaca bischoffi Hacker, 2012 in Yemen). Separating the southernmost population on species rank is not justified based on these minor morphological differences, so cana has herewith been placed within the species aegyptiaca as valid subspecies., Published as part of L��szl��, Gyula M. & Vetina, Alvaro A., 2019, Contribution to the knowledge of the Nolini of the Maputo Special Reserve in South Mozambique with description of two new species and several taxonomic updates (Lepidoptera, Nolidae, Nolinae), pp. 225-246 in Zootaxa 4571 (2) on page 229, DOI: 10.11646/zootaxa.4571.2.3, http://zenodo.org/record/2608276, {"references":["Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612."]}

Genus Afrasura Durante, 2009 Afrasura Durante, 2009, Zootaxa 2280: 30. Type species: Barsine indecisa Walker, 1869, by original designation., Published as part of Volynkin, Anton V. & L��szl��, Gyula M., 2018, A review of the genus Afrasura Durante, 2009 from S��o Tom�� Island with descriptions of a new species and a new subspecies (Lepidoptera: Erebidae: Arctiinae), pp. 237-248 in Zootaxa 4508 (2) on page 238, DOI: 10.11646/zootaxa.4508.2.6, http://zenodo.org/record/2607050, {"references":["Durante, A. (2009) Revision of the Afrotropical species of Asura Walker, 1854 (Lepidoptera: Arctiidae, Lithosiinae), with the description of a new genus. Zootaxa, 2280, 27 - 52."]}

Afrasura discocellularis saotomensis Volynkin & L��szl��, ssp. nov. (Figs. 9, 10, 19, 26) Type material. Holotype (Figs. 9, 19): ♂, ��� S��o Tom��, 482m, Roca Bombaim (montane forest-plantation), 0��14���41.7������N, 6��37���59.8������E, 27���31.I.2018, MV Light Trap, Aristophanous, M., Miles, W. leg., ANHRT:2018.1��� / ���ANHRTUK 00019763��� / slide AV4546 ♂ A. Volynkin ��� (Coll. ANHRT). Paratypes. S��O TOM��: 8 ♂, with the same data as the holotype, slide No.: AV4545 ♂ (coll. ANHRT); 4 ♂, 1303m, Antenna Bom Successo, 00��16���33.7���N, 6��36���19.7���E, 20.X.2016, MV, primary upland forest with a narrow recent plantation margin, C. R. Turner & T. Tasane leg., ANHRT: 2017.21 (coll. ANHRT); 1 ♂, 2 ♀, 174m, Guadalupe, 00��23���12.8������N, 6��37���6.9������E, 22.X.2016, savanna dry mosaic grassland/forest scrub/plantation, MV, C. R. Turner & T. Tasane, leg., ANHRT:2017.21, slide No.: AV4400 ♀ (coll. ANHRT); 1 ♂, 186m, Ponta Furada, 0��14���01.8���N, 6��28���15.5���E, 24.X.2016, MV Light Trap, Turner, C.R. & Tasane, T. leg., slide No.: AV3324 ♂ (coll. ANHRT); 1 ♂, 635m, Ponta Figo, 0��17���12.7���N, 6��34���14.5���E, 27.X.2016, MV Light Trap, Turner, C.R. & Tasane, T. leg. (coll. ANHRT); 1 ♀, 264m, Guadalupe, Antenna, 00��22���49.9���N, 6��38���47.4���E, 31.X.2016, MV, Dry coastal forest and scrub, C. R. Turner & T. Tasane leg. ANHRT 2017.21 (Coll. ANHRT). Diagnosis. Afrasura discocellularis saotomensis ssp. nov. (Figs. 9, 10) differs from the nominate subspecies occuring widespread in West Africa (Figs. 11, 12) by its more yellowish forewing ground colour (while that of A. d. discocellularis is pale orange) and grey forewing pattern what is brown with a reddish suffusion in the mainland taxon. The male genitalia of the two subspecies are very similar, but the distal section of the costa is somewhat narrower and more elongated and the distal costal process is slightly longer in A. d. saotomensis ssp. nov. (Fig. 19) compared to those of the nominate subspecies, in addition the distal saccular process of the new subspecies is apically narrowly rounded, while that is sharply pointed in A. d. discocellularis (Fig. 20). The female genitalia of A. d. saotomensis ssp. nov. (Fig. 26) differs from that of A. d. discocellularis (Fig. 27) by its slightly shorter antrum, and somewhat larger signum bursae. Description. Adult (Figs. 9, 10). Forewing length 10���11 mm in males and 10���12 mm in females. Antennae of both sexes ciliate. Head bright yellow, frons with grey spot; thorax brownish yellow with four grey spots on tegulae; abdomen paler than thorax and head, light yellow. Forewing ground color bright yellow with some brownish shade, pattern grey; basal dot present; subbasal line interrupted, consisted of a medial round spot and a longitudinal streak on costa; antemedial line double, its outer part irregularly dentate, inner part dentate and interrupted into three spots; medial line irregularly dentate, forming a loop inwards the cell spot; postmedial line dentate, curved inwards between costa and cell; subterminal line interrupted, consisted of several diffuse spots; discal spot present, dot-like; cilia brownish yellow. Hindwing considerably lighter than forewing, pale yellow, without pattern; cilia pale yellow. Male genitalia (Fig. 19). Uncus medium long, slender, curved, apically pointed; tuba analis broad, scaphium thin, weakly sclerotized; tegumen medium broad, rather elongate, moderately sclerotized; juxta broad, with two broad basal lobes; vinculum relatively short, V-shaped, apically rounded; transtillae fused, forming a broad plate with two short apical lobes; valva elongated, broadened medially and strongly tapering distally; costa moderately sclerotized, curved medially, its distal section elongated, narrow, with thorn-like apical process; sacculus narrow, with well developed, finger-like, slightly arcuate, apically rounded distal process. Aedeagus tubular, relatively short and broad, coecum rounded, slightly broadened; vesica short, strongly granulated medially, with two granulated ventral diverticula and robust thorn-like cornutus with broad and curved base. Female genitalia (Fig. 26). Ovipositor short and broad; apophyses long and thin, apophyses anteriores slightly longer than apophyses posteriores; ostium bursae very broad; antrum heavily sclerotized, caliciform; ductus bursae very short and broad, membranous; posterior section of corpus bursae relatively narrow, weakly sclerotized, rugose; anterior section of corpus bursae rounded, covered with dense spinulose scobination, signum bursae small, rounded; appendix bursae broad and short, with a broad and long cluster of short spinules and a trigonal, heavily sclerotised process. Distribution. The new subspecies is known so far only from S��o Tom�� Island., Published as part of Volynkin, Anton V. & L��szl��, Gyula M., 2018, A review of the genus Afrasura Durante, 2009 from S��o Tom�� Island with descriptions of a new species and a new subspecies (Lepidoptera: Erebidae: Arctiinae), pp. 237-248 in Zootaxa 4508 (2) on page 240, DOI: 10.11646/zootaxa.4508.2.6, http://zenodo.org/record/2607050

Dendrobaena ressli Zicsi, 1973 Dendrobaena ressli Zicsi, 1973: 220. Csuzdi et al. 2006: 14. Material examined. HNHM /17057 3 ex., No. 25, Yeşilova, Kastamonu, SW of the village, pasture with open oak and Juniperus forest, N41��13.169' E32��59.585', 477 m, leg. L. D��nyi, M.B. Kaydan, 22.0 4.2015., Published as part of Szederjesi, T��mea, D��nyi, L��szl��, Kaydan, Mehmet Bora & Csuzdi, Csaba, 2018, Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili), pp. 160-172 in Zootaxa 4496 (1) on page 164, DOI: 10.11646/zootaxa.4496.1.11, http://zenodo.org/record/1446737, {"references":["Zicsi, A. (1973) Regenwurmer (Oligochaeta: Lumbricidae) aus der Turkei. Acta Zoologica Hungarica, 19, 217 - 232.","Csuzdi, Cs., Zicsi, A. & Misirlioglu, M. (2006) An annotated checklist of the earthworm fauna of Turkey (Oligochaeta: Lumbricidae). Zootaxa, 1175, 1 - 29."]}

Aporrectodea caliginosa caliginosa (Savigny, 1826) Enterion caliginosum Savigny, 1826: 180. Aporrectodea caliginosa (part.): Csuzdi et al. 2006: 4. Material examined. HNHM/17065 1 ex., No. 35, G��zelim (N of the village), Tufanbeyli, Adana, under Pinus and Quercus, N38��08.922' E36��10.718', 1427 m, leg. M.B. Kaydan, 0 1.0 5.2015., Published as part of Szederjesi, T��mea, D��nyi, L��szl��, Kaydan, Mehmet Bora & Csuzdi, Csaba, 2018, Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili), pp. 160-172 in Zootaxa 4496 (1) on page 161, DOI: 10.11646/zootaxa.4496.1.11, http://zenodo.org/record/1446737, {"references":["Savigny, J. C. (1826) Analyse des Travaux de l'Academie royale des Sciences, pendant l'annee 1821, partie physique. Memoires de l'Academie des Sciences de l'Institut de France Paris, 5, 176 - 184.","Csuzdi, Cs., Zicsi, A. & Misirlioglu, M. (2006) An annotated checklist of the earthworm fauna of Turkey (Oligochaeta: Lumbricidae). Zootaxa, 1175, 1 - 29."]}

Lumbricus rubellus Hoffmeister, 1843 Lumbricus rubellus Hoffmeister, 1843: 187. Csuzdi et al. 2006: 23. Material examined. HNHM /17054 2 ex., No. 10, G��lc��k, Kocaeli, S of the city, deciduous oak forest, N40��41.435' E29��52.673', 187 m, leg. L. D��nyi, M.B. Kaydan, 20.0 4.2015. HNHM /17075 6 ex., No. 27, Yukarı��aylı, Dadalı, Kastamonu, W of the village, chestnut orchard, N41��55.527' E33��44.430', 479 m, leg. L. D��nyi, M.B. Kaydan, 23.0 4.2015. HNHM/17088 4 ex., HNHM /17102 4 ex., No. 13, Ak��akoca, D��zce, southern edge of the city, hazelnut plantation, N41��05.119' E31��09.819', 58 m, leg. L. D��nyi, M.B. Kaydan, 21.0 4.2015. HNHM /17098 3 ex., No. 24, Ovacuma, Karab��k, SW of the village, disturbed Acer -pine-oak forest, N41��25.649' E32��44.564', 490 m, leg. L. D��nyi, M.B. Kaydan, 22.0 4.2015. HNHM /17110 3 ex., No. 18, Nebioğlu, Zonguldak, W of the village, old bog-forest, N41��26.026' E32��12.630', 246 m, leg. L. D��nyi, M.B. Kaydan, 21.0 4.2015. HNHM /17119 1 ex., No. 22, Bah��ecik, Bartın, NW of the village, stream bank with Platanus gallery, N41��28.406' E32��38.025', 248 m, leg. L. D��nyi, M.B. Kaydan, 22.0 4.2015. HNHM /17125 1 ex., No. 7, Yenik��y, Bursa, SE of the village, machia of deciduous oaks, N40��18.833' E29��27.800', 360 m, leg. L. D��nyi, M.B. Kaydan, 20.0 4.2015., Published as part of Szederjesi, T��mea, D��nyi, L��szl��, Kaydan, Mehmet Bora & Csuzdi, Csaba, 2018, Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili), pp. 160-172 in Zootaxa 4496 (1) on page 168, DOI: 10.11646/zootaxa.4496.1.11, http://zenodo.org/record/1446737, {"references":["Csuzdi, Cs., Zicsi, A. & Misirlioglu, M. (2006) An annotated checklist of the earthworm fauna of Turkey (Oligochaeta: Lumbricidae). Zootaxa, 1175, 1 - 29."]}

Dendrobaena fridericae uludagi Omodeo & Rota, 1991 Dendrobaena fridericae uludagi Omodeo & Rota, 1991: 179. Csuzdi et al. 2006: 10. Material examined. HNHM /17104 1 ex., No. 13, Ak��akoca, D��zce, southern edge of the city, hazelnut plantation, N41��05.119' E31��09.819', 58 m, leg. L. D��nyi, M.B. Kaydan, 21.0 4.2015. Remarks. The type locality of this subspecies is a bit obscure. In the original description Omodeo & Rota (1991) just gave Loc. 7 (1 adult and 1 juv. spec.) which refers to ��ankiri (10 km North of ��ankiri). However, this was a typing error for Loc. 73 (Emilia Rota pers. comm.) referring to the Northern slope of Uludağ, 10 km before the entrance to the National Park. This is the first report of D. fridericae uludagi since the original description from some 200 km N of the supposed type locality., Published as part of Szederjesi, T��mea, D��nyi, L��szl��, Kaydan, Mehmet Bora & Csuzdi, Csaba, 2018, Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili), pp. 160-172 in Zootaxa 4496 (1) on page 163, DOI: 10.11646/zootaxa.4496.1.11, http://zenodo.org/record/1446737, {"references":["Omodeo, P. & Rota, E. (1991) Earthworms of Turkey. II. Bolletino di Zoologia, 58, 171 - 181. https: // doi. org / 10.1080 / 11250009109355749","Csuzdi, Cs., Zicsi, A. & Misirlioglu, M. (2006) An annotated checklist of the earthworm fauna of Turkey (Oligochaeta: Lumbricidae). Zootaxa, 1175, 1 - 29."]}

Dendrobaena persimilis Omodeo & Rota, 1989 Dendrobaena persimilis Omodeo & Rota, 1989: 193. Csuzdi et al. 2006: 12. Material examined. HNHM /17052 1 ex., No. 9, Hayriye, Kocaeli, N of the village, young deciduous oak forest, N40��38.590' E29��34.514', 284 m, leg. L. D��nyi, M.B. Kaydan, 20.0 4.2015. HNHM/17053 1 ex., HNHM /17053 1 ex., No. 10, G��lc��k, Kocaeli, S of the city, deciduous oak forest, N40��41.435' E29��52.673', 187 m, leg. L. D��nyi, M.B. Kaydan, 20.0 4.2015. HNHM /17078 1 ex., No. 27, Yukarı��aylı, Dadalı, Kastamonu, W of the village, chestnut orchard, N41��55.527' E33��44.430', 479 m, leg. L. D��nyi, M.B. Kaydan, 23.0 4.2015. HNHM /17101 6 ex., No. 14, Dadali, Ak��akoca, D��zce, E of the village, disturbed beech forest patch among agricultural fields, N41��03.860' E31��12.002', 71 m, leg. L. D��nyi, M.B. Kaydan, 21.0 4.2015. Remarks. D. persimilis was described from two localities, Uludağ and Trabzon. Our new samples are from the Black Sea region just in between the two original localities., Published as part of Szederjesi, T��mea, D��nyi, L��szl��, Kaydan, Mehmet Bora & Csuzdi, Csaba, 2018, Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili), pp. 160-172 in Zootaxa 4496 (1) on page 164, DOI: 10.11646/zootaxa.4496.1.11, http://zenodo.org/record/1446737, {"references":["Omodeo, P. & Rota, E. (1989) Earthworms of Turkey. Bolletino di Zoologia, 56, 167 - 199. https: // doi. org / 10.1080 / 11250008909355639","Csuzdi, Cs., Zicsi, A. & Misirlioglu, M. (2006) An annotated checklist of the earthworm fauna of Turkey (Oligochaeta: Lumbricidae). Zootaxa, 1175, 1 - 29."]}

Dendrobaena pavliceki Szederjesi & Csuzdi, sp. nov. (Figure 2) Holotype. HNHM /17260 1 adult ex., No. 40, Uzun Ahmetk��y, Erzurum, NW of the village, pasture, N39��57.710' E41��24.461', 1855 m, leg. L. D��nyi, M.B. Kaydan, 26.0 4.2015. Paratypes. HNHM /17082 4 adult ex., NHMA 1 adult ex., locality and date same as that of the Holotype. Derivatio nominis. The new species is named in honour of our colleague Tom���� Pavl��ček (University of Haifa) who has collected rich earthworm materials all over the world and especially in the East Mediterranean. Diagnosis. Length 36���48 mm, diameter 3���4 mm, setae distant. Colour pale with slight reddish hints on dorsum at the head. First dorsal pore in 5/6. Clitellum on 25, ��25���33, tubercles on 1/n30�����32. Male pores on 15, surrounded by glandular crescents. Nephridial pores irregularly alternate between b and above d. Three pairs of vesicles in 9, 11, 12, the last pair vestigial. Spermathecae two pairs in 9/10, 10/11, open near d. Calciferous glands in ��10���11. Hearts in segments 6���11, nephridial bladders simple, sausage-shaped. External characters. Holotype 42 mm long and 4 mm wide. Number of segments 132. Paratypes 36���48 mm long and 3���4 mm wide. Number of segments 128���140. Colour pale with slight reddish hints in the head region. Prostomium tanylobous. First dorsal pore at intersegmental furrow 5/6. Setae distant. Setal arrangement behind clitellum: aa: ab: bc: cd: dd = 1.6:1.1:1.7:1:4.6 (Fig. 2A). Male pores on segment 15, surrounded by glandular crescents, protruding into the neighbouring segments. Nephridial pores irregularly alternate between setal line b and above d. Clitellum on 25, ��25���33. Tubercula pubertatis on 1/n30, 31�����32. Glandular tumescence on 10, 26��� 27 ab (Fig. 2B). Internal characters (based on holotype and two paratypes). Septa 6/7��� 8/9 and 11/12 ���12/13 slightly thickened, 13/14���14/15 thickened. Testes and funnels paired in 10���11. Three pairs of seminal vesicles in 9, 11 and 12, the last pair vestigial. Spermathecae two pairs in 9/10, 10/11 with external openings near setal line d. Calciferous glands in ��10���11. Last heart in 11, with a pair of extraoesophageal vessels in 12. Nephridial bladders simple, sausage-shaped. Crop in 15���16, gizzard in 17���18. Typhlosolis bilobate. Longitudinal musculature of pinnate type. Remarks. The new species with tubercles in the region of 31���32 seems to be close to D. nevoi Csuzdi & Pavl��ček, 1999 and D. cevdeti Szederjesi, Pavl��ček, Coşkun & Csuzdi, 2014. D. pavliceki differs from D. nevoi first of all in the position of the last pair of hearts (11 vs. 10), the presence of an extraoesophageal vessel, in its colour (pale with slight reddish hints at the head vs. overall dark red) and in the position of the clitellum (25, ��25���33 vs. 23, 24���33). The new species differs from D. cevdeti in the position of the clitellum (25, ��25���33 vs. ��24���34), in the presence of an extraoesophageal vessel and in the head (tanylobous vs. epilobous). With lack of vesicles in 10 and vestigial vesicles in 12, D. pavliceki shows an incomplete metandric reduction., Published as part of Szederjesi, T��mea, D��nyi, L��szl��, Kaydan, Mehmet Bora & Csuzdi, Csaba, 2018, Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili), pp. 160-172 in Zootaxa 4496 (1) on page 165, DOI: 10.11646/zootaxa.4496.1.11, http://zenodo.org/record/1446737, {"references":["Szederjesi, T., Pavlicek, T., Coskun, Y. & Csuzdi, Cs. (2014 a) New earthworm records from Turkey, with description of three new species (Oligochaeta: Lumbricidae). Zootaxa, 3764 (5), 555 - 570. https: // doi. org / 10.11646 / zootaxa. 3764.5.4"]}

Dendrobaena taurica Szederjesi & Csuzdi, sp. nov. (Figure 3) Holotype. HNHM/17261 1 adult ex., No. 29, Arslank��y, Mersin, SW of the town, under Pinus and Cedrus, N36��59.265' E34��14.818', 1555 m, leg. M.B. Kaydan, 0 1.0 5.2015. Paratypes. HNHM /17061 1 adult ex., NHMA 1 adult ex., locality and date same as that of the Holotype. Derivatio nominis. The new species is named after its type locality which is in the Taurus Mts, Turkey. Diagnosis. Length 45���46 mm, diameter 3���3.5 mm, setae distant. Colour pale, pigmentation lacking. First dorsal pore in 5/6. Clitellum on 25���34, tubercles on 30���32. Male pores on 15, surrounded by glandular crescents. Nephridial pores hardly visible. Three pairs of vesicles in 9, 11, 12, the first pair small. Spermathecae two pairs in 9/10, 10/11, open near the mid-dorsal line. Calciferous glands in 11���12. Last heart in 11, nephridial bladders biscuit-shaped. External characters. Holotype 45 mm long and 3 mm wide. Number of segments 143. Paratype 46 mm long and 3���3.5 mm wide. Number of segments 146. Colour pale, pigmentation lacking. Prostomium epilobous �� closed. First dorsal pore at intersegmental furrow 5/6. Setae distant. Setal arrangement behind clitellum: aa: ab: bc: cd: dd = 2.5:1.25:1.62:1:3.12 (Fig. 3A). Male pores on 15, surrounded by glandular crescents, protruding slightly into the neighbouring segments. Nephridial pores hardly seen, the detected ones are aligned just above setal line b. Clitellum on 25���34. Tubercula pubertatis on 30���32. Glandular tumescence on 9, 11 abcd (Fig. 3B). Internal characters (based on holotype and one paratype). No dissepiments notably thickened. Testes and funnels paired in 10���11. Three pairs of seminal vesicles in 9, 11, and 12, the first pair is small. Spermathecae two pairs in 9/10, 10/11 with external openings near the mid-dorsal line (M). Calciferous glands in 11���12. Last hearts in segment 11, with a pair of extraoesophageal vessels in 12. Nephridial bladders biscuit-shaped (octaedra-type). Crop in 15���16, and gizzard in 17���18. Typhlosolis bilobate. Longitudinal musculature is of pinnate type. Remarks. The new species with pale colour, pinnate musculature and the presence of extraoesophageal vessels seems to be most close to D. szalokii Szederjesi, Pavl��ček, Coşkun & Csuzdi, 2014 but differs from it in much smaller body dimensions (45���46 vs. 90���123 mm), the lack of thickened septa, and in the position of the clitellar organs (cl. 25���34, tb. 30���32 vs. cl. 24�����34, tb. ��29�����32)., Published as part of Szederjesi, T��mea, D��nyi, L��szl��, Kaydan, Mehmet Bora & Csuzdi, Csaba, 2018, Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili), pp. 160-172 in Zootaxa 4496 (1) on pages 166-167, DOI: 10.11646/zootaxa.4496.1.11, http://zenodo.org/record/1446737, {"references":["Szederjesi, T., Pavlicek, T., Coskun, Y. & Csuzdi, Cs. (2014 a) New earthworm records from Turkey, with description of three new species (Oligochaeta: Lumbricidae). Zootaxa, 3764 (5), 555 - 570. https: // doi. org / 10.11646 / zootaxa. 3764.5.4"]}

Healyella boluana Omodeo & Rota, 1989 Healyella boluana Omodeo & Rota, 1989: 176. Csuzdi et al. 2006: 20. Material examined. HNHM /17095 3 (2 truncated and 1 intact) ex., No. 15, Ak��a��ren, D��zce, above the lake S of the village, open forest with deciduous oak and Platanus, N40��55.721' E31��17.392', 296 m, leg. L. D��nyi, M.B. Kaydan, 21.0 4.2015. HNHM /17097 1 truncated ex., No. 19, Kazpınarı, Bartın, N of the village, rocky slope with bushes, N41��42.634' E32��21.811', 319 m, leg. L. D��nyi, M.B. Kaydan, 21.0 4.2015. External characters. 62 mm long and 4 mm wide. Number of segments 116. Slight purplish pigmentation dorsally. Prostomium tanylobous. First dorsal pore at intersegmental furrow 6/7. Setae distant. Setal arrangement behind clitellum: aa: ab: bc: cd: dd = 1.27:1.8:1:1.13:1.8. Male pores on 15 between a and b, spherical, slightly invaginated, with large circular, glandular tumescences. Female pore not visible, covered by flat spermatophores. Nephridial pores irregularly alternate between setal line b and above d. Clitellum annular on 26���30. Tubercula pubertatis lacking. Internal characters (based on one specimen). Septa 6/7���14/15 slightly thickened. Testes and funnels paired in 10���11, covered by perioesophageal testic sacs. Two pairs of seminal vesicles in 11 and 12. Spermathecae lacking. Calciferous glands in 10���12 with small diverticula in the posterior part of 10. Last heart in 11, with a pair of extraoesophageal vessels in 12. Nephridial bladders simple, sausage-shaped. Crop in 15���16, gizzard in 17���18, occupying the cavity of 19 as well. Typhlosolis large, anchor-shaped. Longitudinal musculature of pinnate type. Remarks. He. boluana was described on a single specimen from the surroundings of Bolu city (Boludagi Ge��idi). Our specimens are from some 60 (No. 15) and 120 (No. 19) km North-East of the type locality. Our specimens completely agree with the original description, except the position of the nephridial pores: Omodeo & Rota (1989) state that they are aligned just above setal line b, in our specimens they alternate irregularly between b and above d., Published as part of Szederjesi, T��mea, D��nyi, L��szl��, Kaydan, Mehmet Bora & Csuzdi, Csaba, 2018, Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili), pp. 160-172 in Zootaxa 4496 (1) on pages 167-168, DOI: 10.11646/zootaxa.4496.1.11, http://zenodo.org/record/1446737, {"references":["Omodeo, P. & Rota, E. (1989) Earthworms of Turkey. Bolletino di Zoologia, 56, 167 - 199. https: // doi. org / 10.1080 / 11250008909355639","Csuzdi, Cs., Zicsi, A. & Misirlioglu, M. (2006) An annotated checklist of the earthworm fauna of Turkey (Oligochaeta: Lumbricidae). Zootaxa, 1175, 1 - 29."]}

Allolobophora chlorotica chlorotica (Savigny, 1826) Enterion chloroticum Savigny, 1826: 182. Allolobophora chlorotica: Zicsi & Michalis 1981: 248; 1993: 302. Michalis 1982: 349; 1987: 62. Michalis et al. 1985: 38; 1989: 5. Szederjesi & Csuzdi 2012: 29. Allolobophora chlorotica chlorotica: Szederjesi 2017. Widely introduced peregrine species (Csuzdi & Zicsi 2003). Distribution in Greece. Mainland: Thessaloniki, Kastoria, Naoussa, Imathia, Florina, Grevena, Kilki, Velvendos, Aiginio, Katerini (Michalis 1972). Florina (��apkarev 1972). Kaligoni, Patras (Zicsi 1973). Konitsa, Filiates, Metsovo, Megalo Peristeri (Zicsi 1974). Veroia (Michalis 1975a). Trikala, Agiokampos, Sykourion, Karditsa, Mouzaki (Michalis 1976). Lyggiades, Stavraki, Mpizani, Katsikas, Perama, Agios Ioannis, Vryssi Pappa, Eleousa, Kastritsa, Anatoli, Ioannina, Korinthos, Meteora, Deskati, Foteina, Nea Pella, Trikala, Edessa, Agras (Zicsi & Michalis 1981). Kato Milea, Palaeogratsano (Michalis 1983). Agia Foteini, Kerassies, Giannakochori, Agios Nikolaos, Agios Athanasios, Militsa, Rhodochori, Naoussa, Karydies, Fyteia (Michalis et al. 1985). Soufli, between Rizia and Orestiada (Michalis 1987). Aridea, Mavrovouni, Polycarpi, Sevastiana, Agras, Limnochori, Edessa, Logos (Michalis et al. 1989). Meteora, Kastraki, Korydallos, Larissa, Metsovo, Ktima Despoti, Lyggiades, Karyes, Konitsa, Kalpaki, Voidomatis, Nea Apollonia (Zicsi & Michalis 1993). Athamania (Szederjesi & Csuzdi 2012). Lefkas: Kaligoni, Fryni (Zicsi 1973). Kos: Asfendiou (Michalis 1975b, 1983). Crete: Garipa, Skalani (Szederjesi 2017)., Published as part of Szederjesi, T��mea, Vavoulidou, Evangelia, Chalkia, Christina, D��nyi, L��szl�� & Csuzdi, Csaba, 2017, An annotated checklist of earthworms of Greece (Clitellata: Megadrili), pp. 57-82 in Zootaxa 4272 (1) on page 58, DOI: 10.11646/zootaxa.4272.1.3, http://zenodo.org/record/583379, {"references":["Savigny, J. C. (1826) G. Cuvier: Analyse des Travaux de l'Academie royale des Sciences, pendant l'annee 1821, partie physique. Memoires de l'Academie des Sciences de l'Institut de France Paris, 5, 176 - 184.","Zicsi, A. & Michalis, K. (1981) Ubersicht der Regenwurm-fauna Griechenlands (Oligochaeta: Lumbricidae). Acta Zoologica Hungarica, 27, 239 - 264.","Michalis, K. (1982) Katalog der Oligochaetenfauna Griechenlands. Biologia Gallo-Hellenica, 9, 343 - 362.","Michalis, K., Fragoulis, A. & Panidis, S. (1985) Earthworms (Oligochaeta: Lumbricidae) from the montainous mas of Pella province: southwest region, and Imathias province, northwest region: a systematical and ecological study. Ecologia Mediterranea, 11 (4), 37 - 42.","Szederjesi, T. & Csuzdi, Cs. (2012) New and little known earthworm species form Greece (Oligochaeta: Lumbricidae, Acanthodrilidae). Zootaxa, 3304, 25 - 42.","Csuzdi, Cs. & Zicsi, A. (2003) Earthworms of Hungary (Annelida: Oligochaeta; Lumbricidae). In: Csuzdi, Cs. & Mahunka, S. (Eds.), Pedozoologica Hungarica. 1. Hungarian Natural History Museum, Budapest, pp. 1 - 271.","Michalis, K. (1972) Symvoli is tin meletin to Oligochaeton tis Makedonias. PhD Dissertation, University of Thessaloniki, Thessaloniki, 98 pp.","Sapkarev, J. (1972) Beitrage zur Kenntnis der Lumbricidenfauna Griechenlands. Macedonian Academy of Sciences and Arts, 4, 31 - 36.","Zicsi, A. (1973) Regenwurmer (Oligochaeta: Lumbricidae) aus Griechenland. Opuscula Zoologica Budapest, 12, 99 - 103.","Zicsi, A. (1974) Ein neue Dendrobaena - Art (Oligochaeta: Lumbricidae) aus Griechenland. Acta zoologica hungarica, 20, 449 - 451.","Michalis, K. (1975 a) Neue Angaben zur Fauna der Oligochaeten von Mazedonien. Scientific Annals, Faculty of Physics and Mathematics, University of Thessaloniki, 15, 201 - 211.","Michalis, K. (1976) Beitrag zur Kenntnis der Oligochaetenfauna Thessaliens. Scientific Annals, Faculty of Physics and Mathematics, University of Thessaloniki, 16, 153 - 168.","Michalis, K. (1983) New biotopes of the species Allolobophora dofleini (Ude, 1922) and the subspecies Bimastus antiquus bouchei (Zicsi-Michalis, 1981) (Oligochaeta-Lumbricidae). Ecologia Mediterranea, 9, 57 - 61.","Michalis, K. (1987) Contribution to the systematic and ecological study of the earthworms (Oligochaeta, Lumbricidae) of the Western Thrace. Arquivos do Museu Bocage, 1 (5), 59 - 69.","Michalis, K., Fragoulis, A. & Panidis, S. (1989) Notes on the earthworms (Oligochaeta, Lumbricidae) from central west Macedonia. Acta zoologica cracoviensia, 32, 3 - 14.","Zicsi, A. & Michalis, K. (1993) Zwei neue Dendrobaena - Arten aus Griechenland (Oligochaeta: Lumbricidae). Acta Zoologica Hungarica, 39, 301 - 310.","Michalis, K. (1975 b) Die Oligochaeten der Insel Kos. Scientific Annals, Faculty of Physics and Mathematics, University of Thessaloniki, 15, 187 - 200."]}

Octodrilus complanatus (Dug��s, 1828) Lumbricus complanatus Dug��s, 1828: 289. Octodrilus complanatus: Zicsi & Michalis 1981: 256; 1993: 305. Michalis 1982: 357; 1987: 64; 1993: 18. Michalis et al. 1985: 39; 1989: 5. Szederjesi & Csuzdi 2012: 38. Szederjesi 2015: 148; 2017. Octodrilus peleensis Michalis, 1995: 16. (Szederjesi & Csuzdi 2012). Circum-Mediterranean species (Pavl��ček & Csuzdi 2016). Distribution in Greece. Mainland: Mt. Parnes (Michaelsen 1928). Edessa, Kastoria, Pelion, Argalasti, Milies, Parnis, Penteli, Phaliron, Malakasa, Mpogiati, Pikermi, Kessiriani (Tzelepe 1943). Attika (Omodeo 1955). Episkopi, Agios Germanos, Aridaea, Giannitsa, Athytos, Litochoron (Michalis 1972). Pidima, Litochoron, Kamena Vourla, Epidavros, Giannouleika, Paleochorion, Korakolithos, Delphi, Itea, Elaeon, Gravia, Edessa, Serrae, Drama, Milina Magnesia, Platanias, Katerini (Zicsi & Michalis 1981). Agios Nikolaos, Militsa (Michalis et al. 1985). Between rizia and Orestiada, between sitaria and Didymoteichou, Xanthi, between Psathades and Didymoteichou, between Iasmos and Komotini, between Peplos and Soufli (Michalis 1987). Aridea, Rizari (Michalis et al. 1989). Thessaloniki (Michalis 1993). Tempi, Litochoron, Chortiatis (Zicsi & Michalis 1993). Anatoliki Fragista, Vouliasta, Mesimvria, Avantas (Szederjesi & Csuzdi 2012). Loutra Traianoupolis (Szederjesi 2015). Peloponnese: Kalavryta, Argos (Černosvitov 1938). Argos, Kalavryta, Pyrgos (Tzelepe 1943). Kalavryta, Patra (Zicsi 1973). Solomos, Tyruns, Mystras, Mykene, Kaza (Zicsi & Michalis 1981). Magouliana, Korfes, Mystras, Garea, Agios Apostoli, Agios Petros, Planitero (Szederjesi & Csuzdi 2012). Crete: Omalos, Rethymno, Chania, Dafnes, Aselakia, Askifou, Plateau, Tsikalaria, Vizari (Michaelsen 1902). Kristallenia (Cognetti 1906). Rethymno, Heraklion (Tzelepe 1943). Axos, Agii Deka, Sisarcha, Moni Veni, Apostoli, Goulediana, Katharo (Szederjesi 2015). Anogia, Kampanos, Omalos, Piso Moni Preveli, Loutraki Kato Simis, Platanias, Males, Thripti (Szederjesi 2017). Korfu. Samos (Tzelepe 1943). Lefkas. Kefalonia (Zicsi 1973). Lesvos: Neapolis, Panagouda, Vryssa, Mytilini, Geras. Chios. Leros: Partheni. Kos: Asfendiou, Kardamena. Athos (Zicsi & Michalis 1981). New records. HNHM /17170 1 ex., Rhodes Island, Eleousa, W of the vollage, spring among Platanus trees, 36��16.452'N, 27��59.815'E, 336 m a.s.l., leg. L. D��nyi, 0 3.0 4.2016. HNHM /17185 1 ex., Rhodes Island, Salakos, S side of Mt. Kako Vouni, Cupressus - Pinus brutia forest, 36��16.562'N, 27��55.966'E, 673 m a.s.l., leg. L. D��nyi, 0 3.0 4.2016., Published as part of Szederjesi, T��mea, Vavoulidou, Evangelia, Chalkia, Christina, D��nyi, L��szl�� & Csuzdi, Csaba, 2017, An annotated checklist of earthworms of Greece (Clitellata: Megadrili), pp. 57-82 in Zootaxa 4272 (1) on page 72, DOI: 10.11646/zootaxa.4272.1.3, http://zenodo.org/record/583379, {"references":["Duges, A. (1828) Recherche sur la circulation, la respiration, et la reproduction des Annelides setigeres abranches. Annales des Sciences Naturelles Paris, 15, 284 - 336.","Zicsi, A. & Michalis, K. (1981) Ubersicht der Regenwurm-fauna Griechenlands (Oligochaeta: Lumbricidae). Acta Zoologica Hungarica, 27, 239 - 264.","Michalis, K. (1982) Katalog der Oligochaetenfauna Griechenlands. Biologia Gallo-Hellenica, 9, 343 - 362.","Michalis, K., Fragoulis, A. & Panidis, S. (1985) Earthworms (Oligochaeta: Lumbricidae) from the montainous mas of Pella province: southwest region, and Imathias province, northwest region: a systematical and ecological study. Ecologia Mediterranea, 11 (4), 37 - 42.","Szederjesi, T. & Csuzdi, Cs. (2012) New and little known earthworm species form Greece (Oligochaeta: Lumbricidae, Acanthodrilidae). Zootaxa, 3304, 25 - 42.","Szederjesi, T. (2015) New earthworm records from various parts of Greece (Oligochaeta: Lumbricidae, Acanthodrilidae, Megascolecidae, Ocnerodrilidae). Opuscula Zoologica Budapest, 46 (2), 143 - 152.","Michalis, K. (1995) Oligochaeten - Funde aus Thessalien (Griechenland) nebst Beschreibung der Art Octodrilus peleensis sp. nov. Bios (Macedonia, Greece), 3, 15 - 20.","Pavlicek, T. & Csuzdi, Cs. (2016) Chapter 23: Clitellata: Oligochaeta: Earthworms. In: Sparrow, D. J. & Eddie, J. (Eds.), An Introduction to the Wildlife of Cyprus. Terra Cypria, Limassol, pp. 587 - 599.","Michaelsen, W. (1928) Beitrage zur Kenntnis der Fauna Griechenlands, namentlich der agaischen Inseln. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 137, 289 - 290.","Tzelepe, N. D. (1943) Symvoli is tin meletin ton Oligochaeton tis Ellados. Diss. Unir., Athen, 60 pp.","Omodeo, P. (1955) Lombrichi cavernicoli di Grecia e Turchia raccolti dal Dr. K. Lindberg. Annuario dell'Istituto e Museo di Zoologia della Universita di Napoli, 7, 1 - 16.","Michalis, K. (1972) Symvoli is tin meletin to Oligochaeton tis Makedonias. PhD Dissertation, University of Thessaloniki, Thessaloniki, 98 pp.","Michalis, K. (1987) Contribution to the systematic and ecological study of the earthworms (Oligochaeta, Lumbricidae) of the Western Thrace. Arquivos do Museu Bocage, 1 (5), 59 - 69.","Michalis, K., Fragoulis, A. & Panidis, S. (1989) Notes on the earthworms (Oligochaeta, Lumbricidae) from central west Macedonia. Acta zoologica cracoviensia, 32, 3 - 14.","Zicsi, A. & Michalis, K. (1993) Zwei neue Dendrobaena - Arten aus Griechenland (Oligochaeta: Lumbricidae). Acta Zoologica Hungarica, 39, 301 - 310.","Cernosvitov, L. (1938) Zur Kenntnis der Oligochatenfauna des Balkans. VI. Oligochaten aus Griechenland. Zoologischer Anzeiger, 123, 192 - 200.","Zicsi, A. (1973) Regenwurmer (Oligochaeta: Lumbricidae) aus Griechenland. Opuscula Zoologica Budapest, 12, 99 - 103.","Michaelsen, W. (1902) Neue Oligochaeten und neue Fundorte alt-bekannter. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 19, 3 - 53.","Cognetti, L. (1906) Nuovi dati sui Lumbricidi dell'Europa orientale. Bollettino dei Musei di zoologia ed anatomia comparata della R. Universita di Torino, 21 (257), 1 - 18."]}

Dendrobaena balcanica (Černosvitov, 1937) Eisenia veneta var. balcanica Černosvitov, 1937: 81. Dendrobaena balcanica: Zicsi & Csuzdi 1986: 115. Szederjesi & Csuzdi 2012: 31. Balkanic endemic species distributed in Bulgaria and Greece (Szederjesi & Csuzdi 2012). Distribution in Greece. Mainland: Kalikarpos (Szederjesi & Csuzdi 2012)., Published as part of Szederjesi, T��mea, Vavoulidou, Evangelia, Chalkia, Christina, D��nyi, L��szl�� & Csuzdi, Csaba, 2017, An annotated checklist of earthworms of Greece (Clitellata: Megadrili), pp. 57-82 in Zootaxa 4272 (1) on pages 62-63, DOI: 10.11646/zootaxa.4272.1.3, http://zenodo.org/record/583379, {"references":["Cernosvitov, L. (1937) Die Oligochaetenfauna Bulgariens. Mitteilungen aus den Koniglich Naturwissenschaftlichen Instituten in Sofia, 10, 62 - 92.","Zicsi, A. & Csuzdi, Cs. (1986) Regenwurmer aus Bulgarien (Oligochaeta: Lumbricidae). Opuscula Zoologica Budapest, 22, 113 - 121.","Szederjesi, T. & Csuzdi, Cs. (2012) New and little known earthworm species form Greece (Oligochaeta: Lumbricidae, Acanthodrilidae). Zootaxa, 3304, 25 - 42."]}

Aporrectodea longa (Ude, 1885) Allolobophora longa Ude, 1885: 136. Aporrectodea longa: Valchovski 2016: 2. Widely introduced peregrine species (Csuzdi & Zicsi 2003). Distribution in Greece. Mainland: Porto Koufo (Valchovski 2016)., Published as part of Szederjesi, T��mea, Vavoulidou, Evangelia, Chalkia, Christina, D��nyi, L��szl�� & Csuzdi, Csaba, 2017, An annotated checklist of earthworms of Greece (Clitellata: Megadrili), pp. 57-82 in Zootaxa 4272 (1) on page 61, DOI: 10.11646/zootaxa.4272.1.3, http://zenodo.org/record/583379, {"references":["Ude, H. (1885) Uber die Ruckenporen der terricolen Oligochaeten, nebst Beitrage zur Histologie des Leibesschlauches und zur Sytematik der Lumbriciden. Zeitschrift fur Wissenschaftliche Zoologie, 43, 87 - 143.","Valchovski, H. (2016) First earthworm (Oligochaeta: Lumbricidae) record from Sithonia Peninsula (Greece). ZooNotes, 94, 1 - 3.","Csuzdi, Cs. & Zicsi, A. (2003) Earthworms of Hungary (Annelida: Oligochaeta; Lumbricidae). In: Csuzdi, Cs. & Mahunka, S. (Eds.), Pedozoologica Hungarica. 1. Hungarian Natural History Museum, Budapest, pp. 1 - 271."]}

Aporrectodea caliginosa trapezoides (Dug��s, 1828) Lumbricus trapezoides Dug��s, 1828: 289. Allolobophora caliginosa trapezoides: Zicsi & Michalis 1981: 248. Michalis 1982: 351; 1987: 62. Aporrectodea caliginosa: Szederjesi & Csuzdi 2012: 29 (part.). Szederjesi 2015: 144 (part.); 2017. Peregrine subspecies (Stojanović & Milutinović 2014). Distribution in Greece. Mainland: Katovermion (Černosvitov 1938). Vermio, Attika (Tzelepe 1943). Kastoria (Omodeo 1955). Beldento, Doxaton, Korisos Kastarias, Kastoria (Karaman 1972). Florina, Drama, Serrae, Kozani, Veroia (Michalis 1975). Trikala, Alykae Volos, Zagora, Agiokampos, Agia Paraskevi, Sykourion, Nea Anchialos, Amaliapolis, Kalampaka, Platykampos, Megalochori (Michalis 1976). Between Aigeiros and Komotini (Michalis 1987). Crete: Neapoli (Cognetti 1906). Ida Mts, Zakros, Loutraki, Orino, Agios Georgios, Omalos (Szederjesi 2015). Anogia, Piso Moni Preveli, Platanias, Omalos, Seliniotikos Giros, Males (Szederjesi 2017). Peloponnese: Kalavryta (Černosvitov 1938, Tzelepe 1943). Rhodes: Agios Isidoros, Kattavia, Koskinou (Cognetti 1913). Vati, Afandou (Szederjesi 2015). Korfu (Tzelepe 1943). Kos: Asfendiou, Pylion (Michalis 1975b). Naxos: Mt. Zas (Szederjesi 2015). New records. HNHM /17167 1 ex., Rhodes Island, Rhodes, Rodini Park, Platanus trees at a stream, 36��25.635'N, 28��13.209'E, 36 m a.s.l., leg. L. D��nyi, 0 5.0 4.2016. HNHM /17181 1 ex., Karpathos Island, Olympos, in the valley under the village, streambank, 35��44.453'N, 27��10.510'E, 183 m a.s.l., leg. L. D��nyi, 28.0 3.2016., Published as part of Szederjesi, T��mea, Vavoulidou, Evangelia, Chalkia, Christina, D��nyi, L��szl�� & Csuzdi, Csaba, 2017, An annotated checklist of earthworms of Greece (Clitellata: Megadrili), pp. 57-82 in Zootaxa 4272 (1) on page 60, DOI: 10.11646/zootaxa.4272.1.3, http://zenodo.org/record/583379, {"references":["Duges, A. (1828) Recherche sur la circulation, la respiration, et la reproduction des Annelides setigeres abranches. Annales des Sciences Naturelles Paris, 15, 284 - 336.","Zicsi, A. & Michalis, K. (1981) Ubersicht der Regenwurm-fauna Griechenlands (Oligochaeta: Lumbricidae). Acta Zoologica Hungarica, 27, 239 - 264.","Michalis, K. (1982) Katalog der Oligochaetenfauna Griechenlands. Biologia Gallo-Hellenica, 9, 343 - 362.","Szederjesi, T. & Csuzdi, Cs. (2012) New and little known earthworm species form Greece (Oligochaeta: Lumbricidae, Acanthodrilidae). Zootaxa, 3304, 25 - 42.","Szederjesi, T. (2015) New earthworm records from various parts of Greece (Oligochaeta: Lumbricidae, Acanthodrilidae, Megascolecidae, Ocnerodrilidae). Opuscula Zoologica Budapest, 46 (2), 143 - 152.","Stojanovic, M. & Milutinovic, T. (2014) The earthworms (Oligochaeta: Lumbricidae) of the Pannonian region of Serbia, Vojvodina province: Zoogeography and Diversity. North-Western Journal of Zoology, 10 (2), 305 - 313.","Cernosvitov, L. (1938) Zur Kenntnis der Oligochatenfauna des Balkans. VI. Oligochaten aus Griechenland. Zoologischer Anzeiger, 123, 192 - 200.","Tzelepe, N. D. (1943) Symvoli is tin meletin ton Oligochaeton tis Ellados. Diss. Unir., Athen, 60 pp.","Omodeo, P. (1955) Lombrichi cavernicoli di Grecia e Turchia raccolti dal Dr. K. Lindberg. Annuario dell'Istituto e Museo di Zoologia della Universita di Napoli, 7, 1 - 16.","Karaman, S. (1972) Beitrag zur Kenntnis der Lumbricidenfauna von Griechenland. Fragmenta Balcanica Skopje, 9 (11), 109 - 115.","Michalis, K. (1976) Beitrag zur Kenntnis der Oligochaetenfauna Thessaliens. Scientific Annals, Faculty of Physics and Mathematics, University of Thessaloniki, 16, 153 - 168.","Michalis, K. (1987) Contribution to the systematic and ecological study of the earthworms (Oligochaeta, Lumbricidae) of the Western Thrace. Arquivos do Museu Bocage, 1 (5), 59 - 69.","Cognetti, L. (1906) Nuovi dati sui Lumbricidi dell'Europa orientale. Bollettino dei Musei di zoologia ed anatomia comparata della R. Universita di Torino, 21 (257), 1 - 18.","Cognetti, L. (1913) Escursioni zoologiche del Dr. E. Festa nell'Isola di Rodi V. Oligocheti. Bollettino dei Musei di zoologia ed anatomia comparata della R. Universita di Torino, 28 (74), 1 - 6.","Michalis, K. (1975 b) Die Oligochaeten der Insel Kos. Scientific Annals, Faculty of Physics and Mathematics, University of Thessaloniki, 15, 187 - 200."]}

Octodrilus transpadanus (Rosa, 1884) Allolobophora transpadana Rosa, 1884: 45. Octodrilus transpadanus: Zicsi & Michalis 1981: 257; 1993: 305. Michalis 1982: 358; 1993: 18; 1995: 16. Szederjesi & Csuzdi 2012: 39. Szederjesi 2015: 149. Octodrilus transpadanum: Michalis 1987: 64. Trans-Aegean species (Csuzdi & Zicsi 2003). Distribution in Greece. Mainland: Messolongi (Tzelepe 1943). Despilio, Elephtheroupolis, Veroia, Skydra, Edessa, Giannitsa, Agios Germanos, Emmanouil Pappa, Kallithea, Athytos (Michalis 1972). Milina, Kalampaka (Michalis 1976). Ioannina, Katsikas, Perama, Arnissa (Zicsi & Michalis 1981). Between Nea Chili and Alexandroupolis, Soufli, between Aigeiros and Komotini, between Palladio and Komotini, between amvrosia and Komotini, between Arisvi and Komotini, Ferrae, between Ardanio and Ferrae, between Dadia and Soufli, between Psathades and Didymoteichou, between Iasmos and Komotini, between Poros and Ferrae, Komotini, between Sappae and Komotini, between Peplos and Soufli (Michalis 1987). Nea Vathi (Michalis 1993). Kalpaki, Tempi (Zicsi & Michalis 1993). Nestos Delta (Szederjesi & Csuzdi 2012). Loutros, Tsifliki (Szederjesi 2015). Peloponnese: Patras, Kalavryta. Lefkas: Kaligoni. Kefalonia: Sami (Zicsi 1973). Athos (Michalis 1977). Lesvos: Panagouda. Leros: Partheni (Zicsi & Michalis 1981)., Published as part of Szederjesi, T��mea, Vavoulidou, Evangelia, Chalkia, Christina, D��nyi, L��szl�� & Csuzdi, Csaba, 2017, An annotated checklist of earthworms of Greece (Clitellata: Megadrili), pp. 57-82 in Zootaxa 4272 (1) on page 73, DOI: 10.11646/zootaxa.4272.1.3, http://zenodo.org/record/583379, {"references":["Rosa, D. (1884) Lumbricidi del Piemonte. Unione Tipografico-Editrice, Torino, 54 pp.","Zicsi, A. & Michalis, K. (1981) Ubersicht der Regenwurm-fauna Griechenlands (Oligochaeta: Lumbricidae). Acta Zoologica Hungarica, 27, 239 - 264.","Michalis, K. (1982) Katalog der Oligochaetenfauna Griechenlands. Biologia Gallo-Hellenica, 9, 343 - 362.","Szederjesi, T. & Csuzdi, Cs. (2012) New and little known earthworm species form Greece (Oligochaeta: Lumbricidae, Acanthodrilidae). Zootaxa, 3304, 25 - 42.","Szederjesi, T. (2015) New earthworm records from various parts of Greece (Oligochaeta: Lumbricidae, Acanthodrilidae, Megascolecidae, Ocnerodrilidae). Opuscula Zoologica Budapest, 46 (2), 143 - 152.","Michalis, K. (1987) Contribution to the systematic and ecological study of the earthworms (Oligochaeta, Lumbricidae) of the Western Thrace. Arquivos do Museu Bocage, 1 (5), 59 - 69.","Csuzdi, Cs. & Zicsi, A. (2003) Earthworms of Hungary (Annelida: Oligochaeta; Lumbricidae). In: Csuzdi, Cs. & Mahunka, S. (Eds.), Pedozoologica Hungarica. 1. Hungarian Natural History Museum, Budapest, pp. 1 - 271.","Tzelepe, N. D. (1943) Symvoli is tin meletin ton Oligochaeton tis Ellados. Diss. Unir., Athen, 60 pp.","Michalis, K. (1972) Symvoli is tin meletin to Oligochaeton tis Makedonias. PhD Dissertation, University of Thessaloniki, Thessaloniki, 98 pp.","Michalis, K. (1976) Beitrag zur Kenntnis der Oligochaetenfauna Thessaliens. Scientific Annals, Faculty of Physics and Mathematics, University of Thessaloniki, 16, 153 - 168.","Zicsi, A. & Michalis, K. (1993) Zwei neue Dendrobaena - Arten aus Griechenland (Oligochaeta: Lumbricidae). Acta Zoologica Hungarica, 39, 301 - 310.","Zicsi, A. (1973) Regenwurmer (Oligochaeta: Lumbricidae) aus Griechenland. Opuscula Zoologica Budapest, 12, 99 - 103.","Michalis, K. (1977) Die Oligochaetenfauna der Halbinsel von Athos (Chalkidiki). Scientific Annals, Faculty of Physics and Mathematics, University of Thessaloniki, 17, 285 - 298."]}

Eisenia colchidica (Perel, 1967) Eiseniella colchidica Perel, 1967: 101. Zicsi & Michalis 1981: 243. Michalis 1982: 355. Michalis et al. 1985: 39. Eisenia colchidica: Mr��ić 1991: 507. Caucasian-East Mediterranean species distributed in Georgia, Turkey and Greece (Omodeo & Rota 1991, Zicsi & Michalis 1981). Distribution in Greece. Mainland: Verioa (Zicsi & Michalis 1981, Michalis et al. 1985). Kos: Pylion (Zicsi & Michalis 1981)., Published as part of Szederjesi, T��mea, Vavoulidou, Evangelia, Chalkia, Christina, D��nyi, L��szl�� & Csuzdi, Csaba, 2017, An annotated checklist of earthworms of Greece (Clitellata: Megadrili), pp. 57-82 in Zootaxa 4272 (1) on page 67, DOI: 10.11646/zootaxa.4272.1.3, http://zenodo.org/record/583379, {"references":["Perel, T. S. (1967) Die Regenwurmer der Reliktwalder des westlichen Transkaukasien und des Talysch-Gebietes. Pedobiologia, 7, 93 - 120. [in Russian]","Zicsi, A. & Michalis, K. (1981) Ubersicht der Regenwurm-fauna Griechenlands (Oligochaeta: Lumbricidae). Acta Zoologica Hungarica, 27, 239 - 264.","Michalis, K. (1982) Katalog der Oligochaetenfauna Griechenlands. Biologia Gallo-Hellenica, 9, 343 - 362.","Michalis, K., Fragoulis, A. & Panidis, S. (1985) Earthworms (Oligochaeta: Lumbricidae) from the montainous mas of Pella province: southwest region, and Imathias province, northwest region: a systematical and ecological study. Ecologia Mediterranea, 11 (4), 37 - 42.","Mrsic, N. (1991) Monograph on earthworms (Lumbricidae) of the Balkans I-II. Slovenska Akademija Znanosti in Umetnosti, Zazred za Naravoslovne Vede Opera, Ljubljana, 757 pp.","Omodeo, P. & Rota, E. (1991) Earthworms of Turkey. II. Bollettino di Zoologia, 58, 171 - 181. https: // doi. org / 10.1080 / 11250009109355749"]}

Helodrilus patriarchalis (Rosa, 1893) (Figure 1.5) Allolobophora patriarchalis Rosa, 1893: 9. Helodrilus patriarchalis: Zicsi & Michalis 1981: 262. Michalis 1982: 352. Szederjesi et al. 2014: 184. Caucasian-East-Mediterranean species (Pavl��ček & Csuzdi 2016). Distribution in Greece. Crete: Vizari (Michaelsen (1902). Georgioupoli (Szederjesi et al. 2014). Naxos (Michaelsen 1928). Kos: Pylion (Zicsi & Michalis 1981)., Published as part of Szederjesi, T��mea, Vavoulidou, Evangelia, Chalkia, Christina, D��nyi, L��szl�� & Csuzdi, Csaba, 2017, An annotated checklist of earthworms of Greece (Clitellata: Megadrili), pp. 57-82 in Zootaxa 4272 (1) on page 71, DOI: 10.11646/zootaxa.4272.1.3, http://zenodo.org/record/583379, {"references":["Rosa, D. (1893) Viaggio del Dr. E. Festa in Palestina, nel Libano e regioni vicine. - II. Lumbricidi. Bollettino dei Musei di Zoologia ed Anatomia comparata della R. Universita di Torino, 8 (160), 1 - 14.","Zicsi, A. & Michalis, K. (1981) Ubersicht der Regenwurm-fauna Griechenlands (Oligochaeta: Lumbricidae). Acta Zoologica Hungarica, 27, 239 - 264.","Michalis, K. (1982) Katalog der Oligochaetenfauna Griechenlands. Biologia Gallo-Hellenica, 9, 343 - 362.","Szederjesi, T., Angyal, D., Balazs, G. & Danyi, L. (2014) Remarks on the earthworm genus Helodrilus Hoffmeister, 1845 with new epigean and subterranean records (Oligochaeta, Lumbricidae). Opuscula Zoologica Budapest, 45 (2), 181 - 188.","Pavlicek, T. & Csuzdi, Cs. (2016) Chapter 23: Clitellata: Oligochaeta: Earthworms. In: Sparrow, D. J. & Eddie, J. (Eds.), An Introduction to the Wildlife of Cyprus. Terra Cypria, Limassol, pp. 587 - 599.","Michaelsen, W. (1902) Neue Oligochaeten und neue Fundorte alt-bekannter. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 19, 3 - 53.","Michaelsen, W. (1928) Beitrage zur Kenntnis der Fauna Griechenlands, namentlich der agaischen Inseln. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 137, 289 - 290."]}