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Maladera perniciosa (Brenske, 1898) Figures 14 A���E, 93 Autoserica perniciosa Brenske, 1898: 336. Maladera (Cephaloserica) perniciosa: Ahrens 2004a: 276; Krajcik 2012: 155; Ahrens & Fabrizi 2016: 242. Type material examined. See Ahrens (2004a). Additional material examined. China: 1 ♂ ��� Pantiange, Weixi, Yunnan, 29.VII.1981, 2500m, leg. Wang Shuyong��� (IZAS), 2 ♂♂ ��� Huoditang, Ningshan, Shaanxi, 30.VI.1999, 1580m, leg. Yuan Decheng ��� (IZAS), 2 ex. ���China N Sichuan Jiuzhaigou 2000 m 16. Jun 1992 M. Bok lgt.��� (SMNS), 1 ex. ��� China: Yunnan Province, Tengchong, 3.-6.VI.2007 near long distance bus station, 25��00.3���N 098��29.3���E, 1625m, J. Hajek & J. Ruzicka leg.��� (NMPC), 1 ex. ��� China: Yunnan Province, Shanzhi env., 22.-24.VI.2007 Jizu Shan Mt., 25��57.7���N 100��23.6���E, 2180m, J. Hajek & J. Ruzicka leg.��� (NMPC), 5 ex. ��� China: Yunnan Province, Dali old city, 31.V.-2.VI.2007, 25��41.4���N 100��09.8���E, 2030m, J. Hajek & J. Ruzicka leg.��� (NMPC), 106 ex. ��� Tibet: Ma-ma-luo, Zayu county, 15-26 June 2010 local collector of Li Jingke��� (ZFMK), 31 ex. ��� China Yunnan 2000-3000m 27.20N 100.11E Habashan mt. SE slope 10.-13.7. David Kral leg. 92/ coll. Milan Nikodym, Praha ��� (ZFMK, NMPC), 2 ex. ��� China, N-Yunnan 27��18���N 100��13���E Jinsha r. vall. 1900M Daju, Hutiao gorge lgt. D. Kral 15-17/7���90��� (ZFMK), 2 ex. ��� China Yunnan 2000-3000m 27.20N 100.11E Habashan mts. SE slope 10-13/7. Vit Kuban leg. 92��� (ZFMK), 6 ex. ��� China (Yunnan) Dali Bai Aut. Pref., foothills of Diacang Shan W of Dali 2050m 25��42���12������N / 100��08���26������E (stone and gravel pit) 12.VI.2007 D.W. Wrase [34]��� (ZFMK), 12 ex. ��� China: Sichuan; Wolong Reserve, Sigulian Shan, 31��09���N 103��06���E v.2006, 1500- 1800m V. Siniaev ��� (ZFMK), 1 ♂ ��� China Yunnan 27.-28.VI. Daju, 50 km M Lijiang 27��21���N 100��19���E E. Jendek leg. 1992��� (NHMW), 1 ♂ ��� China: E-Yunnan; Damaidi 2500m, Guangnan near Vietnam VII-2003 leg. Li et al. ��� (ZFMK). Vietnam: 1 ex. ��� Tonkin Montes Mauson April, Mai 2-3000��� H. Fruhstorfer ��� (ZMHB), 2 ex. ���N-Vietnam Cao Bang Prov., vic. Tinh Tuc, Nui Pia Oac Nat. Res. 13.V.2013, 22��36���50������N, 105��36���50������E 14-1800m leg. A. Skale ��� (CASH), 2 ex. ���N-Vietnam Vinh Phuc Prov. vic. Tam Dao Town, Tam Dao NP, 02.-05.V.2013, 21��27���N, 105��38���E 700-1000m, A. Skale ��� (CASH), 1 ex. ���N-Vietnam Cao Bang Prov., vic. Vin Den, Nui Pia Oac Nat. Res. 06.-10.V.2013, 22��33���53������N, 105��52���53������E 900-1300m A. Skale ��� (CASH), 2 ex. ���N-Vietnam Cao Bang Prov., vic. Vin Den, Nui Pia Oac Nat. Res. 10.-15.V.2013, 22��33���53������N, 105��52���53������E 900-1300m A. Skale ��� (CASH), 1 ♂ ���Vietnam-N Tuyen Quang pr. SE-E env. Of Na Hang, 22��17���30������N 105��26-28���E, 200-700m L. Dembicky leg., 1.- 12.V.2010 ��� (ZFMK). Thailand: 1 ex. ���Thai-N, Nan Prov., Bo Klua, 700 m, 19��8���N 101��10���E, 28.iv.-1.v.1999, D. Hauck leg.��� (CP), 2 ex. ���N-Thailand 17.VII.1990 Doi Inthanon, leg. Malicky ��� (ZSM), 4 ex. ���N-Thailand V.1990 Doi Inthanon, leg. Malicky ��� (ZSM), 1 ♂ ��� Thailand, Mae Hong Son prov., Kiwlom pass near Soppong, WGS 84; 19��26���N 98��19���E, 23.vi.-2.vii.2002, alt. 1400m + 50m, lgt. Fouque R.+ H. Kozich, J.��� (NMPC), 1 ♂ ���N. Thailand: Angkhai village, Samoeng Dist. Chiang Mai Prov., 9.-11.v.1999 K. Masumoto leg.��� (ZFMK). Laos: 1 ex. ��� Laos: Sayaboury Prov. Sayaboury 5.V.1966 / Native Collector Bishop Museum / Autoserica rubescens Mos. det. G. Frey 1971��� (CF), 2 ex. ���NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20��12���N, 104��01���E; 14.iv.-15.v.2012; 1300-1900m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012/13��� (ZFMK), 2 ex. ���Laos-NE Hua Phan prov., 20��12���N, 104��01���E, Phu Phan Mt., 1500-1900m, 17.5.- 3.6.2007, leg. Vit Kuban ��� (ZFMK), 3 ex. ���NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20��12���N, 104��01���E; 11.iv.-15.v.2012; 1300-1900m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012��� (ZFMK), 1 ex. ���Laos-NE, Houa Phan prov., 20��13���09- 19������N 103��59���54������- 104��00���03������E, 1480-1510m Phou Pane Mt., 22.IV.-14.V.2008 Vit Kuban leg. (NMPC, ZFMK), 1 ♂ ���X-DA4702 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera perniciosa��� (ZFMK), 1 ♂ ���X-DA4918 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera perniciosa ��� (ZFMK), 1 ♂ ���X-DA4923 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera perniciosa ��� (ZFMK), 1 ♂ ���X-DA4699 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.- 30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera perniciosa ��� (ZFMK), 1 ♂ ���X-DA4830 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera perniciosa ��� (ZFMK). Remarks. The species was exhaustively treated (including redescription) by Ahrens (2004a, p. 276). Distribution. See map (Fig. 93) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 69-71, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Brenske, E. (1898) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 43, 205 - 403. https: // doi. org / 10.1002 / mmnd. 47918980303","Ahrens, D. (2004 a) Monographie der Sericini des Himalaya (Coleoptera, Scarabaeidae). Dissertation. de-Verlag im Internet GmbH, Berlin, 534 pp.","Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement, 5, 1 - 278.","Ahrens, D. & Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355.","Fairmaire, M. L. (1888) Coleopteres de l'Interieur de la Chine. Annales de la Societe Entomologique de Belgique, 32, 7 - 46."]}

Maladera paradetersa Ahrens, Fabrizi & Liu, sp. n. Figures 39 I���L, 107 Type material examined. Holotype: ♂ ���Prov. Fo-Kien (China)/ detersa Er. ��� (ZMHB). Paratype: 1 ♀ ���Prov. Fo-Kien (China)/ Autoserica detersa coll. Moser ��� (ZMHB). Description. Length: 7.2 mm, length of elytra: 4.7 mm, width: 4.0 mm. Body oblong-oval, yellowish brown, antenna yellow, shiny, labroclypeus shiny, dorsal face nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and strongly convergent anteriorly, anterior angles moderately rounded, anterior margin weakly emarginate, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex medially, finely, densely punctate, with a few long setae anteriorly; frontoclypeal suture indistinctly incised, weakly curved; smooth area anterior to eye convex, twice as wide as long; ocular canthus short and narrow (1/4 of ocular diameter), finely densely punctate, with a terminal seta. Frons with dense and fine punctures, with a few single setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.67. Antenna with ten antennomeres, club composed of three antennomeres and straight, distinctly shorter than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin straight, with fine marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; anterior and lateral margin sparsely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures. Elytra widest at middle, striae finely impressed, finely and densely punctate, intervals flat, with fine, moderately dense punctures and with minute setae in punctures, odd interval with a few single adpressed setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface shiny, finely and densely punctate, metasternum including disc sparsely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.8. Pygidium moderately convex, shiny, finely and densely punctate, without smooth midline, with a few long but fine setae along apical margin. Legs short and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, without adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, with a few single long setae. Metatibia short and moderately wide, widest at middle, ratio of width/length: 1/2.38, sharply carinate dorsally, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, in basal half with a few short and robust single setae; lateral face longitudinally convex, except a few superficial punctures on sides and base smooth and glabrous; ventral margin finely serrate, with five equidistant robust setae; medial face smooth and glabrous; apex finely serrate, concavely emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 39 I���K. Habitus: Fig. 39L. Diagnosis. Maladera paradetersa Ahrens, Fabrizi & Liu, sp. n. differs from M. liaochengensis Ahrens, Fabrizi & Liu, sp. n. in having the basal and distal lobes of the parameres straight apically rather than being reflexed externally as in M. liaochengensis. Etymology. The name of the new species (noun in apposition) is derrived from the combined Greek prefix ��� para -��� (close by) and the species name detersa with reference to the similarity to M. detersa (adjective in nominative singular). Variation. Female: Length: 6.8 mm, length of elytra: 4.5 mm, width: 3.9 mm. Eyes as large as in male, antennal club slightly shorter. Distribution. See map (Fig. 107) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 176-177, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera aureola (Murayama, 1938) Figures 29 I���L, 102 Aserica aureola Murayama, 1938: 19; Serica aureola: Murayama 1954: 64. Maladera aureola: Kim & Lee 1997: 128; Kim & Kim 2003: 92. Maladera liotibia Nomura, 1974: 104, 110; Ahrens 2002: 84, syn. n. Additional material examined. 1 ♂ ��� Kuatun (2300 m) 27, 40 n. Br. 117,40 ��. L. J. Klapperich 3.5.1938 (Fukien)��� (ZFMK), 1 ♂ ��� Juxian County, Henan, 21.VI.1960 ��� (IZAS), 1 ♂ ��� Nanmu Garden, Changshou, Sichuan, 9.VI. 1994, 450m, leg. Zhang Youwei ��� (IZAS), 1 ♀ ��� Shiping, Fengdu, Sichuan, 3.VI. 1994, 610m, leg. Zhang Youwei ��� (IZAS), 1 ♂ ��� Jiangya Forestry Farm, Cili, Hunan, 26.VIII.1988 ��� (IZAS), 1 ♂ ��� Mts. Baiyunshan, Songxian, Henan, 14- 17.VIII.2008, leg. Ren Guodong, Wu Qiqi etc.��� (HBUM), 1 ♂ ��� Shibanyan Town, Linzhou, Henan, 21-24.VII.2006, leg. Wang Fengyan, Huang Wenjing ��� (HBUM), 1 ♂ ��� China: Hunan; Mupu Mt. 1600m, Pingjiang VIII-2003, leg. Li et al. ��� (ZFMK), 2 ♂♂, 1 ♀ ��� Kuatun, Fukien China 22.7.46 (Tschung Sen.)��� (NMPC), 1 ♂ ��� China, Wang-lungkum, Bergebiet d. Lo-fau-zan/ am Ostfluss in Kuangtung 4.-5.12 Mell S.V.��� (ZMHB, head missing in specimen), 1 ♂ ��� China: Hunan, Jiucai Ling, 25��32���N 111��22���E, 1300m, VII.2006, leg. Viktor & Sveta Siniaev ��� (ZFMK), 1 ♂ ��� Wen Chuan Shien 30mi NNW Kuanshien VII-22 1933 / Szechwan China DC Graham 4600ft. ��� (USNM), 1 ♂, 1 ♀ ��� China: S-Hubei Mupushan Tangshan vi.2004 leg. Wen ��� (ZFMK), 1 ♂ ��� China Tsingtau 1 July 1926 D.C. Graham ��� (USNM), 1 ♂ ��� Foochow China VI-6-1928 / C.R. Kellogg Collector ��� (USNM). Redescription. Length: 8.1 mm, length of elytra: 6.4 mm, width: 5.3 mm. Body oblong-oval, yellowish brown, dull, labroclypeus shiny, with a few long setae on head, pronotum and elytra. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins nearly straight and convergent anteriorly, anterior angles strongly rounded, anterior margin shallowly emarginate, margins moderately reflexed; lateral margin and ocular canthus produce an indistinct angle; surface convexly elevated medially, finely, densely punctate, with a few erect, long setae; frontoclypeal suture distinctly incised, evely curved; smooth area anterior to eye convex, twice as wide as long; ocular canthus short and narrow (1/3 of ocular diameter), finely densely punctate, with a terminal seta. Frons dull, with dense, fine punctures, with a few single setae beside eyes and behind frontoclypeal suture. Eyes small, ratio diameter/ interocular width: 0.56. Antenna with ten antennomeres; club with three antennomeres and straight, slightly shorter than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt, slightly rounded at tip; anterior margin straight, with fine marginal line, base without marginal line; surface moderately finely and densely punctate, with minute setae in punctures and a few long semi-erect setae on sides; anterior and lateral margin finely densely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, moderately dense punctures, impunctate on midline. Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals flat, with fine, sparse punctures and with minute setae in punctures, odd intervals with numerous single long setae; epipleural margin robust, ending at half of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, finely and densely punctate, nearly glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.9. Pygidium strongly convex, dull, coarsely and moderately densely punctate, with narrow smooth midline, with a few long setae on disc and along apical margin. Legs short and wide, dull; femora with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, without adjacent serrated line, anterior row of setae completely reduced; posterior ventral margin smooth, moderately widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.7, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short and robust single setae; lateral face longitudinally convex, superficially and sparsely punctate, along midline widely smooth, glabrous; ventral margin finely serrate, with five equidistant robust setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 29 I���K. Habitus: Fig. 29L. Remarks. The tiny differences in the shape of the parameres within the respective original descriptions of Maladera liotibia Nomura, 1974 and M. aureola Murayama, 1938 correspond to the variation found in M. aureola already noted among Taiwanese specimens. Therefore, we believe these names are synonyms of a single species. Distribution. See map (Fig. 102) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 138-139, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Murayama, J. (1938) Revision des Sericines (Col., Scar.) de la Coree. Annotiones Zoologicae Japonenses, 17, 7 - 21.","Murayama, J. (1954) Icones of the Scarabeid-Beetles from Manchuria and Korea. I. Nihon Gakujutu, Shinko-kai, 163 pp.","Kim, J. I. & Lee, O. J. (1997) Taxonomic study of Korean Sericinae (Melolonthidae, Coleoptera) II, Genus Maladera. Insecta Koreana, 14, 119 - 135.","Nomura, S. (1974) On the Sericini of Taiwan. Toho-Gakuho, 24, 81 - 115.","Ahrens, D. (2002) Notes on distribution and synonymy of sericid beetles from Taiwan, with descriptions of new species (Coleoptera, Scarabaeoidea, Sericini). Annales Historico-Naturales Musei Nationalis Hungarici, 94, 53 - 91."]}

Serica nigrobrunnea Moser, 1926: 198; Ahrens 2006b: 245; 2007b: 36. Type material examined. Lectotype (here designated): ♂ ���Szetschwan Washan/ Serica nigrobrunnea Type Mos./ Typus��� (ZMHB). Paralectotypes (here designated): 1 ♀ ��� Szetschwan Washan / Serica nigrobrunnea Type ♀ Mos./ Typus / Zool. Mus. Berlin ��� (ZMHB), 5 ♂♂, 1 ♀ ��� Szetschwan Washan / Zool. Mus. Berlin ��� (ZMHB). Additional material examined. 1 ♂ ���Szechuen China DC Graham/ Yao Gin nr Mupin 3-4 Jul ���29 7400 ft. ��� (USNM), 1 ♂ ���Szechuen China DC Graham/ Yao Gin nr Mupin 15 Jul ���29 7400 ft. ��� (USNM), 1 ♂ ��� Mt. Omei & vicinity Szechuen China 22 Graham��� (USNM), Redescription. Length: 8.1 mm, length of elytra: 6.4 mm, width: 5.2 mm. Body oblong-oval, reddish brown, antenna yellow, shiny, labroclypeus shiny, dorsal face nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and strongly convergent anteriorly, anterior angles moderately rounded, anterior margin weakly emarginate, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex medially, finely, densely punctate, with a few single erect setae; frontoclypeal suture indistinctly incised, weakly curved; smooth area anterior to eye convex, twice as wide as long; ocular canthus short and narrow (1/4 of ocular diameter), finely densely punctate, with a short terminal seta. Frons with dense and fine punctures, with a few single setae beside eyes. Eyes moderately large, ratio diameter/ interocular width: 0.58.Antenna with ten antennomeres, club composed of three antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest shortly before base, lateral margins evenly convex and convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin straight, with fine, rather indistinct marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; anterior and lateral margin sparsely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures. Elytra widest at posterior third, striae finely impressed, finely and densely punctate, intervals moderately convex, with fine, moderately dense punctures concentrated along striae and with minute setae in punctures, penultimate interval with a few single erect setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface shiny, finely and densely punctate, metasternum including disc sparsely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.3. Pygidium moderately convex, dull, finely and densely punctate, without smooth midline, with numerous long setae on apical half. Legs moderately long and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur in widest part shiny, anterior margin acute, without adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, with a few single long setae. Metatibia short and moderately wide, widest at middle, ratio of width/ length: 1/3.3, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, in basal half with a few fine single setae and a short serrate line beside dorsal margin over basal quarter; lateral face longitudinally convex, except a few sparse punctures on base smooth and glabrous; ventral margin finely serrate, with three equidistant robust setae; medial face smooth and glabrous; apex finely serrate, concavely emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere little shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, tridentate, basal tooth blunt and indistinct; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 75 E���G. Habitus: Fig. 75H. Distribution. See map (Fig. 128) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 318-319, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Moser, J. (1926) Beitrag zur Kenntnis der Melolonthiden. XIV. Stettiner Entomologische Zeitung, 87, 198 - 208.","Ahrens, D. (2006 b) Sericinae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248.","Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1"]}

Maladera egregia (Arrow, 1946) Figures 22 A���D, 97 Aserica egregia Arrow, 1946b: 14. Maladera egregia: Ahrens 2004a: 324. Type material examined. Syntypes: 3 ♂♂ ���N. E. Burma Kambaiti 2000 m 28.5.1934 Malaise / N. E. Burma R. Malaise B. M. 1945-71/ Cotype/ Aserica egregia Arrow co-type��� (BMNH), 1 ♂ ���N. E. Burma Kambaiti 2000 m 20/6.1934 R. Malaise / N. E. Burma R. Malaise B. M. 1945-71/ Cotype��� (BMNH), 1 ♂ ���N. E. Burma Kambaiti 2000 m 4/6.1934 Malaise / N. E. Burma R. Malaise B. M. 1945-71/ Cotype / Aserica egregia Arrow co-type��� (BMNH). Redescription. Length: 8.6 mm, length of elytra: 6.4 mm, width: 5.6 mm. Body oblong-oval, reddish brown, antenna yellow, iridescent and shiny, frons and pronotum somewhat dull, dorsal face nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles moderately rounded, anterior margin distinctly emarginate, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex, finely, densely punctate, with a few long setae anteriorly; frontoclypeal suture finely incised, weakly curved; smooth area anterior to eye convex, 1.5 times as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely densely punctate, without terminal setae. Frons with sparse and fine punctures, with a few fine setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.52. Antenna with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest at base, lateral margins evenly convex and weakly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin convex, with fine marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; lateral margin sparsely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures, impunctate medially. Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, moderately dense punctures concentrated along striae, with minute setae in punctures, penultimate lateral interval with a few fine setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.49. Abdomen in available syntype specimen missing. Legs moderately long and narrow, shiny; femora with two longitudinal rows of setae, coarsely and moderately densely punctate. Metafemur dull, anterior margin acute, with adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 22 A���C. Habitus: Fig. 22D. Remarks. The metatibiae were lacking in the syntype specimen we examined and are thus omitted in the redescription. Distribution. See map (Fig. 97) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 105-107, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Arrow, G. J. (1946 b) Entomological results from the Swedish Expedition 1934 to Burma and British India. Coleoptera: Melolonthidae. Arkiv for Zoologie, 38 A (9), 1 - 33.","Ahrens, D. (2004 a) Monographie der Sericini des Himalaya (Coleoptera, Scarabaeidae). Dissertation. de-Verlag im Internet GmbH, Berlin, 534 pp."]}

Maladera hutiaoensis Ahrens, Fabrizi & Liu, sp. n. Figures 76 E���H, 129 Type material examined. Holotype. ♂ ���[China] Yunnan ca. 2000m 27.15N 100.09E Hutiao gorge Jinsha r. 18.- 22.7.92 leg. David Kral / 561 Sericini Asia spec.��� (ZFMK). Paratypes: 3 ♂♂, 3 ♀♀ ���[China] Yunnan cca 2000m 27.15N 100.09E Hutiao gorge Jinsha r. 18-22/7. Vit Kuban leg. 92��� (ZFMK), 1 ♀ ���[China] Yunnan 3000m 27.05N 100.1E Yulongshan mts. Ganhaizi pass 4/7. Vit Kuban leg. 92��� (ZFMK), 1 ♂ ��� China Yunnan, 24.-28.VI. Daju-Tiger Gorge 27.21N 100.19E lgt. S. Becvar 1992��� (ZFMK), 1 ♂ ��� China Yunnan, 27-28.6. Daju, 27��21���N, 100��19���E E. Jendek leg. 1992/ Coll. P. Pacholatko / CS 68��� (CP). Description. Length: 11.5 mm, length of elytra: 7.7 mm, width: 6.4 mm. Body oval, dark reddish brown, antenna yellowish, dorsal surface slightly iridescent shiny, head shiny, dorsal surface almost glabrous. Labroclypeus wide and trapezoidal, much wider than long, widest at base, lateral margins straight and strongly convergent anteriorly, anterior angles strongly rounded, anterior margin weakly emarginate medially, margins moderately reflexed, lateral margins produce an indistinct angle with the ocular canthus; surface weakly convex, finely and densely punctate, glabrous, with a row of fine erect seta behind the anterior margin; frontoclypeal suture finely incised, not elevated and weakly angled medially; smooth area anterior to eye wide, almost flat, three times as wide as long; ocular canthus moderately long and wide (1/2.5 of ocular diameter), finely densely punctate, with a single terminal seta. Frons with dense, fine punctures, with two distant erect setae beside each eye. Eyes small, ratio diameter/ interocular width: 0.6. Antenna with ten antennomeres, club with three antennomeres and straight, slightly longer than the remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum slightly produced medially, transverse, with deep median emargination. Pronotum moderately transverse, widest shortly before base, lateral margins weakly convex, in basal half nearly subparallel, in anterior half evenly convergent anteriorly, anterior angles distinctly produced and moderately acute, posterior angles strongly rounded; anterior margin convexly produced medially, with a fine and complete marginal line, base of pronotum without marginal line; surface finely and densely punctate, punctures with very minute setae only, otherwise glabrous; lateral anterior and lateral margins setose; hypomeron carinate. Scutellum wide triangular, at apex sharp, with fine, evenly dense punctures, with only very minute setae. Elytra widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and evenly dense punctures, except the very minute setae in the punctures glabrous; epipleural margin robust, ending at strongly curved external apical angle of elytra, elytra along the apical margin convexly elevated; epipleura densely setose, apical border membraneous and with a fine fringe of microtrichomes (100x). Ventral surface dull, coarsely and densely punctate, metepisternum distally impunctate, metasternum sparsely covered with fine, short or very minute setae, metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.4. Pygidium weakly convex and dull, coarsely and densely punctate, without smooth midline, with a few fine white adjacent setae and a few longer setae along the apical margin. Legs wide and moderately long; femur with two longitudinal row of setae, finely and sparsely punctate; metafemur dull, behind the posterior longitudinal row of setae punctures finer and more dense, anterior margin acute, without serrated line behind anterior margin, posterior margin smooth ventrally in apical half, moderately widened, posterior margin smooth dorsally, with just a few short setae basally. Metatibia wide and flattened, moderately long, widest at three quarters of metatibial length, ratio of width/ length: 1/ 2.89, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basally with a few short robust single spines; lateral face longitudinally convex, finely and sparsely punctate, glabrous; ventral margin finely serrated, with four robust equidistant setae; medial face smooth, apex finely serrated, interiorly near tarsal articulation weakly concavely emarginate. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, ventrally robustly densely setose; metatarsomeres ventrally with a strongly serrated ridge, first metatarsomere slightly shorter than the two following tarsomeres combined and as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 76 E���G. Habitus: Fig. 76H. Diagnosis. The species differs from all other Chinese Maladera in having a smooth elytra surface with very indistinct striae and completely flat intervals. Etymology. The species name (adjective in the nominative singular case) is derived from the name of its type locality placed in the Hutiao gorge (Yunnan). Variation. Length: 10.4���12.4 mm, length of elytra: 6.9���8.4 mm, width: 5.6���6.6 mm. Female: eyes smaller than in male, ratio of diameter/interocular width: 0.45; antennal club with three antennomeres, slightly shorter than remaining antennomeres combined; pygidium weakly convex. Distribution. See map (Fig. 129) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 322-323, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera masumotoi Nomura, 1974 Figures 18 A���E, 92 Maladera masumotoi Nomura, 1974: 105, 112. Additional material examined. China: 1 ♂ ���China Kiang-Si T���en-gan/ 1075 Sericini Asia spec.��� (ZFMK). Taiwan: 1 ♂ ���834673/ Taiwan, Lanyu Island (Taitung Ciunty) Yung Hsin Farm N22��01.703 E121��34.642 29.-30.vii.2008, At light H. Mendel & M.V.L. Barclay BMNH(E) 2008-85��� (BMNH). Redescription. Length: 9.6 mm, length of elytra: 6.6 mm, width: 5.2 mm. Body oblong-oval, dark reddish brown, antenna yellowish, surface dull, labroclypeus shiny, except a few short setae on sides of elytra, glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins nearly straight and convergent anteriorly, anterior angles strongly rounded, anterior margin weakly emarginate medially; margins moderately reflexed; lateral margin and ocular canthus produce an indistinct angle; surface flat, with a very indistinct median elevation behind anterior margin, finely, densely punctate, with a few erect setae; frontoclypeal suture distinctly incised, angled medially; smooth area anterior to eye convex, four times as wide as long; ocular canthus short and narrow (1/3 of ocular diameter), finely densely punctate, with a long terminal seta. Frons dull, with moderately dense, fine punctures, with a few single setae beside eyes. Eyes moderately large, ratio diameter/ interocular width: 0.7. Antenna with ten antennomeres; club with three antennomeres and straight, 1.1 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest at base, lateral margins in basal half straigt and weakly convergent anteriorly, in anterior half moderately evenly convex and convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt, moderately rounded at tip; anterior margin weakly convex, with fine complete marginal line, base without marginal line; surface moderately finely and densely punctate, with minute setae in punctures; anterior and lateral margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, moderately dense punctures. Elytra widest at posterior third, striae finely impressed, finely and densely punctate, intervals slightly convex, with fine, moderately dense punctures being except on second interval partly concentrated along striae, with minute setae in punctures and penultimate interval with a few short setae, otherwise glabrous; epipleural margin robust, ending at nearly blunt external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, finely and densely punctate, with dense and fine, adpressed setae, metasternal disc sparsely covered with longer setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.6. Pygidium moderately convex, dull, finely and sparsely punctate, without smooth midline, with a few setae along apical margin. Legs short and wide; femora dull, with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, without adjacent serrated line, anterior row of setae completely reduced; posterior ventral margin smooth, moderately widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.4, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short single spines close to dorsal margin; lateral face longitudinally convex, nearly entirely impunctate, only a few fine punctures beside dorsal and ventral margin, glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 18 A���D. Habitus: Fig. 18E. Distribution. See map (Fig. 92) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 86-88, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Nomura, S. (1974) On the Sericini of Taiwan. Toho-Gakuho, 24, 81 - 115."]}

Maladera teinzoana (Brenske, 1898) group Key to species of the Maladera teinzoana (Brenske, 1898) group occurring in China (����): 1 Eyes small, ratio diameter/ interocular width 0.7............................................................ 3 2 Phallobase with a long lateral apical apophysis. Parameres strongly asymmetrical, long and narrow.......................................................................................... Maladera futschauana (Brenske, 1897) 2��� Phallobase without a long lateral apical apophysis. Parameres subsymmetrical and long.. Maladera obscurata (Moser, 1915) 3 Antennal club in male short, as long as, or only little longer than the remaining antennomeres combined................ 6 3��� Antennal club in male long, at least 1.2 times as long as the remaining antennomeres combined, or longer............... 4 4 Metatibia with a complete, serrate line beside dorsal margin, at least in basal half................................... 5 4��� Metatibia without a complete, serrate line beside dorsal margin.............. M. longruiensis Ahrens, Fabrizi & Liu, sp. n. 5 Antennal club in male straight. Parameres and phallobase asymmetrical, left paramere with a long filiform internal basal lobe........................................................... Maladera shengqiaoae Ahrens, Fabrizi & Liu, sp. n. 5��� Antennal club in male reflexed. Parameres and phallobase symmetrical, parameres without basal lobes........................................................................... Maladera fereobscurata Ahrens, Fabrizi & Liu, sp. n. 6 Basal group of metatibia spines at just before the middle of metatibial length................................................................................................ Maladera parobscurata Ahrens, Fabrizi & Liu, sp. n. 6��� Basal group of metatibia spines well behind the middle of metatibial length........................................ 7 7 Parameres short, less than 1/3 of phallobasal length........................................................... 8 7��� Parameres long, more than half of phallobasal length...................... Maladera saitoi (Niijima & Kinoshita, 1927) 8 Each paramere without a lateral tooth apically. Parameres straight...................... Maladera piceola (Moser, 1915) 8��� Each paramere with an lateral tooth apically. Parameres reflexed..................... Maladera invenusta (Moser, 1918), Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on page 90, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Brenske, E. (1898) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 43, 205 - 403. https: // doi. org / 10.1002 / mmnd. 47918980303","Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310","Niijima, Y. & Kinoshita, E. (1927) Revision of Japanese Melolonthids and allied families. III. Research Bulletin of the College Expermental Forestry Hokkaido Imperial University, 4, 1 - 97.","Moser, J. (1918) Neue Melolonthiden aus der Sammlung des Deutschen Entomologischen Museums zu Berlin Dahlem (Col.). Stettiner Entomologische Zeitung, 79, 209 - 247."]}

Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke, Ahrens, Dirk (2021): A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 4922 (1): 1-400, DOI: https://doi.org/10.11646/zootaxa.4922.1.1

Maladera (subgen. Eumaladera Nomura, 1967) Maladera subgen. Eumaladera Nomura, 1967: 53 (Type species by original designation: Maladera nitididorsis Nomura, 1967); Ahrens 2006b: 230; Ahrens 2007b: 13; Ahrens & Bezděk 2016: 284. Key to species of Maladera (sbg. Eumaladera) occurring in China (����): 1 Body size larger,> 10 mm............................................................................... 2 1��� Body size smaller, Maladera fencli Ahrens, Fabrizi & Liu, sp. n. 2��� Left paramere short (lateral view), ventrally with a blunt tooth (Fig. 5E).......................................... 3 4 Left paramere wide and pointed at apex, subapical teeth more ventrally positioned................................................................................................ Maladera robustula Ahrens, Fabrizi & Liu, sp. n. 4��� Left paramere narrower and blunt at apex, subapical teeth more laterally positioned................................................................................................ Maladera wipfleri Ahrens, Fabrizi & Liu, sp. n. 4 Left paramere elongate (lateral view), ventrally without blunt tooth (Fig. 4A)........... Maladera subrugata (Moser, 1926) 4��� Left paramere short (lateral view), ventrally with a blunt tooth (Fig. 5A)............. Maladera opaciventris (Moser, 1915), Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on page 27, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Nomura, S. (1967) Some new and remarkable species of the Coleoptera from Japan and its adjacent regions, II. Entomological Review of Japan, 19, 52 - 62.","Ahrens, D. (2006 b) Sericinae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248.","Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1","Ahrens, D. & Bezdek, A. (2016) Sericini. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, pp. 281 - 317.","Moser, J. (1926) Beitrag zur Kenntnis der Melolonthiden. XIV. Stettiner Entomologische Zeitung, 87, 198 - 208."]}

Maladera longruiensis Ahrens, Fabrizi & Liu, sp. n. Figures 21 I���L, 96 Type material examined. Holotype: ♂ ���[China] Longrui, Ningming, Guangxi, 25.V. 1984, 180m, leg. Li Fasheng / LW-710��� (CAU). Paratypes: 1 ♂ ��� Burma (Myanmar) SW Shan State Taunggyi J. Rejsek 1.- 18.6.1997 ��� (ZFMK), 1 ♂ ���N-Thailand 27.III-3.IV.1990 Doi Inthanon lg. Malicky / Thai H/ 41 Sericini Asia spec.��� (ZSM), 1 ♂ ��� Thailand bor. occ. 30.04.- 14.05.1991 Mae Hong Song env. Ban Huai Po, 1800 m Jan Farkac leg.��� (NHMB), 1 ♂ ���Thai 1- 8.V.1993 Soppong Pai 1800m Pacholatko & Dembicky ��� (CP), 1 ♂ ���NW Thailand 9.-16.V. Mae Hong Son 1991 Ban Huai Po 1600m leg. P. Pachol��tko / TS 69��� (CP). Description. Length: 6.9 mm, length of elytra: 6.6 mm, width: 5.1 mm. Body oval, dark reddish brown to blackish, antenna yellow, club brown, dorsal surface dull, except some single setae on head dorsal surface nearly glabrous. Labroclypeus wide and trapezoidal, shiny, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles blunt and slightly reflexed, anterior margin moderately emarginate, margins moderately reflexed; lateral margin and ocular canthus produce a blunt angle; surface weakly convex medially, finely and densely punctate, with a few erect setae anteriorly; frontoclypeal suture indistinctly incised, curved; smooth area anterior to eye flat, 1.5 times as wide as long; ocular canthus long and moderately wide (nearly 1/3 of ocular diameter), finely and sparsely punctate, terminal seta lacking. Frons dull, with fine, dense punctures and a few single erect setae beside eyes. Eyes moderately large, ratio diameter/ interocular width: 0.67. Antenna with ten antennomeres; club with three antennomeres and weakly reflexed, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, lateral margins evenly convex and moderately convergent anteriorly, anterior angles moderately produced and right-angled, posterior angles blunt, weakly rounded at tip; anterior margin weakly convex, with fine and complete marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures, otherwise glabrous; lateral and anterior margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, sparse punctures. Elytra widest shortly behind middle, striae distinctly impressed, finely and sparsely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with minute setae in punctures, odd intervals with a few single short setae; epipleural margin robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra membraneous, with narrow rim of microtrichomes (at ca 100x magnification). Ventral surface dull, coarsely and moderately densely punctate, metasternum sparsely covered with sparse fine setae; metacoxa with a few longer setae laterally, otherwise glabrous. Abdominal sternites finely and irregularly densely punctate, glabrous, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.45. Pygidium strongly convex and dull, coarsely and densely punctate, without smooth midline, with a few longer setae along apical margin. Legs moderately wide and short, weakly shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, with anterior margin acute, with continuous serrated line behind anterior margin, setae of anterior row very short; posterior margin smooth and moderately widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia short, widest at apex, ratio of width/length: 1/2.86, sharply carinate dorsally, with two groups of spines, basal group shortly behind middle, apical group at four fifths of metatibial length, with a few short robust setae basally with serrated margin; lateral face weakly longitudinally convex, sparsely punctate on basal half and sides, with minute setae in punctures; ventral margin finely serrate, with three robust setae of which the apical one is more distant; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, sparsely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere slightly longer than dorsal tibial spur, distal metarsomeres lacking in holotype. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 21 I���K. Habitus: Fig. 21L. Female unknown. Diagnosis. Maladera longruiensis Ahrens, Fabrizi & Liu, sp. n. is very similar to M. fereobscurata Ahrens, Fabrizi & Liu, sp. n. and M. shenoqiae Ahrens, Fabrizi & Liu, sp. n., M. longruiensis Ahrens, Fabrizi & Liu, sp. n. being distinguished by lacking a serrated line beside the dorsal margin of the metatibia, and by the shape of the parameres, which have long basal and distal filiform processes, structures that are absent in the two other species. Etymology. The species name (adjective in the nominative singular) is derived from its type locality, Longrui. Variation. Length: 5.9���6.9 mm, length of elytra: 4.2���6.6 mm, width: 3.6���5.1 mm. Distribution. See map (Fig. 96) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 104-105, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera fangchengensis Ahrens, Fabrizi & Liu, sp. n. Figures 6 E���H, 87 Type material examined. Holotype: ♂ ���Fulong, Fangcheng, Guangxi, 25.V. 1999, 200m, leg. He Xin ��� (IZAS). Paratypes: China: 1 ♂ ��� Beidou, Napo, Guangxi, 22.VI. 2000, 550m, leg. Li Wenzhu ��� (ZFMK), 1 ♂ ��� Banbaxiang, Fangcheng, Guangxi, 4.VI. 2000, 550m, leg. Yao Jian ��� (IZAS), 1 ♂ ��� Longrui, Ningming, Guangxi, 25.V. 1984, 180m, leg. Li Fasheng ��� (CAU). Thailand: 1 ♂ ��� Thai 28-31/5 1995 19.27N 98.20E Soppong, 1500 m, Vit Kub��n leg./ TS 43/ 309 Sericini Asia spec.��� (CP), 1 ♂ ���Thai-NE, Loei prov., Phu Kradung N.P., 16��52���N 101��49���E, 16.��� 18.v.1999 1000 m D. Hauck leg.��� (CP). Vietnam: 1 ♂ ��� Yen-Bai Vietnam 28.��� 29.5.1990 J. Seck��y leg.��� (ZFMK), 1 ♂ ���N Vietnam (Tonkin) pr. Hoang Lien Son Yen Bai 10.V.1990 P. Pachol��tko leg./ VS43/ 361 Sericini Asia spec.��� (CP). Description. Length: 6.5 mm, length of elytra: 5.0 mm, width: 4.0 mm. Body short oval, light reddish brown, antenna yellow, labroclypeus moderately shiny, remainder of dorsal surface dull, glabrous except for a few small setae on head and lateral margins of pronotum and elytra. Labroclypeus moderately trapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to strongly rounded anterior angles, lateral margin and ocular canthus producing an indistinct angle, margins weakly reflexed, anterior margin moderately emarginate medially; surface weakly convex medially, shiny, finely and moderately densely punctate, with a few erect setae anteriorly; frontoclypeal suture feebly incised and weakly curved; smooth area anterior to eye twice as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely scarcely punctate, with a single terminal seta. Frons dull, anterior quarter shiny, with fine, sparse punctures, glabrous except for a few setae beside eyes. Eyes small, ratio of diameter/ interocular width: 0.53. Antenna with ten antennomeres; club with three antennomeres, as long as remaining antennomeres combined. Mentum elevated and flattened anteriorly. Pronotum transverse, widest at posterior third, lateral margins evenly convex and narrowed towards strongly produced and sharp anterior angles as well as posteriorly; anterior margin weakly convex, with complete and fine marginal line, posterior angles strongly rounded; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only; anterior and lateral margins setose; hypomeron carinate but not produced ventrally. Scutellum wide, triangular, with fine and moderately dense punctures. Elytra oblong, widest in posterior third, striae weakly impressed, finely and sparsely punctate, intervals weakly convex, with fine, moderately dense and evenly distributed punctures; penultimate lateral interval with a few sparse fine, short setae, remainder of punctures with minute setae only; epipleural margin robust, ending at widely rounded external apical angle of elytra, epipleura sparsely setose, apical border narrowly membranous, apex covered with short microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, sparsely setose; metacoxa glabrous except for numerous long setae laterally; abdominal sternites finely and densely punctate, some punctures with very short or minute setae, each sternite with a distinct transverse row of coarse punctures each bearing a short seta, penultimate sternite apically with a wide shiny smooth chitinous border of half sternite length. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.7. Pygidium weakly convex, dull, finely and moderately densely punctate, glabrous except a few short setae apically. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur wide, shiny, anterior margin acute, lacking an adjacent serrated line, surface very sparsely and superficially punctate, anterior row of setae complete; posterior ventral margin feebly concave medially, moderately widened in apical half and not serrate apically, posterior margin dorsally not serrated, glabrous. Metatibia wide and short, widest at middle, ratio width/length: 1/1.7, dorsal margin sharply carinate, with two groups of spines, basal one at middle, apical one at three quarters of metatibial length, beside dorsal margin basally with one or two fine setae; lateral face weakly longitudinally convex, nearly entirely impunctate; ventral margin finely serrate, with four strong equidistant spines; medial face impunctate, apex shallowly concave interiorly near tarsal articulation. Tarsomeres impunctate dorsally, with sparse, short setae ventrally; metatarsomeres with a strongly serrated longitudinal ridge and a strong, smooth carina beside it, first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 6 E���G. Habitus: Fig. 6H. Female unknown. Diagnosis. Maladera fangchengensis Ahrens, Fabrizi & Liu, sp. n. differs from Indian species of Hemiserica in lacking a ���snout��� on the head and the simply shaped parameres; from the other Chinese species (see below), in the body size being smaller (Etymology. The name of the new species is derived from its occurrence in Fangcheng county (adjective in the nominative singular). Variation. Length: 6.5���7.1 mm, length of elytra: 5.0��� 5.2 mm, width: 4.0��� 4.4 mm. Distribution. See map (Fig. 87) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 37-38, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera yakouensis Ahrens, Fabrizi & Liu, sp. n. Figures 25 I���L, 98 Type material examined. Holotype: ♂ ���[China] Yunnan, Yakou, 2012-V-11 / LW-1289��� (ISAZ). Description. Length: 8.1 mm, length of elytra: 6.8 mm, width: 5.8 mm. Body oblong-oval, dark brown, antenna yellow, shiny iridescent, dorsal face nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles moderately rounded, anterior margin distinctly emarginate, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex medially, finely, densely punctate, with a few long setae anteriorly; frontoclypeal suture finely incised, weakly curved; smooth area anterior to eye convex, 1.5 times as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely densely punctate, with one terminal seta. Frons with dense and fine punctures, with a few fine setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.58. Antenna with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest at base, lateral margins evenly convex and weakly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles moderately rounded; anterior margin convex, with fine complete marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous, with a shallow median longitudinal impression; lateral margins sparsely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures, impunctate medially. Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, moderately dense punctures, with minute setae in punctures, penultimate lateral interval with a few fine setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.4. Pygidium dull, strongly convex, finely and densely punctate, with short smooth midline, with numerous long and short, fine setae in apical half. Legs moderately long and narrow, shiny; femora with two longitudinal rows of setae, coarsely and moderately densely punctate. Anterior margin of metafemur acute, with adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia long and wide, widest at apex, ratio of width/length: 1/3.6, moderately carinate dorsally, with two groups of spines, basal group at three fifths, apical group at four fifths of metatibial length, in basal half with a few short and fine single setae and a continuous serrated line beside entire dorsal margin until the basal group of spines; lateral face longitudinally convex, with sparse fine punctures in basal half, minutely setose; ventral margin finely serrate, with three equidistant robust setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere distinctly longer than following tarsomere and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 25 I���K. Habitus: Fig. 25L. Female unknown. Diagnosis. Maladera yakouensis Ahrens, Fabrizi & Liu, sp. n. differs from all other large species of the M. egregia group in that the fused left and right parameres have two pronounced, pointed tips. Etymology. The name (adjective in the nominative singular case) of the new species is derived from its type locality, Yakou. Distribution. See map (Fig. 98) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 120-121, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera flavipennis Ahrens, Fabrizi & Liu, sp. n. Figures 52 E���H, 116 Type material examined. Holotype: ♂ ���CH-Guizhou NE 27.V.-3.VI. 20 km NW of Jiankou 1995 Fanjing Shan-Kuaichang E. Jendek & O. ��au��a leg/ Coll. P. Pacholatko / CS23/ 562 Sericini Asia spec.��� (CP). Paratypes: 1 ♂, 2 ♀♀ ���CH-Guizhou 21.-26.V. 50 km N of Kaili 1995 Shibing-Yuntai Shan E. Jendek & O. ��au��a leg/ Coll. P. Pacholatko��� (CP, ZFMK). Description. Length: 10.2 mm, length of elytra: 7.3 mm, width: 6.2 mm. Body oval, head, legs and ventral surface black, pronotum, elytra and antenna yellowish brown, dorsal shiny, dorsal surface glabrous. Labroclypeus wide and trapezoidal, much wider than long, widest at base, lateral margins straight and strongly convergent anteriorly, anterior angles moderately rounded, anterior margin weakly emarginate medially, margins moderately reflexed, lateral margins produce an indistinct angle with the ocular canthus; surface strongly convex, coarsely and very densely punctate, glabrous, with a row of fine erect seta behind the anterior margin; frontoclypeal suture indistinctly incised, not elevated and bluntly angled medially; smooth area anterior to eye moderately wide, almost flat, twice as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely densely punctate, with a single and short terminal seta. Frons with dense, irregularly scattered coarse punctures, partly fusing with each other, with a single short seta beside the middle of each eye. Eyes very small, ratio diameter/ interocular width: 0.4. Antenna with ten or nine antennomeres (with tendency of symphysocerie; Krell 1992), club with three antennomeres and straight, distinctly shorter than the remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum slightly produced medially, transverse, with deep median emargination. Pronotum subtrapezoidal, widest at base, lateral margins in basal two thirds straight and strongly convergent, in anterior third weakly convex and convergent, anterior angles distinctly produced and acute, posterior angles blunt, only weakly rounded at the tip; anterior margin weakly convex produced medially, with a very fine and complete marginal line, base of pronotum without marginal line; surface very densely and coarsely punctate, punctures partly fusing with each other, glabrous; anterior and lateral margins glabrous; hypomeron carinate and slightly produced ventrally. Scutellum yellow, wide triangular, at apex sharp, with coarse and very dense punctures, along the middle with a smooth impunctate and slightly elevated carina, glabrous. Elytra widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals moderately convex, with fine and dense punctures concentrated along the striae, except a few single and short setae on lateral intervals glabrous; epipleural margin fine, ending at strongly curved external apical angle of elytra; epipleura sparsely shortly setose, apical border with a fine fringe of microtrichomes (100x). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with fine, short or very minute setae, metacoxa glabrous; abdominal sternites except the ultimate one shiny, finely and densely punctuate, with a transverse row of short setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.24. Pygidium moderately convex and dull, coarsely and densely punctate, without smooth midline, with a few longer setae on the apical part. Legs slender and moderately long; femur with two longitudinal row of setae, finely and sparsely punctate; metafemur shiny, behind the posterior longitudinal row of setae punctures finer and more dense, anterior margin acute, without serrated line behind anterior margin, posterior margin completely finely serrate ventrally, not widened apically, posterior margin finely serrate dorsally, glabrous. Metatibia slender and not flattened, moderately long, widest at apex, ratio of width/ length: 1/ 4.2, moderately carinate dorsally, with a single group of spines at three quarters of metatibial length, the basal one is reduced to a single short spine shortly before the middle of metatibia, basally with just one other short single spine at basal third of metatibia; lateral face longitudinally convex, coarsely and densely punctate, with minute setae in the punctures; ventral margin finely serrated, with three robust equidistant setae; medial face coarsely and densely punctate, glabrous, apex finely serrated, interiorly near tarsal articulation weakly concavely trunctate. Tarsomeres dorsally very sparsely finely punctate, neither laterally nor dorsally carinate, ventrally finely sparsely setose; metatarsomeres ventrally with a strongly serrated ridge, first metatarsomere slightly shorter than the two following tarsomeres combined and a quarter of its length longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 52 E���G. Habitus: Fig. 52H. Diagnosis. Maladera flavipennis Ahrens, Fabrizi & Liu, sp. n. differs from the other species of the M. brunnescens group in being yellow dorsally and blackish ventrally (compared to the reddish brown color of the other species), as well as having short, simple parameres lacking lateral lobes or teeth, and having a pair of apical lateral processes on the phallobase. Etymology. The species name (noun in nominative singular) is derived from the combined Latin words flavus- (yellow) and pennes (wings). Variation. Length: 9.5���10.2 mm, length of elytra: 7.0��� 7.3 mm, width: 5.2���6.2 mm. Female: eyes as large as in male; antennal club with three antennomeres, little shorter than remaining antennomeres combined; pygidium weakly convex and shiny. Distribution. See map (Fig. 116) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 227-229, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Krell, F. - T. (1992) Verschmelzung von Antennomeren (Symphysocerie) als Regelfall bei Temnorhynchus repandus Burmeister, 1847, sowie phylogenetische, taxonomische, faunistische und nomenklaturische Anmerkungen zu diversen Taxa dieser Gattung (Coleoptera. Scarabaeoidea, Melolonthidae, Dynastinae, Pentodontini). Deutsche Entomologische Zeitschrift, 39, 295 - 367. https: // doi. org / 10.1002 / mmnd. 19920390408"]}

Maladera maoershana Ahrens, Fabrizi & Liu, sp. n. Figures 71 I���N, 127 Type material examined. Holotype: ♂ ��� China, NE Guangxi, 5-19.VI. Mao Ershan, 1500-1600m for. Park, 25��55���N 110��28���E Jaroslav Turna leg., 2011/ 1039 Sericini Asia spec.��� (ZFMK). Paratypes: 1 ♂, 3 ♀♀ ��� China, NE Guangxi, 5-19.VI. Mao Ershan, 1500-1600m for. Park, 25��55���N 110��28���E Jaroslav Turna leg., 2011��� (ZFMK), 1 ♂ ���[China] Mt. Mao���ershan, Guangxi, 14.VII.1985, 1900m, leg. Liao Subai / LW-782��� (IZAS), 1 ♂ ���[China] Mts. Maoershan, Guangxi, 15.VII.1985, 1100m, leg. Wang Ziqing / LW-177��� (IZAS). Description. Length: 8.4 mm, Length: of elytra 6.4 mm, width: 5.0 mm. Body oblong oval, dark brown, antenna yellowish, labroclypeus, anterior frons and pronotum shiny, remainder of dorsal surface dull, glabrous except a few long setae on head, pronotum and elytra. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and convergent to rounded anterior angles, lateral margin and ocular canthus producing a blunt angle, margins weakly reflexed; anterior margin moderately emarginate medially; surface flat, shiny, densely, finely punctate, with few long, erect setae anteriorly; frontoclypeal suture weakly elevated, finely incised and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and wide, sparsely and finely punctate, with one terminal seta. Frons posteriorly dull, anteriorly shiny, with fine, irregularly dense punctures, with a few single setae beside eyes and numerous adpressed ones on posterior disc. Eyes small, ratio of diameter/interocular width: 0.55. Antenna with 10 antennomeres; club with three antennomeres, straight, 1.1 times as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum subtrapezoidal, moderately wide, widest at base, lateral margins moderately convex and evenly convergent anteriorly, anterior angles produced and sharp, posterior angles blunt; anterior margin convex, marginal line widely interrupted medially; surface finely and densely punctate, with minute setae in punctures, with a very shallow longitudinal impression along midline; anterior and lateral margins sparsely setose, setae of anterior margin lacking in middle; hypomeron carinate. Scutellum wide, triangular, dull, with fine and dense punctures, each bearing a minute seta, impunctate along middle. Elytra oblong, widest at posterior third, striae weakly impressed, finely and densely punctate, even intervals flat, with fine, dense punctures, odd intervals weakly convex, dense punctures concentrated along striae and with a few fine, erect, short setae, otherwise punctures with minute setae; epipleural margin robust, ending at the strongly rounded external apical angle of elytra, epipleura densely setose, apical border membraneous, with a rim of short microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, metasternal disc densely setose, metacoxa glabrous except for a several long setae laterally. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.27. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, punctures with microscopic setae, penultimate sternite apically with a shiny smooth chitinous border, which is one quarter as long as sternite. Pygidium moderately convex, with fine dense punctures and minute setae in punctures, on apical half with a few long setae. Legs moderately long and narrow; femora with two longitudinal rows of setae, finely and moderately densely punctate. Metafemur shiny, partly dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin medially straight, weakly widened in apical half and not serrate, dorsally not serrated, finely setose. Metatibia moderately wide and long, widest at apex, ratio width/length: 1/4.0, dorsally longitudinally convex, with two groups of spines, basal one at middle, apical one at four fifths of metatibial length, with a few fine spines basally; lateral face longitudinally convex, with moderately dense and fine punctures, with minute setae in punctures; ventral margin serrate, with three equidistant spines; medial face impunctate, apex interiorly near tarsal articulation shallowly concave. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge and smooth subventral longitudinal carina; first metatarsomere distinctly shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 71 I���M. Habitus: Fig. 71N. Diagnosis. Maladera maoershana Ahrens, Fabrizi & Liu, sp. n. is very similar to M. xingkeyangi Ahrens, Fabrizi & Liu, sp. n. and M. shenglongi Ahrens, Fabrizi & Liu, sp. n. in external appearance. However, in M. maoershana, the left paramere is longer, while both lobes of the right paramere are well separated (compared to the other two species); furthermore, the phallobase has two ventral processes producing a central cavity. Etymology. The name of the new species is derived (adjective in the nominative singular case) from its type locality, Mao Ershan. Variation. Length: 8.4���9.2 mm, Length: of elytra 6.4���6.8 mm, width: 5.0��� 5.1 mm. Female: eyes little smaller than in male, ratio of diameter/interocular width: 0.52; antennal club with three antennomeres, little shorter than remaining antennomeres combined; pygidium weakly convex. Distribution. See map (Fig. 127) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 305-306, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera ventralis (Brenske, 1897) Autoserica ventralis Brenske, 1897: 397. Maladera ventralis: Ahrens 2006b: 238; 2007b: 25. Remarks. The species was originally described from China, Kiukiang (Pratt, Mai 1887). The types of this species, which Brenske (1897) indicated were deposited in the former Museum Tring, have not been found in any museum. The type specimens could be labelled ��� ventricosa ��� since Brenske (1897: 398) mentioned that he subsequently changed the name to Autoserica ventralis., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on page 375, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310","Ahrens, D. (2006 b) Sericinae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248.","Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1"]}

Maladera filigraniforceps Ahrens, Fabrizi & Liu, sp. n. Figures 52 A���D, 116 Type material examined. Holotype: ♂ ���Kuatun (2300m) 27,40n.Br. 117,40.��. L. J. Klapperich 7.6.1938 (Fukien)/ 824 Sericini: Asia spec.��� (ZFMK). Paratypes: China: 1 ♂ ���Fukien, S. China Shaowu: Tachulan 1000m. T. Maa / 5.VI.42��� (BMH), 1 ♂ ��� China, W Fujian, 3.-4.VI. Emei Feng, 1200-1500m 27��01���N 117��04���E Jaroslav Turna leg., 2008��� (ZFMK), 3 ♂♂, 1 ♀ ��� China: Fujian; Tongguzhang 1600m, Jianling vii-2004 / Phil 11-2004-2��� (ZFMK), 4 ♂♂ ���[China] Jiugongshan Tongshan, S-Hubei, 1.V.2004, Leg Wen ��� (ZFMK), 1 ♂ ��� China, Guangdong prov. W of Qixing, 1.-3.v.2011 Heishiding (stream; pools) (forested stream valley; at light) 23��27.9���N, 111��54,3���E, 190m M. Fik��ček & J. H��jek leg.��� (NMPC), 1 ♂ ���[China] Huangjing, Luzhou, Sichuan, 16.VII.2002, leg. Bai Ming, Wang Jianfeng ��� (HBUM), 1 ♂ ���[China] Guadun, Mt. Wuyishan, Fujian, 22-23.V.2004, leg. Yuan Caixia, Lijing ��� (HBUM). Vietnam: 1 ♂ ��� Vietnam N (Sa Pa) Lao Cai Prov., 250km from Hanoi bearing 31��, Sa Pa vill. env. Hoang Lien Son Nat. Res. 9.- 15.6.1998 1250m, leg. A. Napolov ��� (Coll. Napolov, Riga), 1 ♂ ��� Vietnam N (Sa Pa) Lao Cai Prov., 250km from Hanoi bearing 31��, Sa Pa vill. env. Hoang Lien Son Nat. Res. 27.V.-3.VI.1998 1250m leg. A. Napolov ��� (CNA), 1 ♂ ���N Vietnam (Tonkin) pr. Vinh Phu 1990 Tam Dao 6.-9. V. P. Pacholatko leg./ VS88��� (CP), 2 ♂♂ ���N-Vietnam Vinh Phuc Prov., vic. Tam Dao town, Tam Dao NP, 02.-5.V.2013, 21��27���N 105��38���E, 700- 1000m, A. Skale ��� (CASH). Laos: 2 ♂♂ ���Laos-NE, Huoa Phan prov., 20��13���09-19������N 103��59���54������- 104��00���03������E, 1480-1510m, Phou Pane Mt., 22.iv.-14.v.2008, Vit Kuban leg.��� (ZFMK), 1 ♂ ���NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20��12���N, 104��01���E; 11.iv.-15.v.2012; 1300-1900m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012/13��� (ZFMK), 1 ♂ ��� Laos: NE; Hua Phan prov. Ban Saluei; Phu Phan Mt., 20��15���N 104��02���E; 1500-2000m; D. Hauck leg.; 26.iv.-11.v.2001 ��� (CP). Description. Length: 10.5 mm, length of elytra: 7.0 mm, width: 6.0 mm. Body oblong, reddish brown, antenna yellow, dorsal surface shiny, almost glabrous. Labroclypeus subrectangular, widest at base, lateral margins straight and weakly convergent to strgonyl convex anterior angles, lateral border and ocular canthus produce a distinctly blunt angle, anterior margin widely emarginate medially, margins weakly reflexed; surface flat and shiny, finely and moderately densely punctate, with numerous coarse punctures in anterior half bearing fine and erect setae; frontoclypeal suture indistinctly incised and medially weakly angled; smooth area in front of eye small, 1.5 times as wide as long; ocular canthus moderately long and slender (1/3 of ocular diameter), densely finely punctate, with two terminal setae. Frons with fine, sparsely scattered punctures, with a few setae beside eyes and behind frontoclypeal suture. Eyes moderately small, ratio of diameter/ interocular width: 0.58. Antenna yellow, with ten antennomeres; club with three segments, as long as remaining segments together. Mentum elevated, anteriorly flattened and glabrous. Pronotum moderately wide, widest at base, lateral margins in basal half straight and weakly convergent anteriorly, in anterior half convex and moderately convergent, anterior angles strongly produced and sharp, posterior angles sharp; anterior margin weakly produced medially, base without marginal line; surface finely and densely punctate, glabrous; anterior and lateral border sparsely setose. Scutellum wide, triangular, with fine and dense punctures, glabrous. Elytra moderately oblong, widest behind middle, striae moderately impressed, finely and densely punctate, intervals moderately convex, with fine and sparsely punctures concentrated along striae, a few punctures on odd intervals with sparse fine, erect setae; epipleural margin robust, ending at strongly curved external apical angle of elytra; epipleura sparsely setose, apical border membraneous, covered with very short microtrichomes. Ventral surface shiny, coarsely and densely punctate, almost glabrous, metacoxal plates laterally with a few long setae; each abdominal sternite with a transversal row of coarse punctures bearing short and fine setae between fine and dense punctation, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as sternite. Mesosternum between mesocoxae a little less wide than mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.4. Pygidium strongly convex, finely and moderately densely punctate, without smooth midline, surface shiny, with numerous fine and long setae at apex; apical margin medially not emarginate. Legs moderately slender; femora shiny, with two longitudinal rows of setae, between the rows only sparsely punctate; anterior margin of metafemur sharply carinate, adjacent to anterior margin without serrated line; posterior margin ventrally in apical half finely serrated, little widened at apex, completely serrated dorsally, in basal half with a few fine setae. Metatibia moderately slender and moderately long, widest at apex, ratio width/ length: 1/ 3.2; dorsal margin moderately carinate, with two groups of spines, basal one shortly before middle, apical one at three quarters of metatibial length, basally with two single setae with punctures having a serrated margin; lateral face longitudinally convex, with moderately dense, superficial and partly longitudinally impressed punctures, glabrous; ventral margin with three robust spines of which the apical two are more distant; internal face sparsely, finely punctate, apex shallowly emarginate interiorly near tarsal articulation. Tarsomeres distinctly and densely punctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside it with an additional longitudinal margin, glabrous, laterally moderately carinate, dorsally with longitudinal wrinkles; first metatarsomere as long as following two tarsomeres combined and about one third longer than dorsal tibial spur. Protibiamoderately long, bidentate; protarsal claws missing in holotype. Aedeagus: Fig. 52 A���C. Habitus: Fig. 52D. Diagnosis. Maladera filigraniforceps Ahrens, Fabrizi & Liu, sp. n. differs from the other unicolored, reddish brown species of the M. brunnescens group in having the phallobase weakly widened apically and the parameres composed of two filiform branches. Etymology. The name of the new species (adjective in the nominative singular) is derived from the combined Latin word ��� filigranus ��� (narrow) and ��� forceps ��� (pins), with reference to the long, filigranous shape of the parameres. Variation. Length: 8.9���10.6 mm, length of elytra: 6.9���7.5 mm, width: 5.8���6.2 mm. Protarsal claws symmetric. Female: eyes smaller than in male, ratio of diameter/interocular width: 0.49; antennal club with three antennomeres, distinctly shorter than remaining antennomeres combined; pygidium weakly convex. Distribution. See map (Fig. 116) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 226-227, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera enigma Ahrens, Fabrizi & Liu, sp. n. Figures 10 I–L, 91 Type material examined. Holotype: ♂ “[China] Jiangxi, Mt. Jinggangshan Songmuping, under the stone of the stream, 2011-V-27, leg. Song Keqing / LW-1160” (ISAZ). Description. Length: 9.4 mm, length of elytra: 7.2 mm, width: 5.2 mm. Body oval, dark brown, antenna yellow, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous. Labroclypeus wide, trapezoidal, lateral margins straight and convergent, producing with ocular canthus an indistinct angle, anterior angles strongly rounded, anterior margin weakly emarginate medially, margins weakly reflexed; surface flat, base narrowly dull, finely and densely punctate, with numerous erect setae in slightly larger punctures; frontoclypeal suture finely incised, bluntly angled medially; smooth area in front of eyes 1.5 times as wide as long; ocular canthus moderately wide and long (equals 1/3 of ocular width), finely and sparsely punctate, without terminal seta. Frons with superficial, moderately dense punctures, with a few single setae beside eyes and behind frontoclypeal suture. Antenna with ten antennomeres, club with three antennomeres, as long as remaining antennomeres combined. Eyes small, ratio diameter/ interocular width: 0.55. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins moderately but evenly convex and convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt; anterior margin nearly straight, with a fine marginal line being widely interrupted medially; basal margin without marginal line; surface finely and moderately densely punctate, setae of lateral and anterior margin fine but sparse, punctures with microscopic setae only. Hypomeron carinate, not ventrally produced. Scutellum wide, triangular, punctation as on pronotum. Elytra widest behind middle, striae distinctly impressed, finely punctate, intervals weakly convex, finely and densely punctate, on odd intervals punctures concentrated along striae and with a few single short setae; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a very fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.67. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium weakly convex, finely and densely punctate, lateral and apical margins with very short setae. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur dull, anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, strongly widened in apical half, as dorsal posterior margin not serrated, glabrous. Metatibia long and moderately wide, ratio width/ length: 1/ 3.1, sharply carinate dorsally, with two groups of spines, basal one at middle, apical one at four fifths of metatibial length, basally with a few robust and short spines; lateral face weakly longitudinally convex, finely and sparsely punctate on sides and basal half; medial face impunctate and glabrous, apex concavely emarginate interiorly near tarsal articulation. Tarsomeres impunctate dorsally, with dense, fine setae ventrally; metatarsomeres ventrally glabrous, with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined, little longer than dorsal tibial spur. Protibia moderately long, bidentate, without blunt external extension at middle; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 10 I–K. Habitus: 10L. Female unknown. Diagnosis. Maladera enigma Ahrens, Fabrizi & Liu, sp. n. resembles other species of Omaladera in the shape of phallobase and and parameres, however, it differs in characteristics of labroclypeus (dull at base, not carinate medially). Etymology. The name of the new species is derived from Greek word, enigma (Engl. enigma ), with reference on to uncertain systematic position of the species (noun in apposition). Distribution. See map (Fig. 91) and Table 1.

Maladera parabrunnescens Ahrens, Fabrizi & Liu, sp. n. Figures 51 E���H, 116 Type material examined. Holotype: ♂ ���[China] Mt. Wuyanling, Taishun, Zhejiang, 28.VII-3.VIII.2005, leg. Ba Yibin ��� (HBUM). Description. Length: 10.2 mm, width: 7.5 mm, length of elytra: 6.1 mm. Body oblong, reddish brown, antenna yellow, dorsal surface shiny, almost glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins slightly curved and strongly convergent to feebly rounded anterior angles, lateral border and ocular canthus produce a distinctly blunt angle, anterior margin widely emarginate medially, margins weakly reflexed; surface distinctly convex medially and shiny, finely and densely punctate, with numerous coarse punctures in anterior half bearing fine and erect setae; frontoclypeal suture very indistinctly incised and medially weakly angled; smooth area in front of eye very small, 1.5 times as wide as long; ocular canthus moderately long and moderately slender, densely finely punctate, with one terminal seta. Frons with fine, sparsely scattered punctures, with a few setae beside eyes and behind frontoclypeal suture. Eyes moderately small, ratio of diameter/ interocular width: 0.64. Antenna yellow, with ten antennomeres; club with three segments, as long as remaining segments together. Mentum elevated, anteriorly flattened and glabrous. Pronotum moderately wide, widest at base, lateral margins in basal half straight and moderately convergent anteriorly, in anterior half convex and strongly convergent, anterior angles strongly produced and sharp, posterior angles sharp; anterior margin weakly produced medially, base without marginal line; surface finely and densely punctate, glabrous; anterior and lateral border sparsely setose. Scutellum wide, triangular, with fine and dense punctures, glabrous. Elytra moderately oblong, widest behind middle, striae moderately impressed, finely and densely punctate, intervals moderately convex, with fine and sparsely punctures concentrated along striae, a few punctures on odd intervals with sparse fine, erect setae; epipleural margin robust, ending at strongly curved external apical angle of elytra; epipleura sparsely setose, apical border membraneous, covered with very short microtrichomes. Ventral surface shiny, coarsely and densely punctate, almost glabrous, metacoxal plates laterally with a few long setae; each abdominal sternite with a transversal row of coarse punctures bearing short and fine setae between fine and dense punctation, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as sternite. Mesosternum between mesocoxae a little less wide than mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.04. Pygidium moderately convex, finely and densely punctate, without smooth midline, surface apically dull and with numerous fine and long setae; apical margin medially distinctly emarginate, beside each side of emargination with a blunt tooth. Legs moderately slender; femora shiny, with two longitudinal rows of setae, between the rows only sparsely punctate; anterior margin of metafemur sharply carinate, adjacent to anterior margin without serrated line; posterior margin ventrally in apical half finely serrated, little widened at apex, completely serrated dorsally, in basal half with a few fine setae. Metatibia moderately slender and moderately long, widest at apex, ratio width/ length: 1/ 3.5; dorsal margin moderately carinate, with two groups of spines, basal one at one third, apical one at three quarters of metatibial length, basally with two single setae with punctures having a serrated margin; lateral face longitudinally convex, with dense, coarse and longitudinally impressed punctures, glabrous; ventral margin with three robust spines of which the apical two are more distant; internal face sparsely, finely punctate, apex shallowly emarginate interiorly near tarsal articulation. Pro-and mesotibia as well as metatarsomeres lacking in holotype. Aedeagus: Fig. 51 E���G. Habitus: Fig. 51H. Female unknown. Diagnosis. Maladera parabrunnescens Ahrens, Fabrizi & Liu, sp. n. differs from the similar M. brunnescens (Frey, 1972) in having relatively straight parameres with a long, median lateral sharp tooth. Etymology. The name of the new species (adjective in the nominative singular) is derived from the combined Greek prefix ��� para -��� (close to) and the species name ��� brunnescens ���, with reference to the strong similarity to Maladera brunnescens (Frey). Distribution. See map (Fig. 116) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 224-226, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera peregoi Ahrens, Fabrizi & Liu, sp. n. Figures 47 F–I, 112 Type material examined. Holotype: ♂ “Birmania, S.S.S. Palaing, 1450m. VI.1935 R. Perego ” (MSNM). Paratypes: Myanmar: 1 ♂ “Birmania, S.S.S. Palaing, 1450m. V.-1935 R. Perego ” (ZFMK). Thailand: 1 ♂ “Thai 11- 15.V.1993 Nan Pha Khab Pacholatko & Dembicky leg./ TS57” (CP), 1 ♂ “Thai-N, Nan prov., 19°13’N 101°7’E, Doi Phukha N.P. Headq., 22.-26.iv.1999, ca 1500m, D. Hauck leg.” (CP). Vietnam: 1 ♂, 1 ♀ “Museum Paris (coll. Parendel) Coll. J. Chatarmy 1914/ Ngap-Son Tonkin” (MNHN), 2 ♂♂, 1 ♀♀ “Museum Paris (coll. Parendel) Coll. J. Chatarmy 1914[/ Ngap-Son Tonkin]” (MNHN). Laos: 21 ♂♂, 25 ♀♀ “NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20°12’N, 104°01’E; 11.iv.-15.v.2012; 1300-1900m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012” (ZFMK), 1 ♀ “Laos-NE, Huoa Phan prov., 20° 13’09-19’’N 103°59’54’’- 104°00’03’’E, 1480-1510m, Phou Pane Mt., 22.iv.-14.v.2008, Vit Kuban leg.” (ZFMK), 1 ♂ “X-DA4723 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20°12’N, 104°01 ’ 15.-30.iv.2014 leg. C. Holzschuh—ZFMK Ankauf 2014 Serica spLaos_ Phupane 41” (ZFMK), 1 ♂ “X-DA4771 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20°12’N, 104°01 ’ 15.-30.iv.2014 leg. C. Holzschuh—ZFMK Ankauf 2014 Serica spLaos_ Phupane 41” (ZFMK), 1 ♂ “X-DA4773 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20°12’N, 104°01 ’ 15.-30.iv.2014 leg. C. Holzschuh—ZFMK Ankauf 2014 Serica spLaos_ Phupane 41” (ZFMK), 1 ♂ “X-DA4931 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20°12’N, 104°01 ’ 15.- 30.iv.2014 leg. C. Holzschuh—ZFMK Ankauf 2014 Serica spLaos_ Phupane 41” (ZFMK), 1 ♂ “X-DA4772 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20°12’N, 104°01 ’ 15.-30.iv.2014 leg. C. Holzschuh—ZFMK Ankauf 2014 Serica spLaos_ Phupane 41” (ZFMK). China: 1 ♂ “[China] Mt. Eshan, VI.1979, No. 301” (IZAS), 1 ♂ “[China] Jingdong, Yunnan, 30.VI.1956, 1170m, leg. Zagulaev ” (IZAS), 1 ♂ “[China] Jingdong, Yunnan, 6.VI.1956, 1170m, leg. Krischanovskna ” (IZAS). Description. Length: 5.2 mm, length of elytra: 3.4 mm, width: 3.0 mm. Body oblong-oval, dorsal face reddish brown, dull, antenna yellow, head shiny, except some single setae on head dorsal surface nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins convex and distinctly convergent anteriorly, anterior angles moderately rounded, anterior margin emarginate medially, margins strongly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface flat, finely and very densely punctate, with a few larger punctures each bearing an erect seta; frontoclypeal suture indistinct, evenly curved; smooth area anterior to eye weakly convex, as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely and densely punctate, with a terminal seta. Frons completely shiny, with very dense, fine punctures, behind frontoclypeal suture and beside eyes with a few long setae in large sparse punctures. Eyes moderately large, ratio diameter/ interocular width: 0.7. Antenna with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest at base, lateral margins evenly convex and strongly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles strongly rounded; anterior margin straight, with complete and fine marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures; lateral anterior and lateral margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures. Elytra widest shortly behind middle, striae finely impressed, finely and sparsely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with minute setae in punctures, odd intervals with a very few single, short setae; epipleural margin robust, ending at weakly curved external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, coarsely and densely punctate, glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.4. Pygidium weakly convex and dull, coarsely and densely punctate, without impunctate midline, glabrous except a few robust setae along apical margin. Legs short and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without adjacent serrated line, anterior row of setae complete; posterior margin smooth, weakly widened at apex and smooth ventrally, not serrate dorsally, finely shortly setose. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.8, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, with a few robust setae basally subparallel to dorsal margin; lateral face longitudinally convex, shiny, sparsely punctate and glabrous; ventral margin finely serrate, with four equidistant, short and robust setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, laterally and dorsally not carinate; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 47 F–H. Habitus: Fig. 47I. Diagnosis. Maladera peregoi Ahrens, Fabrizi & Liu, sp. n. differs from all other other species of the indica group in having the parameres longer than the phallobase and in having a narrow median process on the dorsal apex of phallobase. Etymology. The name (adjective in the nominative singular) of the new species is dedicated to one of the collectors of the species, R. Perego. Variation. Length: 5.0– 5.8 mm, length of elytra: 3.4–3.9 mm, width: 3.0– 3.6 mm. Morphology of the female is the same as that of male, no sexual dimorphism. Female: eyes little smaller than in male, ratio of diameter/interocular width: 0.59; antennal club as in male; pygidium flat. Distribution. See map (Fig. 112) and Table 1.

Maladera songi Ahrens, Fabrizi & Liu, sp. n. Figures 38 E���H, 106 Type material examined. Holotype: ♂ ���[China] Mt. Jianfengling, Hainan, 21.III.1984, leg. Song Shimei ��� (IZAS). Paratypes: 1 ♂ ���[China] Tianchi, Mt. Jianfengling, Hainan, 19.II.1982, leg. Liang Shaoying ��� (LSSYU), 1 ♂ ���[China] Mts. Yinggeling, Hainan, 2-6.XII.2007, leg. Li Jing ��� (ZFMK). Description. Length: 10.7 mm, length of elytra: 8.0 mm, width: 6.9 mm. Body oblong-oval, dark reddish brown, antenna yellow, labroclypeus shiny, dorsal face dull, nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and strongly convergent anteriorly, anterior angles moderately rounded, anterior margin weakly emarginate, margins moderately reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex medially, coarsely, very densely punctate, with a few long setae anteriorly; frontoclypeal suture distinctly incised, weakly curved; smooth area anterior to eye convex, twice as wide as long; ocular canthus moderately long and wide (1/3 of ocular diameter), finely densely punctate, with a terminal seta. Frons dull, with dense and coarse punctures behind frontoclypeal suture, otherwise punctures sparser, with a few single setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.58. Antenna with ten antennomeres; club with three antennomeres and straight, 1.6 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest at base, lateral margins in basal half nearly straight and weakly convergent anteriorly, in anterior half convex and more convergent, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin weakly convex, with fine marginal line, base without marginal line; surface densely and coarsely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; anterior and lateral margin densely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures. Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals flat, with fine, sparse punctures and with minute setae in punctures, penultimate lateral interval with a few single setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.49. Pygidium moderately convex, weakly shiny, coarsely and densely punctate, without smooth midline, with a few long but fine setae along apical margin. Legs moderately long and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, without adjacent continuous serrated line, anterior row of setae complete and duplicate; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, with a few single long setae. Metatibia moderately long and moderately wide, widest shortly before apex, ratio of width/length: 1/2.7, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short and robust single setae and a continuous serrated line beside basal 1/5 of dorsal margin; lateral face longitudinally convex, except a few punctures on sides and base smooth and glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, distinctly concavely emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere distinctly shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 38 E���G. Habitus: Fig. 38H. Female unknown. Diagnosis. Maladera songi Ahrens, Fabrizi & Liu, sp. n. differs from all other species of the Maladera nitens group by the dark and dull surface, anterior margin of the metafemur lacking adjacent serrated line, and the metatibia lacking a serrated line at base beside its dorsal margin. Etymology. The new species is named after one of its collectors, Song Shimei (noun in genitive singular). Variation. Length: 10.1���10.6 mm, length of elytra: 7.5���8.5 mm, width: 6.2���6.9 mm. Distribution. See map (Fig. 106) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 171-173, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Erichson, W. F. (1834) Coleoptera. In: Meyen, F. J. F. (Ed.), Beitrage zur Zoologie gesammelt auf einer Reise um die Erde. 6. Abhandlung. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae, Naturae Curiosorum, 16 (Supplement 1), pp. 219 - 276."]}

Maladera hajeki Ahrens, Fabrizi & Liu, sp. n. Figures 9 A–E, 88 Type material examined. Holotype: ♂ “X-DA2976 China Jiangxi prov. Jinggang Shan Mts., Xiaoxidong (at light), 342m 26°28,0’N, 114°12.9’E 24.iv.2011 M. Fikáček & J. Hájek / 800 Sericini Asia spec.” (ZFMK). Paratype: 1 ♂ “ China Jiangxi prov. 24.iv.2011 Jinggang Shan Mts., Xiaoxidong (at light), 26°28.0’N, 114°12.9’E, 342m M. Fikáček & J. Hájek ” (NMPC). Description. Length: 9.5 mm, length of elytra: 7.3 mm, width: 5.3 mm. Body oval, dark brown, dull, antenna yellow, labroclypeus moderately shiny, glabrous except for a few short setae on head and lateral margins of pronotum and elytra. Labroclypeus trapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing an indistinct angle; margins weakly reflexed; anterior margin moderately emarginate medially; surface weakly convex medially, shiny, finely and densely punctate, with a few erect setae anteriorly; frontoclypeal suture feebly incised and weakly bent medially; smooth area anterior to eye twice as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely scarcely punctate, without terminal seta. Frons dull, with fine, dense punctures, with setae beside eyes and along a transversal line on the posterior part. Eyes small, ratio of diameter/ interocular width: 0.57. Antenna with ten antennomeres; club with three antennomeres, as long as remaining antennomeres combined. Mentum elevated and flattened anteriorly. Pronotum trapezoidal, widest at base, lateral margins weakly evenly convex and narrowed towards strongly produced and sharp anterior angles; posterior angles blunt; anterior margin nearly straight, with complete and fine marginal line; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only; anterior and lateral margins sparsely setose; hypomeron carinate but not produced ventrally. Scutellum wide, triangular, with fine and dense punctures. Elytra oblong, widest in posterior third, striae weakly impressed, finely and sparsely punctate, intervals weakly convex, with fine, sparse and evenly distributed punctures; glabrous, punctures with minute setae only; epipleural margin robust, ending at widely rounded external apical angle of elytra, epipleura sparsely setose; apical border chitinous, without short microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, sparsely setose; metacoxa glabrous except for numerous long setae laterally; abdominal sternites finely and densely punctate, some punctures with very short or minute setae, each sternite with a distinct transverse row of coarse punctures each bearing a short seta, penultimate sternite apically with a wide shiny smooth chitinous border of half sternite length. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.35. Pygidium weakly convex, dull, finely and moderately densely punctate, glabrous except a few longer and shorter setae apically. Legs moderately wide; femora finely and moderately densely punctate, with two longitudinal rows of setae. Metafemur wide, moderately shiny, anterior margin acute, with a fine adjacent and continuously serrated line, surface very sparsely and superficially punctate, anterior row of setae complete; posterior ventral margin moderately widened in apical half and not serrate apically, posterior margin dorsally not serrated, glabrous. Metatibia moderately wide and short, widest at middle, ratio width/length: 1/3.2, dorsal margin sharply carinate, with two groups of spines, basal one at middle, apical one at three quarters of metatibial length, beside dorsal margin basally with a few single fine setae; lateral face weakly longitudinally convex, finely sparsely punctate, along middle impunctate; ventral margin finely serrate, with three strong spines of which the distal one is more distant; medial face impunctate, apex shallowly concave interiorly near tarsal articulation. Tarsomeres impunctate dorsally, with sparse, short setae ventrally; metatarsomeres glabrous ventrally and with a strongly serrated longitudinal ridge as well as a strong, smooth carina beside it, first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 9 A–D. Habitus: Fig. 9E. Female unknown. Diagnosis. Maladera hajeki Ahrens, Fabrizi & Liu, sp. n. is very similar to M. cariniceps (Moser) in shape of the aedeagus; M. hajeki differing in having the right paramere more elongate and having an additional basal, sharply pointed lobe; the left paramere is straight medially but hooked dorsally at apex. Etymology. The new species is dedicated to one of the collectors of the new species, Jiri Hajek (Prague) (noun in the genitive singular case). Variation. Length: 9.2–9.5 mm, length of elytra: 6.5–7.3 mm, width: 5.3–5.6 mm. Distribution. See map (Fig. 88) and Table 1.

Maladera wipfleri Ahrens, Fabrizi & Liu, sp. n. Figures 5 J–N, 86 Type material examined. Holotype: ♂ “Szechuen China DC Graham/ bet Giu Keo Ho & Dawes/ 1300–5000ft Aug. 1–2’25/ 1087 Sericini Asia spec.” (USNM) Description. Length: 10.0 mm, length of elytra: 7.2 mm, width: 5.2 mm. Body oblong-oval, dark reddish brown, antenna yellow, dorsal face shiny, nearly glabrous, ventral face dull. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins straight and convergent anteriorly, anterior angles strongly rounded, anterior margin weakly emarginate, margins moderately reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex, coarsely, very densely punctate, with numerous long erect setae; frontoclypeal suture distinctly incised, weakly curved; smooth area anterior to eye convex, twice as wide as long; ocular canthus moderately long and wide (1/3 of ocular diameter), finely densely punctate, with a terminal seta. Frons shiny, with very dense and coarse punctures, with a few long setae beside eyes and behind frontoclypeal suture. Eyes moderately large, ratio diameter/ interocular width: 0.6. Antenna with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, lateral margins weakly convex and moderately convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin straight, with fine marginal line, base without marginal line; surface very finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; anterior and lateral margin densely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, very dense punctures. Elytra widest in posterior third, striae finely impressed, finely and densely punctate, odd intervals weakly convex, even ones flat, with fine, very dense punctures, with minute setae in punctures, penultimate lateral interval with a few fine setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.5. Pygidium dull, weakly convex, coarsely and densely punctate, without smooth midline, with dense short and numerous long and fine setae. Legs moderately long and wide, shiny; femora with two longitudinal rows of setae, coarsely and moderately densely punctate. Anterior margin of metafemur acute, with adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, moderately widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia short and wide, widest at apex, ratio of width/length: 1/2.4, sharply carinate dorsally, with two groups of spines, basal group at three fifths, apical group at five sixths of metatibial length, in basal half with a few short and robust single setae, with a continuous serrated line beside entire dorsal margin; lateral face longitudinally convex, with dense coarse punctures, punctures setose; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, distinctly deeply emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate and glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere little longer than following tarsomere and as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 5 J–M. Habitus: Fig. 5N. Female unknown. Diagnosis. Maladera wipfleri Ahrens, Fabrizi & Liu, sp. n. is similar to M. opaciventris (Moser), differing in having the first metatarsomere longer and in the shape of the parameres: the left paramere is only half as wide and not sharply pointed at apex. Etymology. The species is named after Benedict Wipfler, technical assistant at the section Coleoptera at the ZFMK, thanking him for his support in the last years (noun in the genitive singular case). Distribution. See map (Fig. 86) and Table 1.

Maladera jiucailingensis Ahrens, Fabrizi & Liu, sp. n. Figures 37 I–L, 105 Type material examined. Holotype: ♂ “ China: Hunan, Jiucai Ling, 25°32’N 111°22’E IV.2006, H~ 1300m Leg. V. Siniaev / 787 Sericini: Asia spec.” (ZFMK). Paratypes. 7 ♂♂, 7 ♀♀ “ China: Hunan, Jiucai Ling, 25°32’N 111°22’E IV.2006, H~ 1300m Leg. V. Siniaev ” (ZFMK), 6 ♂♂, 4 ♀♀ “ China: Fujian Province; Mt. Liang shan-ding, Wuping County, July 2-13, 2009 local collectors Ankauf via Li Jingke 2010” (ZFMK), 23 ♂♂, 11 ♀♀ “ China: Hunan, Jiucai Ling, 25°32’N 111°22’E VII.2006, 1300m leg. Viktor & Sveta Siniaev ” (ZFMK, CP), 4 ♂♂, 4 ♀♀ “ China: Hunan, Jiucai Ling, 25°32’N 111°22’E H~ 1300m VII.2006, Leg./ Coll. Viktor & Sveta Siniaev ” (ZFMK). Description. Length: 8.2 mm, length of elytra: 5.7 mm, width: 4.9 mm. Body oblong-oval, dark reddish brown, elytra reddish brown, antenna yellow, shiny, dorsal face nearly glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins strongly convex and convergent anteriorly, anterior angles strongly rounded, anterior margin widely and distinctly emarginate, margins moderately reflexed; lateral margin and ocular canthus produce a blunt angle; surface flat, coarsely, densely punctate, with a few long setae anteriorly; frontoclypeal suture distinctly incised, weakly curved; smooth area anterior to eye convex, 3 times as wide as long; ocular canthus moderately long and wide (1/3 of ocular diameter), finely densely punctate, with two terminal setae. Frons shiny, with irregularly dense and coarse punctures, with a few single setae beside eyes. Eyes large, ratio diameter/ interocular width: 0.74. Antenna with nine antennomeres; club with three antennomeres and strongly reflexed, 1.8 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, lateral margins in basal half straight and subparallel, in anterior half convex and more convergent, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin weakly convex, with robust marginal line, base without marginal line; surface densely and coarsely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; anterior and lateral margin densely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures. Elytra widest in posterior third, striae finely impressed, finely and densely punctate, odd intervals weakly convex, with fine, sparse punctures concentrated along striae, even intervals flat with even punctures, with minute setae in punctures, penultimate lateral interval with a few single setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface shiny, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.43. Pygidium moderately convex, shiny, coarsely and densely punctate, without smooth midline, with numerous long, fine setae along apical margin. Legs long and moderately wide, shiny; femora with two longitudinal rows of setae, coarsely and densely punctate in basal half, otherwise punctures sparse. Anterior margin of metafemur acute, with adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia moderately long and moderately wide, widest at apex, ratio of width/length: 1/2.8, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short and robust single setae and a continuous serrated line beside basal half of dorsal margin; lateral face longitudinally convex, except a few punctures on base smooth and glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, distinctly concavely emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere distinctly longer than following tarsomere and longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 37 I–K. Habitus: Fig. 37L. Diagnosis. Maladera jiucailingensis Ahrens, Fabrizi & Liu, sp. n. differs from M. nitens (Moser) in having the right paramere concave laterally in basal half, rather than convex as in M. nitens. Etymology. The new species is named after the type locality, Jiucai Ling (adjective in the nominative singular). Variation. Length: 7.2–9.5 mm, length of elytra: 5.1–6.9 mm, width: 4.5–5.6 mm. Female: eyes much smaller than in male, ratio of diameter/interocular width: 0.54; antennal club with three antennomeres, little shorter than remaining antennomeres combined; pygidium weakly convex. Distribution. See map (Fig. 105) and Table 1.

Maladera diaolinensis Ahrens, Fabrizi & Liu, sp. n. Figures 25 A���D, 97 Type material examined. Holotype: ♂ ��� China, W Yunnan Diaolin, 18.VI.1993 E. Jendek & O. ��au��a leg./ 319 Sericini Asia spec.��� (CP) (head of holotype specimen missing). Description. Length: n/a, length of elytra: 6.2 mm, width: 5.1 mm. Body oblong-oval, dark reddish brown, antenna yellow, dull, elytra and labroclypeus iridescent and shiny, dorsal face nearly glabrous; head in holotype missing. Pronotum transverse, widest in posterior third, lateral margins evenly convex and weakly convergent anteriorly and posteriorly, anterior angles distinctly produced and sharp, posterior angles moderately rounded; anterior margin convex, with fine marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; lateral margin densely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures. Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, moderately dense punctures, with minute setae in punctures, penultimate lateral interval with a few fine setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.4. Pygidium dull, moderately convex, finely and densely punctate, with short smooth midline, with numerous long and short, fine setae in apical half. Legs moderately long and narrow, shiny; femora with two longitudinal rows of setae, coarsely and moderately densely punctate. Anterior margin of metafemur acute, with adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia long and wide, widest at apex, ratio of width/length: 1/3.4, moderately carinate dorsally, with two groups of spines, basal group shortly behind the middle, apical group at four fifths of metatibial length, in basal half with a few short and fine single setae and a continuous serrated line beside entire dorsal margin until the basal group of spines; lateral face longitudinally convex, with sparse fine punctures in basal half, minutely setose; ventral margin finely serrate, with five equidistant robust setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere distinctly longer than following tarsomere and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 25 A���C. Habitus: Fig. 25D. Female unknown. Diagnosis. Maladera diaolinensis Ahrens, Fabrizi & Liu, sp. n. differs from M. tiani Ahrens, Fabrizi & Liu, sp. n. in having the fused left and right parameres straight and bearing blunt lateral teeth at middle on each side and the basal group of metatibial setae of is situated shortly behind the middle. Etymology. The name (adjective in the nominative singular case) of the new species is derived from its type locality Diaolin. Distribution. See map (Fig. 97) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 117-119, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera brunnescens (Frey, 1972) group Key to species of the Maladera brunnescens (Frey, 1972) group (³³): 1 Phallobase with an apical lateral process on each side. Body bicolored, dorsal face yellow, ventral face blackish...................................................................... Maladera flavipennis Ahrens, Fabrizi & Liu, sp. n. 1’ Phallobase without apical lateral processes. Body unicolored, reddish brown....................................... 2 2 Phallobase strongly widened at apex. Parameres composed by a single branch (Fig. 51B, F).......................... 3 2’ Phallobase weakly widened at apex. Parameres composed by two filiform branches (Fig. 52B)................................................................................. Maladera filigraniforceps Ahrens, Fabrizi & Liu, sp. n. 3 Parameres strongly curved externally, without a median lateral sharp tooth............ Maladera brunnescens (Frey, 1972) 3’ Parameres straight, with a median lateral sharp tooth............. Maladera parabrunnescens Ahrens, Fabrizi & Liu, sp. n.

Maladera nanpingensis Ahrens, Fabrizi & Liu, sp. n. Figures 63 I���L, 122 Type material examined. Holotype: ♂ ���[China] Milv, Nanping, Shangsi, Guangxi, 3.IV. 2002, 770m, light trap, leg. Xue Huaijun ��� (NKU). Description. Length: 10.6 mm, length of elytra: 8.2 mm, width: 6.2 mm. Body oval, reddish brown, frons slightly darker, antenna yellow, labroclypeus shiny, remainder of dorsal surface dull, with several short setae on the head and at margin of pronotum and elytra. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins convex and convergent to strongly rounded anterior angles, lateral margin and ocular canthus producing a distinct blunt angle, margins moderately reflexed, anteriorly emarginate medially; surface weakly convex, shiny, coarsely and moderately punctate, with moderately long, erect setae; frontoclypeal suture feebly impressed and weakly curved medially; smooth area in front of eye approximately 2.5 times as wide as long; ocular canthus narrow and moderately long, finely punctate, with two terminal setae. Frons dull, with fine, dense punctures, glabrous except for a few setae beside eyes. Eyes moderately large, ratio of diameter/interocular width: 0.53. Antenna with 10 antennomeres; club with three antennomeres, 1.3 times as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum subtrapezoidal, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles produced and sharp, posterior angles blunt; anterior margin weakly convexly produced medially; marginal line medially moderately interrupted; surface finely and densely punctate, with minute setae in punctures only; anterior and lateral margins sparsely setose. Scutellum wide, triangular, with fine and dense punctures, each bearing a single minute seta, impunctate medially. Elytra oblong, widest at middle, striae moderately impressed, finely and densely punctate, intervals moderately convex, with fine, dense punctures concentrated along striae, on odd intervals with a few fine, erect, short setae, otherwise punctures with minute setae; epipleural margin robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border membraneous, with a rim of short microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, sparsely setose, metacoxa glabrous except for a several long setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.7. Abdominal sternites, in addition to fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, punctures with microscopic setae, penultimate sternite apically with a shiny smooth chitinous border, which is one third as long as sternite. Pygidium moderately convex, finely and densely punctate, without smooth midline, punctures with microscopic setae and with a few long setae along apical margin. Legs moderately long and wide; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior margin acute, lacking an adjacent serrated line, anterior row of setae absent, posterior ventral margin straight, weakly widened in apical half and finely serrate, posterior dorsal margin completely serrated, finely setose. Metatibia moderately wide and long, widest at apex, ratio width/length: 1/3.0, dorsally sharply carinate, with two groups of spines, basal one at one third, apical one at three quarters of metatibial length; lateral face with dense and coarse punctures, punctures with minute setae; ventral margin serrate, with three strong spines of which the distal one is more distant; medial face impunctate; apex interiorly near tarsal articulation shallowly concave. Tarsomeres dorsally sparsely finely punctate, with dense setae ventrally; metatarsomeres of holotype missing. Protibia moderately long, bidentate; protarsal claws symmetric. Aedeagus: Fig. 63 I���K. Habitus: Fig. 63L. Female unknown. Diagnosis. Maladera nanpingensis Ahrens, Fabrizi & Liu, sp. n. differs from the closely related and similar Maladera bawanglingana Ahrens, Fabrizi & Liu, sp. n. in having the parameres bifid (split into two branches). Etymology. The new species is named after its type locality, Nanping (adjective in the nominative singular). Distribution. See map (Fig. 122) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 271-272, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera futschauana (Brenske, 1897) Figures 21 E���H, 95 Autoserica futschauana Brenske, 1897: 404. Maladera futschauana: Ahrens 2006b: 237, 2007b: 23. Autoserica atavana Brenske, 1902: 62, syn. n. Autoserica montivaga Moser, 1915: 348, syn. n. Type material examined. Holotype (futschauana): ♀ ��� ♀ / China Futschau/ coll. Brenske / Autoserica futschauana Type Brsk/ futschauana Brsk. ��� (ZMHB). Syntype (atavana): 1 ♂ ���Museum Paris Louang-Prabang A Theng A. Pavie 1888/ Autoserica atavana Typ. ♂ Brenske/ Type��� (MNHN). Syntype (montivaga): 1 ♂ ��� Tonkin Montes Mauson April, Mai 2-3000��� H. Fruhstorfer / Autoserica montivaga Type Mos.��� (ZMHB). Additional material examined. China: 1 ♂ ��� China, E Hubei, 7.-10.V. Dabie Shan, 31.1N 115.8E Wujiashan forest park Jaroslav Turna leg., 2004��� (ZFMK), 1 ♂ ��� Chekiang China / OL Cartwright Collection 1959��� (USNM), 1 ♂ ��� Dujun, Guizhou, 7.VI.1981, leg. Li Fasheng ���(CAU), 1 ♂ ��� Defu, Napo, Guangxi, 5.IV.1998, 1440m, leg. Li Wenzhu ��� (IZAS), 1 ♂ ��� Mt. Eshan, VI.1979, No.310��� (IZAS), 1 ♂ ��� Mt. Yanshan, Guilin, Guangxi, 7.XII.1953 ��� (IZAS), 1 ♂ ��� Moxi, Luding, Sichuan, 13.VI.1983, 1600m, leg. Chai Huaicheng ��� (IZAS), 1 ♂ ��� Yongsheng, Yunnan, VI.1955 ��� (IZAS), 1 ♂ ��� Huangkeng, Jianyang, Fujian, 20.VI.1960, 270- 300m, leg. Zuo Yong ��� (IZAS), 1 ex. (♂) ���Museum Paris Kiang-Si A. David 1875��� (MNHN), 1 ♂ ��� Kuatun (2300 m) 27, 40 n. Br. 117,40 ��. L. J. Klapperich 3.5.1938 (Fukien)��� (ZFMK), 6 ♂♂, 5 ♀♀ ��� China: E-Yunnan; Damaidi 2500m, Guangnan near Vietnam VII-2003 leg. Li et al. ��� (ZFMK), 1 ♂ ��� China, Guangdong prov., Danxia Shan NP, 23.iv.2013 (garden: at light) 25��02.4���N, 113��45.0���E 100 m J. H��jek & J. Růžicka leg.��� (NMPC), 2 ♂♂, 2 ♀♀ ���Museum Paris Se-Tchouen Env. De Ta-Sein-Lou Mo-Sy-Mien P��re Aubert 1902��� (MNHN). Myanmar: 1 ex. ��� Tenasserim Tandong 4000��� Mai Fruhstorfer leg.��� (MNHN), 1 ♂ ���Myanmar N (Burma) 25 km E Putao, H= 800 m Nan Sa Bon vill. 06- 09.05.1998 leg. S. Murzin & V. Sinaev / cf. atavana / 707 Asia Sericini spec.��� (ZFMK). Thailand: 1 ex. ��� N. Thailand 27.III.-3.IV.1990 Doi Inthanon lg. Malicky��� (ZSM), 1 ♂ ��� Thailand bor. Hang Dong 6.1990. lgt. Wimmer��� (ZFMK). Vietnam: 1 ex. ��� N. Vietnam (Tonkin) Tam Dao 12.- 24.5.1989 Pachol��tko leg./ VS 32��� (CP). Cambodia: 1 ex. ��� Cambodia: Kirirom Nat. Park 780 m, 11��21���23������N, 104��4���38������E, 2.-17.I.2000 M. & S. Murzin ��� (ZFMK). Laos: 1 ♂ ���X-DA4704 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera attavana��� (ZFMK), 1 ♂ ���X-DA4925 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera attavana��� (ZFMK), 1 ♂ ���X-DA4821 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera attavana��� (ZFMK), 1 ♂ ���X-DA4824 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera attavana��� (ZFMK), 1 ♂ ���X-DA4670 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera attavana��� (ZFMK). Doubtful record: 1 ♂ ���proxima Burm. Narkondan Ribbe��� (EMAU). Redescription. Length: 6.5 mm, length of elytra: 5.1 mm, width: 4.6 mm. Body oval, dark reddish brown to blackish, antenna yellow, club brown, dorsal surface dull, except some single setae on head dorsal surface nearly glabrous. Labroclypeus wide and trapezoidal, shiny, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles blunt and slightly reflexed, anterior margin moderately emarginate, margins moderately reflexed; lateral margin and ocular canthus produce a blunt angle; surface flat, finely and densely punctate, with a few erect setae anteriorly; frontoclypeal suture indistinctly incised, curved; smooth area anterior to eye flat, 1.5 times as wide as long; ocular canthus long and moderately wide (nearly 1/3 of ocular diameter), finely and sparsely punctate, with a terminal seta. Frons dull, with fine, dense punctures and a few single erect setae beside eyes. Eyes large, ratio diameter/ interocular width: 0.6. Antenna with ten antennomeres; club with three antennomeres and weakly reflexed, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, lateral margins evenly convex and moderately convergent anteriorly, anterior angles moderately produced and right-angled, posterior angles blunt, weakly rounded at tip; anterior margin weakly convex, with fine and complete marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures, otherwise glabrous; lateral and anterior margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, sparse punctures. Elytra widest shortly behind middle, striae distinctly impressed, finely and sparsely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with minute setae in punctures, odd intervals with a few single short setae; epipleural margin robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra membraneous, with narrow rim of microtrichomes (at ca 100x magnification). Ventral surface dull, coarsely and moderately densely punctate, metasternum sparsely covered with sparse fine setae; metacoxa with a few longer setae laterally, otherwise glabrous. Abdominal sternites finely and irregularly densely punctate, glabrous, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.6. Pygidium moderately convex and dull, coarsely and densely punctate, without smooth midline, with a few longer setae along apical margin. Legs moderately wide and short, weakly shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, with anterior margin acute, with continuous serrated line behind anterior margin, setae of anterior row very short; posterior margin smooth and moderately widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia short, widest at apex, ratio of width/length: 1/2.7, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at four fifths of metatibial length, with a few short robust setae basally with serrated margin; lateral face weakly longitudinally convex, sparsely punctate on basal half and sides, with minute setae in punctures; ventral margin finely serrate, with three robust setae of which the apical one is more distant; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, sparsely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 21 E���G. Habitus: Fig. 21H. Distribution. See map (Fig. 95) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 102-104, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310","Ahrens, D. (2006 b) Sericinae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248.","Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1","Brenske, E. (1902) Die Serica - Arten der Erde. VII. Berliner Entomologische Zeitschrift, 47, 1 - 70."]}

Maladera pui Ahrens, Fabrizi & Liu, sp. n. Figures 55 A���D, 118 Type material examined. Holotype: ♂ ���[China] Menghai, Xishuangbanna, Yunnan, 21.VII.1958, 1200-1600m, leg. Pu Fuji / LW-791��� (IZAS). Description. Length: 9.7 mm, length of elytra: 7.2 mm, width: 5.5 mm. Body oval, reddish brown, antenna yellow, dorsal surface dull, except a few small setae on head glabrous. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins convex and convergent to strongly rounded anterior angles, lateral margin and ocular canthus producing a indistinct blunt angle, margins moderately reflexed; anterior margin straight; surface moderately convex medially, coarsely and densely punctate, with a transverse row of long, erect setae anteriorly; frontoclypeal suture feebly impressed and moderately angled medially; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus short and wide, finely punctate, with one terminal seta. Frons dull, with fine, moderately dense punctures, with long setae beside eyes. Eyes small, ratio of diameter/interocular width: 0.5. Antenna with 10 antennomeres; club with three antennomeres, 1.2 times as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum subtrapezoidal, widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles moderately produced and sharp, anterior margin weakly convex, marginal line complete; surface moderately finely and densely punctate, with minute setae in punctures; lateral margins sparsely setose, setae at anterior margin lacking medially; hypomeron carinate. Scutellum wide, triangular, with fine and moderately dense punctures each bearing a single minute seta. Elytra oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals almost flat, with fine, dense punctures, penultimate intervals with a few fine, erect, short setae, otherwise punctures with minute setae; epipleural margin robust, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a fine rim of very short microtrichomes. Ventral surface dull, metasternum with large and dense punctures, densely setose, metacoxa with minute setae in large punctures, with several long setae laterally. Abdominal sternites in addition to generally distributed fine and moderately dense punctures each with a distinct transverse row of coarse punctures each bearing a short and robust seta, otherwise punctures each with a microscopic seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.49. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, punctures with microscopic setae, with a few long setae along apical margin. Legs short and moderately wide; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, anterior margin acute, lacking an adjacent serrated line and anterior longitudinal row of setae, posterior ventral margin medially straight, weakly widened in apical half and not serrate, dorsally not serrated, finely setose. Metatibia moderately wide and long, widest at apex, ratio width/length: 1/3.0, dorsally sharply carinate, with two groups of spines, basal one at middle, apical one at three quarters of metatibial length, without contiguous serrated line basally; lateral face longitudinally convex, superficially densely punctate on basal half; ventral margin finely serrate, with four equidistant spines; medial face impunctate; apex interiorly near tarsal articulation shallowly concave. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside with subventral longitudinal carina; first metatarsomere shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 55 A���C. Habitus: Fig. 55D. Female unknown. Diagnosis. Maladera pui Ahrens, Fabrizi & Liu, sp. n. differs from M. bubengensis Ahrens, Fabrizi & Liu, sp. n. in having the long parameres more narrow and bearing no interior teeth on the medial margin. Etymology. The new species is named after its collector, Pu Fuji (noun in the genitive singular case). Distribution. See map (Fig. 118) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 238-240, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera wupingensis Ahrens, Fabrizi & Liu, sp. n. Figures 53 J���M, 117 Type material examined. Holotype: ♂ ��� China: Fujian Province; Mt. Liang-shan-ding, Wuping County, July 2- 13, 2009 local collector Ankauf via Li Jingke 2010/ 771 Sericini Asia spec.��� (ZFMK). Paratypes: 1 ♂ ��� China: Fujian Province; Mt. Liang-shan-ding, Wuping County, July 2-13, 2009 local collector Ankauf via Li Jingke 2010��� (ZFMK), 1 ♂ ���[China] Mt. Dadongshan, Lianxian County, Guangdong, 5.VI.2001, leg. Pan Lijing ��� (LSSYU), 1 ♂ ��� China, Hunan, Jiucai Ling, 25��32���N 111��22���E, 1300m, VII.2006, leg. Viktor & Sveta Siniaev��� (ZFMK). Description. Length: 11.4 mm, length of elytra: 8.4 mm, width: 6.9 mm. Body wide, oval, dark brown, legs brown, antenna yellow, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous. Labroclypeus moderately wide, trapezoidal, lateral margins slightly convex and convergent, producing with the ocular canthus a nearly straight line, anterior angles moderately rounded, anterior margin distinctly emarginate medially, margins weakly reflexed; surface flat, basal half with dull toment, finely and densely punctate, distance between punctures wider than their diameter, with a few erect setae in slightly larger punctures anteriorly; frontoclypeal suture finely incised, bluntly angled medially; ocular canthus moderately wide and long (equals 1/3 of ocular width), finely and sparsely punctate, with a short terminal seta. Frons with superficial, moderately dense punctures, with a few single setae beside eyes. Antenna with ten antennomeres, club with three antennomeres, distinctly longer than remaining antennomeres combined. Eyes small, ratio diameter/ interocular width: 0.47. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt; anterior margin weakly convex, with a fine marginal line that is widely incomplete medially; basal margin without marginal line; surface finely and moderately densely punctate, setae of lateral and anterior margin fine but sparse, punctures with microscopic setae only. Scutellum wide, triangular, punctation as on pronotum. Elytra widest behind middle, striae distinctly impressed, finely punctate, intervals moderately convex, finely and densely punctate, punctures concentrated along striae; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.78. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium moderately convex, finely and densely punctate, lateral and apical margins with very short setae. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur shiny, anterior margin acute, without adjacent serrated line, anterior longitudinal row of setae reduced; posterior ventral margin almost straight, weakly widened in apical half, ventrally at apical third and dorsally completely serrated, glabrous. Metatibia long and moderately wide, ratio width/ length: 1/ 3.1, sharply carinate dorsally, with two groups of spines, basal one at middle, apical one at four fifths of metatibial length, basally with a few fine and short setae; lateral face weakly longitudinally concave, finely and sparsely punctate, dorsally with a few larger punctures; apex concavely emarginate interiorly near tarsal articulation. Tarsomeres finely densely punctate dorsally, with dense, fine setae ventrally; metatarsomeres ventrally glabrous, with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little longer than following two tarsomeres combined, and a quarter of its length longer than dorsal tibial spur. Protibia moderately long, bidentate, with blunt external extension at middle; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 53 J���L. Habitus: Fig. 53M. Female unknown. Diagnosis. Maladera wupingensis Ahrens, Fabrizi & Liu, sp. n. differs from the other two species of this species group in having the parameres shorter and the dorsoapical median emargination of phallobase shallower. Etymology. The new species is named after its type locality (Wuping) (adjective in nominative singular). Variation. Length: 11.4���11.8 mm, length of elytra: 8.4���8.9 mm, width: 6.9���7.5 mm. Distribution. See map (Fig. 117) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 233-234, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera taiyangheensis Ahrens, Fabrizi & Liu, sp. n. Figures 56 A���D, 119 Type material examined. Holotype: ♂ ��� China, W Hubei / E Chongquing, Shiruguan, 1.-9.VII.2003 pass ~ 5km NW Taiyanghe pitfall traps, 30.65N 109.5E Jaroslav Turna leg., ~ 1700m / 736 Sericini: Asia spec.��� (ZFMK). Paratypes: 4 ♂♂, 43 ♀♀ ��� China, W Hubei / E Chongquing, Shiruguan, 1.-9.VII.2003 pass ~ 5km NW Taiyanghe pitfall traps, 30.65N 109.5E Jaroslav Turna leg., ~ 1700m ��� (ZFMK, CP). Description. Length: 10.9 mm, length of elytra: 8.2 mm, width: 7.2 mm. Body oval, reddish brown, antenna yellow, dorsal surface dull, except a few small setae on head glabrous. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins convex and convergent to strongly rounded anterior angles, lateral margin and ocular canthus producing a indistinct blunt angle, margins strongly reflexed; anterior margin straight; surface convex medially, finely and densely punctate, with a transverse row of long, erect setae anteriorly; frontoclypeal suture feebly impressed and moderately angled medially; smooth area in front of eye approximately as wide as long; ocular canthus short and wide, sparsely finely punctate, with one terminal seta. Frons dull, with fine, dense punctures, with a few long setae beside eyes. Eyes small, ratio of diameter/interocular width: 0.45. Antenna with 10 antennomeres; club with three antennomeres, little longer than remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum subtrapezoidal, widest at base, lateral margins in basal half nearly straight and convergent, in anterior half convex and convergent anteriorly, anterior angles moderately produced and sharp, anterior margin weakly convex, marginal line complete; surface moderately finely and densely punctate, with minute setae in punctures; lateral margins sparsely setose, setae at anterior margin lacking medially; hypomeron carinate. Scutellum wide, triangular, with fine and moderately dense punctures each bearing a single minute seta. Elytra oval, widest at middle, striae distinctly impressed, finely and densely punctate, even intervals almost flat, with fine, moderately dense punctures, odd intervals with a few fine, erect, short setae, otherwise punctures with minute setae; epipleural margin robust, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a fine rim of very short microtrichomes. Ventral surface moderately shiny, metasternum with large and dense punctures, densely setose, metacoxa with minute setae in large punctures, with several long setae laterally. Abdominal sternites in addition to generally distributed fine and moderately dense punctures each with a distinct transverse row of coarse punctures each bearing a short and robust seta, otherwise punctures each with a microscopic seta; 3 rd abdominal sternite medially elevated. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.55. Pygidium weakly convex and moderately shiny, finely and moderately densely punctate, without smooth midline, punctures with microscopic setae, with a few long setae along apical margin. Legs moderately long and wide; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin medially straight, weakly widened in apical half and not serrate, dorsally not serrated, finely setose. Metatibia moderately wide and long, widest at apex, ratio width/length: 1/3.0, dorsally sharply carinate, with two groups of spines, basal one shortly before middle, apical one at three quarters of metatibial length, without contiguous serrated line basally; lateral face longitudinally convex, finely sparsely punctate; ventral margin finely serrate, with five equidistant spines; medial face impunctate; apex interiorly near tarsal articulation shallowly concave. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge and glabrous, beside with subventral longitudinal carina; first metatarsomere shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 56 A���C. Habitus: Fig. 56D. Diagnosis. Maladera taiyangheensis Ahrens, Fabrizi & Liu, sp. n. differs from M. tengchongensis Ahrens, Fabrizi & Liu, sp. n. in having the parameres symmetrical and the 3 rd abdominal sternite elevated medially. Etymology. The new species is named after its type locality, Taiyanghe (adjective in the nominative singular). Variation. Length: 10.9���13.9 mm, length of elytra: 8.2���10.0 mm, width: 7.2���8.2 mm. Female: eyes only little smaller than in male, ratio of diameter/interocular width: 0.42; antennal club with three antennomeres, as long as remaining antennomeres combined; pygidium weakly convex. Distribution. See map (Fig. 119) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 241-243, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera guangzhaishanica Ahrens, Fabrizi & Liu, sp. n. Figures 41 E���H, 108 Type material examined. Holotype: ♂ ��� China, SW Fujian, 28.iv.-24.VII. Guang Zhai Shan 25��42���N 116��47���E, 450-480m Jaroslav Turna leg., 2006/ 783 Sericini Asia spec.��� (ZFMK). Paratype: 1 ♂ ���[China] Baimianshan Yanling, F-Henugu, 1.X.2003, Leg. Li et al. ��� (ZFMK), 1 ♂ ���[China] Mt. Lushan, Jiangxi, 10.V.1977, leg. Zhang Youwei��� (IZAS). Description. Length: 10.2 mm, length of elytra: 7.9 mm, width: 6.4 mm. Body wide, oval, reddish brown, antenna yellowish brown, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous. Labroclypeus wide, lateral margins weakly convex and moderately convergent, lateral margins produce with ocular canthus an indistinct angle, anterior angles convex, anterior margin shallowly emarginate medially, margins weakly reflexed; surface weakly shiny, widely dull at base, surface convex medially, finely and densely punctate, distance between punctures smaller their diameter, with a few long erect setae; smooth area in front of eyes three times as wide as long; frontoclypeal suture finely incised, bluntly angled medially; ocular canthus moderately wide and short (1/3 of ocular diameter), finely and densely punctate, without a short terminal seta. Eyes small, ratio diameter/ interocular width: 0.58. Frons with fine and moderately dense punctures, with a few single setae beside eyes. Antenna with ten antennomeres, club with three antennomeres and straight, distinctly shorter than remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins weakly convex and weakly convergent anteriorly, anterior angles sharp and distinctly produced; anterior margin convex, with widely interrupted marginal line; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only, setae of lateral and anterior margin sparse; hypomeron carinate. Scutellum wide, triangular, punctation as in pronotum. Elytra (left elytron slightly deformed in holotype) widest at posterior third, striae finely impressed, finely punctate, intervals flat, finely and densely punctate, with minute setae in punctures, odd intervals with a few erect short setae; epipleura ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Mesosternum between mesocoxae wide, as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.71. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium strongly convex, coarsely and densely punctate, long setae on apical margin absent. Legs moderately long and wide; femora superficially and sparsely punctate, with two longitudinal rows of setae. Metafemur shiny, anterior margin acute, without adjacent serrated line, anterior longitudinal row of setae absent; posterior ventral margin almost straight, not widened in apical half, neither ventrally nor dorsally serrated but smooth, glabrous. Metatibia short and wide, flattened, ratio width/length: 1/2.9, sharply carinate dorsally, with two groups of spines, basal one at basal third, apical one at three quarters of metatibial length, basally with a few robust and short setae; lateral face weakly longitudinally convex, with a very superficial fine punctures; medial face impunctate, apex shallowly and concavely emarginate interiorly near tarsal articulation. Tarsomeres sparsely finely punctate dorsally, with dense, fine setae ventrally; metatarsomeres ventrally glabrous, with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere as long as following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate. Anterior claws symmetrical. Aedeagus: Fig. 41 E���G. Habitus: Fig. 41H. Female unknown. Diagnosis. Maladera guangzhaishanica Ahrens, Fabrizi & Liu, sp. n. is very similar to Maladera guanxiamensis Ahrens, Fabrizi & Liu, sp. n. in habitus and shape of the aedeagus. The former is distinguished in having the left paramere wider (lateral view) and the right paramere shorter and strongly pointed (both in lateral view). Etymology. The name of the new species is derived from its type locality, Guang Zhai Shan (adjective in the nominative singular). Variation. Length: 9.9���10.2 mm, length of elytra: 7.0��� 7.9 mm, width: 5.6���6.4 mm. Distribution. See map (Fig. 108) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 184-185, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera renardi (Ballion, 1870) Figures 1 E���H, 83 Serica renardi Ballion, 1870: 339; Brenske 1897: 371; Murayama 1938: 11, 1954: 54. Serica sibirica Brenske, 1897: 372 (syn by Nomura 1973: 128; Nikolajev 1982: 285); Maladera sibirica: Dalla Torre 1912: 18. Maladera renardi: Reitter 1896: 187; Nomura 1967: 52, 1973: 128; Kalinina 1989: 419; Kim & Lee 1997: 123; Kim & Kim 2003: 84; Ahrens 2007b: 21. Serica delicta Brenske, 1897: 372, syn. n. Type material examined. Lectotype (delicta, here designated): ♂ ���Wladivostok Koltze/ Serica delicta Type Brks./ type/ delicta Brsk. ��� (ZMHB). Lectotype (sibirica, here designated): ♂ ���Wladivostok Koltze/ sibirica Typ. Brsk. / Kein Forceps, unausgef��rbt/ Coll. Brenske ��� (ZMHB) (aedeagus in lectotype missing). Paralectotypes (sibirica, here designated): 1 ♀ ���Siberia or. Chabarofka Koltze F./ ♀ / sibirica typ. Brsk. / Coll. Brenske ��� (ZMHB), 1 ♀ ���Wladivostok/ sibirica Typ. Brsk. / coll. Brenske ��� (ZMHB). Additional material examined. Russia: 1 ♂ ��� Sibiria Stgr. / renardi v. sibirica m./ Coll. Brenske ��� (ZMHUB), 1 ♂ ��� Russia: primorskyi Kray, Ryazanovka 14km SW Slavyanka 42.48N 131.12E 16.VI.1993, 50m, leg. L. Zerche et al. ��� (DEIC), 1 ♂ ���SU-SE (Primor���e) ca. 50km SWW Vladivostok Bamburovo vill. env. 17.07.1990 Lg. A. Napolov ��� (CNA), 1 ♂ ���SU-SE (Primor���e) ca. 50km SWW Vladivostok Bamburovo vill. 28.7.1990 Lg. A. Napolov ��� (CNA), 1 ♂ ���SU-Far East (Primor���e) ca. 50km SWW Vladivostok Bamburovo vill. 6.8.1990 Lg. A. Napolov ��� (CNA), 1 ♂, 2 ♀♀ ���SU-Far East (Primor���e) ca. 50km SWW Vladivostok Bamburovo vill. env. 17.8.��� 6.9.1991 Lg. A. Napolov ��� (CNA), 1 ♂, 1 ♀ ���SU-Far East (Primor���e) ca. 50km SWW Vladivostok Bamburovo vill. env. 6.��� 10.9.1991 Lg. A. Napolov ��� (CNA), 3 ♂ ��� Vladivostok Sibir. Or. Ussuursk Dr. Jureček 1919��� (ZFMK), 3 ♂♂, 11 ♀♀ ��� Russia or. Primorskij reg. Black mts. Ananjevka A. Plutenko leg. 21.���30.5.94��� (ZFMK), 1 ♂ ��� Russia Prim. Partizansk distr. Alexeseyevsky distr. 20 km E Sergeyevka / Forests near Andreyevska river 400 m, 26.���29.VII.1995 P��tz & Wrase��� (ZFMK), 1 ♂ ��� Sutschan Ussuri ��� (ZFMK), 1 ♂, 2 ♀♀ ��� Russia: Far East Khabarovsk reg. Amur distr. Obluchje vill., 18.vi.2001 ��� (ZFMK), 2 ♂♂, 3 ♀♀ ��� Far East of Russia S. Primorje reg. Kondratenovka v. 12.VII.1998 ��� (ZFMK), 1 ♂, 1 ♀ ��� Rossia or. VI.1992 Cemyia gory Mts. Griaznaja riv. J. Farkac leg.��� (ZFMK). China: 12 ♂♂, 18 ♀♀ ��� China: Liaoning prov. Mt. Wulongshan, Dadaong city, June 11���15, 2001 local collector/ Via J. Jingke ��� (ZFMK), 2 ♂♂ ��� China: Inner Mongolia Mt. Daguangdingzishan (2061m) Chifeng 2.12. vii.2009 local coll. Ankauf via Li Jingke 2010��� (ZFMK), 2 ♂♂ ��� Qingyuan, Liaoning, 1, 7.IV.1954 ��� (IZAS). Japan: 1 ♀ ��� Tashiro Chikugo 1.ix.1952 Y. Miyake ��� (ZFMK). Redescription. Length: 8.1 mm, length of elytra: 6.5 mm, width: 4.9 mm. Body oblong, blackish brown, antenna yellowish, surface dull, labroclypeus and anterior frons shiny, except a few short setae on head and sides of elytra, glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins convex and convergent anteriorly, anterior angles moderately rounded; anterior margin weakly emarginate medially; margins moderately reflexed; lateral margin and ocular canthus produce an indistinct angle; surface flat, coarsely, densely punctate, with a few single erect setae; frontoclypeal suture finely incised, angled medially; smooth area anterior to eye convex, as wide as long; ocular canthus short and narrow (1/3 of ocular diameter), finely densely punctate, terminal seta present. Frons posteriorly dull, with moderately dense, irregular, fine punctures, with a few single setae beside eyes. Eyes moderately large, ratio diameter/ interocular width: 0.5. Antenna with ten antennomeres; club with three antennomeres and straight, 1.5 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum narrow, widest at base, lateral margins in basal half straight and weakly convergent, in anterior half moderately evenly convex, moderately convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles moderately rounded; anterior margin weakly convex, with fine marginal line being medially narrowly interrupted, base without marginal line; surface finely and densely punctate, with minute setae in punctures; anterior and lateral margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine and dense punctures. Elytra widest at posterior third, striae finely impressed, finely and densely punctate, intervals flat, with fine, moderately dense punctures, odd intervals with a few single short, erect setae, otherwise with minute setae in punctures; epipleural margin robust, ending at convex external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, finely and densely punctate, with dense and fine, adpressed setae, metasternal disc sparsely covered with longer setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.18. Pygidium moderately convex, dull, finely and densely punctate, without smooth midline, with short setae throughout and a few long setae along apical margin. Legs moderately long and wide; femora shiny, with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, with adjacent serrated line; posterior ventral margin smooth, not widened at apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia long and moderately wide, widest at middle, ratio of width/length: 1/3.9, sharply carinate dorsally, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, in basal half with a serrated line beside dorsal margin and few short single spines close to dorsal margin; lateral face longitudinally convex, finely sparsely punctate, glabrous; ventral margin finely serrate, with three robust setae of which the distal one is more distant; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres glabrous and with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere slightly longer than following tarsomere and slightly shorter than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 1 E���G. Habitus: Fig. 1H. Remarks. Since the male type specimen of Serica sibirica has no aedeagus, the status/ synonymy of the latter species is a little uncertain. Kim & Kim (2003) considered M. motschulskyi and M. spissigrada as synonyms of M. renardi, which is not correct, both are valid species. Distribution. See map (Fig. 83) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 15-16, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Ballion, E. (1870) Eine Centurie neuer Kafer aus der Fauna des russischen Reiches. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 43, 320 - 353.","Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310","Murayama, J. (1938) Revision des Sericines (Col., Scar.) de la Coree. Annotiones Zoologicae Japonenses, 17, 7 - 21.","Murayama, J. (1954) Icones of the Scarabeid-Beetles from Manchuria and Korea. I. Nihon Gakujutu, Shinko-kai, 163 pp.","Nomura, S. (1973) On the Sericini of Japan. I. Toho-Gakuho, 23, 119 - 152.","Nikolajev, G. V. (1982) Material on the taxonomy of Sericinae (Coleoptera, Scarabaeidae) from Mongolia and adjacent countries. Nasekomye Mongolii, 8, 284 - 289.","Dalla Torre, K. W. (1912) Scarabaeidae: Melolonthinae I. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, 45, pp. 1 - 84.","Reitter, E. (1896) Uebersicht der mir bekannten palaearktischen, mit der Coleopteren-Gattung Serica verwandten Gattungen und Arten. Wiener Entomologische Zeitung, 15, 180 - 188. https: // doi. org / 10.1002 / mmnd. 48018960133","Nomura, S. (1967) Some new and remarkable species of the Coleoptera from Japan and its adjacent regions, II. Entomological Review of Japan, 19, 52 - 62.","Kalinina, O. I. (1989) Podsem. Sericinae. In: Lehr, P. A. (Ed.), Opredelitel nasekomykh Dal'nego Vostoka SSSR. Tom 3. Zhestkokrylye ili zhuki. Part 1. Nauka, Leningrad, pp. 416 - 421.","Kim, J. I. & Lee, O. J. (1997) Taxonomic study of Korean Sericinae (Melolonthidae, Coleoptera) II, Genus Maladera. Insecta Koreana, 14, 119 - 135.","Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1"]}

Maladera aptera Ahrens, Fabrizi & Liu, sp. n. Figures 69 E–H, 126 Type material examined. Holotype: ♂ “ China-Yunnan Habashan -Habashan Mts. 7.- 10.6.2002, alt. 3500-4000m WGS84 27°19’N, 100°08’E lgt. Becvar & Fouque R.+H./ 584A” (ZFMK). Paratypes: 2 ♂♂, 1 ♀ “ China: Yunnan; 1995; Habashan mts., 2500-3800m 27.20N 100.11E; 3.-6.vi.; SE slope; S. Becvar leg.” (CP), 2 ♀♀ “China-Yunnan Habashan -Habashan Mts. 7.- 10.6.2002, alt. 3500-4000m WGS84 27°19’N, 100°08’E lgt. Becvar & Fouque R.+H.” (ZFMK), 1 ♀ “ China: Yunnan; 2,5-3,8km 27,20N; 100,11E Habashan mts., SE slope, 3.-6.vi.; SE slope; lgt. S. Becvar, 1995” (NMPC), 1 ♀ “[China] Yunnan; 3800-4600m 27.19N 100.08E Habashan mts. E slope, 15.7.92 David Kral leg.” (NMPC), 1 ♀ “[China] Yunnan; 2000-3000m 27°20’N 100°11’E Habashan mts., SE slope, 10.[-] 13.7.1992, D. Kral lgt.” (NMPC), 2 ♀♀ “[China] Yunnan 2000-3000m 27.20N 100.11E Habashan mts. SE slope 10-13/7. Vit Kuban leg. 92” (CP), 1 ♀ “ China: Yunnan prov., 1.3-2.0 km S of Haba, 17.-20.VI.2007, Haba Xueshan Mts., 2830-3000 m, 27°22.1‘N, 100°08.2‘E leg. J. Hajek & J. Ruzicka / individually collected on soil surface and on plants and shrubs, mixed forest (with dominant Pinus) near the brook (Ch32-35)/ XDA-1626” (ZFMK). Description. Length: 10.6 mm, length of elytra: 6.7 mm, width: 6.1 mm. Body egg-shaped, black, shiny, nearly glabrous except lateral setae of pronotum and elytra and a few setae on head. Labroclypeus transverse, trapezoidal, widest at base, lateral margins weakly convex and strongly convergent anteriorly, being nearly in straight line with ocular canthus, not incised laterally before labrum; anterior angles moderately convex, anterior margin weakly emarginate medially, margins weakly reflexed; surface weakly convex medially, densely and coarsely punctate, distance between punctures partly smaller their diameter, punctures rarely fused with each other, on anterior part intermixed with a few larger punctures bearing each a robust erect seta. Frontoclypeal suture finely incised, weakly bent medially; smooth area in front of eyes as wide as long; ocular canthus very short and wide (1/3 of ocular diameter), finely and densely punctate, with a short terminal seta. Frons finely and moderately densely punctate, with a few short setae beside eyes and behind frontoclypeal suture. Eyes very small, ratio diameter/ interocular width: 0.29. Antenna with ten antennomeres, club with three antennomeres, half as long as remaining antennomeres combined. Mentum convexly elevated, flattened anteriorly. Pronotum moderately wide, widest shortly before base, lateral margins evenly moderately curved and convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles strongly rounded; anterior margin nearly straight, fine marginal line widely lacking medially; surface with fine, sparse to moderately dense punctures, with minute setae in punctures, anterior and lateral margins with sparse and moderately long setae. Scutellum extremely short, triangular, nearly vanishing under pronotum, punctation not visible. Elytra convex, widest at middle, humeri small, external apical angles strongly rounded, apex not truncate; striae finely impressed, finely and densely punctate; intervals weakly convex, finely and sparsely punctate, only with minute setae in punctures; epipleural margin robust and convex, ending slightly before external apical angle; epipleura finely and sparsely setose; apical margin chitinous, without a rim of microtrichomes. Ventral face coarsely and densely punctate, moderately densely setose; mesosternum between mesocoxae as wide as mesofemur. Metasternum short, distance between meso-and metacoxae equal to width: of metafemur. Metacoxa glabrous, laterally with a few robust setae. Ratio of length of metepisternum/ metacoxa: 1/ 1.29. Abdominal sternites shiny, finely densely punctate, with a row of robust punctures each bearing a short seta. Pygidium apically weakly convex, coarsely and moderately densely punctate, punctures partly with a longitudinal impression, with a few robust setae at apex. Legs long and moderately wide, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with acute anterior margin, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, neither ventrally nor dorsally serrated but smooth, glabrous. Metatibia long and moderately wide, widest at apex, ratio width/ length: 1/ 3.18; dorsal margin longitudinally convex, with two groups of spines, basal one at first third, apical one at three quarters of metatibial length, with a few fine robust setae basally; lateral face longitudinally convex, with very sparse and fine punctures; ventral margin without serrate carina, with three strong nearly equidistant spines; medial face smooth and glabrous; apex interiorly near tarsal articulation shallowly and concavely emarginate. Tarsi in cross section circular, without lateral carina, impunctate dorsally, ventrally finely setose; metatarsomeres robustly setose ventrally, with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere distinctly shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia moderately long, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Wings are reduced in length. Aedeagus: Fig. 69 E–G. Habitus: Fig. 69H. Diagnosis. Maladera aptera Ahrens, Fabrizi & Liu, sp. n. differs from the related M. erlangensis Ahrens, Fabrizi & Liu, sp. n. in the shape of aedeagus, in having the body black shiny, as well as lacking setae on the frons. Etymology. The name of this new species (adjective in the nominative singular case) is derrived from the Latin adjective apterus (without wings), with reference on the lacking wings in this species. Variation. Length: 9.6–10.6 mm, length of elytra: 6.4–6.7 mm, width: 6.0– 6.1 mm. Female: Eyes smaller than in male, ratio diameter/ interocular width: 0.27; antennal club half as long as remaining antennomeres combined; pygidium weakly convex Distribution. See map (Fig. 126) and Table 1.

Maladera fengyangshanica Ahrens, Fabrizi & Liu, sp. n. Figures 63 A���D, 122 Type material examined. Holotype: ♂ ���[China] Mt. Fengyangshan, Longquan, Zhejiang, 25.VII-1.VIII.2007, leg. Liu Haoyu, Zou Zhenhua��� (HBUM). Description. Length: 8.9 mm, length of elytra: 6.9 mm, width: 4.9 mm. Body oval, dark brown, antenna brown, labroclypeus weakly shiny, remainder of dorsal surface dull, glabrous. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and strongly convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed, anterior margin weakly emarginate medially; surface weakly convex medially, finely and densely punctate, distance between punctures less their diameter, with few long, erect setae anteriorly; frontoclypeal suture weakly impressed and angled medially; smooth area in front of eye approximately three times as wide as long; ocular canthus short and wide, finely punctate, with a short terminal seta. Frons with fine, dense punctures, glabrous except for a few setae beside eyes. Eyes small, ratio of diameter/interocular width: 0.53. Antenna with 10 antennomeres; club with three antennomeres, slightly shorter than remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum subtrapezoidal, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles moderately produced and rounded, anterior margin weakly convex, marginal line widely interrupted medially; surface finely and densely punctate, glabrous; anterior and lateral margins sparsely setose; hypomeron carinate. Scutellum wide, triangular, with fine and dense punctures, each bearing a single minute seta. Elytra oblong, widest shortly behind middle, striae weakly impressed, finely and densely punctate, intervals weakly elevated, with fine, moderately dense punctures concentrated along striae, odd intervals with a few fine, erect, short setae, otherwise punctures with a very minute seta; epipleural margin robust, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a rim of very short microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, sparsely setose; metacoxa glabrous except for a several long setae laterally. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, punctures with microscopic setae, penultimate sternite apically with a shiny smooth short chitinous border. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.39. Pygidium small, weakly convex, finely densely punctate, with a few short setae beside apical margin. Legs moderately long and wide; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin medially straight, weakly widened in apical half and entirely strongly serrate, dorsally also serrate, finely setose. Metatibia moderately wide and long, widest at apex, ratio width/length: 1/3.75, dorsally sharply carinate, with two groups of spines, basal one at middle, apical one at four fifths of metatibial length, in basal half with a blunt subdorsal margin, beside it with a few punctures, and short spines; lateral face longitudinally convex, with moderately dense and fine punctures, punctures with minute setae; ventral margin serrate, with two widely distant spines; medial face finely sparsely punctate, apex interiorly near tarsal articulation shallowly concave. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally glabrous, with a strongly serrated ridge, and a subventral longitudinal carina beside it; first metatarsomere shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 63 A���C. Habitus: Fig. 63D. Female unknown. Diagnosis. Maladera fengyangshanica Ahrens, Fabrizi & Liu, sp. n. differs from the very similar M. excisilabrata Ahrens, Fabrizi & Liu, sp. n. in the longer lateral apophysis of the phallobase and the parameres being longer and curved ventrally. Etymology. The new species is named after its type locality, Fengyangshan Mts (adjective in the nominative singular). Distribution. See map (Fig. 122) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 268-270, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera fuscipes (Moser, 1915) Figures 27 A���D, 100 Autoserica fuscipes Moser, 1915: 353. Maladera fuscipes Ahrens 2006b: 237, 2007b: 23. Type material examined. Lectotype (here designated): ♂ ��� Tchao Pin-io Chine / Autoserica fuscipes Mos. Type ♂ ��� (ZMHB). Paralectotypes (here designated): 1 ♂, 2 ♀♀ ��� Tchao Pin-io Chine ��� (ZMHB), 1 ♀ ��� Kiukiang China / Autoserica fuscipes Mos. Type ♂ ��� (ZMHB), 4 ♂♂, 7 ♀♀ ��� Kiukiang China ��� (ZMHB), 1 ♂ ��� Kiukiang China ��� (CF). Additional material examined. China: 4 ex. ��� Museum Paris Kouy-Tch��ou P. Cavalerie 1910��� (MNHN), 2 ♂♂ ��� Bazhai, Maguan, Yunnan, 19.VII.2006, leg. Mao Benyong etc.��� (HBUM), 1 ♂ ��� Mt. Daqingshan, Longzhou, Guangxi, 19,20. IV. 1963, 360m, leg. Wang Shuyong, Wang Chunguang ��� (IZAS), 1 ♂ ��� Changpotou, Jinping, Yunnan, 23.V.1956, 1200m, leg. Huang Keren etc.��� (IZAS), 1 ♂ ���Jingdong, Yunnan, 31.V.1956, 1170m, leg. Polov ��� (IZAS), 2 ♂♂ ���Damenglong, Xishuangbanna, Yunnan, 21.IV. 1958, 650m, leg. Pu Fuji ��� (IZAS), 1 ♂ ��� Mt. Dushan, Guizhou, 7.VI.1981, 1000m, leg. Li Fasheng ��� (CAU), 1 ♂ ���Huangguoshu, Guizhou, 8.VII.1983, leg. Zou ��� (NKU), 1 ♂ ��� China, Fujian Province, Mt. Liang-shan-ding, Wuping Country, 2-13.VII.2009 local collectors, Ankauf via Li Yingke 2010 ex coll. Ahrens ��� (ZFMK). Vietnam: 1 ex. ��� Tonkin Hoa-Bin leg. A. de Cooman, VII.40��� (IZAS), 1 ex. ��� Tonkin R��gion de Hoa-Binh / Museum Paris 1934 A. de Cooman ��� (MNHN), 1 ♂ ��� Hoa-Binh, Tonkin, VII.1940, leg. A. de Cooman ��� (IZAS), 2 ♂♂ ��� Vietnam, Cuc Phong, 100km SE Hanoi V-1993; leg. Michio Hori ��� (ZFMK), 2 ex. ��� Vietnam N (Sa Pa) Lao Cai Prov., 250 km from Hanoi bearing 31��, Sa Pa vill. env., Hoang Lien Son Nat. Res. 9.- 15.5.1998 1250 m leg. A. Napolov ��� (CNA), 1 ex. ��� Vietnam N (Sa Pa) Lao Cai Prov., 250 km from Hanoi bearing 31 ��, Sa Pa vill. env., Hoang Lien Son Nat. Res. 1.- 6.7.1998 1250 m leg. A. Napolov ��� (CNA). Laos: 18 ex. ���NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20��12���N, 104��01���E; 14.iv.-15.v.2012; 1300-1900m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012/13��� (ZFMK), 4 ex. ���Laos-NE Hua Phan prov., 20��12���N, 104��01���E, Phu Phan Mt., 1500-1900m, 17.5.- 3.6.2007, leg. Vit Kuban ��� (ZFMK), 58 ex. ���NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20��12���N, 104��01���E; 11.iv.-15.v.2012; 1300-1900m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012��� (ZFMK), 2 ex. ���Laos (west): Oudomxao province, Beng, 20.20��N 101.80��E, May 1-14.2010 local collectors of Li Jingke��� (ZFMK), 1 ex. ���Laos-NE, Houa Phan prov., 20��13���09-19������N 103��59���54������- 104��00���03������E, 1480-1510m Phou Pane Mt., 22.IV.-14.V.2008 Vit Kuban leg.��� (NMPC, ZFMK), 1 ♂ ���X-DA4844 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera spLaos_Phupane25��� (ZFMK), 1 ♂ ���X-DA5152 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.- 30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera spLaos_Phupane25��� (ZFMK), 1 ♂ ���X-DA018 Laos Hua Phan prov.; Ban Saleui, Phou Pan (Mt.) 20��12���N, 104��01 ��� 15.-30.iv.2014 leg. C. Holzschuh���ZFMK Ankauf 2014 Maladera spLaos_ Phupane 25��� (ZFMK). Redescription. Length: 5.4 mm, length of elytra: 3.6 mm, width: 3.6 mm. Body oval, black to dark reddish brown, club of antenna yellowish, parts of pronotum and head with greenish shine, dorsal surface dull, except some single setae on head dorsal surface nearly glabrous. Labroclypeus wide and trapezoidal, shiny, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles strongly rounded, anterior margin distinctly emarginate, margins strongly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex, finely and densely punctate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely incised, medially angled; smooth area anterior to eye weakly convex, as wide as long; ocular canthus short and wide (one third of ocular diameter), finely and sparsely punctate, with a terminal seta. Frons with fine, sparse punctures and a few single erect setae beside eyes. Eyes very small, ratio diameter/ interocular width: 0.36. Antenna with nine or ten antennomeres; club with three antennomeres and straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest at base, lateral margins evenly convex, distinctly convergent anteriorly, anterior angles moderately produced and rectangular, posterior angles blunt, weakly rounded at tip; anterior margin convex, with fine and complete marginal line, base without marginal line; surface finely and densely punctate, glabrous; lateral and anterior margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, sparse punctures, smooth at middle. Elytra widest at middle, striae distinctly impressed, finely and sparsely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with only a few short, setae on odd intervals; epipleural margin robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra not membraneous, without visible rim of microtrichomes (at ca 100x magnification). Ventral surface dull, coarsely and moderately densely punctate, metasternum sparsely covered with sparse fine setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and irregularly densely punctate, glabrous, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.62. Pygidium moderately convex and dull, coarsely and densely punctate, without smooth midline, glabrous except a few robust setae along apical margin. Legs wide and short, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, with anterior margin acute, without serrated line behind anterior margin, setae of anterior row very short, posterior margin smooth and widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia short, widest at apex, ratio of width/length: 1/2.6, sharply carinate dorsally, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, with a few short robust setae basally; lateral face longitudinally convex, sparsely punctate, glabrous; ventral margin finely serrate, with four robust equidistant setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, sparsely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 27 A���C. Habitus: Fig. 27D. Distribution. See map (Fig. 100) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 128-130, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Ahrens, D. (2006 b) Sericinae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248.","Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1"]}

Maladera bubengensis Ahrens, Fabrizi & Liu, sp. n. Figures 57 A���D, 118 Type material examined. Holotype: ♂ ���[China] Bubeng, Xishuangbanna, Yunnan, 14.IX. 1993, 700m, leg. Yang Longlong ��� (IZAS). Description. Length: 10.4 mm, length of elytra: 7.4 mm, width: 5.1 mm. Body oval, dark brown, antenna yellow, labroclypeus shiny, remainder of dorsal surface dull with some iridescent greenish sine, glabrous except a few short setae on head. Labroclypeus subtrapezoidal, widest at base, lateral margins straight and convergent to strongly rounded anterior angles, lateral margin and ocular canthus producing a distinct blunt angle, margins moderately reflexed; anteriorly margin straight; surface moderately convex medially, shiny, finely densely punctate, with a few single setae; frontoclypeal suture not impressed and weakly angled medially; smooth area in front of eye approximately as wide as long; ocular canthus short and wide, impunctate, without terminal seta. Frons dull with iridescent shine, with fine, dense punctures, glabrous except for a few setae beside eyes. Eyes small, ratio of diameter/interocular width: 0.56. Antenna with 10 antennomeres; club with three antennomeres, slightly longer than remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum subtrapezoidal, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles produced and sharp; anterior margin convex, marginal line complete; surface finely and densely punctate, with very minute setae in the punctures only; anterior and lateral margins sparsely setose; hypomeron carinate. Scutellum wide, triangular, with fine and dense punctures each bearing a minute seta. Elytra oblong, widest at middle, striae weakly impressed, finely and densely punctate, intervals almost flat, with fine, dense punctures, with a few fine, erect, short setae, punctures with minute setae; epipleural margin robust, ending before strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a fine rim of short microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, sparsely setose, metacoxa glabrous except for a several long setae laterally. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, punctures with microscopic setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.81. Pygidium weakly convex, finely and densely punctate, without smooth midline, punctures with microscopic setae, long setae along apical margin lacking. Legs short and wide; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior margin acute, lacking an adjacent serrated line, anterior longitudinal row of setae absent, posterior ventral margin strongly widened in apical half and not serrate, glabrous; posterior dorsal margin smooth, with a few fine setae. Metatibia moderately wide and long, widest at middle, ratio width/length: 1/2.2, dorsal margin sharply carinate, with two groups of spines, basal one at middle, apical one at four fifths of metatibial length, without a contiguous serrated line basally; lateral face longitudinally convex, without punctures; ventral margin serrate, with four equidistant spines; medial face impunctate, apex interiorly near tarsal articulation shallowly concave. Tarsomeres impunctate dorsally, with a few short setae ventrally; metatarsomeres ventrally with a strongly serrated ridge, and with a further subventral longitudinal carina; first metatarsomere slightly longer than dorsal tibial spur, subsequent metatarsomeres of holotype missing. Aedeagus: Fig. 57 A���C. Habitus: Fig. 57D. Female unknown. Diagnosis. Maladera bubengensis Ahrens, Fabrizi & Liu, sp. n. differs from all other larger Chinese Maladera species in having symmetrical parameres that have asymmetrically distributed small teeth along the internal margins of both parameres. Etymology. The new species is named after the type locality, Bubeng (adjective in the nominative singular). Distribution. See map (Fig. 118) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 244-246, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera nitens (Moser, 1915) comb. n. Figures 36 A–D, 106 Serica nitens Moser, 1915c: 117; Ahrens 2006b: 245; 2007b: 36. Type material examined. Syntypes: 1 ♂ “Prov. Fo-Kein (China)/ Serica nitens Type ♂ Moser ” (ZMHB), 1 ♀ “Prov. Fo-Kein (China)./ Serica nitens Type ♀ Moser ” (ZMHB). Additional material examined. 1 ♂ “ Chine 8.IV.46 Kuatun, Fukien leg. Tschung-Sen ” (MHNG), 1 ♂ “ Fukien S. China Shaowu Tachulan 1000m T. Maa ” (DEIC), 1 ex. “ Kuatun, Fukien China 25.5.46 (Tschung Sen.)/ ex. Coll. V. Balthasar Natinal Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun 2300 m 27, 40 n. Br. 117,40 ö. L. J. Klapperich 12.5.1938 (Fukien)/ ex. Coll. V. Balthasar Natinal Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun 2300 m 27, 40 n. Br. 117,40 ö. L. J. Klapperich 15.4.1938 (Fukien)/ ex. Coll. V. Balthasar Natinal Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun 2300 m 27, 40 n. Br. 117,40 ö. L. J. Klapperich 30.4.1938 (Fukien)/ ex. Coll. V. Balthasar Natinal Museum Prague, Czech Republic ” (NMPC), 1 ex. (♀) “ Kuatun 2300 m 27, 40 n. Br. 117,40 ö. L. J. Klapperich 7.4.1938 (Fukien)/ ex. Coll. V. Balthasar Natinal Museum Prague, Czech Republic ” (NMPC), 1 ex. (♂) “ China, NE Jiangxi, 18.IV. Sanqingshan, 450-600m, 28°52‘N, 118°04‘E Jaroslav Turna leg., 2006” (ZFMK), 37 ex. “ China, E Fujian, 1.-2. V. Shiniushan, 1350m 25°38‘N 118°28‘E Jaroslav Turna leg., 2008” (ZFMK), 1 ♂ “ China: Fujian; Tongguzhang 1600m, Jianling vii-2004 / Phil 11-2004-2” (ZFMK), 1 ♂ “ Guadun, Mt. Wuyishan, Fujian, 22-23.V.2004, leg. Yuan Caixia, Lijing ” (HBUM), 2 ♂♂ “ China: Zhejiang Prov. Qinqliangfeng Qianqingfang 30°18’N, 119°07’E alt. 1000-1100m 16-V-2012 Chen, Ma & Zhao leg.” (SNUC). Redescription. Length: 8.4 mm, length of elytra: 6.4 mm, width: 5.2 mm. Body oblong-oval, reddish brown, antenna yellow, shiny, dorsal face nearly glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins strongly convex and convergent anteriorly, anterior angles strongly rounded, anterior margin distinctly emarginate, margins moderately reflexed; lateral margin and ocular canthus produce a blunt angle; surface flat, coarsely, very densely punctate, with a few long setae anteriorly; frontoclypeal suture distinctly incised, weakly curved; smooth area anterior to eye convex, 2.5 times as wide as long; ocular canthus moderately long and wide (1/3 of ocular diameter), finely densely punctate, with a terminal seta. Frons shiny, with irregularly dense and coarse punctures, partly completely impunctate, with a few single setae beside eyes. Eyes large, ratio diameter/ interocular width: 0.85. Antenna with nine antennomeres; club with three antennomeres and strongly reflexed, 1.8 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, lateral margins in basal half straight and weakly convergent anteriorly, in anterior half convex and more convergent, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin weakly convex, with fine marginal line, base without marginal line; surface densely and coarsely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; anterior and lateral margin densely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures. Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, sparse punctures concentrated along striae, with minute setae in punctures, penultimate lateral interval with a few single setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface shiny, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.43. Pygidium moderately convex, shiny, coarsely and densely punctate, without smooth midline, with a few long but fine setae along apical margin. Legs long and moderately wide, shiny; femora with two longitudinal rows of setae, coarsely and densely punctate in basal half, otherwise punctures sparse. Anterior margin of metafemur acute, with adjacent continuous serrated line, anterior row of setae complete and duplicated; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia moderately long and moderately wide, widest at apex, ratio of width/length: 1/2.8, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short and robust single setae and a continuous serrated line beside basal half of dorsal margin; lateral face longitudinally convex, except a few punctures on base smooth and glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, distinctly concavely emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere distinctly longer than following tarsomere and longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 36 A–C. Habitus: Fig. 36D. Distribution. See map (Fig. 106) and Table 1.

Maladera brunnescens (Frey, 1972) comb. n. Figures 51 A–D, 116 Serica brunnescens Frey, 1972a: 166; Ahrens 2006b: 245; 2007b: 36. Type material examined. Holotype: ♂ “Kuatun (2300 m) 27,40 n.Br. 117,40 ö. L. J. Klapperich 11.6. 1938 (Fukien)/ Type Serica brunnescens G. Frey 1972 ” (ZMFK). Paratypes: 1 ♀ “Kuatun (2300 m) 27,40 n.Br. 117,40 ö.L. J. Klapperich 17.6. 1938 (Fukien)/ Type Serica brunnescens G. Frey 1972 ” (ZMFK), 1 ♀ “Kuatun (2300 m) 27,40 n.Br. 117,40 ö. L. J. Klapperich 8.6. 1938 (Fukien)/ Type Serica brunnescens G. Frey 1972 ” (ZMFK). Additional material examined. 1 ex. “ Kuatun (2300 m) 27, 40 n.Br. 117,40 ö. L. J. Klapperich 11.6. 1938 (Fukien)/ ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun (2300 m) 27, 40 n.Br. 117,40 ö. L. J. Klapperich 8.6. 1938 (Fukien)/ ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 43 ex. “ China, W Fujian, 3.-4.VI. Emei Feng, 1200-1500m 27°01‘N 117°04E Jaroslav Turna leg., 2008” (ZFMK), 10 ex. “ China, W Fujian, 1.-2.VI. Emei Feng, 1200-1500m 27°01‘N 117°04E Jaroslav Turna leg., 2007” (ZFMK). Redescription. Length: 10.1 mm, width: 6.0 mm, length of elytra: 7.0 mm. Body oblong, reddish brown, antenna yellow, dorsal surface shiny, except head almost glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins slightly curved and strongly convergent to feebly rounded anterior angles, lateral border and ocular canthus produce a distinctly blunt angle, only anterior margin weakly reflexed, anteriorly indistinctly but widely emarginate medially; surface distinctly convex medially and shiny, finely and moderately densely punctate, with numerous coarse punctures bearing fine and erect setae; frontoclypeal suture very indistinctly incised and medially weakly angled; smooth area in front of eye very small, as wide as first antennal joint and approximately 1.5times as wide as long; ocular canthus moderately long and moderately slender, densely finely punctate, with one terminal seta. Frons with fine, sparsely scattered punctures, with a few setae bent backwards. Eyes moderately small, ratio of diameter/ interocular width: 0.57. Antenna yellow, 10-segmented; club with three segments, in male as long as remaining segments together. Mentum elevated, anteriorly flattened and glabrous. Pronotum narrow, widest at base, lateral margins weakly curved and strongly narrowed anteriorly, behind the middle feebly convexly emarginate, anterior angles strongly produced and sharp, anterior margin without marginal line and weakly produced medially, posterior angles sharp; surface moderately finely and densely punctate, punctures denser basally, glabrous; anterior and lateral border setaceous. Scutellum wide, triangular, with fine and very dense punctures, glabrous. Elytra moderately oblong, widest behind the middle, striae strongly impressed and finely and densely punctate, intervals distinctly convex, with fine and sparsely scattered punctures, mostly concentrated along striae, a few punctures with sparse fine, short, erect setae; epipleural margin robust, ending at strongly curved external apical angle of elytra, epipleura sparsely setaceous, apical border chitinous, covered with very short microtrichomes. Ventral surface shiny, coarsely and densely punctate, almost glabrous, metacoxal plates laterally with a few long setae; each abdominal sternite with a transversal row of coarse punctures bearing short and fine setae between fine and dense punctation, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as sternite. Mesosternum between mesocoxae a little less wide than mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.41. Pygidium moderately convex, finely and densely punctate, without smooth midline, surface apically dull and with numerous fine and long setae. Legs moderately slender; femora shiny, with two longitudinal rows of setae, between the rows only sparsely punctate; metafemur anteriorly sharply carinate, behind anterior margin without serrated line, posterior margin in apical half dorsally finely serrated, ventrally completely serrated, in basal half with a few fine setae. Metatibia moderately slender and moderately long, widest at apex, ratio width/ length: 1/ 3.33, dorsally moderately carinate, with two groups of spines, basal one at one third, apical one at three quarters of metatibial length, basally with two single setae with punctures having a serrated margin; lateral face longitudinally convex, with moderately dense, coarse and longitudinally impressed punctures and glabrous; ventral margin with three strong spines, of which the apical two are very distant, inside sparsely scattered, finely punctate, apex interiorly near tarsal articulation shallowly concavely emarginate. Tarsomeres distinctly and densely punctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a moderate longitudinal margin, segments one to four ventrally glabrous, first metatarsomere distinctly shorter than the two following segments combined and about one third longer than dorsal tibial spur. Protibia in holotype are lacking. Aedeagus: Fig. 51 A–C. Habitus: Fig. 51D. Variation. Antennal club in female a little shorter than remaining antennal joints together. Distribution. See map (Fig. 116) and Table 1.

Maladera jaintiaensis Ahrens & Fabrizi, 2016 Figures 32 I–M, 103 Maladera jaintiaensis Ahrens & Fabrizi, 2016: 224. Additional material examined. China: 1 ♂ “ Pingshiang Süd-China Dr. Kreyenberg ” (DEIC), 1 ♂ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 9.4.1938 (Fukien) / Paratype Autoserica fusca G. Frey 1972 “ (ZFMK), 1 ♂ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 29.4.1938 (Fukien)” (CF). Laos: 1 ♂ “ Laos: Bolikhamsai pr. 18°16’N 103°11’E, 70 km NEE Vientiane, 27.-30.iv. 1997, 150m, Vit Kuban leg./ 266 Sericini Asia spec.” (CP). Remarks. The species was originally described from northeastern India and China (Ahrens & Fabrizi 2016). Distribution. See map (Fig. 103) and Table 1.

Maladera jizuana Ahrens, Fabrizi & Liu, sp. n. Figures 44 E���H, 111 Type material examined. Holotype: ♂ ���[China] Yunnan, 2300m, Jizu Mts. 18.-20. Jul. 1995 Bolm lgt./ 837 Sericini Asia spec.��� (CP). Paratypes: 1 ♂ ���N Vietnam (Tonkin) pr. Vinh Phu 1990 Tam Dao 17.-21.v. P. Pacholatko leg./ 346 Sericini Asia spec.��� (CP), 1 ♂ ���[China] Yunnan, Lincang, Mt. Wulaoshan, 2010-VII-31, N: 23.90648, E: 100.15944, 1807m / LW-1302��� (IZAS), 1 ♂ ���[China] Mts. Linaoshan, Langping, Tianlin, Guangxi, 28.V.2002, 1400m, leg. Jiang Guofang/ LW-688��� (IZAS). Description. Length: 7.5 mm, length of elytra: 5.8 mm, width: 4.6 mm. Body oblong-oval, dark reddish brown, dull, antenna and labroclypeus yellow, labroclypeus shiny, except some single setae on head dorsal surface nearly glabrous. Labroclypeus narrow and subtrapezoidal, widest at base, lateral margins weakly convex and slightly convergent, anterior angles moderately rounded, anterior margin weakly emarginate medially, margins distinctly reflexed; lateral margin and ocular canthus produce a distinct angle; surface flat, finely and densely punctate, with a few larger punctures each bearing an erect seta; frontoclypeal suture indistinctly incised, evenly curved; smooth area anterior to eye weakly convex, twice as wide as long; ocular canthus moderately long and narrow (1/4 of ocular diameter), finely and densely punctate, with a terminal seta. Frons dull, behind frontoclypeal suture narrowly shiny, with sparse, fine punctures, with a few long setae beside eyes. Eyes very large, ratio diameter/ interocular width: 0.84. Antenna missing in holotype. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest shortly before base, lateral margins evenly convex and strongly convergent anteriorly, anterior angles moderately produced and nearly right-angled, posterior angles strongly rounded; anterior margin convex, with complete robust marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures; lateral anterior and lateral margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures. Elytra widest shortly behind middle, striae distinctly impressed, finely and sparsely punctate, odd intervals slightly convex, even intervals flat, with fine and moderately dense punctures being concentrated along striae on odd intervals, with minute setae in punctures, odd intervals with a very few short setae; epipleural margin robust, ending at strongly curved external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, coarsely and densely punctate, glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Pygidium weakly convex and dull, coarsely and densely punctate, without impunctate midline, glabrous except a few robust setae along apical margin. Legs moderately long and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without adjacent serrated line, anterior row of setae reduced to a few single setae; posterior margin smooth, weakly widened at apex and smooth ventrally, not serrate dorsally, finely shortly setose. Metatibia moderately long and wide, widest at middle, ratio of width/length: 1/2.9, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, with a few robust setae basally subparallel to dorsal margin; lateral face longitudinally convex, shiny, sparsely punctate and glabrous; ventral margin finely serrate, with three long and robust setae of which distal one is more distant; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally sparsely and finely punctate, punctures with short setae, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, laterally and dorsally carinate; first metatarsomere slightly shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 44 E���G. Habitus: Fig. 44H. Female unknown. Diagnosis. In external morphology and the shape of the genitalia, Maladera jizuana Ahrens, Fabrizi & Liu, sp. n. is similar to M. sedula Ahrens & Fabrizi, 2016, from India. The new species differs from M. sedula in having the apical phallobase with a fine longitudinal carina in front of the dorsal median emargination that projects into the emargination; additionally, the parameres of the new species are narrower (lateral view). Etymology. The name (adjective in the nominative singular) of the new species is derived from the occurrence of the species in the Jizu mountains. Variation. Length: 7.5���8.5 mm, length of elytra: 5.8���6.1 mm, width: 4.4���4.6 mm. Antenna with ten antennomeres; club with three antennomeres and strongly reflexed, 1.8 times as long as remaining antennomeres combined. Distribution. See map (Fig. 111) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 195-196, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Ahrens, D. & Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355."]}

Maladera euphorbiae (Burmeister, 1855) Figures 3 J���M, 85 Serica euphorbiae Burmeister, 1855: 177; Brenske 1897: 362. Amaladera euphorbiae: Reitter 1896: 186; Dalla Torre 1912: 19; Medvedev 1952: 145. Maladera (Amaladera) euphorbiae: Ahrens 2006b: 234; 2007b: 18. Serica fusca Ballion, 1870: 339; syn by Reitter 1896: 186; Brenske 1897: 362. Additional material examined. China: 1 ♂ ��� Ining, Xinjiang, VI.1984 ��� (IZAS), 2 ♂♂ ��� Yining, Xinjiang, mid of VI.1983, leg. Wang Aijing ��� (IZAS). Redescription. Length: 7.5 mm, length of elytra: 5.2 mm, width: 4.1 mm. Body wide, oval, yellowish brown, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous. Labroclypeus moderately wide, lateral margins straight and moderately convergent, lateral margins produce with ocular canthus a blunt angle, anterior angles moderately convex, anterior margin distinctly emarginate medially, margins weakly reflexed; surface flat and moderately shiny, coarsely and densely punctate, distance between punctures smaller their diameter, with numerous long erect setae; smooth area in front of eyes twice as wide as long; frontoclypeal suture finely incised, slightly elevated and bluntly angled medially; ocular canthus moderately wide and short (1/3 of ocular diameter), finely and densely punctate, with a short terminal seta. Eyes moderately large, ratio diameter/ interocular width: 0.65. Frons with fine and moderately dense punctures, with a few single setae beside eyes. Antenna with nine antennomeres, club with three antennomeres and straight, 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins in basal half straight and weakly convergent anteriorly, in anterior half moderately convex and convergent anteriorly, anterior angles nearly blunt and weakly produced, posterior angles blunt; anterior margin convex, with complete and fine marginal line; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only, setae of lateral and anterior margin sparse; hypomeron carinate. Scutellum wide, triangular, punctation as in pronotum. Elytra convex, widest at posterior third, striae finely impressed, finely punctate, intervals weakly convex, finely and densely punctate, punctures concentrated along striae, with minute setae in punctures, odd intervals with a few erect short setae; epipleura ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Distance between mesocoxae little less than width of mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.3. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta; penultimate sternite medially retracted, ultimate sternite twice as long as penultimate. Pygidium moderately convex, coarsely and densely punctate, on apex with a few long setae. Legs moderately long and wide; femora superficially and sparsely punctate, with two longitudinal rows of setae. Metafemur shiny, anterior margin acute, without adjacent serrated line, anterior longitudinal row of setae present; posterior ventral margin almost straight, little widened in apical half and smooth, and dorsal posterior margin also smooth, densely setose. Metatibia moderately long and wide, dorsal and ventral margin subparallel, ratio width/length: 1/3.6, sharply carinate dorsally, with two groups of spines, basal one at basal third, apical one at two thirds of metatibial length, basally with a few single fine and short setae; lateral face weakly longitudinally convex, with fine and sparse punctures; ventral margin with three widely separated spines, of which the distal one is more distant; medial face impunctate, apex shallowly and concavely emarginate interiorly near tarsal articulation. Tarsomeres dorsally very sparsely and finely punctate, neither laterally nor dorsally carinate, ventrally finely sparsely setose; metatarsomeres ventrally with a strongly serrated ridge, first metatarsomere slightly longer than following tarsomere and as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 3 J���L. Habitus: Fig. 3M. Distribution. See map (Fig. 85) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 26-27, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Burmeister, H. (1855) Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Fortsetzung. 2. Abteilung, Coleoptera Lamellicornia Phyllophaga chaenochela. T. C. F. Enslin, Berlin, x + 569 pp.","Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310","Reitter, E. (1896) Uebersicht der mir bekannten palaearktischen, mit der Coleopteren-Gattung Serica verwandten Gattungen und Arten. Wiener Entomologische Zeitung, 15, 180 - 188. https: // doi. org / 10.1002 / mmnd. 48018960133","Dalla Torre, K. W. (1912) Scarabaeidae: Melolonthinae I. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, 45, pp. 1 - 84.","Medvedev, S. I. (1952) Fauna USSR: Coleoptera 10 Nr. 1: Lamellicornia (Scarabaeidae), subfam. Melolonthinae. Part 2. Izdatelstvo Akademii Nauk SSSR, Moskva, Leningrad, 276 pp.","Ahrens, D. (2006 b) Sericinae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248.","Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1","Ballion, E. (1870) Eine Centurie neuer Kafer aus der Fauna des russischen Reiches. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 43, 320 - 353."]}

Maladera riberai Ahrens, Fabrizi & Liu, sp. n. Figures 13 A–D, 93 Type material examined. Holotype ♂ “ China—Shaanxi, Daba Shan, Shou Man vill., 32°14’N, 108°34’E, 25.v.- 14.vi.2000, 1000m, Siniaev & Plutenko leg./ 1072 Sericini Asia spec.” (CP). Description. Length: 9.6 mm, length of elytra: 6.5 mm, width: 6.0 mm. Body oval, dark brown, antenna yellow, labroclypeus shiny, remainder of dorsal surface dull, glabrous, except for a few small setae on the head and elytra. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and convergent to strongly rounded anterior angles, lateral margin and ocular canthus producing a distinct blunt angle, margins moderately reflexed; anterior margin very shallowly emarginate medially; surface convex medially, weakly shiny, coarsely punctate, with a few short, erect setae; frontoclypeal suture feebly impressed and weakly curved medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and moderately wide, finely punctate, with a short terminal seta. Frons with fine, moderately dense punctures, glabrous except for a few setae beside eyes and frontoclypeal suture as well as on disc. Eyes small, ratio diameter/ interocular width: 0.57. Antenna yellow, with 10 antennomeres; club with three antennomeres, little longer than remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum widest at base, lateral margins weakly and evenly convex and convergent anteriorly, anterior angles distinctly produced and sharp, anterior margin weakly convex, marginal line widely interrupted medially; surface moderately finely and densely punctate, with minute setae in punctures; anterior and lateral margins sparsely setose. Scutellum wide, triangular, with fine and moderately dense punctures, each bearing a single very minute seta. Elytra oblong, widest at middle, striae distinctly impressed, finely and densely punctate, odd intervals moderately convex, with a few erect setae, even intervals nearly flat, with fine, moderately dense punctures concentrated along striae, with minute setae in punctures; epipleural margin robust, ending at the weakly convex external apical angle of elytra, epipleura densely setose, apical border membraneous, with a very fine rime of short microtrichomes. Ventral surface dull, thorax and metacoxa with large dense punctures, sparsely setose, metacoxa glabrous except for a few long setae laterally. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, with a distinct transverse row of coarse punctures each bearing a short and robust seta, some punctures with microscopic setae, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as sternite. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.66. Pygidium moderately convex, finely and densely punctate, without smooth midline, punctures with minute setae and with a few long setae along apical margin. Legs moderately narrow and long; femora with two longitudinal rows of setae, finely and moderately densely punctate. Metafemur dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin medially straight, weakly widened in apical half and not serrate, posterior dorsal margin not serrated, finely setose. Metatibia moderately wide and long, widest at apex, ratio width/ length: 1/ 3.4, dorsally sharply carinate, with two groups of spines, basal one at middle, apical one at four fifths of metatibial length, basally with a few punctures, each bearing short single spines; lateral face shiny and longitudinally convex, with sparse and fine punctures, with minute setae in punctures; ventral margin serrate, with four strong equidistant spines; medial face impunctate, apex interiorly near tarsal articulation shallowly concave. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside it with smooth subventral longitudinal carina; first metatarsomere shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia short, bidentate. Protarsal claws asymmetrical, basal tooth of interior claw convexly rounded at tip, basal tooth of external claw with normally developed sharply truncate basal tooth. Aedeagus: Fig. 13 A–C. Habitus: Fig. 13D. Female unknown. Diagnosis. Maladera riberai Ahrens, Fabrizi & Liu, sp. n. is very similar to M. xingkei Ahrens, Fabrizi & Liu, sp. n. in shape of the aedeagus, differing in having the right paramere straight and the left paramere fused with the apex of the phallobase. Etymology. The new species is dedicated to Ignacio Ribera Galán (Barcelona), well known beetle specialist and friend, who passed away this year to much early (noun in the genitive singular case). Distribution. See map (Fig. 93) and Table 1.

Maladera yongrenensis Ahrens, Fabrizi & Liu, sp. n. Figures 45 A���D, 112 Type material examined. Holotype: ♂ ��� China: N-Yunnan, Baiyungshan (Rai Railing Mts.) 2400m, Yong Ren, VII-2003 leg. Ying et al. / 776 Sericini Asia spec.��� (ZFMK). Paratypes: 1 ♂ ��� China, W Yunnan prov., mts. 20km SW Baoshan, 2400m, 23.-25.v.2006, S. Murzin & I. Shokhin ��� (CP), 1 ♂ ���[China] Yunnan 2000-3000m 27.20N 100.11E Habanshan mts. SE slope 10-13/7. Vit Kuban leg. 92��� (CP), 2 ♂♂ ��� China Yunnan 2000-3000m 27��20���N 100��11���E Habashan Mts. SE slope, 10.[-] 13.7.1992. D. Kral lgt.��� (NMPC), 1 ♂ ���[China] Qujing, Yunnan, 10.VII.1982, leg. Song Shimei ��� (IZAS), 1 ♂ ���[China] Ninglang, Yunnan, 13-15.VII.2004, leg. Yuan Caixia, Lijing ��� (HBUM), 1 ♂ ��� China, Garze / Sichuan Yajiang env., W Yalong river, 30.01N, 100.98E 2800-3000m, 12.-21.VI.2016, leg. Reuter ��� (ZFMK). Description. Length: 7.6 mm, length of elytra: 5.1 mm, width: 3.7 mm. Body oblong-oval, dorsal face dark brown, dull, ventral face reddish brown, antenna yellow, labroclypeus moderately shiny, except some single setae on head dorsal surface nearly glabrous. Labroclypeus narrow and subtrapezoidal, widest at base, lateral margins weakly convex and narrowed anteriorly, anterior angles strongly rounded, anterior margin weakly emarginate medially, margins moderately reflexed; lateral margin and ocular canthus produce a distinct angle; surface flat, finely and densely punctate, with a few larger punctures each bearing an erect seta; frontoclypeal suture indistinctly incised, evenly curved; smooth area anterior to eye weakly convex, twice as wide as long; ocular canthus moderately long and narrow (1/4 of ocular diameter), finely and densely punctate, with a terminal seta. Frons dull, anterior third shiny, with moderately dense, fine punctures, behind frontoclypeal suture and beside eyes with a few long setae in large sparse punctures. Eyes large, ratio diameter/ interocular width: 0.81. Antenna with ten antennomeres; club with three antennomeres and straight, 1.3 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest in posterior third, lateral margins evenly convex and strongly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles strongly rounded; anterior margin convex, with complete robust marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures; lateral anterior and lateral margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures. Elytra widest shortly behind middle, striae finely impressed, finely and sparsely punctate, intervals slightly convex, with fine and dense punctures concentrated along striae, with minute setae in punctures, odd intervals with a very few single, short setae; epipleural margin robust, ending at strongly curved external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, coarsely and densely punctate, glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.59. Pygidium moderately convex and dull, coarsely and densely punctate, without impunctate midline, glabrous except a few robust setae along apical margin. Legs moderately long and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without adjacent serrated line, anterior row of setae reduced to a few single setae; posterior margin smooth, weakly widened at apex and smooth ventrally, not serrate dorsally, finely shortly setose. Metatibia moderately long and wide, widest at middle, ratio of width/length: 1/3.5, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, with a few robust setae basally subparallel to dorsal margin; lateral face longitudinally convex, shiny, sparsely punctate and glabrous; ventral margin finely serrate, with four equidistant, long and robust setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally sparsely and finely punctate, punctures with short setae, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, laterally and dorsally not carinate; first metatarsomere slightly shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 45 A���C. Habitus: Fig. 45D. Female unknown. Diagnosis. Maladera yongrenensis Ahrens, Fabrizi & Liu, sp. n. differs from the other species of the M. indica group that possess a dorsomedian phallobasal emargination covered by a wide and large dorsal process and the parameres without basal lobe in having the dorsomedian process of the phallobase shallowly incised medially. Etymology. The name (adjective in the nominative singular) of the new species is derived from the type locality, Yong Ren. Variation. Length: 7.1���7.6 mm, length of elytra: 4.9���5.1 mm, width: 3.7���4.1 mm. Distribution. See map (Fig. 112) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 198-199, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Autoserica straminea Moser, 1915c: 156. Maladera straminea: Ahrens 2006b: 238, 2007b: 24. Type material examined. Syntype: 1 ♂ ��� Kiukiang China / Autoserica straminea Type Mos./ straminea ��� (ZMHB) (head and prothorax missing in only available syntype). Additional material examined. China. 4 ex. ��� Yunnan 1800-2500 m 25.10N 100.21E Weishan mts. 22.-25.6.92 David Kral lgt.��� (CK), 2 ex. ���China: Yunnan; 1800-2500 m, 25.10N 100.21E Weishan Mt. D. Kr��l lgt.��� (ZFMK), 1 ex. ��� Yunnan 2000-2800 m 25.11N 100.24E Weibaoshan mts. W slope 25-28/6.92 Vit Kuban leg.��� (CN), 3 ex. (♀) ��� Yunnan 2000-2500 m 25.42N 100.08E Cangshan mts. E slope 21/6.92 Vit Kuban leg.��� (CN, CA), 1 ♂ ���Luoxiang, Jinxiu, Guangxi, 23.V. 1999, 200m, leg. Zhang Xuezhong��� (IZAS), 1 ♂ ���China, SE���Yunnan 1300-1600m, Maguan Wenshan karst region 23��11-15���/ 104��18-25���/ 18.-19.5.95 L.+R. Businsky lgt.��� (CP). Vietnam. 1 ♂ ���Vietnam N (Sa Pa) Lao Cai Prov., 250 km from Hanoi bearing 31�� Sa Pa vill. env. Hoang Lien Son Nat. Res., 9.- 15.6.1998 1250 m leg. A. Napolov��� (CNA), 1 ♂ ���Vietnam N 1990 Sa-Pa 11-19.VI. 2500 m Hoang Lien Son prov. Strnad Jan lgt.��� (CP), 1 ex. ���Vietnam-N (Ben En) 180 km SSW Hanoi, 40 km SW Than Hoa, Ben En Nat. Park 23.7.- 20.8.1997 50 m lg. A. Napolov��� (ZFMK), 1 ex. ���Vietnam N (Sa Pa) Lao Cai Prov., 250 km from Hanoi bearing 31�� Sa Pa vill. env. Hoang Lien Son Nat. Res., 21.- 23.6.1998 1250 m leg. A. Napolov��� (CNA), 3 ex. ���Vietnam N (Sa Pa) Lao Cai Prov., 250 km from Hanoi bearing 31�� Sa Pa vill. env. Hoang Lien Son Nat. Res., 16.- 20.6.1998 1250 m leg. A. Napolov��� (CNA), 1 ex. ���Vietnam N (Sa Pa) Lao Cai Prov., 250 km from Hanoi bearing 31�� Sa Pa vill. env. Hoang Lien Son Nat. Res., 27.V.-15.VI.1995 1250 m leg. A. Napolov��� (CNA), 2 ex. ���Vietnam-N (Na Hang) 160 km NW Ha Noi, NE env. of Na Hang 3.6.- 14.6.1996 150-200 m lg. A. Napolov & I. Roma��� (CNA), 6 ex. ���N-Vietnam, Bac Giang Prov., Tay Yen Tu Nat. Res., Thanh So���n, N 21�� 12.812��� E 106�� 45.846���, 86m 18.5.-21.5.5.2015, leg. A. Skale��� (CASH), 2 ex. ���N. Vietnam Bac Kan Prov., Ba B�� NP (entry), 16.-20.V.2014, 22��25���07������N, 105��38���09������E, 180-200m, leg. A. Skale ��� (CASH), 2 ex. ���N-Vietnam Cao Bang Prov., vic. Vin Den, Nui Pia Oac Nat. Res. 06.-10.V.2013, 22��33���53������N, 105��52���53������E 900-1300m A. Skale��� (CASH), 1 ex. (♀) ���N-Vietnam Cao Bang Prov., vic Tinh Tuc, Son Dong, Nui Pia Oac Nature Reserve, 9.-15.V.2014 22��37���55������N, 105��52���98������E 850-1300m leg. A. Skale��� (CASH), 1 ex. ���N-Vietnam, P: Thai Nguyen, vic. Ngoc Than, Me Linh (IEBR station), 13.V.2017, 21��23���3������N, 105��42���44������E, 65m NN, leg. A. Weigel, LFF��� (NME). Redescription. Length: 8.1 mm, length of elytra: 6.4 mm, width: 5.0 mm. Body oblong-oval, dark yellowish brown, dull, labroclypeus shiny, dorsal face nearly glabrous, only with a few erect long setae on pronotum, head and elytra. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles moderately rounded, anterior margin shallowly emarginate and distinctly reflexed medially, margins moderately reflexed; lateral margin and ocular canthus produce an indistinct angle; surface convexly elevated along middle, finely, densely punctate, with a few erect, long setae; frontoclypeal suture distinctly incised, bluntly angled medially; smooth area anterior to eye very small and flat, 3 times as wide as long; ocular canthus short and wide (1/3 of ocular diameter), finely densely punctate, with a terminal seta. Frons dull, with very sparse and superficial punctures, with a few single setae beside eyes and behind frontoclypeal suture. Eyes moderately large, ratio diameter/ interocular width: 0.58. Antenna with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at middle, lateral margins evenly convex and convergent towards angles, anterior angles moderately produced and blunt, posterior angles blunt, rounded at tip; anterior margin straight, with fine marginal line, base without marginal line; surface moderately finely and densely punctate, with minute setae in punctures and a very few long setae on lateral disc; anterior and lateral margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, sparse punctures. Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals flat, with fine, sparse punctures and with minute setae in punctures, odd intervals with a very few long single setae (often abraded); epipleural margin fine, ending behind middle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, finely and densely punctate, nearly glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/2.0. Pygidium moderately convex, dull, coarsely and moderately densely punctate, without smooth midline, with a few long setae beside apical margin. Legs short and wide, dull; femora with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, without adjacent serrated line, anterior row of setae completely reduced; posterior ventral margin smooth, moderately widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.3, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short and robust single setae; lateral face longitudinally convex, superficially and sparsely punctate, along midline widely smooth, glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 30 A���C. Habitus: Fig. 30D. Distribution. See map (Fig. 103) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 139-141, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Moser, J. (1915 c) Beitrag zur Kenntnis der Melolonthiden. VI. Deutsche Entomologische Zeitschrift, 1915, 113 - 151. https: // doi. org / 10.1002 / mmnd. 48019150204","Ahrens, D. (2006 b) Sericinae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248.","Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1","Nomura, S. (1974) On the Sericini of Taiwan. Toho-Gakuho, 24, 81 - 115.","Fairmaire, M. L. (1887) Coleopteres de l'Interieur de la Chine. Annales de la Societe Entomologique de Belgique, 31, 87 - 136."]}

Maladera pallida (Burmeister, 1855) comb.n. Figures 47 J–M, 112 Serica pallida Burmeister, 1855: 167; Ahrens 2006b: 245; 2007b: 36. Amaladera pallida: Medvedev 1952: 144. Maladera ludipennis Miyake, Yamaguchi & Aoki 2002: 52, syn. n. Type material examined: Lectotype (pallida, here designated): ♂ “703 5/9/51/ Hongkong ” (ZMHB). Paralectotype (pallida, here designated): 1 ♀ “703 9/5/51/ pallida Br. Hongk. Dhn. ” (MLUH). Holotype (ludipennis): @ “Ban Lao Kram Mukdahan (light) E-Thailand 23. Feb. 1999 S. Yamaguchi & T. Aoki leg./ Holotype Maladera ludipennis Y. Miyake, Yamaguchi et Akiyama 2002” (RIEBT). Additional material examined. China: 1 ♀ [Syntype?; without abdomen and proper labels] (MLUH), 1 ex. “China”, 1 ex. “ Hongkong ” (ZMHB), 1 ex. (♀) “Mus. Westerm./ Hongkong” (ZMUC), 1 ♂ “ China Tsonglok ” (HNHM), 3 ex. “ Hongkong / China / Coll. Reitter ” (HNHM). Vietnam: 8 ex. “ Tonkin Dap ban/ contracta Brsk. [Handschr. Moser]” (ZMHB). Laos: 1 @ “ Laos centr., 27.IV.-1.V.1997 70km NE Vientiane, Ban Phabat env., 150m, N 18°16.1’, E 103°10.9’ E. Jendek & O. Šauša leg./ Coll. P. Pacholatko / 360 Sericini: Asia spec.” (CP). Doubtful record: 8 ex. “ Bengalen / Coll. C. Felsche Kauf 20, 1918” (SMTD). Redescription. Length: 4.9 mm, length of elytra: 3.8 mm, width: 3.0 mm. Body oblong-oval, yellowish brown, dull, antenna yellow, head shiny, except some single setae on head dorsal surface nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins convex and distinctly convergent anteriorly, anterior angles moderately rounded, anterior margin emarginate medially, strongly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface flat, finely and densely punctate, with a few larger punctures each bearing an erect seta anteriorly; frontoclypeal suture indistinct, evenly curved; smooth area anterior to eye weakly convex, as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely and densely punctate, with a terminal seta. Frons completely shiny, with very dense, fine punctures, behind frontoclypeal suture and beside eyes with a few long setae in large sparse punctures. Eyes small, ratio diameter/ interocular width: 0.6. Antenna with ten antennomeres; club with three antennomeres and straight, distinctly shorter than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest shortly before base, lateral margins evenly convex and strongly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles strongly rounded; anterior margin straight, with complete and fine marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures; lateral anterior and lateral margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures. Elytra widest shortly behind middle, striae finely impressed, finely and sparsely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with minute setae in punctures, penultimate interval with a very few single, short setae; epipleural margin robust, ending at weakly curved external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, coarsely and densely punctate, glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.35. Pygidium moderately convex and dull, coarsely and densely punctate, without impunctate midline, glabrous except a few short setae along apical margin. Legs short and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without adjacent serrated line, anterior row of setae complete; posterior margin smooth, weakly widened at apex and smooth ventrally, not serrate dorsally, finely shortly setose. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.7, sharply carinate dorsally, with two groups of spines, basal group at one quarter, apical group at two thirds of metatibial length, with a few robust setae basally subparallel to dorsal margin; lateral face longitudinally convex, shiny, sparsely punctate and glabrous; ventral margin finely serrate, with four equidistant, short and robust setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, laterally and dorsally not carinate; first metatarsomere little shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 47 J–L. Habitus: Fig. 47M. Variation. Pronotum in male slightly wider, pygidium wider and more convex; antennal club in both sexes of same length. Remarks. The shape of the aedeagus in this species is very similar that of Cephaloserica vulpecula Arrow, 1946 from Myanmar. Distribution. See map (Fig. 112) and Table 1.

Maladera orientalis (Motschulsky, 1858) Figures 10 E���H, 89 Serica orientalis Motschulsky, 1858: 33; Brenske 1897: 429; Niijima & Kinoshita 1923: 21; Murayama 1938: 10, 1954: 48. Maladera orientalis: Reitter 1896: 187; Nakane & Baba 1960: 5; Nomura 1963: 124, 1969: 79; Kim & Lee 1997: 124; Kim & Kim 2003: 88. Amaladera cavifrons Reitter, 1896: 188; Dalla Torre 1912: 19; Serica cavifrons: Brenske 1897: 391 (syn by Ahrens 2006b: 22). Amaladera diffinis Reitter, 1896: 188; Dalla Torre 1912: 19; syn by Nikolaev 1977: 269. Serica famelica Brenske, 1897: 391; Niijima & Kinoshita 1923: 24. Serica pekingensis Brenske, 1897: 366. Serica salebrosa Brenske, 1897: 428, syn. n. Autoserica davidis Brenske, 1898: 213, syn. n. Serica mirabilis Brenske, 1894: 9, 41; Autoserica mirabilis: Brenske, 1897: 40; Maladera mirabilis: Ahrens 2007b: 24, syn. n. Type material examined. Lectotype (orientalis, here designated): ♂ ��� SYNTYPUS ♂ Serica orientalis Motschulsky, 1858 A.A. Gusakov des. 2013/ Zoomuzeij MGU (Moskva, ROSSIA) N�� ZMMU Col 03052 Zool. Mus. Mosq. Univ. (Mosquae, ROSSIA) ex coll. V. I. Motschulsky��� (ZMMU). Paralectotypes (orientalis, here designated): 1 ex. (not sexable) ��� SYNTYPUS Serica orientalis Motschulsky, 1858 A.A. Gusakov des. 2013/ Zoomuzeij MGU (Moskva, ROSSIA) N�� ZMMU Col 03051 Zool. Mus. Mosq. Univ. (Mosquae, ROSSIA) ex coll. V. I. Motschulsky��� (ZMMU), 1 ex. (not sexable) ��� Serica orientalis Motsch. Japonia / SYNTYPUS Serica orientalis Motschulsky, 1858 A.A. Gusakov des. 2013/ Zoomuzeij MGU (Moskva, ROSSIA) N�� ZMMU Col 03050 Zool. Mus. Mosq. Univ. (Mosquae, ROSSIA) ex coll. V. I. Motschulsky��� (ZMMU). Holotype (diffinis): ♂ ���Shan-Si 1884 G. Patanin/ Coll. Reitter/ Holotypus 1896 Amaladera (Omaladera) diffinis Reitter ��� (HNHM). Syntypes (pekingensis): 2 ♀♀ ��� Peking 1879 D. Kraatz/ Typus/ pekingensis Brsk. / pekingensis Type Brsk.��� (MSNG), 1 ♂ ��� Peking 1879 D. Kraatz/ 198. Brsk.��� (MSNG). Syntypes (famelica): 1 ♂ ��� China Chang Yang/ Serica famelica type Brsk./ Coll. Brenske/ Typus/ Maladera orientalis (Motsch.) Nikolaev det.��� (ZMHB), 1 ♂ ��� Shang-hai / Brenske vidit/ Serica famelica m��� (MNHN), 1 ♀ ��� China Donck. v./ Serica famelica Brsk. / Coll. Brenske/ Typus��� (ZMHB), 1 ♀ ��� China Tche-fou/ famelica Type Brsk./ Coll. Brenske/ Typus��� (ZMHB), 1 ♂ ���Nord China Chefoo/ tschefuana Type Brsk./ Typus/ Maladera orientalis (Motsch.) Nikolaev det.��� (ZMHB). Lectotype (salebrosa, here des.): ♂ ��� Japan. (Heyne)/ salebrosa Type. Brsk./ Coll. Brenske��� (ZMHB). Paralectotypes (salebrosa, here des.): 2 ♂♂ ��� Japan. (Heyne)/ salebrosa Type. Brsk./ Coll. Brenske��� (ZMHB), 3 ♀ ���Hakane Sch��nfeldt/ salebrosa type Brsk./ Coll. Brenske��� (ZMHB). Syntype (davidis): 1 ♀ ���Mou-Pin Thib. or. David/ davidis type. Brsk./ Davidis Brs��� (ZMHB). Holotype (mirabilis): ♀ ���Coll. R. I. Sc. N. B. Chine / Schang-hai/ 8968/ Coll. J. Thomson/ S. mirabilis type. Brsk./ Type��� (ISNB). Additional material examined. Japan: 1 ♂ ���Near Tokyo Japan/ Serica orientalis Motsch. ��� (NMPC), 1 ♂ ���Peking Westberge Exp. St��tzner��� (NMPC), 1 ♂ ���Japan/ coll. Kraatz/ Serica salebrosa Brsk. J. Moser det.��� (DEIC), 1 ♂ ���Kuzukawa, Aomori Pref. Honshu, Japan 1.5.1951 leg. K. Shimoyama��� (DEIC), 2 ♂♂ ���Kuzukawa, Aomori Pref. Honshu, Japan 25.6.56 leg. K. Shimoyama��� (DEIC), 1 ♂ ���Kuzukawa, Aomori Pref. Honshu, Japan 30.5.56 leg. K. Shimoyama��� (DEIC), 1 ♂ ���Kuzukawa, Aomori Pref. Honshu, Japan 30.5.1945 leg. K. Shimoyama��� (DEIC), 1 ♂ ���Kuzukawa, Aomori Pref. Honshu, Japan 31.5.49 leg. K. Shimoyama��� (DEIC), 4 ♂♂ ���Kuzukawa, Aomori Pref. Honshu, Japan 1.6.1951 leg. K. Shimoyama��� (DEIC), 2 ♂♂ ���Kuzukawa, Aomori Pref. Honshu, Japan 3.6.1955 leg. K. Shimoyama��� (DEIC). 1 ♂ ���Kuzukawa, Aomori Pref. Honshu, Japan 16.6.1951 leg. K. Shimoyama��� (DEIC), 1 ♂ ���Kuzukawa, Aomori Pref. Honshu, Japan 20.6.56 leg. K. Shimoyama��� (DEIC). Russia: 1 ♀ ���Sutschan Ussuri/ Collectio Coleopterorum Johan Albert Hultgren��� (MZLU), 1 ♂ ���SU, Far East (Primor���e) 80km E Vladivostok Tigroviy vill. env. 05.07.1989, Lg. A. Napolov��� (CNA). China: 1 ♂ ��� Peking / ex. Irig. Samlg. J. Breit Wien��� (CF), 1 ♂ ���Shohchow 34 Shansi China��� (CF), 38 ex. ��� Tientsin. May 1905 F.M. Thomson 1907-272.��� (BMNH), 1 ex. ��� Peking China/ Collectio Coleopterorum Johan Albert Hultgren��� (MZLU), 1 ex. ���China Peking V.19 Schoede S.G.��� (ZMHB), 26 ex. ���China: Hubei; Dahongshan 1700 m, Shuizhou VI-2003 leg. Ying et al. ��� (ZFMK), 7 ex. ���China, Bejing Shi, 27.IV. Songshan 40.55N 115.85E Jaroslav Turna leg., 2005��� (ZFMK), 1 ex. (♂) ���Tokio P. Savolainen/ 25.4.54/ japonica��� (MZHF), 1 ex. (♂) ���Taiynanfu/ Serica spec.��� (ZSM), 4 ex. ���Shansi N. China��� (ZSM), 1 ex. ���China: S-Shaanxi (Qinling Shan) river bank at rd. Zhouzhi-Foping. 95 km WSW Xi���an, 1000 m, 33��53���N 108��01���E leg. M. Sch��lke [C01-04]��� (CA-MSNG), 1 ex. ���China-N. Sechuan Shangliang 24- 29.5.2002 lgt. E. Kucera��� (CP), 1 ex. ���China, 1000-1300m, Shaanxi, Qinling mts., Xunyangba (6km E) 23.v.-13.vi.1998 I.H. Marshal leg.��� (CP), 1 ex. ���C-China, Shaanxi prov., Qinling Shan 500m, Lianghekou, 18-19.vi.2000 leg. C. Holzschuh��� (CP), 10 ex. ���China: E Hebei 17.VI.2001 Qinglong env. 40,4��N 118,9��E O. Safranek lgt.��� (CP), 11 ex. ���China, E Hebei, 17.VI. 40.4N 118.9E 2001 Qinglong S env. Jaroslav Turna leg.��� (CP), 1 ex. (♀)���C-China, Shaanxi prov., 110km E of Xi���an, 500m Huashan env. 23-24.VI.2000 leg. C. Holzschuh��� (CP), 2 ex. (♂) ���Kiautschou China��� (ZMHB), 2 ex. (♂) ���Inn Shan Mongol.��� (ZMHB), 1 ex. ���Mongolei Inn Shan��� (ZMHB), 1 ex. (♂) ���Thibet Kuku-Nor 3200m F. Hauser 1898.��� (ZMHB), 1 ex ���China Tche-fou��� (ZMHB), 1 ♂ ���China n. IV-VII.1992 Mongol Ziziqu pr. Baotou env./ ex coll. Milan Nikodym, Praha��� (ZFMK), 1 ♂ ���China bor.occ. 9.5. Shanxi reg. 1990 Shen Tou env. Z. Dolezal lgt.��� (ZFMK), 32 ex. ���China: Inner Mongolia Mt. Daquangdingzishan (2061m) Chifeng 2-12.vii.2009 local coll. Ankauf via Li Jingke 2010��� (ZFMK), 10 ex. ���China, 28 May 2000 Badaling env., 60km NW Bejing Zd. Jindra lgt.��� (ZFMK), 1 ex. ���China, 4 June 2000 Yunmeng Shan Forest Park 30 km N of Huairou (80 km N of Bejing) Zd. Jindra lgt.��� (ZFMK), 6 ex. ���China, 3 June 2000 Fengning env. 1200M NW slope of Yunwu Shan Zd. Jindra lgt.��� (ZFMK), 2 ex. ���Chekiang China/ OL Cartwright Collection 1959��� (USNM), 1 ex. ���Chin-ling Mts. Shensi, E.B. Apr.-May, 1904��� (USNM), 1 ex. ���China��� (USNM), 2 ex. ��� Peking China Summer ���13/ F.N. Meyer collector��� (USNM), 3 ex. ��� Peking Chili, China/ A.P. Jacot May 24 1921 ��� (USNM), 1 ♂ ���Longmenhe, Xingshan, Hubei, 8.VI.1995, 1200m, leg. Wang Shuyong��� (IZAS), 2 ♂♂ ���Chine A. Davis��� (Mus. Paris), 1 ♂ ���Museum Paris Nord Pekin A. Davis 1865/ Jehol��� (MNHN), 1 ♂ ���Museum Paris 1906 Coll. Leon Fairmaire/ Serica orientalis Mot. Pekin ��� (MNHN), 1 ♀ ���Museum Paris 1906 Coll. Leon Fairmaire/ Type / Serica orientalis Mot. Pekin ��� (MNHN), 1 ♀ ��� Type / Museum Paris 1906 Coll. Leon Fairmaire/ Serica diffinis Frm. Pekin ��� (MNHN), 4 ♂♂ ���China Yunnang��� (ZMHB), 3 ♂♂ ���Mongol. Or-Chan-heou��� (ZMHB), 3 ♂♂, 2 ♀♀ ���Kansou or. K���ing-iang-fou��� (ZMHB), 1 ♂ ���Shi-wan-tsze Hiu-mou kiangkeou��� (ZMHB), 3 ♂♂, 3 ♀♀ ���Kiautschougebiet Tsingtau Hass S.G.��� (ZMHB), 1 ♂ ��� Yunnan 2000-3000m 27.20N 100.11E Habashan mts. SE slope 10-13/7. Vit Kuban leg. 92��� (CP), 2 ex. ���China���Shaanxi Lueang env. 15Km NW 26.5.- 31.5.2001 leg. E. Kucera��� (CP), 1 ex. ���China, N-Sichuan 30km W Nanping 11., 16.VI.1992 Jiuzhaigou, 2100m, Jaroslav Turna leg.��� (CP), 46 ex. ���China, Beijing Shi, 28.V. 60Km NE Beijing 2000 Badaling 40.3N 116.0E Jaroslav Turna leg.��� (CP), 1 ex. ���China, Yunnan; Haba Mts., Bailakou pass; h= 3300m, 9.vii.2002 leg. S. Murzin & I. Shokin��� (CP), 1 ex. ���China Yunnan Liujang, 8-24.VI.1994 leg. B. Siska & T. Spevar��� (CP), 1 ♂, 6 ♀♀ ��� Hunan SC. China Luki: Pushih 5.IV.1939 T.C. Maa��� (BMH), 1 ♂ ���Kiagsi, China Shangjao 27.IV.1939 T.C. Maa��� (BMH), 1 ♀ ���China, Beijing Olympic Forest Park, 40.0131N, 116.3914E J. Prena 7.V.2013 ��� (ZFMK), 1 ♂ ���Ta-tsien-Lou Chasseurs Thibetains 1896��� (MNHN). S. Korea: 1 ♂ ���Korea Gonsan��� (ZMHB), 1 ex. (♂) ���Korea��� (ZMHB). N. Korea: 1 ♂ ���N. Korea 18.7.1990 Ondouri 20 km W of Pu-Tshou lgt. S. Murzin��� (ZFMK). Redescription. Length: 7.6 mm, length of elytra: 5.4 mm, width: 4.7 mm. Body oval, dark brown, dull, antenna yellow, labroclypeus moderately shiny, glabrous except for a few short setae on head and lateral margins of pronotum and elytra. Labroclypeus trapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing an indistinct angle; margins weakly reflexed; anterior margin moderately emarginate medially; surface moderately convex medially, shiny, coarsely and densely punctate, partly rugose, with numerous erect setae; frontoclypeal suture feebly incised and weakly bent medially; smooth area anterior to eye twice as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely scarcely punctate, with one to four setae. Frons dull, with fine, dense punctures, with a few short setae beside eyes, behind frontoclypeal suture and along a transverse line on posterior part. Eyes small, ratio of diameter/ interocular width: 0.4. Antenna with ten antennomeres; club with three antennomeres, 0.9 times as long as remaining antennomeres combined. Mentum elevated and flattened anteriorly. Pronotum trapezoidal, widest at base, lateral margins weakly evenly convex and narrowed towards strongly produced and sharp anterior angles; posterior angles blunt; anterior margin nearly straight, with complete and fine marginal line; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only; anterior and lateral margins sparsely setose; hypomeron carinate but not produced ventrally. Scutellum wide, triangular, with fine and dense punctures. Elytra oblong, widest in posterior third, striae weakly impressed, finely and sparsely punctate, intervals weakly convex, with fine, sparse and evenly distributed punctures; odd intervals with a few single adjacent short setae, otherwise punctures with minute setae only; epipleural margin robust, ending at widely rounded external apical angle of elytra, epipleura sparsely setose; apical border membraneous, with a rim of short microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, sparsely setose; metacoxa glabrous except for numerous long setae laterally; abdominal sternites finely and densely punctate, some punctures with very short or minute setae, each sternite with a distinct transverse row of coarse punctures each bearing a short seta, penultimate sternite apically with a wide shiny smooth chitinous border of half sternite length. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.56. Pygidium moderately convex, dull, finely and moderately densely punctate, glabrous except a few longer and shorter setae apically. Legs moderately wide; femora finely and moderately densely punctate, with two longitudinal rows of setae. Metafemur wide, moderately shiny, anterior margin acute, without adjacent, continuously serrated line, surface sparsely and superficially punctate, anterior row of setae complete; posterior ventral margin moderately widened in apical half and not serrate apically, posterior margin dorsally not serrated, glabrous. Metatibia moderately wide and short, widest at middle, ratio width/length: 1/3.1, dorsal margin sharply carinate, with two groups of spines, basal one shortly before middle, apical one at three quarters of metatibial length, beside dorsal margin basally with a few single fine setae; lateral face weakly longitudinally convex, finely sparsely punctate, along middle impunctate; ventral margin finely serrate, with three strong spines of which the distal one is more distant; medial face impunctate, apex shallowly concave interiorly near tarsal articulation. Tarsomeres impunctate dorsally, with sparse, short setae ventrally; metatarsomeres glabrous ventrally and with a strongly serrated longitudinal ridge as well as a strong, smooth carina beside it, first metatarsomere distinctly shorter than following two tarsomeres combined and little shorter than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 10 E���G. Habitus: Fig. 10H. Remarks. Unfortunately, the paralectotypes of Serica orientalis are severly damaged, with both head and pronotum lacking, and the abdomen mostly destroyed; consequently, the most intact male specimen possessing an aedeagus was chosen as lectotype. Both Serica mirabilis and Autoserica davidis were described from female specimens and they are virtually identical externally to females of Maladera orientalis. Syntypes of M. diffinis, M. salebrosa, M. famelica and M. pekingensis all fit with in the range of morphological variation of M. orientalis, and are thus confirmed as synonyms. Distribution. See map (Fig. 89) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 54-56, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Motschulsky, V. (1858) Entomologie speciale. Insectes du Japon. Etudes Entomologiques, 6, 25 - 41. [1857]","Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310","Niijima, Y. & Kinoshita, E. (1923) Die Untersuchungen u ¨ ber japanische Melolonthiden. II. Research Bulletin of the College Experimental Forestry Hokkaido Imperial University, 2, 1 - 253.","Murayama, J. (1938) Revision des Sericines (Col., Scar.) de la Coree. Annotiones Zoologicae Japonenses, 17, 7 - 21.","Murayama, J. (1954) Icones of the Scarabeid-Beetles from Manchuria and Korea. I. Nihon Gakujutu, Shinko-kai, 163 pp.","Reitter, E. (1896) Uebersicht der mir bekannten palaearktischen, mit der Coleopteren-Gattung Serica verwandten Gattungen und Arten. Wiener Entomologische Zeitung, 15, 180 - 188. https: // doi. org / 10.1002 / mmnd. 48018960133","Nakane, T. & Baba, K. (1960) Scarabaeidae of Niigata Pref., Japan. In: Insects of the Niigata Prefecture 4. Bulletin of the Nagaoka Science Museum, 1960, pp. 1 - 9.","Nomura, S. (1963) Some new species of the Coleoptera from Loochoo Is. and its adjacent regions. Entomological Review of Japan, 15, 37 - 42.","Kim, J. I. & Lee, O. J. (1997) Taxonomic study of Korean Sericinae (Melolonthidae, Coleoptera) II, Genus Maladera. Insecta Koreana, 14, 119 - 135.","Dalla Torre, K. W. (1912) Scarabaeidae: Melolonthinae I. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, 45, pp. 1 - 84.","Ahrens, D. (2006 b) Sericinae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248.","Nikolaev, G. V. (1977) Zametki po sinonimii plastinchatousych zhukov Mongolii i sopredelnych territorii. Nasekomye Mongolii, 5, 268 - 271.","Brenske, E. (1898) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 43, 205 - 403. https: // doi. org / 10.1002 / mmnd. 47918980303","Brenske, E. (1894) Die Melolonthiden der palaearktischen und orientalischen Region im Koniglichen naturhistorischen Museum zu Br ʾ ssel. Beschreibung neuer Arten und Bemerkung zu bekannten. Memoire de la Societe Entomologique de Belgique, 2, 3 - 87.","Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1"]}

Maladera laocaiensis Ahrens, Fabrizi & Liu, sp. n. Figures 24 A���E, 97 Type material examined. Holotype: ♂ ���N-Vietnam, Prov. Lao Cai, Sa Pa, 1600-1700 m, N22��19���52������; E103��50���; 23.-27.V.1999 leg. Ahrens, J��ger, Fabrizi/ 1044 Sericini Asia spec.��� (ZFMK). Paratypes: 1 ♂ ���N Vietnam (Tonkin) pr. Vinh Phu 1990 Tam Dao 17.-21.v. P. Pacholatko leg./ VS 79��� (CP), 1 ♂ ��� Caiyanghe Nature Reserve, Pu���er, Yunnan, 28-29.VII.2007, leg. Li Jiahui / LW-118��� (LSSYU). Description. Length: 9.4 mm, length of elytra: 6.6 mm, width: 5.6 mm. Body oval, dark reddish brown, antenna yellow, dorsal surface iridescent shiny, dorsal face nearly glabrous. Labroclypeus wide and trapezoidal, widest at base, lateral margins straight and strongly convergent anteriorly, anterior angles strongly rounded, anterior margin distinctly emarginate, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex, finely, densely punctate, with a few long setae anteriorly; frontoclypeal suture finely incised, weakly curved; smooth area anterior to eye convex, 1.5 times as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely densely punctate, with one terminal seta. Frons with sparse and fine punctures, with a few fine setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.57. Antenna with ten antennomeres; club with three antennomeres and straight, 1.1 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest at base, lateral margins evenly convex and weakly convergent anteriorly, anterior angles moderately produced and right-angled, posterior angles convex; anterior margin weakly convex, with fine marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; lateral margin densely setose; hypomeron carinate, not produced ventrally. Scutellum triangular, with fine, dense punctures. Elytra widest inposterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, moderately dense punctures concentrated along striae, with minute setae in punctures, odd intervals with a few short and fine setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra convex before apex, membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.37. Pygidium dull, convexly elevated at middle before apex, finely and densely punctate, without smooth midline, with numerous long and short, fine setae in apical half. Legs moderately long and narrow, shiny; femora with two longitudinal rows of setae, coarsely and moderately densely punctate. Anterior margin of metafemur acute, with adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia long and moderately wide, widest at apex, ratio of width/length: 1/3.3, moderately carinate dorsally, with two groups of spines, basal group shortly behind middle, apical group at four fifths of metatibial length, in basal half with a few short and fine single setae and a continuous serrated line beside entire dorsal margin until the basal group of spines; lateral face longitudinally convex, with sparse, fine punctures, minutely setose; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere distinctly longer than following tarsomere and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 24 A���D. Habitus: Fig. 24E. Female unknown. Diagnosis. Maladera laocaiensis Ahrens, Fabrizi & Liu, sp. n. differs from the similar Maladera diaolinensis Ahrens, Fabrizi & Liu, sp. n. and M. tiani Ahrens, Fabrizi & Liu, sp. n. in having the fused left and right parameres sharply carinate ventrally. From the syntopically occurring M. tiachinensis Ahrens, Fabrizi & Liu, sp. n., it differs in being iridescent and shiny dorsally and the parameres being shorter and straight in distal half (lateral view), while in M. tiachinensis the dorsal surface if the body is dark and dull and its parameres are curved ventrally in distal half (lateral view). Etymology. The new species is named (adjective in the nominative singular case) after its occurrence in Lao Cai province. Variation. Length: 8.6���9.4 mm, length of elytra: 6.5���6.6 mm, width: 5.1���5.6 mm. Distribution. See map (Fig. 97) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 113-115, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316

Maladera jinghongensis Ahrens, Fabrizi & Liu, sp. n. Figures 35 E���H, 104 Type material examined. Holotype: ♂ ��� China 9.-14.7. Prov. Yunnan mer. Jinghong 1990 R. Cervenka lgt./ 795Sericini: Asia spec.��� (ZFMK). Paratypes: 1 ♂, 4 ♀♀ ��� China 9.-14.7. Prov. Yunnan mer. Jinghong 1990 R. Cervenka lgt.��� (ZFMK), 1 ♂ ���S. China 10.- 14.7.1990 Jinghong pr. Yunnan lgt. S. Becvar ��� (ZFMK). Description. Length: 6.8 mm, length of elytra: 4.7 mm, width: 3.7 mm. Body oblong-oval, reddish brown, dull, labroclypeus shiny, dorsal face nearly glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles moderately rounded, anterior margin shallowly emarginate medially, margins strongly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex, coarsely, densely punctate, with a few erect, long setae; frontoclypeal suture finely incised, medially angled; smooth area anterior to eye small and flat, twice as wide as long; ocular canthus short and wide (1/3 of ocular diameter), finely densely punctate, with a terminal seta. Frons dull, anterior quarter shiny, with dense and fine punctures, with a few single setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.6. Antenna with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at middle, lateral margins in basal half straight and subparallel, evenly convex and convergent in anterior half, anterior angles distinctly produced and sharp, posterior angles blunt, rounded at tip; anterior margin straight, with fine marginal line, base without marginal line; surface moderately finely and densely punctate, with minute setae in punctures and a very few long setae on lateral disc; anterior and lateral margin setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, sparse punctures. Elytra widest at middle, striae finely impressed, finely and densely punctate, intervals flat, with fine, sparse punctures and with minute setae in punctures, penultimate lateral interval with a very few long single setae; epipleural margin fine, ending at external apical angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface moderately shiny, coarsely and densely punctate, nearly glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.78. Pygidium moderately convex, shiny, coarsely and densely punctate, without smooth midline, with a few long setae beside apical margin. Legs short and wide, dull; femora with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, without adjacent serrated line, anterior row of setae complete; posterior ventral margin smooth, strongly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.3, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short and robust single setae; lateral face longitudinally convex, on base sparsely punctate, otherwise smooth and glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 35 E���G. Habitus: Fig. 35H. Diagnosis. Maladera jinghongensis Ahrens, Fabrizi & Liu, sp. n. differs from the similar M. taiwana Nomura, 1974 by the parameres being distinctly longer than the phallobase and the long basal lobe of the left paramere exceeding half the paramere length. Etymology. The new species is named after its type locality, Jinghong (adjective in the nominative singular). Variation. Length: 6.8���7.5 mm, length of elytra: 4.7���5.0 mm, width: 3.7���4.2 mm. Female: eyes only little smaller than in male, ratio of diameter/interocular width: 0.58; antennal club with three antennomeres, distinctly shorter than remaining antennomeres combined; pygidium evenly moderately convex. Distribution. See map (Fig. 104) and Table 1., Published as part of Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), pp. 1-400 in Zootaxa 4922 (1) on pages 160-161, DOI: 10.11646/zootaxa.4922.1.1, http://zenodo.org/record/4496316, {"references":["Nomura, S. (1974) On the Sericini of Taiwan. Toho-Gakuho, 24, 81 - 115."]}

Maladera rotunda (Arrow, 1946) comb.n. Figures 80 E–H, 130 Cephaloserica rotunda Arrow, 1946b: 16. Type material examined. Syntypes: 1 ♂ “ Burma ca. 200 m Washaung, 20 km East of Myitkyina 14.VII.34 Malaise/ N. E. Burma R Malaise B. M. 1945-71/ ♂ / Cotype / Cephaloserica rotunda Arr. co-type” (BMNH), 1 ♀ “N. East Burma Sadon 1200 m 28.6.-5.7. Malaise / N. E. Burma R Malaise B. M. 1945-71/ Cotype / Cephaloserica rotunda Arr. co-type” (BMNH), 1 ♂ “ Burma ca. 200 m Washaung, 20 km East of Myitkyina 14.VII.34 Malaise / Cephaloserica rotunda type Arrow / Typus / 3733 E91” (NHRS), 1 ♀ “ N. East Burma Sadon 1200 m 28.6.-5.7. Malaise / Cephaloserica rotunda n. sp. Arrow / rotunda Arrow / 3735 E91” (NHRS), 1 ♂ “ Burma 1934 Myitkyina, 175m. 15- 30/7 Malaise / Paratypus / 3734 E91” (NHRS). Additional material examined. China: 1 ♂ “ Jinggu, Yunnan, 24.IV. 1955, 930m, leg. Vuschik ” (IZAS), 1 ♂ “ Liuku, Yunnan, 13.VI. 1981, 900m, leg. Liao Subai ” (IZAS), 1 ♂ “ Jinggu, Yunnan, 22.IV. 1955, 930m, leg. Krischnovsknja” (IZAS). Myanmar: 1 ex. (♂) “ Birmania S. S. S. Monti Cariani Bihtan 5-1937 R. Perego” (MSNM), 4 ♂♂ “ Coll. R.I. Sc. N. B. Birmanie / Birmanie 1911/ R. Ley coll. Le Moult vendit” (ISNB), 1 ♂ “ Burma (Myanmar) SW Shan State Taunggyi J. Rejsek 1.- 18.6.1997 ” (ZFMK). Laos: 1 ♂ “ Vientiane I.60 Laos (Baudon)/ 416 Sericini Asia spec.” (NHMB). Thailand: 1 ♂ “ Coll. R.I.Sc.N.B. Thailand / Siam N. Xieng-Sen 24.II.1932 Prince Leopold” (ISNB), 1 ♂ “ Chiang Mai N. Thailand V 1985 N. Koyama leg.” (ZFMK). Redescription. Length: 8.8 mm, length of elytra: 6.4 mm, width: 5.8 mm. Body oval, dark brown, antenna yellowish, surface dull, labroclypeus shiny, except a few short setae on head and sides of elytra, glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and strongly convergent anteriorly, anterior angles blunt; anterior margin weakly emarginate medially; margins moderately but angled and midpoint of median anterior emargination tooth-like reflexed; lateral margin and ocular canthus produce an indistinct angle; surface flat, coarsely, densely punctate, with a transverse row of erect setae behind anterior margin; frontoclypeal suture finely incised, angled medially; smooth area anterior to eye convex, three times as wide as long; ocular canthus short and narrow (1/3 of ocular diameter), finely densely punctate, terminal setae absent. Frons dull, with dense, fine punctures, with a few single short setae beside eyes. Eyes moderately large, ratio diameter/ interocular width: 0.69. Antenna with ten antennomeres; club with three antennomeres and straight, little longer than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum trapezoidal, widest at base, lateral margins evenly convex, strongly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles moderately convex; anterior margin weakly convex, with fine complete marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures; anterior and lateral margin finely sparsely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine and dense punctures. Elytra convex, widest at posterior third, striae finely impressed, finely and densely punctate, intervals flat, with fine, moderately dense punctures, odd intervals with a few single short setae, otherwise with minute setae in punctures only; epipleural margin robust, ending at convex external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a fine rim of microtrichomes (visible at ca 100x magnification). Ventral surface dull, finely and densely punctate, with dense and fine, adpressed setae, metasternal disc sparsely covered with longer setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.38. Pygidium weakly convex, dull, finely and densely punctate, without smooth midline, with a few setae along apical margin. Legs moderately short and wide; femora shiny, with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, with adjacent continuously serrated line; posterior ventral margin smooth, strongly widened at apex, dorsal posterior margin smooth, neither serrate, finely setose. Metatibia long and moderately wide, widest at middle, ratio of width/length: 1/2.2, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half a few short single spines; lateral face longitudinally convex, finely densely punctate on sides, impunctate along midline, punctures with minute setae; ventral margin finely serrate, with six robust equidistant setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres glabrous and with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere slightly longer than following tarsomere and as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 80 E–G. Habitus: Fig. 80H. Distribution. See map (Fig. 130) and Table 1.