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Ptychoptera xanthopleura Dvořák, Oboňa & Manko, sp. nov. from Azerbaijan and Georgia, and Ptychoptera staryi Dvořák, Oboňa & Manko, sp. nov. from Bulgaria are described. P. xanthopleura sp. nov. differs from the other member of the lacustris group mainly by having almost completely yellow pleurae, and by the shape of the epandrium and gonocoxites. The diagnostics of P. staryi sp. nov. and P. incognita Török, Kolcsár & Keresztes, 2015 based on male genitalia are provided.

Pneumia fuehzulii sp. nov. urn:lsid:zoobank.org:act: 07B400D8-6775-4C2E-8B0B-E2BB2D7D4818 Figs 1–20 Differential diagnosis Pneumia fuehzulii sp. nov. resembles P. canescens (Meigen, 1804) in body size, as well as wings, and less distinctly visible characters, such as the shape of aedeagal complex. It clearly differs by stronger CuA 2 (Fig. 13), M3, CuA1 and CuA2 not touched basally. Hypandrium with two swellings in the middle and a deep cleft between them (Figs 8, 16). Gonostyli almost straight (Figs 8, 17–18), distiphallus straight, pointed, extending a little beyond the parameres (dorsal view). Parameres sickle-shaped (Figs 19–20). Apertures of the epandrium small, circular. Remainders of ventral epandrial plate V-shaped, conspicuous (Figs 7, 15). Pneumia canescens is readily distinguishable by CuA 2 not strengthened (Jung 1956: 235, fig. 128). M 3, CuA 1 and CuA 2 touched basally. Hypandrium broader in the middle (Jung 1956: 235, fig. 130; Vaillant 1979: tab. lxxxiv, fig. 9). Gonostyli bent (Vaillant 1979: tab. lxxxiv, fig. 9), distiphallus straight, pointed, as long as the parameres, not sickle-shaped. Basal two apertures of epandrium conspicuous (Vaillant 1979: tab. lxxxiv, fig. 9). Remainders of ventral epandrial plate not detected. Etymology This species is named in honour of the famous medieval Turkish-Azerbaijan poet Məhəmməd Süleyman oğlu Füzuli. Material examined Holotype AZERBAIJAN • 1 ♂; Qəbələ district, Durca; 41.037750° N, 47.886583° E; 1310 m a.s.l.; 9 May 2019; J. Oboňa et al. leg.; Loc. AZE 28; sweep netting, forest edge and brook; slide with a dissected specimen, Cat. No. 34949, Inv. No. 26006; NMPC. Paratypes AZERBAIJAN • 3 ♂♂; same collection data as for holotype; slide, some specimens dissected, Cat. No. 34950 to 34952, Inv. No. 26007 to 26009; NMPC • 1 ♂; Balakən distr., Balakən, Mahamalar; 41.7370700° N, 46.4372111° E; 465 m a.s.l.; 9 May 2019; J. Oboňa et al. leg.; Loc. AZE 25; sweep netting, open small brook (clay, micro-, mesolithal) and marsh, pasture/forest; slide, Cat. No. 34953, Inv. No. 26010; NMPC. Description Male HEAD. As long as broad (Fig. 1); vertex conically inflated with numerous setae, alveoli almost irregularly spaced over its entire surface, with 3–4 supraocular bristles on dorsal margins of eyes on extreme lateral sides of head. Horizontal border of setae above the upper apices of the eyes developed, slightly convex in the middle. Eyes separated (Figs 1, 9); interocular suture conspicuously sclerotized, inverted V-shaped, sclerotized parts gradually tapering from eyes to median, not fused medially, connected by thin ligament. Eye bridge with 5 facet rows (Fig. 9), divided by the approximate width of 4 facet diameters. Ratio of distance of apices of eyes (tangential points) to minimum width of frons approximately 3.5:1. Frontoclypeus (Fig. 1) with a large almost oval central patch of irregular insertions of setae with narrow medial lobe prolonged to interocular suture. Antennae with scape, pedicel and 14-segmented flagellum (Figs 2, 10) haired. Sensory filaments (ascoids) not observed. Scape a little prolonged, barrel-shaped, 1.2 times as long as ovoid pedicel. Both parts with many narrow scales (for size, see Fig. 2). First flagellomere cylindrical (1.1 times as short as pedicel), medial flagellomeres decreasing in size, spindleshaped. Apical progressively bulbous, with a globular apiculus (digit), excentric, 2.1 times as short as the bases bearing them. Mouthparts hardly extend beyond basal palp segments. Length ratios of palpus segments 1.0:1.0:1.2:1.8; last palp segment annulate (Fig. 3). For the terminal lobes of labium, as shown in Fig. 4, lines of spines between both lobes are developed, tongue protuberances in between distally are inconspicuous. Cibarium proximally with a pair of conspicuous bent sclerotized arms, ratio of maximum length of cibarium to length of epipharynx 2.4:1 (Fig. 11), labrum pointed. THORAX. Anepisternum setae patch almost semicircular between spiracle and wing insertion (Fig. 5). Spiracles set high on mesothorax. No thoracic allurement organs, anepimeron with triangular setose patch. Wings (Fig. 13) ovoid, 3.3 mm long in holotype, 3.0– 3.1 mm in paratypes, rounded distally, inconspicuously expanded at posterior margin. Membrane of wing generally clear; however, sometimes distal part of cell, fork areas, ends of all veins excl. Sc and middle part of CuA 2 clouded. Strengthened parts of veins: Sc, R 1 (not the start), R 2+3, basal cell, R 5, whole CuA 2 and almost all wing veins distally. Radial and medial forks arise at the same level, both forks conspicuously in a basal position to end of CuA 2. R5 ends slightly beyond wing apex. Bases of M 3, CuA 1 and CuA 2 are not connected. Wing index (wing length to width) 2.5. Halteres (Fig. 12) stick-shaped, symmetrical in longitudinal axis, bent and haired, with mass of sensory scales subapically. Ratio of maximum length of halteres to their maximum width 2.7:1. Length ratios of femora, tibiae and first tarsomeres: P 1 2.1:2.3:1.0, P 2 2.3:2.8:1.1, P 3 2.7:3.5:1.2. Paired tarsal claws of P 1 conspicuously backward hooked (Fig. 6) with ventral teeth. MALE GENITALIA. Basiphallus of the male aedeagal complex (Figs 8, 19–20) straight from dorsal view (incl. some small differences – Fig. 19), proximally spoon-shaped, triangular, narrowed and edging distally (Figs 8, 19), with longitudinal rib proximally, a little arched in lateral view (Fig. 20). Spatula modified from two broad rectangular bases to a parallel inner deep cleft (Figs 8, 19). Distiphallus straight, pointed, bent dorsally in lateral view (Fig. 20), extending a little beyond the parameres (however, distiphallus sometimes two times longer – see paratype Cat. No. 34953 and Inv. No. 26010 from AZE Loc. 25). Parameres sickle-shaped (Figs 19–20). Ratio of length of gonocoxites to length of aedeagal complex (incl. basiphallus) 1:1.9. Gonocoxites cylindrical (Figs 8, 17), approximately as long as the gonostyli from dorsal view. Gonostyli with circular base, almost straight, waving subapically (Fig. 18), pointed. Epandrium (Figs 7, 14) approximately quadrilateral, caudal half with numerous setae on both sides. Basal two apertures circular, divided, conspicuously bordered by sclerotized folds. Remainders of ventral epandrial plate V-shaped (Figs 7, 15), sclerotized laterally on both sides, with an approximately oval soft part inside, distally with two conspicuous external protrudent corners and near base of the epipropct with two miniature ones. Hypandrium narrow in connection with epandrium; however, the former broad, bar-shaped, with two swellings in the middle and a deep cleft between them (Figs 8, 16). Epiproct relatively small, with an almost semicircular caudal ending or fold, covered with short microsetae (Figs 7, 14–15). Hypoproct large, triangular or tongue-shaped, 0.4 times as long as the epandrial appendages, haired caudally (Figs 7, 14–15). Epandrial appendages (Figs 7, 14) about 1.8 times as long as the epandrium, curved slightly in the middle, and tenacula (numbers 9–12 are stable) restricted to clusters near the rounded apex, gradually becoming shorter towards top of epandrial appendages. Tenacula endings not fringed. Female Unknown. Bionomics Unknown. Adults (males) were collected in montane or hilly habitats, specifically swampy areas of pasture or forest brooks. Distribution Currently recorded only from Azerbaijan., Published as part of Ježek, Jan, Oboňa, Jozef & Manko, Peter, 2022, New synonymy and two new species of Caucasian moth flies (Diptera, Psychodidae, Psychodinae) from Azerbaijan and Georgia, pp. 48-63 in European Journal of Taxonomy 823 (1) on pages 50-54, DOI: 10.5852/ejt.2022.823.1813, http://zenodo.org/record/6628047, {"references":["Jung H. F. 1956. Beitrage zur Biologie, Morphologie und Systematik der europaischen Psychodiden (Diptera). Deutsche Entomologische Zeitschrift 3: 97 - 257."]}

Thornburghiella salihi sp. nov. urn:lsid:zoobank.org:act: B37FA190-53E9-4F00-9937-1DCA00B54BA0 Figs 21–42 Differential diagnosis Thornburghiella salihi sp. nov. resembles T. erinacea (Krek, 1970) comb. nov. in body size, as well as wing shape. Thornburghiella erinacea is readily distinguishable by a scapus 2.4 times as long as the postpedicel. Flagellomere 1 (postpedicel) cucumber-shaped, straight, 3.2 times as long as the following flagellomere (Krek 1970a: 89, fig. 5). Postpedicel with 8 conspicuous, strong bristles arranged in a row. Terminal flagellomere twice as long as the foregoing one, with a little prolonged digit in the axis (Krek 1970a: 89, fig. 3). Gonostyli are with a conspicuous bulbous base (Krek 1970a: 89, fig. 1, sic!) and bent. Distiphallus narrower in the middle; for details, see Krek (1970a: 89, figs 1–2). Table 1 with a morphological comparison shows that Ulomyia erinacea and the new species suggest their belonging to Thornburghiella rather than to Ulomyia. Etymology This species is named in honour of Salih Krek (University of Sarajevo), a famous taxonomist and collector of Psychodidae in the area of the Balkan Peninsula. Material examined Holotype AZERBAIJAN • 1 ♂; Qəbələ district, Durca, Dəmiraparançay river; 41.0541119° N, 47.8868831° E; 1615 m a.s.l.; 10 May 2019; J. Oboňa et al. leg.; Loc. AZE 27; sweep netting, waterfall in canyon and steep fast brook (mega-, macrolithal) passing to a brook in a broad alluvium in a U-shaped valley; slide with a dissected specimen, Cat. No. 34954, Inv. No. 26011; NMPC. Paratypes AZERBAIJAN • 3 ♂♂; same collection data as for holotype; slides, some specimens dissected, Cat. No. 34955, 34956, Inv. No. 26012, 26013; NMPC. GEORGIA • 1 ♂; Batsara Nature Reserve, Samkura River and Khadori waterfall on its tributary; 42.274323° N, 45.351664° E; 1250 m a.s.l.; 2 May 2019; J. Oboňa et al. leg.; Loc. GEO 33; sweep netting; Cat. No. 34957, 34958, Inv. No. 26014, 26015; NMPC • 1 ♂; Mtskheta-Mtianeti region, below Mejilaurni stream; 42.323233° N, 44.646100° E; 1210 m a.s.l., 28 Jun. 2019; J. Oboňa et al. leg.; Loc. GEO 12; sweep netting; Cat. No. 34959, Inv. No. 26016; NMPC • 1 ♂; Ukanamkhari; 42.332017° N, 44.607283° E; 1565 m a.s.l., 28 Jun. 2019; J. Oboňa et al. leg.; Loc. GEO 16; torrent; Cat. No. 34960, Inv. No. 26017; NMPC • 1 ♂; Gveleti village, Gveleti Waterfalls and Tibistskali stream; 42.704444° N, 44.620833° E; 1570 m a.s.l.; 6 Jul. 2019; P. Manko leg.; Loc. GEO 214; sweep netting; Cat. No. 34961, Inv. No. 26018; NMPC. Description Male HEAD. Hardly as long as broad. Vertex conspicuously conically inflated dorsally, forming a V-inverted sclerotized hood with a cut top. The numerous setae alveoli are spaced almost regularly over the entire surface of the upper half of head, with 3–4 supraocular bristles near dorsal margins of the eyes on the extreme lateral sides, despite scar-free striped areas above C-shaped compound eyes medially and laterally. Dorsolateral margins of the eyes are undulant. Eyes separated, interocular suture broadly V-shaped, more sclerotized basally, with a tongue-shaped inconspicuous ligament (Figs 21, 35); eye bridge formed by five facet rows; inner rows of the eyes are reduced to three facets. Minimum distance between eyes corresponds approximately to 2.4 facet diameters; index of distance from tangential points of eye apices to minimum of frons 3.2. Setae alveoli of the frontoclypeus arranged in an almost semicircular, centrally placed patch near the base of the antennae, tapering to a dorsoventral irregular strip of hairs close below the frontal suture (Figs 21, 35). Antenna with 16 articles; scape club-shaped, cylindrical (Fig. 2), somewhat widened apically, 2.7 times as long as its maximum width, narrowed at base, 6.0 times as long as its minimum width. Pedicel almost globular, symmetrical. Flagellomere 1 (postpedicel) pear-shaped, as long as the two following flagellomeres together (Fig. 22), or shorter (Fig. 24, paratype Cat. No. 34955, Inv. No. 26012 from Loc. AZE 27), 1.3 times as long as the second flagellomere. Postpedicel with 3–6 conspicuous, strong bristles arranged in a row. Scape and pedicel with stiletto-shaped scales in contrast to needle-shaped macrosetae of flagellomeres. Flagellomeres 2–12 ovoid, with needle-shaped paired ascoids (Fig. 22), a little bent, shorter than the flagellomeres in which they are inserted; apical pear-shaped flagellomere a little longer than the previous one (Fig. 23), sometimes the last flagellomeres are fused with a long apiculus in the longitudinal axis (Fig. 25, paratype Cat. No. 34956, Inv. No. 26013 from Loc. AZE 27). Length ratio of maxillary palp segments 1.0:1.3:1.4:1.6; apical segment annulated (Fig. 26). Terminal labial lobes (Fig. 36) with diverging rows of spines between them, tongue distal protuberances are inconspicuous. Ratio of maximum length of cibarium (Fig. 37) to length of epipharynx approximately 1.4:1. THORAX. Anepisternum setae patch almost semicircular between spiracle and wing insertion (Fig. 27), anepimeron with a striped setae patch. Spiracles set high on the mesothorax. Patagium not developed. Wings (Fig. 38) lanceolate, 2.7 mm in holotype, 2.4–3.1 mm in paratypes, rounded distally, a little expanded at the posterior margin. Ending of R 5 is beyond the tip of the wing. Wing membrane slightly infuscated only between Sc, R 1 and C. The following veins or their parts strengthened: Sc with conspicuously marked origin and end, R 1 (not the start), R 2, R 5, basal field, CuA 1 (conspicuously basally) and CuA 2. Radial fork complete in contrast to medial one. Both forks and the ending of CuA 2 are in one line (almost central area of wing). Wing index 2.5. Bases of M 3, CuA 1 and CuA 2 are not connected. Knob of halteres hardly globular, a little cut apically, with close sensory microsetae subapically, a prolonged stem as usually developed (Fig. 28). Ratio of maximum length of halteres to their maximum width approximately 2.5:1. Ratios of lengths of femora, tibiae and first tarsal segments P 1 2.2:2.4:1.0, P 2 2.4:2.8:1.2, P 3 2.6:3.4:1.3. Paired tarsal claws of P 1 almost straight, gradually tapering, hooked only subapically (Fig. 29). MALE GENITALIA. Basiphallus almost straight from dorsal view (Figs 31, 41), only sometimes strangulated proximally (Fig. 33), a little undulated from lateral view (Fig. 42). Spatula at the base of aedeagal complex shortly bilobed (Figs 31, 41), distiphallus keg-shaped, prolonged, with inner pipe bordered by indefinable parallel folds and a telescopic gonoporus strengthened by sclerotized, distally connected rounded folds (Figs 31, 34, 41–42). Gonocoxites almost cylindrical, without bulbose base (Figs 31–32, 40), as long as the gonostyli. Epandrium (Figs 30, 39) rectangular, with two divided areas of insertions of hairs distally). Basal paired apertures conspicuous, mostly separated with irregular margins; however, may be connected in a single oval aperture, see paratype from the site Loc. AZE 27, Cat. No. 34955, Inv. No. 26012 (only one male). Ventral epandrial plate not developed (Figs 30, 39). Hypandrium narrow without a lobulus in the middle (Figs 31, 41). Epiproct inconspicuous, triangular, covered with microsetae; hypoproct conspicuous, tongue-shaped, setose, rounded apically from dorsal view (Figs 30, 39), 1.1 times as long as the epandrium. Epandrial appendages strong, cylindrical and straight from dorsal view, enlarged a little basally in contrast to the top (Fig. 39), bent from lateral view (Fig. 30). Tenacula (numbers 9–12) restricted to a cluster near the rounded apex and gradually becoming shorter towards apex of the epandrial appendages (Figs 30, 39). Female Unknown. Comments Krek´s Sijaricia (as subgenus) must be synonymized with Thornburghiella because T. erinacea was established by him as type species (monotypy) of the genus Ulomyia. Bionomics Unknown; males were collected near montane waterfalls and streams or confluences of rivers, approximately 1200–1600 m a.s.l. Distribution Currently recorded only from Georgia and Azerbaijan., Published as part of Ježek, Jan, Oboňa, Jozef & Manko, Peter, 2022, New synonymy and two new species of Caucasian moth flies (Diptera, Psychodidae, Psychodinae) from Azerbaijan and Georgia, pp. 48-63 in European Journal of Taxonomy 823 (1) on pages 54-59, DOI: 10.5852/ejt.2022.823.1813, http://zenodo.org/record/6628047, {"references":["Krek S. 1970 a. Description de cinq especes nouvelles de Psychodidae de Yougoslavie. Bolletino della Associazione Romana di Entomologia 25 (4): 87 - 95.","Vaillant F. 1982. Some comments on the classification of the Psychodidae Psychodinae (Diptera). Bulletin de la Societe entomologique de France 87: 292 - 301.","Enderlein G. 1935. Zur Klassifikation der Psychodinen. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1935: 246 - 249."]}

Genus Pneumia Enderlein, 1935 Pneumia Enderlein, 1935: 247. Type species: Pericoma palustris Meigen, 1804, original designation. Satchelliella Vaillant, 1979: 249. Type species Pericoma mutua Eaton, 1893, subsequent designation p. 255. (Satchelliella Vaillant, 1971, was unavailable without a designation of a type species). Pneumia – Enderlein 1937: 28. — Duckhouse 1987a: 231 (Satchelliella = syn. of Pneumia). — Ježek 1997: 63. — Wagner & Beuk 2002: 86. — Ježek & Omelková 2007: 252. — Salmela 2008: 13. — Svensson 2009: 186. Satchelliella – Salamanna & Raggio 1984: 29. — Wagner 1990: 35. — Wagner & Masteller 1992: 105. — Krek 1999: 7, 304. — Bernotienè 2002: 4. Only selected data are quoted here, for a complete list of synonymies, see Omelková & Ježek 2012, incl. differential diagnosis., Published as part of Ježek, Jan, Oboňa, Jozef & Manko, Peter, 2022, New synonymy and two new species of Caucasian moth flies (Diptera, Psychodidae, Psychodinae) from Azerbaijan and Georgia, pp. 48-63 in European Journal of Taxonomy 823 (1) on page 50, DOI: 10.5852/ejt.2022.823.1813, http://zenodo.org/record/6628047, {"references":["Enderlein G. 1935. Zur Klassifikation der Psychodinen. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1935: 246 - 249.","Vaillant F. 1971 - 1983. Psychodidae - Psychodinae (not finished). In: Lindner E. (ed.) Die Fliegen der palaearktischen Region 287 (1971): 1 - 48; 291 (1972): 49 - 78; 292 (1972): 79 - 108; 305 (1974): 109 - 142; 310 (1975): 143 - 182; 313 (1977): 183 - 206; 317 (1978): 207 - 238; 320 (1979): 239 - 270; 326 (1981): 271 - 310; 328 (1983): 311 - 357.","Enderlein G. 1937. Klassifikation der Psychodiden (Dipt.). Deutsche Entomologische Zeitschrift 3 - 4 (1936): 81 - 112.","Duckhouse D. A. 1987 a. A revision of Afrotropical Setomima, elucidation of their genealogical relationships and descriptions of other Afrotropical Psychodinae (Diptera: Psychodidae). Annals of the Natal Museum 28 (2): 231 - 282.","Jezek J. 1997. New and interesting taxa of moth flies (Diptera, Psychodidae) from different moist biotopes of the Palaearctic region. Casopis Narodniho Muzea, Rada Prirodovedna 166: 105 - 122.","Wagner R. & Beuk P. L. T. 2002. Family Psychodidae. In: Beuk P. L. T. (ed.) Checklist of the Diptera of the Netherlands: 85 - 88. KNNV Uitgeverij, Utrecht.","Jezek J. & Omelkova M. 2007. Faunistic records of the Czech Republic and Slovakia. Psychodidae. Dipterologica Bohemoslovaca 14: 250 - 253.","Salmela J. 2008. Semiaquatic fly (Diptera, Nematocera) fauna of fens, springs, headwater streams and alpine wetlands in the northern boreal ecoregion, Finland. W-album 6: 3 - 63.","Svensson B. W. 2009. Fjarilsmyggfaunan i ett hagmarksomrade och en ladugard i ostra Blekinges skogsland. Med en oversikt av familjen Psychodidae: s morfologi, systematik och utforskande, samt sarskilt de svenska Psychoda s. l. - arternas biologi. Entomologisk Tidskrift, Uppsala 130 (3 - 4): 185 - 208.","Salamanna G. & Raggio S. 1984. Synopsis of the Psychodinae from the Ligurian Alps and Apennines. Annali del Museo Civico di Storia Naturale di Genova 85: 1 - 42.","Wagner R. 1990. Family Psychodidae. In: Soos A. & Papp L. (eds) Catalogue of Palaearctic Diptera 2: 11 - 65. Akademiai Kiado, Budapest and Elsevier Science Publishers, Amsterdam.","Wagner R. & Masteller E. C. 1992. Long term population changes of Psychodidae (Diptera) from a stream recovering from sewage effluent and a control stream. Limnologica 22 (2): 105 - 113.","Krek S. 1999. Psychodidae (Diptera Insecta) Balkanskog Poluotoka [Psychodidae (Diptera Insecta) of the Balkan Peninsula]. Federacija Bosne i Hercegovine, Ministarstvo Obrazovanja, Nauke, Kulture i Sporta, Studentska Stamparija Univerziteta Sarajevo, Sarajevo. [In Croatian.]","Bernotiene R. 2002. Moth flies (Diptera, Psychodidae) new for Lithuanian fauna. Ekologija, Vilnius 2: 4 - 8.","Omelkova M. & Jezek J. 2012. Two new species of Pneumia Enderlein (Diptera, Psychodidae, Psychodinae) from the Palaearctic region. Zootaxa 3180 (1): 1 - 18. https: // doi. org / 10.11646 / zootaxa. 3180.1.1"]}

2. Boreus hyemalis (Linnaeus, 1767) Boreus Hiemalis L [sic!]: Nowicki 1864: 61; 1865: 61 [faunistics] Boreus hiemalis L.: Dziędzielewicz 1867: 162; 1891: 47 [faunistics]; Fudakowski 1923: 202 [distribution] Boreus hiemalis Lin. [sic!]: Majewski 1882: 16; 1885: 1; Dziędzielewicz 1914: 43 [faunistics]; Łomnicki 1914: 110 [faunistics]; Dziędzielewicz 1918: 40 [distribution, ecology] Boreus hyemalis (Linné): Svensson 1972: 27; Penny 1977: 204 [taxonomy, distribution] Boreus hyemalis (Linnaeus) [sic!]: Penny 1977: 204 [taxonomy, distribution] Published records: Lviv [Klepariv], under Poland (Nowicki 1864, 1865; Dziędzielewicz 1867, 1891, 1914, 1918; Majewski 1882, 1885; Penny 1977; Łomnicki 1914; Fudakowski 1923; Svensson 1972). Reviewed material: SMNH collection, 1 ♂, E21.03.01.02/01: “Kleparów Koło Lwowa 19. XII ”; “1”; “ Boreus hiemalis Linnaeus ”, [19.12.?, Lviv]; 1 ♂, E21.03.01.02/02: “ Kleparów Koło Lwowa 19. XII ”; “2”; “ Boreus hiemalis Linnaeus ”, [19.12.?, Lviv]; 1 ♂, E21.03.01.02/03: “ Kleparów Koło Lwowa. 21.11.1897.”; “3”; “ Boreus hiemalis Linnaeus ”, [12.11.1897, Lviv]; 1 ♀, E21.03.01.02/04: “ Czarnohora Foreszczynka 2.-20. XII.1911.”; “4”; “ Boreus hiemalis Linnaeus ”, [2.- 20.12.1911,?]. Remarks: Majewski (1885) reported this species as “pospol. w całym kraju” [common throughout the country], meaning former territories of the Crown of the Kingdom of Poland, which were separated in that time between the Russian and Austro-Hungarian Empires (including Halychyna within the current territory of Ukraine). Penny (1977) mentioned this species from the former territory of Poland, presumably based on previously published historical data. This idea is also confirmed by a published map on figure 81, where one of the distributional points is situated in the western part of Ukraine, close to the Carpathians. All data mentioned by Fudakowski (1923) are based on J. Dziędzielewicz’s and E. Majewski’s reports (the same for B. westwoodi). Mentioned by Dziędzielewicz (1918), specimens of P. hyemalis are preserved in the SMNH collection, except the specimen from Lviv marked in the paper as “19.I” (missing). An additional specimen of a female was collected on the Chornohora Mountains., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on page 463, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Linnaeus, C. (1767) s. n. In: Systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Regnum Animale. 2. Laurentii Salvii, Holmiae [Stockholm], pp. 533 - 1327. https: // doi. org / 10.5962 / bhl. title. 156772","Nowicki, M. (1864) Przyczynek do owadniczej fauny Galicyi. Drukarnia Uniwersytecka w Krakowie, Krakow, 87 pp.","Nowicki, M. (1865) Insecta Haliciae Musei Dzieduszyckiani. Typis Universitatis Jagellonicae, Cracoviae, 87 pp.","Dziedzielewicz, J. (1867) Wykaz owadow siatkoskrzydlych (Neuroptera). Sprawozdania Komisji Fizjograficznej c. k. Towarzystwa naukowego Krakowskiego. Materyaly do fizyografii Galicyi, Krakow, 1, 158 - 165.","Dziedzielewicz, J. (1891) Przeglad fauny krajowej owadow siatkoskrzydlych (Neuroptera, Pseudoneuroptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 26, 26 - 151.","Fudakowski, J. (1923) Znane i nowe stanowiska Borens hiemalis L. i B. westwoodi Hag. (Neuropteroidea - Mecoptera). (Neue und bekannte Standorte von Boreus hiemalis L. u. B. westwoodi Hag. in Polen). Polskie Pismo Entomologiczne, 2 (4), 202 - 204.","Majewski, E. (1882) Systematyczny wykaz owadow Zylkoskrzydlych polskich u nas dotad znalezionych. Insecta Neuroptera Polonica. Ksiegarnia Gebethnera i Wolffa, Warszawa, 42 pp.","Majewski, E. (1885) Owady Zylkoskrzydle (Neuroptera Polonica). Systematyczny wykaz krajowych sieciarek i prasiatnic. Materyaly do fauny krajowej. Nakladem ksiegarni Teodora Paprockiego i S-ki, Warszawa, 1 - 2, 1 - 39.","Dziedzielewicz, J. (1914) O owadzie Posniezku (Boreus), zyjacym na ziemiach Polski. Kosmos, 29 (1 - 3), 42 - 45.","Lomnicki, J. (1914) Zarys fauny okolic Lwowa. Rozprawy i Wiadomo s ci z Muzeum im. Dzieduszyckich. Lwow, 1 (1 - 2), 109 - 111.","Dziedzielewicz, J. (1918) Owady siatkoskrzydlowate ziem Polski (Insecta neuropteroidea Poloniae terrarum). Rozprawy i Wiadomo s ci z Muzeum im. Dzieduszyckich, Lwow, 4 (1 - 4), 38 - 40.","Svensson, S. A. (1972) Boreus Latreille, 1825 (Mecoptera). A Synopsis of Described Species. Insect Systematics & Evolution, 3 (1), 26 - 32. https: // doi. org / 10.1163 / 187631272 X 00049","Penny, N. D. (1977) A systematic study of the family Boreidae (Mecoptera). The University of Kansas science bulletin, 51, 141 - 217."]}

8. Panorpa connexa MacLachlan, 1869 Panorpa connexa MacLachlan, 1869: Martynova 1957: 732 [morphology, taxonomy, key, distribution] Published records: Kropyvnytskyi (Martynova 1957). Remarks: Martynova (1957) reported the presence of two specimens of P. connexa housed in collection of ZIN RAS, labelled as “ЕлиЗаветград [ныне Кировоград], Херс. г. 20 v 91, Яцентковский” [Elisavethrad town [now Kirovograd], Khersonska huberniya, 20.05.1891, leg. A.V. Yatsentkovskii] and concluded that the label is correct. Willmann (1975, 1978a) reported P. connexa only from the Caucasus and northern Turkey along the Caspian Sea. The presence of P. connexa in Ukraine should be confirmed., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on page 470, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Martynova, O. M. (1957) Skopionnitsy (Mecoptera) fauny SSSR. II Semeistvo Panorpidae. [Scorpion-flies (Mecoptera) of USSR fauna. II The family Panorpidae]. Entomologicheskoe obozrenie, 36 (3), 721 - 747. [in Russian]","Willmann, R. (1975) Die Gattung Panorpa in der Turkei. Annalen des Naturhistorischen Museums in Wien, 79, 543 - 564.","Willmann, R. (1978 a) Bemerkungen zu anatolischen Mecopteren. Annalen des Naturhistorischen Museums in Wien, 81, 517 - 523."]}

7. Panorpa communis Linnaeus, 1758 Panorpa communis L.: Belke 1859: 83; Dziędzielewicz 1867: 162; 1884: 2; 1891: 48; Wierzejski 1883: 253 [faunistics]; Lezhenina et al. 2009: 98; Chaplygina 2016: 103 [distribution] Panorpa Communis L [sic!]: Nowicki 1864: 61; 1865: 61 [faunistics] Panorpa Leopoliensis: Dziędzielewicz 1867: 162 nomen nudum [faunistics] Panorpa communis Lin.: Nowicki 1867: 118; 1870: 14; Łomnicki 1877a: 147; Majewski 1882: 16; 1885: 1 [faunistics]; Dziędzielewicz 1918: 38 [distribution, ecology] Panorpa communis: Łomnicki 1877b: 188; Kopytko 2009: 91 [faunistics] Panorpa Leopoliensis Dziędz. [sic!]: Majewski 1882: 16; 1885: 1 [faunistics] Panorpa communis. L. [sic!]: Dziędzielewicz 1898: 193 [faunistics] Panorpa communis Lin. var. diffinis M’L. (vulgaris Imhoff): Dziędzielewicz 1905: 113 [faunistics] Panorpa leopoliensis Dz.: Dziędzielewicz 1891: 49 [faunistics] Panorpa communis Linné: Martynova 1957: 724 [morphology, taxonomy, key, distribution] Panorpa communis Linnaeus, 1758: Dvořák et al. 2017: 15 [faunistics, distribution] Published records: Anheliv, Borshchiv, Buchach, Chornohora, Ciscarpathian region, Drohobych, Dzvenyhorod, Halychyna, Ivano-Frankove, Kamyanets-Podilskyi, Kharkiv, Lviv, Melnytsya-Podilska, Mukachevo, Penyaky, Poland, Siverskyi Donets river in Kharkivska Region, Stanisławów, Ukraine, Uzhhorod, Yaremcha, Zakarpatska Region (Belke 1859; Nowicki 1864, 1865, 1867, 1970; Dziędzielewicz 1867, 1884, 1989, 1891, 1905, 1918; Łomnicki 1877a, 1877b; Majewski 1882 *, 1885 *; Wierzejski 1883; Martynova 1957; Kopytko 2009; Lezhenina et al. 2009; Chaplygina 2016; Dvořák et al. 2017, 2019). Reviewed material: SMNH collection: 1 ♀, E21.01.01.03/02: “ 17.-6.-1907. Worochta.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [17.6.1907, Vorokhta]; 1 ♀, E21.01.01.03/03: “ Lwów okolice.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [?, Lviv]; 1 ♀, E21.01.01.03/04: “Chomiak. P. Weredyk. 25.-7.-1907.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [25.7.1907,?]; 1 ♂, E21.01.01.03/05: “Galicja”; “ Panorpa communis Linnaeus, 1758 det. WK”, [?, Galicia]; 1 ♂, E21.01.01.03/07: “Worochta. Okolice. 2.-8.-1907.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [2.8.1907, Vorokhta]; 1 ♂, E21.01.01.03/08: “Janów 31.5.1903 ”; “ Panorpa communis Linnaeus, 1758 det. WK”, [31.5.1903, Ivaniv]; 1 ♀, E21.01.01.03/09: “Wertełka. 26. V.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [26.5.?, Vertelka]; 1 ♀, E21.01.01.03/10: “Mikuliczyn 31.VII.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [31.7.?, Mykulychyn]; 1 ♂, E21.01.01.03/11: “Worochta. Okolice. 2.-8.-1907.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [2.8.1907, Vorokhta]; 1 ♂, E21.01.01.03/12: “Galicja”; “ Panorpa communis Linnaeus, 1758 det. WK”, [?, Galicia]; 1 ♀, E21.01.01.03/14: “Mikuliczyn. Polonica. 5.-VII”; “ Panorpa communis Linnaeus, 1758 det. WK”, [5.7.?, Mykulychyn]; 1 ♀, E21.01.01.03/15: “ 7.5.1903 Wulka park.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [7.5.1903,?]; 1 ♀, E21.01.01.03/16: “Worochta. Okolice. 2.-8.-1907.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [2.8.1907, Vorokhta]; 1 ♂, E21.01.01.03/17: “Galicja?”; “ Panorpa communis Linnaeus, 1758 det. WK”, [?, Galicia]; 1 ♂, E21.01.01.03/18: “Mikuliczyn. Polonica. 5.-VII”; “ Panorpa communis Linnaeus, 1758 det. WK”, [5.7.?, Mykulychyn]; 1 ♂, E21.01.01.03/19: “Worochta. Okolice. 2.-8.-1907.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [2.8.1907, Vorokhta]; 1 ♂, E21.01.01.03/20: “Mikuliczyn. Polonica. 5.VIII.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [5.7.?, Mykulychyn]; 1 ♀, E21.01.01.03/21: “Mikuliczyn. Polonica. 5.-VII”; “ Panorpa communis Linnaeus, 1758 det. WK”, [5.7.?, Mykulychyn]; 1 ♂, E21.01.01.03/22: “Brzuchowice 23. V.”; “ Panorpa communis Linnaeus, 1758 det. WK”. Other three specimens from SMNH were collected in current territory of Poland labeled as follows: 1 ♂, E21.01.01.03/01: “Tatry”; “ Panorpa communis Linnaeus, 1758 det. WK”, [?, Tatry Mountains?]; 1 ♂, E21.01.01.03/06: “Okolice Warszawy. Zb. Oserloff.”; “ Panorpa communis Linnaeus, 1758 det. WK”, [?, Warsaw]; 1 ♂, E21.01.01.03/13: “Krzeszowice 4.IX”; “ Panorpa communis Linnaeus, 1758 det. WK”, [?, Krzeszowice]. New material examined: 1 ♂, Zakarpatska Region, Perechyn district, Kamyanetskyi, 48°48’37.4”N 22°28’33.3E ”, sweep netting, 11.08.2015; 1 ♂, ibid, Uzh upstream Uzhhorod, 48°36’37.5”N 22°13’49.6”E, sweep netting, 10.08.2015; 3 ♂, 1 ♀, Mukacheve district, near “Pearl of the Carpathians” resort, 48°31’25.8”N 22°52’06.7”E, beer traps in the forest and in trees at the stream, 06.05.2018 – 20.05.2018; ibid, Uzhhorod, 48°39’22.1”N 22°19’50.2”E, beer traps in the garden, apple trees, 01.05.2018 – 31.05.2018 (1 ♂); ibid 20.08.2018 – 26.08.2018 (8 ♀); 2 ♂, 3 ♀, ibid, Uzhhorod, 48°39’22.4”N 22°19’52.2”E, beer traps in oak forest (10 meters at the road), 01.05.2018 – 31.05.2018; 1 ♀, Klyucharky, forest, 48°24’23.5”N 22°39’39.7”E, beer trap, 13.– 28.07.2020; 7 ♂, 12 ♀, Klyucharky, abandoned field, 48°24’23.159”N 22°39’5.414”E, beer trap, 13.– 28.07.2020; 3 ♂, 1 ♀, Klyucharky, abandoned field, 48°24’23.159”N 22°39’5.414”E, beer trap, 27.8.– 11.09.2020. Remarks: The species name P. leopoliensis [with author’ remark “ mihi in litt. Lwów (w maju) ”] was used for the first time in the paper published by (Dziędzielewicz 1867: 162) and should be recognized as a nomen nudum. Later, Dziędzielewicz (1891: 49) reported the similarity of P. leopoliensis with P. communis, and opted to not describe the first species because the specimens were soured. In later contribution, Dziędzielewicz (1905: 113) added more information about ecology and distribution of P. leopoliensis in the Gorgany Mountains. Almost four decades later, Farbotko (1929b: 4) suggested the identity of P. leopoliensis and P. communis based on analysis of J. Dziędzielewicz’s publication from 1891. In Majewski (1882, 1885) a few different species of Panorpa most probably are treated under the single species name P. communis. The record from Kamyanets-Podilskyi in Dziędzielewicz (1891) is published based on Belke (1859). It is probable that many older records, as well as part of the material in this work from SMNH, belong to P. vulgaris (see comments under that species)., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on pages 469-470, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Linnaeus, C. (1758) Systema natura per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Vol. 1. 10 th Edition. Salvii, Holmiae, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542","Belke, G. (1859) Rys historyi naturalnej Kamienca podolskiego [Esquisse l'histoire naturelle de Kamienietz Podolski]. Drukarnia Gazety Codziennej, Warszawa, 1859, 1 - 114.","Dziedzielewicz, J. (1867) Wykaz owadow siatkoskrzydlych (Neuroptera). Sprawozdania Komisji Fizjograficznej c. k. Towarzystwa naukowego Krakowskiego. Materyaly do fizyografii Galicyi, Krakow, 1, 158 - 165.","Dziedzielewicz, J. (1884) Sieciwki (Neuroptera) i Prasiatnice (Pseudoneuroptera) zebrane na Pokuciu w cigu lata 1883. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 18, 1 - 5 (223 - 229).","Dziedzielewicz, J. (1891) Przeglad fauny krajowej owadow siatkoskrzydlych (Neuroptera, Pseudoneuroptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 26, 26 - 151.","Wierzejski, A. (1883) Dodatek do fauny sieciowek (Neuroptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 17, 253 - 255.","Chaplygina, A. B. (2016) Konsortyvni zvyazky kropyvyanky chornoholovoi (Sylvia atricapilla L.) u lisovykh tsenozakh livoberezhnoi Ukrainy. [The consortial relations of Eurasian blackcap (Sylvia atricapilla L.) in the forest cenoses of left bank Ukraine]. Studia Biologica, 10 (1), 99 - 110. [in Ukrainian, English summary] https: // doi. org / 10.30970 / sbi. 1001.464","Nowicki, M. (1864) Przyczynek do owadniczej fauny Galicyi. Drukarnia Uniwersytecka w Krakowie, Krakow, 87 pp.","Nowicki, M. (1865) Insecta Haliciae Musei Dzieduszyckiani. Typis Universitatis Jagellonicae, Cracoviae, 87 pp.","Nowicki, M. (1867) Instrukcya dla dostrzegaczy pojawow w swiecie zwierzecym. Sprawozdania Komisji Fizjograficznej c. k. Towarzystwa naukowego Krakowskiego. Materyaly do fizyografii Galicyi, Krakow, 1, 1 - 29.","Nowicki, M. (1870) Zapiski faunicze. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 4, 1 - 28.","Lomnicki, M. (1877 a) Sprawozdanie z wycieczki zoologicznej odbytej na Podolu w r. 1876 pomiedzy Seretem, Zbruczem a Dniestrem. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 11, 128 - 156.","Majewski, E. (1882) Systematyczny wykaz owadow Zylkoskrzydlych polskich u nas dotad znalezionych. Insecta Neuroptera Polonica. Ksiegarnia Gebethnera i Wolffa, Warszawa, 42 pp.","Majewski, E. (1885) Owady Zylkoskrzydle (Neuroptera Polonica). Systematyczny wykaz krajowych sieciarek i prasiatnic. Materyaly do fauny krajowej. Nakladem ksiegarni Teodora Paprockiego i S-ki, Warszawa, 1 - 2, 1 - 39.","Dziedzielewicz, J. (1918) Owady siatkoskrzydlowate ziem Polski (Insecta neuropteroidea Poloniae terrarum). Rozprawy i Wiadomo s ci z Muzeum im. Dzieduszyckich, Lwow, 4 (1 - 4), 38 - 40.","Lomnicki, M. (1877 b) Spostrzezenia pojawow w swiecie zwierzecym w r. 1876. 3. W Stanislawowie przez Prof. Lomnickiego. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 11, 188.","Kopytko, U. (2009) Konsorty Astrantia major L. v Chornohori (Ukrainski Karpaty). [The consorts of Astrantia major L. in Chornohora (Ukrainian Carpathians). Visnyk Lviv University, Serie Biology, 51, 89 - 92. [in Ukrainian, English summary]","Dziedzielewicz, J. (1898) Wiadomosci o owadach siatkoskrzydlych (Neuroptera) zawarte w roczniku z r. 1896 czasopisma ni- emieckiego: Illustrite Wochenschrift fur Entomologie Neudamm i porownanie spostrzezen o pojawie odnosnych gatunkow w krajach Polski a w szczegolnosci Galicyi. Kosmos, 22, 190 - 197.","Dziedzielewicz, J. (1905) Sieciarki (Neuroptera genuia) i Prasiatnice (Archiptera) zebrane w ciagu lat 1902 i 1903. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 38, 104 - 125.","Martynova, O. M. (1957) Skopionnitsy (Mecoptera) fauny SSSR. II Semeistvo Panorpidae. [Scorpion-flies (Mecoptera) of USSR fauna. II The family Panorpidae]. Entomologicheskoe obozrenie, 36 (3), 721 - 747. [in Russian]","Dvorak, L., Haenni, J. - P., Maca, J., Mariychuk, R. & Obona, J. (2017) Some insects (Dermaptera, Diptera, Mecoptera) from beer traps in Uzhhorod City (Ukraine). Acta Universitatis Presoviensis, Folia Oecologica, 9 (2), 11 - 17.","Lezhenina, I. P., Gramma, V. N., Chaplygina, A. B. & Savinskaya, N. A. (2009) Chlenistonogie v gnezdakh mukholovki-beloshei- ki v nagornoi dubrave (Kharkovskaya oblast). [Arthropods in nests of Muscicapa albicollis in highland oak-grove (Khar- kov Region)]. Nauchnyie vedomosti Belgorodskogo gosudarstvennogo universiteta. Seriya: Yestestvennyie nauki, 3 (58), 95 - 100. [in Russian, English summary]","Dvorak, L., Haenni, J. - P., Dvorakova, K., Kameneva, E. P., Mariychuk, R., Manko, P., Obona, J. & Korneyev, V. A. (2019) Some insects from beer traps in Westernmost Ukraine. Ukrainska Entomofaunistyka, 10 (2), 1 - 6. https: // dx. doi. org / 10.5281 / zenodo. 3584947","Farbotko, J. (1929 b) Materjaly do znajomosci aparatu kopulacyjnego wojsilek (Panorpa) = Materialien zur Kenntnis der Geni- talapparates der Panorpaten. Prace Towarzystwa przyjaciol nauk w Wilnie, 5 (16), 1 - 43."]}

4. Bittacus italicus (Müller, 1766) Bittacus tipularius Latr.: Belke 1859: 83 [faunistics] Bittacus tipularius Fab.: Dziędzielewicz 1868: 153; 1889: 114 [faunistics]; Dziędzielewicz 1918: 40 [distribution, ecology] Bittacus tipularius F.: Łomnicki 1877a: 147 [faunistics] Bittacus tipularius Fb. Ltr. [sic!]: Majewski 1882: 17 [faunistics] Bittacus tipularius Fb. (B. italicus. Klug.) [sic!]: Majewski 1885: 2 [faunistics] Bittacus italicus Müll.: Kuntze 1937 –1938: 9 [faunistics, distribution] Bittacus italicus Müller, 1766 [sic!]: Martynova 1959: 197 [morphology, taxonomy, key, distribution] Bittacus italicus (Müller, 1786) [sic!]: Ermolenko 1994a: 142; Verves et al. 1999: 51 [protection]; Rizun et al. 2000: 26; Ermolenko & Korneev 2009: 138; Holoborodko 2011: 131; Mosin 2017a: 112; Zamoroka et al. 2017: 84; Kletyonkin 2018: 358 [distribution, protection] Bittacus italicus (Müller, 1766): Krivokhatskii & Prokopov 2015: 145 [distribution, protection] Bittacus italicus Mull. [sic!]: Yanytskyi et al. 2008 [distribution, protection]; Chaplygina et al. 2015: 78 [distribution]; Krainyk et al. 2018: 390 [distribution, protection] Published records: Alushta, Bilohirsk and Vyshenne, Carpathians, Cherkaska Regions [near Kiliberdy], Cherkaska, Chortkiv, Crimea, Dnipro river-valley, Dnipropetrovska and Khersonska Regions, Dnister, Donetska Region [Olkhovatka near Enakieve], Hadyach in Poltavska Region, Halychyna, Horodok, Ivano-Frankivsk, Kamyanets- Podilskyi, Khersonska, Khudykivtsi, Melnytsya-Podilska, Mezhyhirtsi near Halych, Mykhailykiv, Mykhalche, Novomlynsk and Krasne Pershe, Verbizh near Kolomyya, Opillya and Pokuttya, Podillya, Poltavska, Prut near Kolomyya, Prysamarya [Dnipropetrovska Region], Pylypche, Rosokhach near Chortkiv, Skala-Podilska, Sumska, Transnistria, Ustya, Velyki Kuchuhury [Kakhovske reservoir], Western Ukraine (Belke 1859; Dziędzielewicz 1868; Łomnicki 1877a; Majewski 1885; Dziędzielewicz 1889, 1918; Kuntze 1937 –1938; Martynova 1959; Ermolenko 1994a; Verves et al. 1999; Rizun et al. 2000; Yanytskyi et al. 2008; Ermolenko & Korneev 2009; Holoborodko 2011; Chaplygina et al. 2015; Krivokhatskii & Prokopov 2015; Mosin 2017a; Dobosz & Hadaś 1999; Rizun et al. 2000; Zamoroka et al. 2017; Kletyonkin 2018; Krainyk et al. 2018). Reviewed material: SMNH collection: 1 ♂, E21.02.01.01/01: “Mielnica nad Dniestrem. 1.-VIII.”; “ Bittacus italicus (Müller, 1786) ”; “1”, [1.8.?, Dnister]; 1 ♀, E21.02.01.01/02: “Rosochacz. 1.VIII”; “ Bittacus italicus (Müller, 1786) ”; “2”, [1.8.?, Rosokhach near Chortkiv]; 1 ♀, E21.02.01.01/03: “Podole.”; “ Bittacus italicus (Müller, 1786) ”; “3”, [?, Podillya]; 1 ♂, E21.02.01.01/04: “Rosochacz. 13.-VIII.”; “ Bittacus italicus (Müller, 1786) ”; “4”, [1.8.?, Rosokhach near Chortkiv]; 1 ♂, E21.02.01.01/05: “Międzyhorce ad Halicz 11.VII 1933 ”; “ Bittacus tipularius ”; “5”; “ Bittacus italicus (Müller, 1786) ” [11.7.1933, Mezhyhirtsi near Halych]. Remarks: Majewski’s (1882) report is based on a previous paper of Dziędzielewicz (1867). At the same time, a few records published by Dziędzielewicz (1868) belong to the personal communication of A. Wiezejski. Majewski (1885) reported this species from “Naddniestrze” [Transnistria] meaning the Podillya Region. Martynova (1959) mentioned this species from Ukraine based on specimens housed in the collection of the Zoological Institute, Russian Academy of Sciences (former ZIN, AS USSR). Distributional data reported by Rizun et al. (2000) are based on faunistical information published by Belke (1859), Dziędzielewicz (1868, 1889), Wierzejski (1868) and Łomnicki (1877a), as well as results of SMNH collection determination. Altogether five specimens were mentioned from this collection: (1) from Podillya region (catalogue number Е 21.01.01.01/3); (2) from Melnytsya-Podilska (21.01.01.01/1); (3–4) from Rosokhach (Е 21.01.01.01/2 and Е 21.01.01.01/4) [all collected by J. Dziędzielewicz]; (5) from Mezhyhirtsi (Е 21.01.01.01/5) [collected by an anonymous author in 11.VIII.1933] (Rizun et al. 2000)., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on pages 466-467, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Muller, O. F. (1766) Manipulus insectorum Taurinensium a Carolo Allionio editus. Miscellania Taurinensia, 3, 185 - 198.","Belke, G. (1859) Rys historyi naturalnej Kamienca podolskiego [Esquisse l'histoire naturelle de Kamienietz Podolski]. Drukarnia Gazety Codziennej, Warszawa, 1859, 1 - 114.","Dziedzielewicz, J. (1868) Dodatek do zeszlorocznego wykazu sieciowek (Neuroptera). Sprawozdania Komisji Fizjograficznej c. k. Towarzystwa naukowego Krakowskiego. Materyaly do fizyografii Galicyi, Krakow, 2, 153.","Dziedzielewicz, J. (1889) Nowy dodatek do fauny owadow siatkoskrzydlych. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 23, 112 - 118.","Dziedzielewicz, J. (1918) Owady siatkoskrzydlowate ziem Polski (Insecta neuropteroidea Poloniae terrarum). Rozprawy i Wiadomo s ci z Muzeum im. Dzieduszyckich, Lwow, 4 (1 - 4), 38 - 40.","Lomnicki, M. (1877 a) Sprawozdanie z wycieczki zoologicznej odbytej na Podolu w r. 1876 pomiedzy Seretem, Zbruczem a Dniestrem. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 11, 128 - 156.","Majewski, E. (1882) Systematyczny wykaz owadow Zylkoskrzydlych polskich u nas dotad znalezionych. Insecta Neuroptera Polonica. Ksiegarnia Gebethnera i Wolffa, Warszawa, 42 pp.","Majewski, E. (1885) Owady Zylkoskrzydle (Neuroptera Polonica). Systematyczny wykaz krajowych sieciarek i prasiatnic. Materyaly do fauny krajowej. Nakladem ksiegarni Teodora Paprockiego i S-ki, Warszawa, 1 - 2, 1 - 39.","Kuntze, R. (1937 - 1938) Charakterystyka entomofaunistyczna scianki wolczynieckiej pod Stanislawowem. Polskie pismo entomologiczne. [Bulletin entomologique de la Pologne], 16 - 17, 1 - 15.","Martynova, O. M. (1959) Skopionnitsy (Mecoptera) fauny SSSR. III Semeistvo Bittacidae. [Scorpion-flies (Mecoptera) of USSR fauna. III The family Bittacidae]. Entomologicheskoe obozrenie, 38 (1), 196 - 199. [in Russian]","Ermolenko, V. M. (1994 a) Bitak italiiskyi, komarivka-bitak italiiska. Bittacus italicus (Muller, 1786). In: Shcherbak, M. M. (Ed.), Chervona knyha Ukrainy. Tvarynnyi svit [Red Book of Ukraine. Fauna]. Kyiv, Ukrainska entsyklopediya, pp. 142. [in Ukrainian]","Verves, Ju. G., Khrokalo, L. A., Pavlyuk, R. S. & Balan, P. G. (1999) Do pryntsypiv doboru bezkhrebetnykh tvaryn u Chervonu knyhu Ukrainy. [Criteria of estimating species of invertebrate animals for including in the red book of Ukraine]. Zapovidna sprava v Ukraini, 5, 48 - 58. [in Ukrainian, English summary]","Rizun, V. B., Konovalova, I. B. & Yanytskyi, T. P. (2000) Ridkisni i znykayuchi vydy komakh v entomolohichnykh kolektsiyakh Derzhavnoho pryrodoznavchoho muzeyu. [Rare and endangered insect species of Ukraine in the entomological collections of the State Museum of Natural History]. SMNH, Lviv, 71 pp. [in Ukrainian, English summary]","Korneev, V. O. & Kotenko, A. G. (2009) Lodovychnyk vestvuda. Boreus westwoodi Hagen 1866. In: Akimov, I. A. (Ed.), Chervona knyha Ukrainy. Tvarynnyi svit. [Red Book of Ukraine. Fauna]. Hlobalkonsaltynh, Kyiv, pp. 139. [in Ukrainian]","Holoborodko, K. K. (2011) Bittacus italicus (Muller, 1786). KOМAРIVKA ItAЛIйсЬKA (BItAK ItAЛЬiнсKIй). Chervona knyga Dnipropetrovskoi oblasti. Tvarynnyj svit. [The red book of Dnipropetrovsk region. Animals]. New Print, Dnipropetrovsk, 131 pp. [in Ukrainian]","Mosin, H. H. (2017 a) Komarivka italiiska. Bittak italyanskii. Bittacus italicus (Muller, 1786). In: Zalevskii, V. D. (Ed.), Chernova knyha Donetskoi oblasti: tvarynnyi svit. Naukovo-informatsiinyi dovidnyk. [R ed Book of Donetska Region: fauna. Scientific and informative guide]. Vinnytska oblasna drukarnya, Vinnytsya, pp. 1 - 112. [in Ukrainian]","Zamoroka, A. M., Bidychak, R. M., Geryak, Yu. M., Glotov, S. V., Kaprus, I. Ya., Kozoriz, Yu. H., Martynov, A. V., Mykhayliuk- Zamoroka, O. V., Pushkar, T. I., Rizun, V. B., Slobodian, O. M., Smirnov, N. A., Utevsky, S. Yu. & Shparyk, V. Yu. (2017) Rozpovsyudzhennya ridkisnykh vydiv bezkhrebetnykh tvaryn, zanesenykh do Chernovoi knyhy Ukrainy, v Ivano-Frankivskii oblasti. [Distribution of rare invertebrate animals listed in the Red Data Book of Ukraine in Ivano-Frankivsk Region]. Ukrainian Entomological Journal, 2 (13), 77 - 94. [in Ukrainian, English summary]","Kletyonkin, V. H. (2018) Znakhidky tvaryn Chervonoi knyhy Ukrainy u Dvorichanskomu raioni Kharkivskoi oblasti. [The findings of animals of the Red book of Ukraine in the Dvorichanskyi district of the Kharkivska Region]. Materialy do 4 - ho vydannya Chervonoi knyhy Ukrainy. Tvarynnyi svit. Series: \" Conservation Biology in Ukraine \", 1 (7), 357 - 363. [in Ukrainian]","Krivokhatskii, V. A. & Prokopov, G. A. (2015) Komarovka italyanskaya (bittak italyanskii) Bittacus italicus (Muller, 1786). In: Ivanov, S. P. & Fateryga, A. V. (Eds.), Krasnaya kniga Respubliki Krym. Zhivotnye. [Red Book of the Republic of Crimea. Animals]. IT \" Arial \", Simferopol, pp. 145. [in Russian]","Yanytskyi, T. P., Godunko, R. J. & Konovalova, I. B. (2008) Formuvannya ekolohichnoi merezhi i zberezhennya entomotsenoziv zakhidnoho regionu Ukrainy. [Formation of ecological network and conservation of entomological cenoses in Western Region of Ukraine]. Naukovyi visnyk Uzhhorodskoho universytetu. Seriya Biolohiya, 23, 249 - 253. [in Ukrainian, English summary]","Chaplygina, A. B., Gramma, V. N., Bondarets, D. I. & Savynska, N. O. (2015) Chlenystonohi u trofotsenotychnii strukturi konsortsii mukholovky biloshyinoi v umovakh lisovykh biogeozenoziv Pivnichno-Skhidnoi Ukrainy. [Arthropods in trophiccenosis structure of collared flycatcher consortium in conditions of forest ecosystems of North-Eastern Ukraine]. Visnyk of Dnipropetrovsk University. Biology, ecology, 23 (1), 74 - 85. [in Ukrainian, English summary] https: // doi. org / 10.15421 / 011511","Krainyk, Yu. M., Busel, V. A., Holovko, O. S. & Horbenko, E. I. (2018) Fauna Chervonoi knyhy Ukrainy Natsionalnoho pryrod- noho parku \" Velykyi Luh \". [The fauna of the Red book of Ukraine of the National Nature Park \" Velykyi Luh \"]. Materialy do 4 - ho vydannya Chervonoi knyhy Ukrainy. Tvarynnyi svit. Series: \" Conservation Biology in Ukraine \", 1 (7), 389 - 393. [in Ukrainian]","Ermolenko, V. M. & Korneev, V. O. (2009) Komarivka italiiska. Bittacus italicus (Muller, 1786). In: Akimov I. A. (Ed.), Chervona knyha Ukrainy. Tvarynnyi svit. [Red Book of Ukraine. Fauna]. Kyiv, Hlobalkonsaltynh, pp 138. [in Ukrainian]","Dobosz, R. & Hadas, T. B. (1999) Bittacus italicus (O. F. Muller, 1766) (Mecoptera: Bittacidae) w Polsce i na Ukrainie. Polskie Towarzystwo Entomologiczne, 17 (3 - 4), 145 - 150.","Dziedzielewicz, J. (1867) Wykaz owadow siatkoskrzydlych (Neuroptera). Sprawozdania Komisji Fizjograficznej c. k. Towarzystwa naukowego Krakowskiego. Materyaly do fizyografii Galicyi, Krakow, 1, 158 - 165.","Wierzejski, A. (1868) Przyczynek do fauny owadow blonkoskrzydlych (Hymenoptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 2, 108 - 120."]}

─ Panorpa picta Hagen, 1863 Panorpa picta Hag.: Martynova 1957: 731; Dorokhova & Martynova 1987: 104 [morphology, taxonomy, key, distribution] Remarks: The occurrence in the Crimea was published by Martynova (1957) and repeated by Dorokhova & Martynova (1987). As Willmann (1975) stated, Panorpa picta is a nomen dubium, and the data from Crimea are unclear because they are based on females only., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on page 471, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Hagen, H. (1863) Die Odonaten- und Neuropteren-Fauna Syriens und Klein-Asiens. Wiener entomologische Monatsschrift, 7, 193 - 199.","Martynova, O. M. (1957) Skopionnitsy (Mecoptera) fauny SSSR. II Semeistvo Panorpidae. [Scorpion-flies (Mecoptera) of USSR fauna. II The family Panorpidae]. Entomologicheskoe obozrenie, 36 (3), 721 - 747. [in Russian]","Dorokhova, G. I. & Martynova, O. M. (1987) Otryad Mecoptera. Skorpionovye mukhi. [The order Mecoptera. Scorpion-flies. Key of the insects of the European part of SSSR]. Opredelitel nasekomykh Evropeiskoi chasti SSSR, 4 (VI), 97 - 107. [in Russian]","Willmann, R. (1975) Die Gattung Panorpa in der Turkei. Annalen des Naturhistorischen Museums in Wien, 79, 543 - 564."]}

9. Panorpa germanica Linnaeus, 1758 Panorpa Montana Brauer [sic!]: Nowicki 1864: 61; 1865: 61 [faunistics] Panorpa montana Brauer: Dziędzielewicz 1867: 162; Łomnicki 1877a: 147 [faunistics] Panorpa germanica L.: Dziędzielewicz 1867: 162; 1891: 48; Majewski 1882: 16 [faunistics]; Dziędzielewicz 1918: 39 [distribution, ecology] Panorpa germanica Lin. (P. montana Brau.) [sic!]: Majewski 1885: 1 [faunistics] Panorpa montana. Bran. [sic!]: Dziędzielewicz 1898: 193 [faunistics] Published records: Carpathians, Dzvenyhorod, Halychyna, Holohory, Horodok, Ivano-Frankove near Lviv, Lviv, Naddnistryanshchyna [within Podillya Region], Prut, Ratyshchi, Sambir, Uzhhorod (Nowicki 1864, 1865; Dziędzielewicz 1867, 1989, 1918; Łomnicki 1877a; Majewski 1882, 1885; Wierzejski 1883; Dvořák et al. 2019). Reviewed material: SMNH collection: Panorpa germanica germanica Linnaeus, 1758: 1 ♂, E21.01.01.04/01: “ Lwów okolice.”; “ Panorpa germanica Linnaeus, 1758 det. WK”, [?, Lviv]; Panorpa germanica graeca Lauterbach, 1972: 1 F, E21.01.01.07/01: “Hołosko wielkie 18.5.”; “ Panorpa germanica graeca Lauterbach det. WK”, [18.5.?, Holosko]; 1 ♂, E21.01.01.07/02: “26.5. Hołosko wielkie”; “ Panorpa germanica graeca Lauterbach det. WK”, [26.5.?, Holosko]; 1 ♂, E21.01.01.07/03: “Hołosko”; “ Panorpa germanica graeca Lauterbach det. WK”, [?, Holosko]; 1 ♂, E21.01.01.07/04: “Pieniaki. 28.V.”; “ Panorpa germanica graeca Lauterbach det. WK”, [28.5.?, Penyaky]; 1 ♀, E21.01.01.07/05: “Mikuliczyn VIII.”; “ Panorpa germanica graeca Lauterbach det. WK”, [?.8.?, Mykulychyn]. New material examined: 1 ♂, Zakarpatska Region, Perehinskyi district, Kamyanytsya village upstream of Sukhyi Stream; 48°42’46.2”N 22°25’24.9”E, sweep netting, 12.8.2015. Remarks: A few records published by Majewski (1882, 1885) most probably belong to P. hybrida (see below); the same author reported P. germanica predominantly based on older publications., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on pages 470-471, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Linnaeus, C. (1758) Systema natura per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Vol. 1. 10 th Edition. Salvii, Holmiae, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542","Nowicki, M. (1864) Przyczynek do owadniczej fauny Galicyi. Drukarnia Uniwersytecka w Krakowie, Krakow, 87 pp.","Nowicki, M. (1865) Insecta Haliciae Musei Dzieduszyckiani. Typis Universitatis Jagellonicae, Cracoviae, 87 pp.","Dziedzielewicz, J. (1867) Wykaz owadow siatkoskrzydlych (Neuroptera). Sprawozdania Komisji Fizjograficznej c. k. Towarzystwa naukowego Krakowskiego. Materyaly do fizyografii Galicyi, Krakow, 1, 158 - 165.","Lomnicki, M. (1877 a) Sprawozdanie z wycieczki zoologicznej odbytej na Podolu w r. 1876 pomiedzy Seretem, Zbruczem a Dniestrem. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 11, 128 - 156.","Dziedzielewicz, J. (1891) Przeglad fauny krajowej owadow siatkoskrzydlych (Neuroptera, Pseudoneuroptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 26, 26 - 151.","Majewski, E. (1882) Systematyczny wykaz owadow Zylkoskrzydlych polskich u nas dotad znalezionych. Insecta Neuroptera Polonica. Ksiegarnia Gebethnera i Wolffa, Warszawa, 42 pp.","Dziedzielewicz, J. (1918) Owady siatkoskrzydlowate ziem Polski (Insecta neuropteroidea Poloniae terrarum). Rozprawy i Wiadomo s ci z Muzeum im. Dzieduszyckich, Lwow, 4 (1 - 4), 38 - 40.","Majewski, E. (1885) Owady Zylkoskrzydle (Neuroptera Polonica). Systematyczny wykaz krajowych sieciarek i prasiatnic. Materyaly do fauny krajowej. Nakladem ksiegarni Teodora Paprockiego i S-ki, Warszawa, 1 - 2, 1 - 39.","Dziedzielewicz, J. (1898) Wiadomosci o owadach siatkoskrzydlych (Neuroptera) zawarte w roczniku z r. 1896 czasopisma ni- emieckiego: Illustrite Wochenschrift fur Entomologie Neudamm i porownanie spostrzezen o pojawie odnosnych gatunkow w krajach Polski a w szczegolnosci Galicyi. Kosmos, 22, 190 - 197.","Wierzejski, A. (1883) Dodatek do fauny sieciowek (Neuroptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 17, 253 - 255.","Dvorak, L., Haenni, J. - P., Dvorakova, K., Kameneva, E. P., Mariychuk, R., Manko, P., Obona, J. & Korneyev, V. A. (2019) Some insects from beer traps in Westernmost Ukraine. Ukrainska Entomofaunistyka, 10 (2), 1 - 6. https: // dx. doi. org / 10.5281 / zenodo. 3584947","Lauterbach, K. E. (1972) Zur Kenntnis der Skorpionsfliegen des Balkans (Mecoptera, Panorpidae). Deutsche Entomologische Zeitschrift, 19, 109 - 140. https: // doi. org / 10.1002 / mmnd. 19720190112"]}

6. Panorpa cognata Rambur, 1842 Panorpa fasciata F. (?) [sic!]: Majewski 1882: 16; 1885: 2 [faunistics] Panorpa cognata Ramb.: Wierzejski 1883: 253 [faunistics]; Dziędzielewicz 1891: 49; 1918: 39 [distribution, ecology] Panorpa cognata Ramb. (P. germanica Brau.) [sic!]: Majewski 1885: 1 [faunistics] Panorpa cognata. Ramb. [sic!]: Dziędzielewicz 1898: 193 [faunistics] Panorpa cognata Rambur, 1842: Martynova 1957: 732 [morphology, taxonomy, key, distribution] Published records: Anheliv, Chernihivska Region, Dzvenyhorod, Halychyna, Kamyanets-Podilskyi, Lviv [Holoko], Potorytsya (Belke 1859; Majewski 1882, 1885; Wierzejski 1883; Dziędzielewicz 1891, 1898, 1918; Martynova 1957). Reviewed material: SMNH collection: 1 ♂, E21.01.01.02/01: “Poturzyca Lom. 8-9. wielki las.”; “ Panorpa cognata Rambur, 1842 det. WK”, [?, Potorytsya l]; 1 ♀, E21.01.01.02/02: “Hołosko wielkie 30.6.1867.”; “ Panorpa cognata Rambur, 1842 det. WK”, [30.6.1867, Holosko]; 1 ♂, E21.01.01.02/03: “Hołosko”; “ Panorpa cognata Rambur, 1842 det. WK”, [?, Holosko]. Remarks: Majewski (1882, 1885) indicated a questionable record of “ Panorpa fasciata F. (?)” previously published by Belke (1859) together with other representatives of the genus Panorpa. Dziędzielewicz (1891) also reported “ P.? fasciata Fab. ” as a questionable species based on G. Belke’s information, and suggested that the record belongs to P. cognata. At present, the name P. fasciata is a synonym of P. americana Swederus, which is restricted to the U.S.A. (Penny & Byers 1979). The specimens from SMNH collection were mentioned in Dziędzielewicz (1918: 39)., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on pages 468-469, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Rambur, M. P. (1842) Histoire Naturelle des Insectes, Nevropteres. Fain et Thunot, Paris, 534 pp.","Majewski, E. (1882) Systematyczny wykaz owadow Zylkoskrzydlych polskich u nas dotad znalezionych. Insecta Neuroptera Polonica. Ksiegarnia Gebethnera i Wolffa, Warszawa, 42 pp.","Majewski, E. (1885) Owady Zylkoskrzydle (Neuroptera Polonica). Systematyczny wykaz krajowych sieciarek i prasiatnic. Materyaly do fauny krajowej. Nakladem ksiegarni Teodora Paprockiego i S-ki, Warszawa, 1 - 2, 1 - 39.","Wierzejski, A. (1883) Dodatek do fauny sieciowek (Neuroptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 17, 253 - 255.","Dziedzielewicz, J. (1891) Przeglad fauny krajowej owadow siatkoskrzydlych (Neuroptera, Pseudoneuroptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 26, 26 - 151.","Dziedzielewicz, J. (1918) Owady siatkoskrzydlowate ziem Polski (Insecta neuropteroidea Poloniae terrarum). Rozprawy i Wiadomo s ci z Muzeum im. Dzieduszyckich, Lwow, 4 (1 - 4), 38 - 40.","Dziedzielewicz, J. (1898) Wiadomosci o owadach siatkoskrzydlych (Neuroptera) zawarte w roczniku z r. 1896 czasopisma ni- emieckiego: Illustrite Wochenschrift fur Entomologie Neudamm i porownanie spostrzezen o pojawie odnosnych gatunkow w krajach Polski a w szczegolnosci Galicyi. Kosmos, 22, 190 - 197.","Martynova, O. M. (1957) Skopionnitsy (Mecoptera) fauny SSSR. II Semeistvo Panorpidae. [Scorpion-flies (Mecoptera) of USSR fauna. II The family Panorpidae]. Entomologicheskoe obozrenie, 36 (3), 721 - 747. [in Russian]","Belke, G. (1859) Rys historyi naturalnej Kamienca podolskiego [Esquisse l'histoire naturelle de Kamienietz Podolski]. Drukarnia Gazety Codziennej, Warszawa, 1859, 1 - 114.","Penny, N. D. & Byers, G. W. (1979) A Check-List of the Mecoptera of the World. Acta Amazonica, 9 (2), 365 - 388. https: // doi. org / 10.1590 / 1809 - 43921979092365"]}

3. Boreus westwoodi Hagen, 1866 Boreus Westwoodi Hag. [sic!]: Dziędzielewicz 1914: 44; Dziędzielewicz 1918: 40 [faunistics, distribution, comparison with P. hyemalis]; Fudakowski 1923: 203 [distribution]; Svensson 1972: 29 [taxonomy, distribution]; Ermolenko 1994b: 144 [protection]; Verves et al. 1999: 51 [protection]; Rizun et al. 2000: 27 [distribution, protection]; Korneev & Kotenko 2009: 139 [distribution]; Mateleshko 2011: 92 [distribution]; Zhuravchak 2011: 65 [distribution, protection]; Mosin 2017b: 113 [distribution, protection] Bareus [sic!] westwoodi Hagen: Krainyk et al. 2018: 390 [distribution, protection] Published records: Bilske near Sarny [Rivnenskyi Nature Reserve], Breskul, Carpathians [Zakarpatska, Ivano- Frankivska and Chernivetska Regions], Chornohora, Donetska Region [“Svyati Hory”], Foreshchynka, Prut, Turkul, Ukrainian Carpathians, Ukrainian Polissya [Zhytomyrska, Kyivska and Chernihivska Regions], Ukrainian Polissya, near Irpin [Kyivska Region], vicinity of Skelky [Vasylivskyi district, Zaporizhska Region], Zavoielya (Dziędzielewicz 1914, 1918; Fudakowski 1923; Svensson 1972; Ermolenko 1994b; Korneev & Kotenko 2009; Verves et al. 1999; Rizun et al. 2000; Zhuravchak 2011; Mateleshko 2011; Mosin 2017b; Krainyk et al. 2018). Reviewed material: SMNH collection: 1 ♀, E21.03.01.01/01: “Czarnohora Turkuł. 7. VI 1911. 1880m.”; “1”; “ Boreus westwoodi Hagen, 1866 ”, [7.6.1911, Chornohora-Turkul]; 1 ♂, E21.03.01.01/02: “Czarnohora Foreszczynka w grudniu (XII)”; “2”; “ Boreus westwoodi Hagen, 1866 ”, [?, Chornohora-Foreshchynka]; 1 ♂, E21.03.01.01/03: “Czarnohora Foreszczynka 2-20.XII-1911.”; “3”; “ Boreus westwoodi Hagen, 1866 ” [2- 20.12.1911, Chornohora- Foreshchynka]; 1 ♂, E21.03.01.01/04: “Czarnohora Foreszczynka I (styczeń) 1911.”; “4”; “ Boreus westwoodi Hagen, 1866 ”, [1911, Chornohora-Foreshchynka]; 1 ♂, E21.03.01.01/05: “Czarnohora. Koźmieska. 10.-1.-1909 zeb. H. Huppenthal.”; “5”; “ Boreus westwoodi Hagen, 1866 ”, [10.1.1909, Chornohora]; 1 ♀, E21.03.01.01/06: “W kosówce nad Niesamowitym 15.IX.1924.”; “ Boreus Hag. westwoodi ”; “ Boreus westwoodi Hagen, 1866 ”, [15.9.1924,?]; 1 ♂, E21.03.01.01/07: “Pod Breskul 11.IX.03.”; “7”; “ Boreus westwoodi Hagen, 1866 ” [11.9.1903, Breskul]. Remarks: The determination of materials listed by Dziędzielewicz (1914) was confirmed by František Klapálek. All records mentioned by Svensson (1972) are based on the summarized publication of Dziędzielewicz (1918) and listed under Poland. Penny (1977) did not report this species from Ukraine on published distributional maps. Fudakowski (1923) reported two additional localities of this species, one from the Tatra Mountains and the second from the western part of the Roztochchya Region within the current territory of Poland. B. westwoodi also occurs in the eastern part of Roztochchya belonging to Ukraine. Distributional data reported by Rizun et al. (2000) are based on faunistical information published in Red Book of Ukraine (1994), and results of SMNH collection determination; altogether seven specimens were mentioned (see also above): (1) Chornohora (catalogue number Е 21.02.01.01/7); (2) Kozmeshchyk (21.02.01.01/5); (3–5) Foreshchynka (Е 21.02.01.01/2, Е 21.02.01.01/3 and Е 21.02.01.01/4); (6) Turkul (Е 21.02.01.01/1) [collected by J. Dziędzielewicz and H. Huppenthal]; (7) Nesamovyte (Е 21.02.01.01/6) [collected by an anonymous author in 15.IX.1924]., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on pages 465-466, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Hagen, A. H. (1866) Synopsis of the genus Boreus. Entomologists Monthly Magazine, 3, 132","Dziedzielewicz, J. (1914) O owadzie Posniezku (Boreus), zyjacym na ziemiach Polski. Kosmos, 29 (1 - 3), 42 - 45.","Dziedzielewicz, J. (1918) Owady siatkoskrzydlowate ziem Polski (Insecta neuropteroidea Poloniae terrarum). Rozprawy i Wiadomo s ci z Muzeum im. Dzieduszyckich, Lwow, 4 (1 - 4), 38 - 40.","Fudakowski, J. (1923) Znane i nowe stanowiska Borens hiemalis L. i B. westwoodi Hag. (Neuropteroidea - Mecoptera). (Neue und bekannte Standorte von Boreus hiemalis L. u. B. westwoodi Hag. in Polen). Polskie Pismo Entomologiczne, 2 (4), 202 - 204.","Svensson, S. A. (1972) Boreus Latreille, 1825 (Mecoptera). A Synopsis of Described Species. Insect Systematics & Evolution, 3 (1), 26 - 32. https: // doi. org / 10.1163 / 187631272 X 00049","Ermolenko, V. M. (1994 b) Lodovychnyk vestvuda. Lodovychnyk zvychainyi. Boreus westwoodi Hagen, 1866. In: Shcherbak, M. M. (Ed.), Chervona knyha Ukrainy. Tvarynnyi svit [Red Book of Ukraine. Fauna]. Ukrainska entsyklopediya, Kyiv, pp. 144. [in Ukrainian]","Verves, Ju. G., Khrokalo, L. A., Pavlyuk, R. S. & Balan, P. G. (1999) Do pryntsypiv doboru bezkhrebetnykh tvaryn u Chervonu knyhu Ukrainy. [Criteria of estimating species of invertebrate animals for including in the red book of Ukraine]. Zapovidna sprava v Ukraini, 5, 48 - 58. [in Ukrainian, English summary]","Rizun, V. B., Konovalova, I. B. & Yanytskyi, T. P. (2000) Ridkisni i znykayuchi vydy komakh v entomolohichnykh kolektsiyakh Derzhavnoho pryrodoznavchoho muzeyu. [Rare and endangered insect species of Ukraine in the entomological collections of the State Museum of Natural History]. SMNH, Lviv, 71 pp. [in Ukrainian, English summary]","Korneev, V. O. & Kotenko, A. G. (2009) Lodovychnyk vestvuda. Boreus westwoodi Hagen 1866. In: Akimov, I. A. (Ed.), Chervona knyha Ukrainy. Tvarynnyi svit. [Red Book of Ukraine. Fauna]. Hlobalkonsaltynh, Kyiv, pp. 139. [in Ukrainian]","Mateleshko, O. Yu. (2011) Lodovychnyk Vestvuda. Boreus westwoodi Hagen, 1866. In: Mateleshko, O. Yu. & Potish, L. A. (Eds.), Chervona knyha Ukrainskykh Karpat. Tvarynnyi svit. [Red Book of the Ukrainian Carpathians. Fauna]. Uzhhorod, Karpaty, pp. 92. [in Ukrainian]","Zhuravchak, R. O. (2011) Stan vyvchennya ta rarytetna skladova entomofauny Rivnenskoho pryrodnoho zapovidnyka ta sumizhnykh terytorii. [The condition of the studying and rare component of entomofauna of Rivnenskyi nature reserve and adjacent territories]. Naukovyi visnyk Uzhhorodskoho universytetu. Seriya Biolohiya, 31, 62 - 67. [in Ukrainian, English summary]","Mosin, H. H. (2017 b) Lodovychnyk Vestvuda. Lednichnik Vestvuda. Boreus westwoodi Hagen, 1866. In: Zalevskii, V. D. (Ed.), Chernova knyha Donetskoi oblasti: tvarynnyi svit. Naukovo-informatsiinyi dovidnyk. [R ed Book of Donetska Region: fauna. Scientific and informative guide]. Vinnytska oblasna drukarnya, Vinnytsya, 113 pp. [in Ukrainian]","Krainyk, Yu. M., Busel, V. A., Holovko, O. S. & Horbenko, E. I. (2018) Fauna Chervonoi knyhy Ukrainy Natsionalnoho pryrod- noho parku \" Velykyi Luh \". [The fauna of the Red book of Ukraine of the National Nature Park \" Velykyi Luh \"]. Materialy do 4 - ho vydannya Chervonoi knyhy Ukrainy. Tvarynnyi svit. Series: \" Conservation Biology in Ukraine \", 1 (7), 389 - 393. [in Ukrainian]","Penny, N. D. (1977) A systematic study of the family Boreidae (Mecoptera). The University of Kansas science bulletin, 51, 141 - 217."]}

10. Panorpa hybrida McLachlan, 1882 Panorpa gibberosa M’ L [sic!]: Dziędzielewicz 1883: 4; 1884: 2 [faunistics] Panorpa Gibberosa M’ L [sic!]: Majewski 1885: 2 [faunistics] Panorpa gibberosa ML.: Dziędzielewicz 1891: 48 [faunistics] Panorpa hybrida Mc‘ Lach. [sic!]: Dziędzielewicz 1918: 39 [distribution, ecology] Published records: Bryukhovychi near Lviv, Dnister, Dobryanychi, Dzvenyhorod, Eastern Carpathians near Kolomyya, Horodok, Kolomyya, Lviv, Molodyatyn, mountain and premountain regions of the Carpathians, Prut, Ratyshchi, Rava-Ruska, Sambir, Svirzh river-valley in Prybyn near Bibrka, Ustechko, Yablunytsya, Yaremcha (Dziędzielewicz 1883, 1984, 1891, 1918; Majewski 1885). Reviewed material: SMNH collection: 1 ♂, E21.01.01.05/01: “Brzuchowice 20. V.1908.”; “ Panorpa hybrida McLachlan, 1882 det. WK”, [20.5.1908, Bryukhovychi]; 1 ♂, E21.01.01.05/02: “Przybyń. 15. V.”; “ Panorpa hybrida McLachlan, 1882 det. WK”, [15.5.?,?]; 1 ♀, E21.01.01.05/03: “Popów koło Kołomyji.”; “ Panorpa hybrida McLachlan, 1882 det. WK”, [?, Kolomyya]; 1 ♂, E21.01.01.05/04: “Brzuchowice. 14. V.1910 ”; “ Panorpa hybrida McLachlan, 1882 det. WK”, [14.5.1910, Bryukhovychi]; 1 ♂, E21.01.01.05/05: “Przybyń Świrż. 16. V.”; “ Panorpa hybrida McLachlan, 1882 det. WK”, [16.5.?, Svirzh]; 1 ♂, E21.01.01.05/06: “Brzuchowice 20. V.1908.”; “ Panorpa hybrida McLachlan, 1882 det. WK”, [15.5.1908, Bryukhovychi]; 1 ♀, E21.01.01.05/07: “Kołomyja maj.”; “ Panorpa hybrida McLachlan, 1882 det. WK”, [?, Kolomyya]; 1 ♀, E21.01.01.05/08: “Brzuchowice 20. V.1908.”; “ Panorpa hybrida McLachlan, 1882 det. WK”, [20.5.1908, Bryukhovychi]; 1 ♂, E21.01.01.05/09: “Przybyń przy rzece Świrż 15. V.”; “ Panorpa hybrida McLachlan, 1882 det. WK”, [15.5.?,?]; 1 ♂, E21.01.01.05/10: “Kołomyja maj.”; “ Panorpa hybrida McLachlan, 1882 det. WK”, [?, Kolomyya]. Remarks: All specimens from the SMNH collection were mentioned in a generalized contribution published by Dziędzielewicz (1918); data previously published as P. gibberosa (now P. germanica ssp. gibberosa) belong to P. hybrida based on reinvestigation performed by F. Klapálek as reported in J. Dziędzielewicz’s paper., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on page 471, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["McLachlan, R. (1882) A new European Panorpa. Entomologists Monthly Magazine, 19, 130 - 132.","Dziedzielewicz, J. (1883) Sieciwki (Neuroptera) zebrane w okolicach Koomyi i nad Dniestrem w r. 1882. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 17, 1 - 9.","Dziedzielewicz, J. (1884) Sieciwki (Neuroptera) i Prasiatnice (Pseudoneuroptera) zebrane na Pokuciu w cigu lata 1883. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 18, 1 - 5 (223 - 229).","Majewski, E. (1885) Owady Zylkoskrzydle (Neuroptera Polonica). Systematyczny wykaz krajowych sieciarek i prasiatnic. Materyaly do fauny krajowej. Nakladem ksiegarni Teodora Paprockiego i S-ki, Warszawa, 1 - 2, 1 - 39.","Dziedzielewicz, J. (1891) Przeglad fauny krajowej owadow siatkoskrzydlych (Neuroptera, Pseudoneuroptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 26, 26 - 151.","Dziedzielewicz, J. (1918) Owady siatkoskrzydlowate ziem Polski (Insecta neuropteroidea Poloniae terrarum). Rozprawy i Wiadomo s ci z Muzeum im. Dzieduszyckich, Lwow, 4 (1 - 4), 38 - 40."]}

— Panorpa caucasica MacLachlan, 1869 Panorpa caucasica Sel. [sic!]: Dziędzielewicz 1907: 12 [faunistics] Aulops caucasica Mc’ Lach. [sic!]: Dziędzielewicz 1918: 39 [distribution, ecology; Aulops in generic rank] Published records: Chomyak, Eastern Carpathians, Mykulychyn, Tatariv and Vorokhta (Dziędzielewicz 1907, 1918). Remarks: Dziędzielewicz (1907) reported one female marked as P. caucasica in the vicinity of Mykulychyn village (Ukrainian Carpathians). The specimen was determined by F. Klapálek. Later, Dziędzielewicz (1918) listed a few localities for this species within Gorgany and Chornohora Mountains. We did not find any specimens belonging to P. caucasica in SMNH collection for additional investigation, because material is lost (V.B. Rizun, personal communication; collection curator at SMNH). However, the record is very probably based on misidentification with another species. Martynova (1957) and later Willmann (1978b) knew P. caucasica from the Caucasus only; no recent records of this species are known., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on page 468, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Dziedzielewicz, J. (1907) Sieciarki (Neuroptera genuina) i Prasiatnice (Archiptera) zebrane w ciagu lat 1904 i 1905. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 42, 13 - 25.","Dziedzielewicz, J. (1918) Owady siatkoskrzydlowate ziem Polski (Insecta neuropteroidea Poloniae terrarum). Rozprawy i Wiadomo s ci z Muzeum im. Dzieduszyckich, Lwow, 4 (1 - 4), 38 - 40.","Martynova, O. M. (1957) Skopionnitsy (Mecoptera) fauny SSSR. II Semeistvo Panorpidae. [Scorpion-flies (Mecoptera) of USSR fauna. II The family Panorpidae]. Entomologicheskoe obozrenie, 36 (3), 721 - 747. [in Russian]","Willmann, R. (1978 b) Neubeschreibung der Typen von Panorpa caucasica Mac Lachlan, 1869. Bulletin de l'Institut royal des sciences naturelles de Belgique, 52 (9), 1 - 4."]}

5. Panorpa alpina Rambur, 1842 Panorpa alpina Rambur, 1842: 330 Panorpa Variabilis Brauer [sic!]: Nowicki 1864: 61; 1865: 61 [faunistics] Panorpa variabilis Brauer: Dziędzielewicz 1867: 162 [faunistics] Panorpa alpine Rbr. [sic!]: Majewski 1882: 16; 1885: 1; Dziędzielewicz 1883: 4; 1891: 47; Wierzejski 1883: 253 [faunistics] Panorpa alpina Ramb.: Dziędzielewicz 1905: 112 [distribution, ecology] Aulops alpina Ramb. (Panorpa alpina Ramb.) [sic!]: Dziędzielewicz 1918: 39 [distribution, ecology; Aulops in generic rank]; Farbotko 1929a: 3 [distribution, ecology] Taxonomic remarks: P. pseudoalpina was described by Nagler (1970) from Romania, and no records were published from other countries. Ward (1979) synonymised Panorpa variabilis Brauer, 1857, Panorpa antiporum Nagler, 1968, Panorpa pseudoalpina Nagler, 1970, Panorpa pura Klapálek 1906, Panorpa susteri Nagler 1970, and Panorpa plitvicensis Lauterbach, 1972 with P. alpina. This synonymy was not accepted by Willmann (2013) who treated P. pseudalpina as a subspecies of P. alpina. In this paper, we are following Ward’s results, as no scientific paper with contradictory opinion was published. On the other hand, WK found in SMNH collection several specimens which are in agreement with Nagler’s description of P. pseudoalpina. As the authors have insufficient material to study the variability in P. pseudoalpina and P. alpina in Ukraine, a conclusion regarding the identity of the specimens presented here cannot be made. The records are kept separate from the P. alpina records for other experts who may wish to carry out comparative studies. Published records: Angeliv, Anheliv, Anheliv, Bibrka, Buchach [Hai Buchatski], Buchach, Carpathians, Chomyak, Ciscarpathian region, Dnister, Hai Buchatski, Ivano-Frankove, Kolomyya, Lisna Slobidka, Lviv and Mykulychyn vicinities, Molodyatyn, Nezvysko, Peremyshlyany, steppic part of Podillya Region, mountain and premontane regions of the Carpathians, Ustechko, Varatyk near Pechenizhyn and south of Kolomyya, Yaremcha, Zubra near Lviv (Nowicki 1864, 1865; Dziędzielewicz 1867, 1883, 1891, 1905, 1918; Majewski 1882, 1885; Wierzejski 1883; Farbotko 1929a). Reviewed material: SMNH collection: 1 ♀, E21.01.01/01: “Czornohora. Foreszczynka. 1.- VI.-1909. zeb. Fiarowicz”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [1.6.1909, Chornohora]; 1 ♂, E21.01.01/02: “Subrawka (koło Kołomyi) 14.VII.1907.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [14.7.1907, Kolomyya]; 1 ♂, E21.01.01/03: “Słobódka leśna. 25. V.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”; 1 M, E21.01.01/04: “Wertełka. 26. V.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [25.5.?, Lisna Slobidka]; 1 ♀, E21.01.01/05: “Słbódka leśna. 25. V.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”; 1 ♂, E21.01.01/06: “Worochta. Okolice. 6.-6.-1908.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [6.6.1908, Lisna Slobidka]; 1 ♀, E21.01.01/07: “Subrawka (koło Kołomyi) 14.VII.1907.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [14.7.1907, Kolomyya]; 1 ♂, E21.01.01/09: “ Lwów. Park stryjski 6.-VII.-1906.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [6.7.1906, Lviv]; 1 ♂, E21.01.01/10: “Korzelice 17. V.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [17.5.?, Korelychi]; 1 ♂, E21.01.01/12: “Zubra.—7. VI.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [7.6.?, Zubra near Lviv]; 1 ♂, E21.01.01/13: “Słobódka leśna 25. V.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [25.5.?, Lisna Slobidka]; 1 ♂, E21.01.01/14: “ 17.-6.-1907. Worochta.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [17.6.1907, Vorokhta]; 1 ♀, E21.01.01/15: “Janów. 14.VII.”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [14.7.?, Ivaniv]; 1 ♂, E21.01.01/16: “Galicja”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [?, Galicja Wschodnia]; 1 ♂, E21.01.01/17: “Hołosko”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK” [?, Holosko]. Other two specimens from SMNH were collected in current territory of Poland labelled as follow: 1 ♀, E21.01.01/08: “Tatry”; “ Panorpa (Aulops) alpina Rambur, 1842 det.WK”, [Tatry Mountains?,?]; 1 ♂, E21.01.01/11: “Tatry”; “ Panorpa (Aulops) alpina Rambur, 1842 det. WK”, [Tatry Mountains?,?]. Reviewed material: Material identified as Panorpa pseudoalpina in SMNH collection: 1 ♂, E21.01.01.06/01: “ 17.-6.-1907. Worochta.”; “ Panorpa (Aulops) pseudoalpina Nagler det. WK”, [17.6.1907, Vorokhta]; 1 ♂, E21.01.01.06/02: “Worochta. Okolice. 6.-6.-1908.”; “ Panorpa (Aulops) pseudoalpina Nagler det. WK”, [6.6.1908, Vorokhta]; 1 ♂, E21.01.01.06/03: “ 17.-6.-1907. Worochta.”; “ Panorpa (Aulops) pseudoalpina Nagler det. WK”, [17.6.1907, Vorokhta]. New material examined: 2 ♂, 1 ♀, Zakarpatska Region, Perechynskyi district, Voevodyn, Turya Polyana, 48°47’7.82”N 22°50’42.45”E, sweep netting, 27.05.2016; 1 ♂, ibid, Voevodyn, 48°46’22.3”N 22°52’30.8”E, sweep netting, 27.05.2016; 2 ♂, 2 ♀, Mukachivskyi district, 48°30’51.9”N 22°51’24.4”E, beer traps in the forest and in trees at the stream, 6.05.2018 – 20.05.2018; 1 ♀, ibid, near “ Pearl of the Carpathians ” resort, 48°31’25.8”N 22°52’06.7”E, beer traps in the forest and in trees at the stream, 6.05.2018 – 20.05.2018; 1 ♂, 5 ♀, ibid, near Latorytsya, 48°31’16.0”N 22°52’13.1”E, beer traps in the garden, by the stream, 6.05.2018 – 20.05.2018. Remarks: Majewski (1885) reported this species based on citations of previous papers published by Dziędzielewicz (1867, 1877, 1883), Nowicki (1864, 1865) and Wierzejski (1883). This species, as well as other representatives of the genus Panorpa (e.g. P. communis, P. cognata and P. germanica), was also reported from the Western Carpathians by Dziędzielewicz (1911). Mocsáry (1875: 177) recorded P. alpina from Humenné [orig. Homonna] and Vihorlat [current territory of Slovakia], close to the border of Ukraine., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on pages 467-468, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Rambur, M. P. (1842) Histoire Naturelle des Insectes, Nevropteres. Fain et Thunot, Paris, 534 pp.","Nowicki, M. (1864) Przyczynek do owadniczej fauny Galicyi. Drukarnia Uniwersytecka w Krakowie, Krakow, 87 pp.","Nowicki, M. (1865) Insecta Haliciae Musei Dzieduszyckiani. Typis Universitatis Jagellonicae, Cracoviae, 87 pp.","Dziedzielewicz, J. (1867) Wykaz owadow siatkoskrzydlych (Neuroptera). Sprawozdania Komisji Fizjograficznej c. k. Towarzystwa naukowego Krakowskiego. Materyaly do fizyografii Galicyi, Krakow, 1, 158 - 165.","Majewski, E. (1882) Systematyczny wykaz owadow Zylkoskrzydlych polskich u nas dotad znalezionych. Insecta Neuroptera Polonica. Ksiegarnia Gebethnera i Wolffa, Warszawa, 42 pp.","Majewski, E. (1885) Owady Zylkoskrzydle (Neuroptera Polonica). Systematyczny wykaz krajowych sieciarek i prasiatnic. Materyaly do fauny krajowej. Nakladem ksiegarni Teodora Paprockiego i S-ki, Warszawa, 1 - 2, 1 - 39.","Dziedzielewicz, J. (1883) Sieciwki (Neuroptera) zebrane w okolicach Koomyi i nad Dniestrem w r. 1882. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 17, 1 - 9.","Dziedzielewicz, J. (1891) Przeglad fauny krajowej owadow siatkoskrzydlych (Neuroptera, Pseudoneuroptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 26, 26 - 151.","Wierzejski, A. (1883) Dodatek do fauny sieciowek (Neuroptera). Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 17, 253 - 255.","Dziedzielewicz, J. (1905) Sieciarki (Neuroptera genuia) i Prasiatnice (Archiptera) zebrane w ciagu lat 1902 i 1903. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 38, 104 - 125.","Dziedzielewicz, J. (1918) Owady siatkoskrzydlowate ziem Polski (Insecta neuropteroidea Poloniae terrarum). Rozprawy i Wiadomo s ci z Muzeum im. Dzieduszyckich, Lwow, 4 (1 - 4), 38 - 40.","Farbotko, J. (1929 a) Przyczynek do znajomosci wojsilek polnocno-wschodniej Polski. Prace Towarzystwa przyjaciol nauk w Wilnie, 5 (17), 1 - 8.","Nagler, C. (1970) Specii noi de Panorpidae (Ord. Mecoptera). Revista Mezeelor, 7, 54 - 57.","Ward, P. H. (1979) Structural variation in the genitalia of the Panorpa alpina - complex (Mecoptera). Systematic Entomology, 4 (1), 71 - 79. https: // doi. org / 10.1111 / j. 1365 - 3113.1979. tb 00612. x","Brauer, F. M. & Low, F. (1857) Neuroptera austriaca. Carl Gerold's Sohn, Wien, 80 pp.","Nagler, C. (1968) Les panorpides de la collection du Mesee \" Grigore Antipa \" - Bucarest, Trav. Mus. Hist. Nat. \" Gr. Antipa \", 8 (2), 813 - 826.","Lauterbach, K. E. (1972) Zur Kenntnis der Skorpionsfliegen des Balkans (Mecoptera, Panorpidae). Deutsche Entomologische Zeitschrift, 19, 109 - 140. https: // doi. org / 10.1002 / mmnd. 19720190112","Willmann, R. (2013) Fauna Europaea: Mecoptera. Fauna Europaea version 2.6.2. Available from: http: // www. faunaeur. org (accessed 7 September 2020)","Dziedzielewicz, J. (1877) Wykaz Prasiatnic i Sieciarek na porzeczach Prutu po Kolomyje i Bystrzycy nadwornianskiej. Pamietnik Towarzystwa Tatrzanskiego, 2, 68 - 75.","Dziedzielewicz, J. (1911) Owady siatkoskrzydle (Neuropteroidea), zebrine w zachodnich Karpatach w roku 1909. Sprawozdania Komisji Fizjograficznej Akademii Umiejetnosci w Krakowie, 45, 39 - 44.","Mocsary, S. (1875) Adatok Zemplen es Ung megyek faunajahoz - Neuroptera. MTA Mathemathika es Termeszettudomanyi Kozlony, 176 - 177."]}

11. Panorpa vulgaris Imhoff & Labram, 1845 Published records: Uzhhorod and Mukachevo env. (Dvořák et al. 2019). New material examined: 1 ♀, Zakarpatska Region, Mukachivskyi district, near “Pearl of the Carpathians” resort, 48°31’25.8”N 22°52’06.7”E, beer traps in the forest and in trees at the stream, 6.05.2018 – 20.05.2018; 3 ♀, ibid, near Latorytsya, 48°31’16.0”N 22°52’13.1”E, beer traps in the garden, by the stream, 6.05.2018 – 20.05.2018; 2 ♂, 3 ♀, Zakarpatska Region, Uzhhorod, 48°39’22.1”N 22°19’50.2”E, beer traps in the garden, apple trees, 1.05.2018 – 31.05.2018; ibid, Uzhhorod, 48°39’22.4”N 22°19’52.2”E, beer traps in oak forest (10 meters at the road), 1.05.2018 – 31.05.2018 (1 ♂, 3 ♀), 20.08.2018 – 26.08.2018 (2 ♀); 1 ♀, Klyucharky, forest, 48°24’23.5”N 22°39’39.7”E, beer trap, 13.– 28.07.2020; 7 ♂, 12 ♀, Klyucharky, abandoned field, 48°24’23.159”N 22°39’5.414”E, beer trap, 13.– 28.07.2020; 3 ♂, 1 ♀, Klyucharky, abandoned field, 48°24’23.159”N 22°39’5.414”E, beer trap, 27.8.– 11.09.2020. Remarks: Penny & Byers (1979) listed this species as a junior synonym of P. communis. Nevertheless, Sauer & Hensle (1977) regarded P. vulgaris as valid, a conclusion widely accepted in recent literature, including the European checklist (Willmann 2013). Unfortunately, many of the records published before the 1980s under the name P. communis belong in fact to the species P. vulgaris. As was shown by Dvořák et al. (2019) and also here in the new material examined, P. vulgaris is probably common and widespread in Ukraine.Also, part of the material in the SMNH collections presented here as P. communis may belong to P. vulgaris., Published as part of Dvořák, Libor, Mariychuk, Ruslan, Manko, Peter, Oboňa, Jozef, Krzemiński, Wiesław & Godunko, Roman J., 2022, Present knowledge on Mecoptera of Ukraine, pp. 459-483 in Zootaxa 5141 (5) on page 472, DOI: 10.11646/zootaxa.5141.5.3, http://zenodo.org/record/6593148, {"references":["Imhoff, L. & Labram, J. D. (1845) Insekten der Schweiz; die vorzuglichsten Gattungen je durch eine Art bildlich dargestellt von J. D. Labram nach Anleitung und mit Text von Dr. L. Imhoff. Basel, 4 & 5. [unknown pagination]","Dvorak, L., Haenni, J. - P., Dvorakova, K., Kameneva, E. P., Mariychuk, R., Manko, P., Obona, J. & Korneyev, V. A. (2019) Some insects from beer traps in Westernmost Ukraine. Ukrainska Entomofaunistyka, 10 (2), 1 - 6. https: // dx. doi. org / 10.5281 / zenodo. 3584947","Penny, N. D. & Byers, G. W. (1979) A Check-List of the Mecoptera of the World. Acta Amazonica, 9 (2), 365 - 388. https: // doi. org / 10.1590 / 1809 - 43921979092365","Sauer, K. P. & Hensle, R. (1977) Reproduktive Isolation, okologische Sonderung und morphologische Differenz der Zwilling- sarten Panorpa communis L. und P. vulgaris IMHOFF und LABRAM (Insecta, Mecoptera). Eine vergleichend biologische und evolutionsokologische Studie. Zeitschrift fur zoologische Systematik und Evolutionsforschung, 1, 169 - 207. https: // doi. org / 10.1111 / j. 1439 - 0469.1977. tb 00537. x","Willmann, R. (2013) Fauna Europaea: Mecoptera. Fauna Europaea version 2.6.2. Available from: http: // www. faunaeur. org (accessed 7 September 2020)"]}

The first annotated checklist of Mecoptera of Ukraine is presented. Altogether, 11 species of scorpionflies are known with a confirmed or probable occurrence in Ukraine. This list is based on previously published data, as well as on recently collected materials and specimens housed in the collection of the State Museum of Natural History NAS Ukraine, Lviv. This collection belongs to the oldest one in Ukraine and comprises specimens collected by Józef Dziędzielewicz, the founder of mecopteran investigations in Eastern Europe in the 19th and 20th centuries. For each listed species, its distribution in Ukraine is given together with notes on the history of investigations, and synonymies of the respective taxa. Detailed information on the distribution of selected species from the Red Book of Ukraine is presented.

Thornburghiella montana Ježek, Oboňa & Manko sp. nov. (Figs 1���21) Description. Male. Head hardly as long as broad (Fig. 1), 1.2 times broader. Vertex conically a little inflated dorsally (Figs 1) with a cut top. Numerous setae alveoli are almost regularly spaced over the entire surface in spite of scar free areas above C-shaped compound eyes laterally. Eyes separated, interocular suture arcuate (Figs 1, 12), eye bridge formed by five facet rows, frontal marginal rows are reduced to four facets. Minimum distance between eyes corresponds roughly to six facet diameters; index of distance from tangential points of eye apices to minimum of frons 3.2. Setae alveoli of frontoclypeus arranged almost in a triangular centrally placed patch near the base of antennae, tapering to a dorsoventral stripe of hairs close below frontal suture (Figs 1, 12). Patagia cylindrical, bag-shaped, constricted and contracted in one third, bent, covered with microsetae, see Fig. 11. Antenna with 15 articles; scape club-shaped (Fig. 2), somewhat widened apically, 2.5 times as long as its maximum width, narrowed et base, 4.9 times as long as its minimum width. Pedicel pitcher-shaped, symmetrical. Flagellomere 1 (postpedicel) cylindrical, hardly as long as three following flagellomeres together (Fig. 2). Postpedicel with six conspicuous, strong bristles arranged in a row, sometimes is the longest distal bristle doubled (from the same insertion). Scape and pedicel with stiletto-shaped scales in contrast to needle-shaped macrosetae of flagellomeres. Flagellomeres 2���12 ovoid, with needle-shaped paired ascoids, a little bent, shorter than flagellomeres in which are inserted; apical flagellomere twice as long as the previous one including digital apiculus placed a little out of longitudinal axis (Fig. 13). Length ratio of maxillary palpus segments 1.0:1.2:1.5:2.1; apical segment annulated (Fig. 3). Terminal labial lobes (Fig. 14) with diverging rows of spines between them. Ratio of maximum length of cibarium (Fig. 4) to length of epipharynx 1.7:1. Thorax. Anepisternum setae patch is almost trapezoid, anepimeron with triangular setose patch (Fig. 15). Spiracles set low on mesothorax. Wings (Fig. 16) lanceolate, 3.3 mm in holotype, 2.9���3.4 mm in paratypes, rounded distally, a little expanded at the posterior margin. The ending of R 5 beyond the tip of wing. Wing membrane slightly infuscated between Sc, R 1 and C and ends of all veins are a little strengthened distally with dark spots. Following veins or their parts strengthened: Sc with conspicuously marked origin and end, R 1 in distal three quarters, R 2, R 5, basal field, cross vein m 1 ��� m 2, CuA 1 and CuA 2 (conspicuously basally). Radial fork complete, medial fork in a form of a cross vein, their position see on Fig. 16. Both forks and the ending of CuA 2 are in one line (almost central area of wing). Wing index 2.4. Knob of halteres globular, with three close sensory microsetae ventrally, a prolonged stem as usually developed (Fig. 5). Ratio of maximum length of halteres to their maximum width approximately 2.8:1. Ratios of lengths of femora, tibiae and first tarsal segments P 1 2.0:2.3:1.0, P 2 2.1:2.8:1.1, P 3 2.4:3.2:1.2. Paired tarsal claws of P 1 gradually tapering, bent distad (Fig. 6). Male genitalia. Ejaculatory apodeme almost straight, only inconspicuously bent proximally and contracted distally (Figs 8, 21), aedeagal complex with paired sclerotized boomerang-shaped ribs diverged laterally and converged caudally. The basis of distiphallus is braced by gonocoxal apodeme ��� a chitinized stripe with three prolonged lobes (arms) of different shape and length (triangular and pale-shaped) on both sides (Fig. 8). Gonocoxites almost hemisphaerical (Figs 8���10), gonostyli ovoid basally, with irregular margins, conspicuously scelrotized, distal parts V-shaped, forked in two protuberances: sickle-shaped thin longer arm and thicker shorter saw-shaped one with numerous teeth (Figs 18���19). Epandrium (Figs 7, 17) almost semicircular in dorsal view, hardly rectangular from lateral one, not bare, (see two divided areas of insertions of hairs distally), posterior margin conspicuously sclerotized, emarginate, with a deep cleft. Basal paired apertures conspicuous, crevice-shaped, connected. Ventral epandrial plate reduced (Fig. 17). Hypandrium narrow with a lobulus in the middle (Fig. 8). Epiproct inconspicuous, as a rounded fold, covered with microsetae and dark structures inside; hypoproct conspicuous, setose, tongueshaped, rounded apically from dorsal view (Figs 7, 17). Epandrial claspers (surstyli) strong, enlarged basally in contrast to the top, almost straight from dorsal view (Fig. 17), bent at about one-third from lateral view (Fig. 7). Tenacula are numerous (30���35), formed in longitudinal rows on inner sides of clasping lobes, apically frayed. Female. Unknown. Differential diagnosis. Thornburghiella montana sp. nov. resembles T. kovari Ježek, 1993 in body size, as well as wing venation. The new species have head vertex a little inflated dorsally (Fig. 1); frons with a dorsoventral stripe of hairs (Figs 1, 12); postpedicel not constricted subapically (Fig. 2); hypandrium narrow with a lobulus in the middle (Fig. 8); gonostyli with two protuberances. (Figs 8���10, 18, 19); aedeagal complex with paired sclerotized boomerang-shaped ribs diverged laterally and converged caudally (Figs 8, 20, 21). Thornburghiella kovari is readily distinguishable by vertex of head, conspicuously elevated dorsally; frons without dorsoventral stripe of hairs; postpedicel constricted subapically; hypandrium stripe-shaped of the same width; gonostyli with three quite different bizarre protuberances; aedeagal complex with two parallel almost spatula-shaped cut protuberances and inner two linear ribs diverged caudally by conspicuous sclerotized hooks protruded outline of distiphallus. Type material. Holotype male: Transcaucasia, Georgia, Mtskheta ��� Mtianeti region, above the village Snotskali, a tributary of the Snotskali river, 1900 m a.s.l., 42��35���49.0���N 44��38���26.0���E (Fig. 38), 5.vii.2019, by sweep netting, Manko leg. Slide with a dissected specimen, Cat. No. 34899, Inv. No. 25956 (NMPC). Paratypes of 15 males (slides, some specimens dissected): The same locality, method, collectors and date, Cat. No. 34900-34909, Inv. No. 25957-25966 (NMPC); Gveleti, a stream beneath small waterfall, 1630 m a.s.l., 42��42���08.4���N 44��37���09.7���E, 12.vii.2019, by sweep netting, Kov��cs, Mur��nyi and Vin��on leg., Cat. No. 34910- 34913, Inv. No. 25967-25970 (NMPC); Pansketi, the Snotskali River at its confluence with the Terek River, 1745 m a.s.l., 42��38���14.0���N 44��37���56.0���E, 12.vii.2019, by sweep netting, Kov��cs, Manko, Mur��nyi and Vin��on leg., Cat. No. 34914, Inv. No. 25971 (NMPC). 6. Tarsal claw of P 1, lateral view. 7. Epandrium and epandrial claspers, lateral view. 8. Aedeagal complex and gonopod, dorsal view. 9. Gonopod, lateral view. 10. Same, caudal view. 11. Patagium. [Scale: 1-5, 7-11 = 0.2 mm; 6 = 0.05 mm] Type locality. Georgia, Mtskheta ��� Mtianeti region, Snotskali. Etymology. The specific epithet is derived from the Latin word ���montanus ��� a ��� um��� (adjective) = montane (mountain); it refers to the high elevation of the studied habitats of this species. Bionomics. Unknown, males were collected near montane waterfalls and streams or confluences of rivers, 1630���1900 m a.s.l. Distribution. Currently recorded only from Georgia., Published as part of Je��ek, Jan, Obo��a, Jozef & Manko, Peter, 2021, Two new Palaearctic species of moth flies (Diptera, Psychodidae, Psychodinae) from the Caucasus Mts., pp. 582-594 in Zootaxa 4985 (4) on pages 583-587, DOI: 10.11646/zootaxa.4985.4.11, http://zenodo.org/record/4964044, {"references":["Jezek, J. (1993) Thornburghiella kovari sp. n. (Diptera, Psychodidae, Psychodinae) from Tajikistan. Casopis Narodniho Muzea, Rada Prirodovedna, 161 (1992), 32 - 34."]}

Thornburghiella montana Ježek, Oboňa & Manko sp. nov. (Figs 1–21) Description. Male. Head hardly as long as broad (Fig. 1), 1.2 times broader. Vertex conically a little inflated dorsally (Figs 1) with a cut top. Numerous setae alveoli are almost regularly spaced over the entire surface in spite of scar free areas above C-shaped compound eyes laterally. Eyes separated, interocular suture arcuate (Figs 1, 12), eye bridge formed by five facet rows, frontal marginal rows are reduced to four facets. Minimum distance between eyes corresponds roughly to six facet diameters; index of distance from tangential points of eye apices to minimum of frons 3.2. Setae alveoli of frontoclypeus arranged almost in a triangular centrally placed patch near the base of antennae, tapering to a dorsoventral stripe of hairs close below frontal suture (Figs 1, 12). Patagia cylindrical, bag-shaped, constricted and contracted in one third, bent, covered with microsetae, see Fig. 11. Antenna with 15 articles; scape club-shaped (Fig. 2), somewhat widened apically, 2.5 times as long as its maximum width, narrowed et base, 4.9 times as long as its minimum width. Pedicel pitcher-shaped, symmetrical. Flagellomere 1 (postpedicel) cylindrical, hardly as long as three following flagellomeres together (Fig. 2). Postpedicel with six conspicuous, strong bristles arranged in a row, sometimes is the longest distal bristle doubled (from the same insertion). Scape and pedicel with stiletto-shaped scales in contrast to needle-shaped macrosetae of flagellomeres. Flagellomeres 2–12 ovoid, with needle-shaped paired ascoids, a little bent, shorter than flagellomeres in which are inserted; apical flagellomere twice as long as the previous one including digital apiculus placed a little out of longitudinal axis (Fig. 13). Length ratio of maxillary palpus segments 1.0:1.2:1.5:2.1; apical segment annulated (Fig. 3). Terminal labial lobes (Fig. 14) with diverging rows of spines between them. Ratio of maximum length of cibarium (Fig. 4) to length of epipharynx 1.7:1. Thorax. Anepisternum setae patch is almost trapezoid, anepimeron with triangular setose patch (Fig. 15). Spiracles set low on mesothorax. Wings (Fig. 16) lanceolate, 3.3 mm in holotype, 2.9–3.4 mm in paratypes, rounded distally, a little expanded at the posterior margin. The ending of R 5 beyond the tip of wing. Wing membrane slightly infuscated between Sc, R 1 and C and ends of all veins are a little strengthened distally with dark spots. Following veins or their parts strengthened: Sc with conspicuously marked origin and end, R 1 in distal three quarters, R 2, R 5, basal field, cross vein m 1 – m 2, CuA 1 and CuA 2 (conspicuously basally). Radial fork complete, medial fork in a form of a cross vein, their position see on Fig. 16. Both forks and the ending of CuA 2 are in one line (almost central area of wing). Wing index 2.4. Knob of halteres globular, with three close sensory microsetae ventrally, a prolonged stem as usually developed (Fig. 5). Ratio of maximum length of halteres to their maximum width approximately 2.8:1. Ratios of lengths of femora, tibiae and first tarsal segments P 1 2.0:2.3:1.0, P 2 2.1:2.8:1.1, P 3 2.4:3.2:1.2. Paired tarsal claws of P 1 gradually tapering, bent distad (Fig. 6). Male genitalia. Ejaculatory apodeme almost straight, only inconspicuously bent proximally and contracted distally (Figs 8, 21), aedeagal complex with paired sclerotized boomerang-shaped ribs diverged laterally and converged caudally. The basis of distiphallus is braced by gonocoxal apodeme – a chitinized stripe with three prolonged lobes (arms) of different shape and length (triangular and pale-shaped) on both sides (Fig. 8). Gonocoxites almost hemisphaerical (Figs 8–10), gonostyli ovoid basally, with irregular margins, conspicuously scelrotized, distal parts V-shaped, forked in two protuberances: sickle-shaped thin longer arm and thicker shorter saw-shaped one with numerous teeth (Figs 18–19). Epandrium (Figs 7, 17) almost semicircular in dorsal view, hardly rectangular from lateral one, not bare, (see two divided areas of insertions of hairs distally), posterior margin conspicuously sclerotized, emarginate, with a deep cleft. Basal paired apertures conspicuous, crevice-shaped, connected. Ventral epandrial plate reduced (Fig. 17). Hypandrium narrow with a lobulus in the middle (Fig. 8). Epiproct inconspicuous, as a rounded fold, covered with microsetae and dark structures inside; hypoproct conspicuous, setose, tongueshaped, rounded apically from dorsal view (Figs 7, 17). Epandrial claspers (surstyli) strong, enlarged basally in contrast to the top, almost straight from dorsal view (Fig. 17), bent at about one-third from lateral view (Fig. 7). Tenacula are numerous (30–35), formed in longitudinal rows on inner sides of clasping lobes, apically frayed. Female. Unknown. Differential diagnosis. Thornburghiella montana sp. nov. resembles T. kovari Ježek, 1993 in body size, as well as wing venation. The new species have head vertex a little inflated dorsally (Fig. 1); frons with a dorsoventral stripe of hairs (Figs 1, 12); postpedicel not constricted subapically (Fig. 2); hypandrium narrow with a lobulus in the middle (Fig. 8); gonostyli with two protuberances. (Figs 8–10, 18, 19); aedeagal complex with paired sclerotized boomerang-shaped ribs diverged laterally and converged caudally (Figs 8, 20, 21). Thornburghiella kovari is readily distinguishable by vertex of head, conspicuously elevated dorsally; frons without dorsoventral stripe of hairs; postpedicel constricted subapically; hypandrium stripe-shaped of the same width; gonostyli with three quite different bizarre protuberances; aedeagal complex with two parallel almost spatula-shaped cut protuberances and inner two linear ribs diverged caudally by conspicuous sclerotized hooks protruded outline of distiphallus. Type material. Holotype male: Transcaucasia, Georgia, Mtskheta – Mtianeti region, above the village Snotskali, a tributary of the Snotskali river, 1900 m a.s.l., 42°35’49.0”N 44°38’26.0”E (Fig. 38), 5.vii.2019, by sweep netting, Manko leg. Slide with a dissected specimen, Cat. No. 34899, Inv. No. 25956 (NMPC). Paratypes of 15 males (slides, some specimens dissected): The same locality, method, collectors and date, Cat. No. 34900-34909, Inv. No. 25957-25966 (NMPC); Gveleti, a stream beneath small waterfall, 1630 m a.s.l., 42°42’08.4”N 44°37’09.7”E, 12.vii.2019, by sweep netting, Kovács, Murányi and Vinçon leg., Cat. No. 34910- 34913, Inv. No. 25967-25970 (NMPC); Pansketi, the Snotskali River at its confluence with the Terek River, 1745 m a.s.l., 42°38’14.0”N 44°37’56.0”E, 12.vii.2019, by sweep netting, Kovács, Manko, Murányi and Vinçon leg., Cat. No. 34914, Inv. No. 25971 (NMPC). 6. Tarsal claw of P 1, lateral view. 7. Epandrium and epandrial claspers, lateral view. 8. Aedeagal complex and gonopod, dorsal view. 9. Gonopod, lateral view. 10. Same, caudal view. 11. Patagium. [Scale: 1-5, 7-11 = 0.2 mm; 6 = 0.05 mm] Type locality. Georgia, Mtskheta – Mtianeti region, Snotskali. Etymology. The specific epithet is derived from the Latin word “montanus – a – um“ (adjective) = montane (mountain); it refers to the high elevation of the studied habitats of this species. Bionomics. Unknown, males were collected near montane waterfalls and streams or confluences of rivers, 1630–1900 m a.s.l. Distribution. Currently recorded only from Georgia.

Genus Thornburghiella Vaillant, 1982 Thornburghiella Vaillant, 1973: 361 (unavailable name, type species not designated). Thornburghiella Vaillant, 1982: 299; type species: Psychoda albitarsis Banks, 1895 (by subseq. des.). Ulomyia sensu Krek 1999: 286, partim, nec Walker, 1856; type species: Psychoda fuliginosa Meigen, 1818: 107, by monotypy. Sijaricia as subgenus of Ulomyia sensu Krek 1999: 281; type species Pericoma erinacea Krek, 1970 (by monotypy) = syn. nov.: as Pericoma erinacea as well Krek 1970a: 87; 1970b: 100; 1974: tab. 7, sp. 29. A new suggestion: Thornburghiella erinacea (Krek, 1970) comb. nov. Thornburghiella – Vaillant 1983: 324. — Ježek 1992: 367; 1993: 32; 1994: 69; 2001: 64. — Wagner 1994: 75; 2003: 107. — Wagner & Joost 1985: 171. Ulomyia sensu Krek 1999 – Krek et al. 1976: 31 (U. erinacea, see below). Thornburghiella (Stupkaiella Vaillant, 1973, as subgenus), partim – Duckhouse 1987b: 85., Published as part of Ježek, Jan, Oboňa, Jozef & Manko, Peter, 2022, New synonymy and two new species of Caucasian moth flies (Diptera, Psychodidae, Psychodinae) from Azerbaijan and Georgia, pp. 48-63 in European Journal of Taxonomy 823 (1) on page 54, DOI: 10.5852/ejt.2022.823.1813, http://zenodo.org/record/6628047, {"references":["Vaillant F. 1982. Some comments on the classification of the Psychodidae Psychodinae (Diptera). Bulletin de la Societe entomologique de France 87: 292 - 301.","Vaillant F. 1973. Some new Psychodidae Psychodinae from the United States (Diptera). Annales de la Societe entomologique de France 9 (2): 345 - 379.","Krek S. 1999. Psychodidae (Diptera Insecta) Balkanskog Poluotoka [Psychodidae (Diptera Insecta) of the Balkan Peninsula]. Federacija Bosne i Hercegovine, Ministarstvo Obrazovanja, Nauke, Kulture i Sporta, Studentska Stamparija Univerziteta Sarajevo, Sarajevo. [In Croatian.]","Meigen J. W. 1818. Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Erster Teil. F. W. Forstmann, Aachen. https: // doi. org / 10.5962 / bhl. title. 12464","Krek S. 1970 a. Description de cinq especes nouvelles de Psychodidae de Yougoslavie. Bolletino della Associazione Romana di Entomologia 25 (4): 87 - 95.","Krek S. 1970 b. Fauna Psychodidae (Diptera) u podrucju planina Maglic, Volujak i Zelengora [Fauna of Psychodidae (Diptera) of the mountains Maglic, Volujak and Zelengora]. Glasnik Zemaljskog muzeja Bosne i Hercegovine u Sarajevu 9: 93 - 106. [In Croatian.]","Krek S. 1974. Ekoloska klasifikacija i cenoticki odnosi Psychodidae u tekucicama jugoistocne Bosne [Ecological classification and coenotic relationships of Psychodidae in running waters of southeastern Bosnia]. Godisnjak Bioloskog Instituta Univerziteta u Sarajevu (1973) 26: 57 - 95. [In Croatian.]","Jezek J. 1992. New and little known Palaearctic Thornburghiella species (Diptera, Psychodidae). Acta Entomologica Bohemoslovaca 89 (5): 367 - 382.","Jezek J. 1993. Thornburghiella kovari sp. n. (Diptera, Psychodidae, Psychodinae) from Tajikistan. Casopis Narodniho Muzea, Rada Prirodovedna 161 (1992): 32 - 34.","Jezek J. 1994. Three new species of moth flies (Diptera, Psychodidae) from Asia. Dipterologica Bohemoslovaca 6: 1 - 126.","Jezek J. 2001. New Palaearctic taxa of moth flies (Diptera: Psychodidae) from very small accidental spirituous samples of insects. Acta Universitatis Carolinae Biologica 45 (1 - 2): 53 - 66.","Wagner R. 1994. On a collection of Psychodidae (Diptera) from the Far East of Russia. Studia Dipterologica 1: 75 - 92.","Wagner R. 2003. Psychodiden aus Mittelasien und angrenzenden Gebieten (Insecta, Diptera, Psychodidae). Beitrage zur Entomologie 53 (1): 107 - 121. https: // doi. org / 10.21248 / contrib. entomol. 53.1.107 - 121","Wagner R. & Joost W. 1985. Bemerkungen zur Psychodiden-Fauna der Mongolischen Volksrepublik (Insecta, Diptera). Entomologische Nachrichten und Berichte 29 (4): 171 - 174.","Krek S., Kacanski D. & Tanasijevic M. 1976. Biocenoloska analiza naselja insekata (Ephemeroptera, Plecoptera, Simuliidae i Psychodidae) sliva rijeke Sutjeske [Biocoenological analysis of insects (Ephemeroptera, Plecoptera, Simuliidae and Psychodidae) of the basin of Sutjeska River]. Godisnjak Bioloskog Instituta Univerziteta u Sarajevu 29: 23 - 54. [In Croatian.]","Duckhouse D. A. 1987 b. Observations on the Himalayan species of Thornburghiella (Diptera; Psychodidae). Aquatic Insects 9 (2): 85 - 92. https: // doi. org / 10.1080 / 01650428709361276"]}

Pericoma inopinata Ježek, Oboňa & Manko sp. nov. (Figs 22���37) Description. Male. Head roundish, as long as wide (Fig. 22), with prominent cut vertex. Eyes separated, compound, kidney or C-shaped laterally, upper apices of eyes narrower in contrast to lower ones. Posteriolateral margins of eyes with 2���4 insertions of supraocular bristles on both sides. Setae alveoli are almost regularly spaced over the medial surface in spite of lateral patches with sporadic insertions and gore-shaped scar free areas laterally above eyes. Interocular suture bow-shaped, eye bridge formed generally by four facet rows, frontal marginal rows are reduced to three facets (Fig. 30). Minimum distance between eyes corresponds roughly to 3.4 facet diameters; index of distance from tangential points of eye apices to minimum of frons 8.0. Setae alveoli of frontoclypeus arranged almost in rectangular, centrally placed patch near base of antennae, tapering abruptly to a dorsoventral very narrow stripe of hairs close below frontal suture (Figs 22, 30). Antennae (Figs 23, 31) of 16 articles. Scape cylindrical, as long as globular pedicel, but narrower, first flagellomeres spindle-shaped, last ones ovoid; terminal flagellomere with an excentrical digital apiculus. Ascoids of flagellomeres 2-13 needle-shaped, a little bent, paired, twice shorter than flagellomeres in which are inserted. At the labellum, as shown in Fig. 32, parallel lines of miniature spines between both lobes absent. Labellum bulbose, without conspicuous digital protuberances in between (Fig. 32). Length ratio of maxillary palps 1.0:1.6:1.6:2.9, segment 4 annulate (Fig. 24). Ratio of maximum length of cibarium to length of epipharynx 1.3:1 (Fig. 25), labrum pointed. Thorax. Anepisternum setae patch almost semicircular, with circular thoracic spiracle anteriorly; anepimeron approximately with trapezoid setose patch (Fig. 26). Spiracles narrowly separated from protuberant anterior region of anepisternite and set on low on mesothorax. No thoracic allurement organs. Wings (Fig. 34) lanceolate, a little expanded et the posterior margin, 2.0 mm in holotype, 1.9���2.1 mm, in paratypes, rounded distally, the ending of R 5 a little beyond the tip of wing. Wing membrane not infuscated. Following veins or their parts strengthened: Sc (conspicuously on both ends), R 1, R 5, CuA 1, CuA 2 (markedly in one quarter basally). Radial fork in contrast to medial one complete (Fig. 34). Both forks and the ending of CuA 2 are not in one line; medial fork positioned somewhat to basal wing cell. Wing index 2.7. Knob of halteres (Fig. 33) globular, slightly asymmetrical, tapering to a prolonged stem with a basal fold. Scales are narrow, lanceolate, in contrast to three sensory microsetae ventrally arranged in a vertical row. Ratio of maximum length of halteres (measured to the fold of stem) to their maximum width 2.3: 1. Ratios of lengths of femora, tibiae and first tarsal segments: P 1 1.9:2.0:1.0, P 2 2.0:2.4:1.1, P 3 2.1:2.7:1.2. Paired tarsal claws of P 1 only a little bent in their last third, pointed, haired basally. Male genitalia with hypandrium stripe-shaped of the same width (Fig. 29). Aedeagal complex with basiphallus (ejaculatory apodeme) almost straight and narrow in dorsal view (Fig. 29), inconspicuously swollen and rounded proximally and conspicuously inflated distally to bul-shaped form; apodeme is weakened in the middle (lateral view, Fig. 37), strangulated by two clefts in one third basally and bent in second third. Distiphallus consisting of two phallomeres forming a spatula with divergent and pointed long lateral V-shaped protuberances only a little shorter than two parameral sclerites pointed and touched distally. Parameral joints with a transverse slightly curved wave band-like sclerite proximally connected by two narrow stripes with a sclerotized circle, and distally with divergent bolt-shaped protuberances, 0.5 time as long as phallomeres. Gonocoxal condyles apparently fused with parameral sheath. Gonopods see Figs 28, 29. Gonocoxites are almost cylindrical, stout, as long as gonostyli, at first of the same width, then rapidly tapering to the conspicuously sclerotized prolonged tips (Fig. 29). Gonocoxites have two long setae on the inner side basally, hardly as long as phallomeres. Epandrium (Figs 35, 36) quadrate with two patches of hairs on both sides caudally and a deep notch distally. Central aperture elliptical, doubled, strangulated in the middle. Ventral plate was not observed, only perhaps elliptical remnants. Hypoproct is tongue-shaped, triangular; epiproct only as a fold with a conical inner basis, narrowed medially. Both parts haired. Epandrial claspers almost cylindrical, haired, straight (dorsal view, Fig. 35), a little widened at basis and bent (lateral view, Fig. 36), 1.3-times as long as epandrium, with 5 tenacula subapically; ends are not frayed. Female. Unknown. Differential diagnosis. Pericoma inopinata can hardly be distinguished at first sight from the similar species P. calcilega Feuerborn, 1923, P. pingarestica Vaillant, 1978, P. tonnoiri Vaillant, 1978; and P. vestita Vaillant & Withers, 1993 (a part of Vaillant���s group trifasciata ��� see Vaillant 1978, part 317, p. 214, calcilega subgroup of Vaillant & Withers 1993 and the key of Kvifte & Ivkovic 2018), with a distiphallus consisting of two phallomeres forming a spatula with divergent and pointed or rounded long lateral V-shaped protuberances. In the new species, the two parameral sclerites are not pointed; they are blunt and separated distally. The gonostyli are at first almost globular (swollen) and bent in almost right angle to very prolonged, thin, needle-shaped protuberances. P. inopinata has two parameral sclerites tapering and touched caudally in one point, and the gonostyli are, moreover, at first cylindrical at first and then rapidly tapering to the conspicuously sclerotized prolonged tips, almost straight or only a little arched (Fig. 29). Type material. Holotype male: Transcaucasia, Georgia, Mtskheta ��� Mtianeti region, Mejilaurni, forest and bushy springs, outlets, 1270 m a.s.l., 42��19���25.4���N 44��38���43.9���E, 13.vii.2019, by sweep netting, Kov��cs, Manko, Mur��nyi and Vin��on leg (Fig 39). Slide with a dissected specimen, Cat. No. 34878, Inv. No. 25935 (NMPC). Paratypes 22 males (slides): 20 males of the same locality, method, collectors and date, Cat. No. 34879-34898, Inv. No. 25936-25955 (NMPC); two males Azerbaijan, Qax district, L��kit, Mamırlı waterfall and springs, 600 m a.s.l., 41��29���34.0���N 46��51���32.1���E, 7.v.2019, by sweep netting, Oboňa and Manko leg. Slide with a dissected specimen, Cat. No. 34876-34877, Inv. No. 25933-25934 (NMPC). Type locality. Georgia, Mtskheta ��� Mtianeti region, Mejilaurni. Etymology. Inopinatus ��� a, ��� um from Lat. (adjective) = unexpected; refers to a scientific bombshell in spring areas of Transcaucasia. Bionomics. Unknown, males were collected near montane waterfalls, springs and outlets. Distribution. Currently recorded only from Azerbaijan and Georgia., Published as part of Je��ek, Jan, Obo��a, Jozef & Manko, Peter, 2021, Two new Palaearctic species of moth flies (Diptera, Psychodidae, Psychodinae) from the Caucasus Mts., pp. 582-594 in Zootaxa 4985 (4) on pages 587-592, DOI: 10.11646/zootaxa.4985.4.11, http://zenodo.org/record/4964044, {"references":["Feuerborn, H. J. (1923) Die Larven der Psychodiden oder Schmetterlingsmucken. Ein Beitrag zur Oekologie des \" Feuchten \". Verhandlungen Internationale Vereinigung fur Theoretische und Angewandte Limnologie, Kiel, Stuttgart, 1 (1922), 181 - 213. https: // doi. org / 10.1080 / 03680770.1923.11896457","Vaillant, F. & Withers, P. (1993) Quelques Pericoma du groupe trifasciata (Diptera: Psychodidae, Psychodinae). Mitteilungen der Schweizerischen entomologischen Gesellschaft, 66 (1 - 2), 99 - 112.","Kvifte, G. M. & Ivkovic, M. (2018) New species and records of the Pericoma trifasciata group from Croatia (Diptera: Psychodidae). Zootaxa, 4486 (1), 76 - 82. https: // doi. org / 10.11646 / zootaxa. 4486.1.5"]}

Genus Pericoma Walker, 1856 Type species: Trichoptera trifasciata Meigen, 1804 (by subseq. des.), Published as part of Je��ek, Jan, Obo��a, Jozef & Manko, Peter, 2021, Two new Palaearctic species of moth flies (Diptera, Psychodidae, Psychodinae) from the Caucasus Mts., pp. 582-594 in Zootaxa 4985 (4) on page 587, DOI: 10.11646/zootaxa.4985.4.11, http://zenodo.org/record/4964044, {"references":["Meigen, J. V. (1804) Klassifikazion und Beschreibung der europaischen zweiflugligen Insekten (Diptera L.). I. Braunschweig, K. Reichard, 152 pp."]}

Leuctra martynovi Zhiltzova, 1960 Ş��ki-Zaqatala region: 20: 1♀ (LMEE); 25: 1♀ (LMEE); 26: 1♂, 1 exuviae (HNHM); 33: 26♂ 29♀ (HNHM), 3♂ 4♀ (LMEE); 34: 1♀ (HNHM). Remarks. A moderately common species in the Greater Caucasus (Georgia, Krasnodar Krai, North Ossetia) and Lesser Caucasus (Armenia, Georgia), rare in Anatolia (Zhiltzova 2003, Darilmaz et al. 2016). It emerges in spring and summer, we found it at a few, fast mountain streams in the Azerbaijani Greater Caucasus. New for the fauna of Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 67, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Zhiltzova, L. A. (1960) Contribution a l'etude des Plecopteres du Caucase. 4. Nouvelles especes de la famille Leuctridae. Entomologicheskoe obozrenie, 39 (1), 156 - 171. [in Russian]","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1"]}

Protonemura sp. (aculeata?) Şəki-Zaqatala region: 33: 1♂ (HNHM) 1♂ (LMEE). Remarks. One of these two, relatively large sized males (the HNHM specimen) was found to be infected by nematode worms of the family Mermithidae (Figs. 15–17). The presence of mermithid worms in stoneflies were reported by Ravizza & Zwick (2006) from Italy and Germany, where they found several infected specimens belonging to Isoperla rivulorum (Pictet, 1841), Protonemura caprai (Aubert, 1954) and another but unidentifiable Protonemura species. Lesions caused by the nematode infection show remarkable similarity between the Azerbaijan specimen and those figured and described by Ravizza & Zwick (2006). These include the fusion of the ventral vesicle with the hypoproct, shortening and fusion of paraproct lobes, and the shortening of epiproct (Figs 11–14). The other specimen (kept in the LMEE), was found to be free of mermithids, and its terminalia shows no lesions but apparently it was conspecific with the infected specimen. They belong to the praecox group, and cannot convincingly be attributed to any of the described species. Since Zhiltzova (1988) reported high geographical variability of P. aculeata within the Caucasian region, we will consider our specimens as a probable variant of P. aculeata, rather than a new species. Needed are additional specimens in good condition, additional character states in the form of DNA sequences, and behavioural characters derived from bioacoustic studies., Published as part of Murányi, Dávid, Manko, Peter, Kovács, Tibor, Vinçon, Gilles, Žiak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Oboňa, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on pages 68-69, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Ravizza, C. & Zwick, P. (2006) First records of Mermithidae (Nematoda) parasitic in Plecoptera. Bollettino della Societa entomologica italiana, 138 (2), 89 - 95.","Pictet, F. J. (1841) Histoire naturelle des insectes Nevropteres. Famille des Perlides. Kessmann-Bailliere, Geneve-Paris, 423 pp.","Aubert, J. (1954) Note sur quelques Plecopteres de Piemont et de Ligurie avec la description de deux especes nouvelles. Bollettino della Societa Entomologica Italiana, 84, 107 - 113.","Zhiltzova, L. A. (1988) New and little known species of Protonemura (Plecoptera, Nemouridae) from the Caucasus. Aquatic Insects, 10 (4), 215 - 219. https: // doi. org / 10.1080 / 01650428809361332"]}

Leuctra furcatella Martynov, 1928 Ş��ki-Zaqatala region: 18: 1♂ 1♀ (HNHM), 2♂ 1♀ (LMEE); 19: 5♂ 4♀ (HNHM); 22: 3♂ 3♀ (HNHM), 1♀ (LMEE); 32: 1♂ 2♀ (HNHM), 1♀ (LMEE); 33: 13♂ 21♀, 1 exuviae (HNHM); 34: 4♂ 3♀ (HNHM), 1♂ (LMEE). Remarks. A common spring and summer species in the Greater Caucasus (Georgia, Krasnodar Krai, Karachay-Cherkessia, North Ossetia), Lesser Caucasus (Armenia, Georgia), the Pontic and central region of Anatolia and the Alborz of Iran (Aubert 1964, Zhiltzova 2003, Darilmaz et al. 2016). In the Azerbaijani Greater Caucasus, it was found mostly along forest brooks and streams, sometimes in huge numbers. New for the fauna of Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 67, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Martynov, A. V. (1928) Zur Kenntnis der Plecopteren des Kaukasus. I: Nemouridae und Leuctridae des Zentral-Kaukasus. Travaux de la Station Biologique du Caucase du Nord, 2 (2 - 3), 18 - 42.","Aubert, J. (1964) Plecopteres du nord de l'Iran. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 37 (1 - 2), 69 - 80.","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1"]}

Perla marginata (Panzer, 1799) It was reported by Kasymov (1965) from the River Kura, later also from Agri��ay Stream by ��ki (��ki-Zaqatala region) (Kasymov 1972). Its case is similar to P. abdominalis, and so it must be omitted from the checklist., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 62, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Panzer, G. W. F. (1799) Fauna Insectorum Germanicae initia. Nurnberg, Felssecker, pp. 1 - 71.","Kasymov, A. G. (1965) Gidrofauna Nizney Kury i Mingechaurskogo vodohranilisa. AN Azerb. SSR, Baku, 372 pp.","Kasymov, A. G. (1972) Presnovadnaya fauna Kavkaza. Elm, Baku, 288 pp."]}

Plesioperla sakartvella (Zhiltzova, 1956) Ş��ki-Zaqatala region: 32: 1♂ (LMEE). Remarks. A rare montane species emerging in spring and summer, hitherto known only from the Lesser Caucasus of Georgia, and the eastern half of the Pontus (Zhiltzova & Cherchesova 2003, Teslenko & Zhiltzova 2009, Darilmaz et al. 2016). The Azerbaijani specimen was found along a fast mountain stream. New for the fauna of Azerbaijan, and our specimen constitutes the first occurrence in the Greater Caucasus., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 75, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Zhiltzova, L. A. (1956) Contribution a l'etude des Plecopteres du Caucase. 1. Nouvelles especes de la faune des Plecopteres (Taeniopterygidae et Chloroperlidae) des montagnes Trialetzky. Entomologicheskoe obozrenie, 35 (3), 659 - 670. [in Russian]","Zhiltzova, L. A. & Cherchesova, S. K (2003) Description of the larva of genus Perla Geoff. (Plecoptera, Perlidae) from the Caucasus. Entomologicheskoe obozrenie, 82 (2), 321 - 326. [in Russian]","Teslenko, V. A. & Zhiltzova, L. A. (2009) Key to the stoneflies (Insecta, Plecoptera) of Russia and adjacent countries. Imagines and nymphs (in Russian). Dalnauka, Vladivostok, 382 pp.","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1"]}

Nemoura irani Aubert, 1964 G��nc��-Qazax region: 39: 1♂ (HNHM); 42: 2♂ 1♀ (MM); 43: 1♂ (SNM); Nax��ıvan : 36: 1♂ (HNHM). Remarks. This species was known only from the Alborz of Iran, where it is a widespread summer and autumnal emerging stonefly (Aubert 1964, Theischinger 1976). This peculiar species shares an isolated position within the genus Nemoura, together with N. persica Zwick, 1980 (also endemic to the Alborz) and N. monae Joost, 1977 (known from the Greater Caucasus (Krasnodar Krai) and the Lesser Caucasus (Georgia) ��� Zhiltzova 2003). In Azerbaijan, we collected the species both in the northeast (G��nc��-Qazax region) and the southwest (Nax��ıvan) of the Lesser Caucasus. It was found in single or few specimens at high elevation seeps covered by tall sedges and other herbs. New for the fauna of Azerbaijan, and the whole of the Caucasian region., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 73, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Aubert, J. (1964) Plecopteres du nord de l'Iran. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 37 (1 - 2), 69 - 80.","Theischinger, G. (1976) Steinfliegen (Plecoptera) aus Anatolien und Iran. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 27 (3 / 4), 105 - 114.","Zwick, P. (1980) Notes on Plecoptera (2) Nemoura persica sp. n. Aquatic Insects, 2 (3), 170. https: // doi. org / 10.1080 / 01650428009361023","Joost, W. (1977) Nemoura monae n. sp. - eine neue Steinfliege (Plecoptera, Nemouridae) aus dem Einzugsgebiet der Teberda (UdSSR, Westkaukasus). Entomologische Nachrichten, 21 (2), 27 - 31."]}

Isoperla sp. Nax��ıvan: 36: 3♀ (HNHM). Remarks. These females from the Lesser Caucasus probably belong to Isoperla armeniaca Zhiltzova, 1961, and not the above I. grammatica sensu lato. Their specific identity must be confirmed with the capture of males., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 75, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Zhiltzova, L. A. (1961) On the study of the fauna of Plecoptera of the Caucasus V. Plecoptera of Armenia (in Russian). Entomologicheskoe obozrenie, 40 (4), 872 - 880."]}

Protonemura aculeata Theischinger, 1976 Ş��ki-Zaqatala region: 20: 4♀ (HNHM); 25: 1♀ (HNHM), 1♂ 1♀ (LMEE); 34: 1♂ (HNHM). Remarks. A common species in the Greater Caucasus (Georgia, Krasnodar Krai, Karachay-Cherkessia, Kabardino-Balkaria, North Ossetia), Lesser Caucasus (Armenia, Georgia), the Pontic region of Anatolia and the Alborz of Iran (Theischinger 1976, Zhiltzova 1988, Vin��on & Zhiltzova 2004). It emerges in spring, and was reported to be highly variable in the whole of its range (Zhiltzova 1988, Vin��on & Zhiltzova 2004). The Azerbaijan specimen was found along fast, often cascading streams of the Greater Caucasus in May. New for the fauna of Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 67, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Theischinger, G. (1976) Steinfliegen (Plecoptera) aus Anatolien und Iran. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 27 (3 / 4), 105 - 114.","Zhiltzova, L. A. (1988) New and little known species of Protonemura (Plecoptera, Nemouridae) from the Caucasus. Aquatic Insects, 10 (4), 215 - 219. https: // doi. org / 10.1080 / 01650428809361332","Vincon, G. & Zhiltzova, L. A. (2004) The genus Protonemura in Turkey (Plecoptera, Nemouridae). Nouvelle Revue d'Entomologie, 21 (2), 171 - 195."]}

Nemoura sp. Lənkəran region: 2: 1♀ (HNHM); Gəncə-Qazax region: 38 : 1♀ (MM); 40: 2♀ (HNHM); 43: 1♀ (LMEE). Remarks. These female Nemoura specimens were collected without males, and their specific identity are uncertain. However, the females from Göygöl and Gədəbəy (sites 38, 40, 43) are probably N. irani, while the single female from Azərüd (site 2) may belong to N. irani, or N. caspica Aubert, 1964, another Alborz endemic., Published as part of Murányi, Dávid, Manko, Peter, Kovács, Tibor, Vinçon, Gilles, Žiak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Oboňa, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 73, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Aubert, J. (1964) Plecopteres du nord de l'Iran. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 37 (1 - 2), 69 - 80."]}

Leuctra klapperichi Mur��nyi, 2005 L��nk��ran region: 15: 7♂ (HNHM). Remarks. This species was hitherto known only from the holotype male, collected a half century before in the Alborz of Iran (Mur��nyi 2005). This second locality is in the Talysh Mts, forming the Azerbaijani section of the Alborz range (Hyrcanian forest ecoregion). The present specimens were collected at a single stream (Fig. 7), despite sampling efforts in several similar habitats of the region. We collected it together with more Leuctra fusca and a single L. collaris specimens. Unfortunately, no female was found, so the female of L. klapperichi remains undescribed. New for the fauna of Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 67, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Muranyi, D. (2005) Leuctra klapperichi, a new stonefly species (Plecoptera: Leuctridae) from Iran with geographic affinities of related species. Zootaxa, 1078 (1), 59 - 64. https: // doi. org / 10.11646 / zootaxa. 1078.1.5"]}

Brachyptera transcaucasica transcaucasica Zhiltzova, 1956 Ş��ki-Zaqatala region: 20: 2♀ (HNHM), 1♂ 2♀ (LMEE); 26: 1♀ exuviae (HNHM), 1♀ exuviae (LMEE); 33: 1♂ 1♀, 3♀ exuviae (HNHM), 1♂ (LMEE). Remarks. The species is known from the Caucasian region, Anatolia and the Aegean Isles (Darilmaz et al. 2016, Karaouzas et al. 2016), the nominal subspecies is from the Greater Caucasus (Georgia, Krasnodar Krai, Karachay-Cherkessia, Kabardino-Balkaria, North Ossetia), Lesser Caucasus (Armenia, Georgia), the Turkish Pontus and Central Anatolia (Zhiltzova 2003, Cherchesova et al. 2012, Cherchesova & Zhiltzova 2013, Darilmaz et al. 2016). A spring emerging, montane species, Azerbaijan specimens were found in low number along fast mountain brooks and a small mountain river. New country record for the fauna of Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 66, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Zhiltzova, L. A. (1956) Contribution a l'etude des Plecopteres du Caucase. 1. Nouvelles especes de la faune des Plecopteres (Taeniopterygidae et Chloroperlidae) des montagnes Trialetzky. Entomologicheskoe obozrenie, 35 (3), 659 - 670. [in Russian]","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1","Karaouzas, I., Andriopoulou, A., Kouvarda, T. & Muranyi, D. (2016) An annotated checklist of the Greek stonefly fauna (In- secta: Plecoptera). Zootaxa, 4111 (4), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 4111.4.1","Cherchesova, S. K., Shioloshvili, M. N., Yakimov, A. V., Nemno, E. V., Lvov, V. D. & Kovilyaeva, N. E. (2012) Stoneflies (Insecta, Plecoptera) of Kabarda-Balkarian Republic (The Central Caucasus). Illiesia, 8 (19), 174 - 181.","Cherchesova, S. K. & Zhiltzova, L. A. (2013) Opredelitel vesnyanok (Plecoptera) Kavkaza. RGAU-MI K. A. T, Moskva-Vladikavkaz, 113 pp."]}

Protonemura microstyla Martynov, 1928 Ş��ki-Zaqatala region: 33: 2♂ 2♀ (HNHM), 8♂ 2♀ (LMEE). Remarks. This species is distributed in the Greater Caucasus (Georgia, Krasnodar Krai, Karachay-Cherkessia, North Ossetia, Stavropol Krai), Lesser Caucasus (Armenia, Georgia), and the eastern Pontic of Anatolia (Zhiltzova 2003, Darilmaz et al. 2016). It emerges in spring and summer, the Azerbaijan specimens inhabiting a single high elevation stream of the Greater Caucasus. New for the fauna of Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 69, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Martynov, A. V. (1928) Zur Kenntnis der Plecopteren des Kaukasus. I: Nemouridae und Leuctridae des Zentral-Kaukasus. Travaux de la Station Biologique du Caucase du Nord, 2 (2 - 3), 18 - 42.","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1"]}

Protonemura brachystyla Zhiltzova, 1988 It was described from Georgia, later recorded also from Turkey (Vin��on & Zhiltzova 2004). In the monograph of the Euholognatha of the former Soviet Union, Zhiltzova (2003) reported its distribution as ��� Georgia and, probably, Azerbaijan���. However, no exact Azerbaijan location was provided., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 61, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Zhiltzova, L. A. (1988) New and little known species of Protonemura (Plecoptera, Nemouridae) from the Caucasus. Aquatic Insects, 10 (4), 215 - 219. https: // doi. org / 10.1080 / 01650428809361332","Vincon, G. & Zhiltzova, L. A. (2004) The genus Protonemura in Turkey (Plecoptera, Nemouridae). Nouvelle Revue d'Entomologie, 21 (2), 171 - 195."]}

Protonemura sp. AZE-1 (temporary name) Lənkəran region: 12: 5♀ adults, 1♂ 1♀ larvae (HNHM). Description of the female. Medium sized species, macropterous, forewing length 9.0– 10.8 mm. Head brown with indistinct pattern, antenna brown, palpi yellowish. Thorax brown, pronotum pale brown and trapezoid with slightly rounded corners, wider than long, rugosities dark brown and distinct. Cervical gills off white and moderately long, bear fine setation, without subterminal constriction, shorter than forecoxa. Legs pale brown, but outer edge of tibiae and all tarsi are dark; wing membranes pale brown to brown, venation brown. Pilosity short and indistinct. Female abdomen (Figs. 18–19): Terga 1–8 light brown, full membranous; tergum 9 weakly sclerotized in apical third, tergum 10 full sclerotized and brown with pale pattern; epiproct lightly sclerotized. Sterna 1–8 light brown, membranous with paired small, anterolateral sclerites lacking distal to sternum 3. Sternum 7 with weakly developed, pale pregenital plate that is rounded and slightly overhangs sternum 8 anterior portion. Subgenital plate moderately large, its width is about two thirds of the segment width; lateral sides strongly diverging, posterior edge widely rounded, medial portion weakly depressed; colour of the plate pale brown, with paired darker patches; posterior portion slightly rounded in lateral view. Vaginal lobes large, rounded and pale, stretching about halfway between subgenital plate lateral edge and segment edges. Sterna 9–10 brown, with mediolateral dark patches on sternum 9. Paraprocts brown, apex rounded; cerci short and simple. Inner genitalia: a wide, bulbous and weakly sclerotized structure originates between the subgenital plate and vaginal lobes, genital opening is free; the structure is darker on basolateral portions where attached to basal part of vaginal lobes; apex of the structure joins to the spermathecal ductus with membranous folds, bearing paired, dark drop-like sclerites. Description of the matured larva (Figs. 20, 22–27). Body relatively slender, body length without antennae and cerci 6.2–7.0 mm. General colour light brown, with distinct and characteristic dark dorsal pattern: head mostly pale but with transverse dark pattern connecting ocelli and M-line, and brown patches on occiput; antennae and palpi light brown; pronotum dark brown with indistinct rugosities but with six pale spots; meso-, metanotum and wing pads mostly pale with brown, linear patches; legs pale, apex of femora and base of tibiae darker; abdominal terga mostly pale, but with dark lateral sides, and apical terga with a medial dark patch increasing in size towards tergum 10; cerci pale. Ventral aspect of the body entirely pale. Setation sparse but dark and distinct. Legs moderately long, width of hind femora about one-third of their length; tarsi with shortened basitarsus, as long as one third of the length of metatarsus; claws large. The pronotum is rounded trapezoidal, distinctly wider than long, as wide as the head. Cervical gills long and slightly curved, without subterminal constriction, about as long as forecoxa. Wing pads more than twice as long as the corresponding segments. Abdomen relatively long, integument light matt brown, first 6 abdominal segments fully divided by pleura. Posterior margin of sternum 9 of the male larva short triangular, with nipple-like apex; sternum 8 of the female larva widely incised. Paraproct elongated triangular, with blunt apex in both sexes. Cerci moderately long, with 25–30 slightly clubbed segments; length of the 15th segment is equal to its width. Setation of the larva: Head, antennae and palpi with short setae, postocular setae stronger. Pronotum covered with short setae; marginal setae distinct and acute, row laterally discontinuous, corners have setae as long as 1/12th of pronotum width. Setae on meso- and metanotum with marginal setae slightly shorter than longest marginal setae on pronotum; wing pads with short, blunt setae. Legs with sparse but distinct setae, all tibiae bear sparse swimming hairs shorter than the tibia width. Longest acute setae of mid- and hind femora are about as long as one third of the femur width, those on the fore femur reach half of the femur width; not arranged in line but restricted to apical half. Tergal segments sparsely covered with short setae; posterior margin with row of 14–16 acute and erect setae, of various length, no paired setae can be distinguished; longest reaches less than half of segment length. Cercal segments bald besides the apical whorl; apical whorl of setae consists of 10–14 acute and pale setae, about of equal length on the given segment; setae on cercomeres 14–16 are about as long as segment length. Remarks. Despite that the terminalia of the female is not so distinctive, it does not agree with any of the species known from the Caucasian region, Anatolia, Iran or the Levant. It can be either a new species, or the unknown female and larva of the Alborz endemic Protonemura waliabadi Aubert, 1964, the sole regional Protonemura of which the female sex remained undescribed. The larva is distinctive by its exceptional, contrasting color pattern and sparse setation. However, larvae of most of the regional Protonemura remained unknown (Zhiltzova et al. 2012). The species probably has true autumnal emergence, since last instars but not yet pharate larvae were present by the end of September. The larvae were collected from vertical hygropetric habitats in the splash zone of a waterfall, a habitat rarely occupied by West Palaearctic stoneflies (Fig. 21)., Published as part of Murányi, Dávid, Manko, Peter, Kovács, Tibor, Vinçon, Gilles, Žiak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Oboňa, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on pages 70-73, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Aubert, J. (1964) Plecopteres du nord de l'Iran. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 37 (1 - 2), 69 - 80.","Zhiltzova, L. A., Cherchesova, S. K. & Dzhioeva, I. E. (2012) Description of the larva of the Caucasian species Protonemura microstyla Martynov (Plecoptera, Nemouridae). Illiesia, 8 (17), 158 - 161."]}

Perlodes sp. Kasymov (1972) reported this species from Yuxarı-Qarabağ region (Şuşa, Turşsu), probably as larvae., Published as part of Murányi, Dávid, Manko, Peter, Kovács, Tibor, Vinçon, Gilles, Žiak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Oboňa, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 63, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Kasymov, A. G. (1972) Presnovadnaya fauna Kavkaza. Elm, Baku, 288 pp."]}

Capnopsis schilleri archaica Zwick, 1984 The subspecies was described on the basis of specimens from Armenia, Azerbaijan, Georgia and Russia (Zwick 1984). Azerbaijani specimens include the holotype and 18 paratypes, collection data are enumerated as ��� Azerbeidshan, Kjurakčai, 15.3.1970, leg. A. Kownacki���. This locality probably refers on K��r��k��ay River, a tributary of the Kura River in G��yg��l district (G��nc��-Qazax region). The species is known from the whole West Palaearctic, while the subspecies is restricted to the Caucasus and North Anatolia (Darilmaz et al. 2016)., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 61, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Zwick, P. (1984) Geographische Rassen und Verbreitungsgeschichte von Capnopsis schilleri (Plecoptera, Capniidae). Deutsche Entomologische Zeitschrift, 31 (1 / 3), 1 - 7. https: // doi. org / 10.1002 / mmnd. 19840310102","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1"]}

Chloroperla grammatica Scopoli Kasymov (1972) reported a species under this name from localities in G��nc��-Qazax (G��nc����ay) and Yuxarı-Qarabağ regions (Şuşa). It is not clear what species was meant under this name. The work of Scopoli (1763) contains Phryganea grammatica, but Scopoli clearly references Poda (1761) in regards to this species, as the original description. The name Phryganea grammatica Poda, 1761 was stabilized with neotype designation as a member of the genus Isoperla Banks, 1906 (Rupprecht 1984). Kasymov (1972) included Isoperla grammatica (Poda, 1761) in his paper among Isoperlidae (sic!), while Chloroperla grammatica Scopoli is listed among Chloroperlidae. Since the specimens are lost, the identity of this species remains dubious and must be omitted from the checklist., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 63, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Kasymov, A. G. (1972) Presnovadnaya fauna Kavkaza. Elm, Baku, 288 pp.","Scopoli, J. A. (1763) Phrygaenea. In: Entomologia Carniolica exhibens insecta Carnioliae indigena et distributa in ordines, genera, species, varietates, methodo Linneana. Trattner, Vindobonae, pp. 265 - 270. https: // doi. org / 10.5962 / bhl. title. 119976","Poda von Neuhaus, N. (1761) Insecta Musei Graecensis, quae in ordines, genera et species juxta Systema Naturae Linnaei digessit. Graecii, Widmanstad, 127 pp.","Banks, N. (1906) On the perlid genus Chloroperla. Entomological News, Lancaster, 17 (5), 174 - 175.","Rupprecht, R. (1984) Isoperla grammatica Poda, 1761 - Beschreibung eines Neotypus (Plecoptera). Annales de Limnologie, 20 (1 - 2), 81 - 90. https: // doi. org / 10.1051 / limn / 1984026"]}

Perla pallida Gu��rin-M��neville, 1843 The species was described without exact locality, mentioned only as ���Caucasus��� (Gu��rin-M��neville 1843). Walker (1851) found its types in the Natural History Museum, London (NHM), though they were lacking from the latest catalog (Kimmins 1970) and, presumably lost during the 19th century. Klap��lek (1923) redescribed the species on the basis of specimens from Azerbaijan, Georgia and Turkey. Though some question the conspecificity of these specimens, they have become de facto neotypes on which the present concept of the species is based (e.g., Sivec & Stark 2002). Klap��lek���s Azerbaijan specimen was a male collected in G��nc�� (G��nc��-Qazax region, original labels: Elisabethop; Kolenati; caucasica / det. Kempny; pallida / Klap��lek) and still exists in the Natural History Museum of Wien (NHMW). One of us (DM) studied and compared Klap��lek���s specimens with our recent specimens from Georgia. Besides the NHMW specimen from the 19th century, Kasymov (1972) reported the species from several sites in G��nc��-Qazax (D��st��fur), K��lb��c��r-La��ın (K��lb��c��r, T��rt��r��ay River), L��nk��ran (Sahgadzhi), Ş��ki-Zaqatala (Katex��ay River) and Yuxarı-Qarabağ regions (Şuşa, Xank��ndi). Sivec & Stark (2002) considered P. pallida to be a species complex that is distributed in the Caucasus, Anatolia, the Balkans and the Carpathians., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 62, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Guerin-Meneville, F. E. (1843) Genre Perle. In: Guerin-Meneville, F. E. (Ed.), Iconographie du Regne Animal de G. Cuvier. Insectes, 1843, pp. 393 - 395.","Walker, T. (1851) Sub-order 2. Perlides. In: Gray, J. E. (Ed.), Catalogue of the specimens of Neuropterous Insects in the Collection of the British Museum. Part I. Phryganides-Perlides. Trustees, London, pp. 136 - 192.","Kimmins, D. E. (1970) A list of type specimens of Plecoptera and Megaloptera in the British Museum (Natural History). Bulletin of the British Museum (Natural History), 24 (8), 337 - 361. https: // doi. org / 10.5962 / bhl. part. 1524","Klapalek, F. (1923) Plecopteres II. Fam. Perlidae. Subfam. Perlinae, Subfam. Neoperlinae. Collections Zoologiques du Baron Edm. de Selys Longchamps, 4 (2), 1 - 193.","Sivec, I. & Stark, B. P. (2002) The species of Perla (Plecoptera: Perlidae): evidence from egg morphology. Scopolia, 49, 1 - 33.","Kasymov, A. G. (1972) Presnovadnaya fauna Kavkaza. Elm, Baku, 288 pp."]}

Protonemura bacurianica bacurianica Zhiltzova, 1957 Ş��ki-Zaqatala region: 20: 1♂ pharate, 2♀ larvae (HNHM); 21: 2♂ 1♀ (HNHM); 22: 4♀ (HNHM), 1♂ 2♀ (LMEE); 24: 1♂ (LMEE); 28: 1♂ (HNHM); 29: 1♂ 3♀, 1♂ 2♀ larvae (HNHM), 1♂ (LMEE); 34: 1♂ (HNHM), 2♂ 2♀ larvae (LMEE). Remarks. A late spring and summer species, it occurs in the Greater Caucasus (Georgia, North Ossetia), Lesser Caucasus (Armenia, Georgia), most of Anatolia, and the Alborz of Iran (Theischinger 1976, Zhiltzova 2003, Darilmaz et al. 2016). Most of the range is inhabited by the nominal subspecies, but in the Central Taurus by P. bacurianica adana Vin��on & Zhiltzova, 2004. In the Greater Caucasus of Azerbaijan, specimens were collected in moderate numbers along forest brooks of medium high elevations. Females from the Mamirli Waterfall (site 22) have a flat subgenital plate, reminiscent of P. bacurianica adana Vin��on & Zhiltzova, 2004. New for the fauna of Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on pages 69-70, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Zhiltzova, L. A. (1957) Contribution a l'etude des Plecopteres du Caucase. 2. Nouvelles especes de la famille Nemouridae dans la faune des Plecopteres des montagnes Trialetzky. Entomologicheskoe obozrenie, 36 (3), 659 - 670. [in Russian]","Theischinger, G. (1976) Steinfliegen (Plecoptera) aus Anatolien und Iran. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 27 (3 / 4), 105 - 114.","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1","Vincon, G. & Zhiltzova, L. A. (2004) The genus Protonemura in Turkey (Plecoptera, Nemouridae). Nouvelle Revue d'Entomologie, 21 (2), 171 - 195."]}

Perla kiritschenkoi Zhiltzova, 1961 L��nk��ran region: 4: 5♂ 2♀ larvae (LMEE); 8 : 2♂ 1♀ larvae (LMEE). Remarks. As mentioned above, this species was described on the basis of syntypes from Armenia and the Talysh Mts (L��nk��ran region). Though the larva of this species is not yet described (Zhiltzova & Cherchesova 2003, Teslenko & Zhiltzova 2009), we have larvae with matured eggs from Iran (Mur��nyi et al. in preparation). On the basis of their morphology, our Azerbaijan specimens are identical. Our specimens from the Talysh were found in small, fast forest streams., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 74, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Zhiltzova, L. A. (1961) On the study of the fauna of Plecoptera of the Caucasus V. Plecoptera of Armenia (in Russian). Entomologicheskoe obozrenie, 40 (4), 872 - 880.","Zhiltzova, L. A. & Cherchesova, S. K (2003) Description of the larva of genus Perla Geoff. (Plecoptera, Perlidae) from the Caucasus. Entomologicheskoe obozrenie, 82 (2), 321 - 326. [in Russian]","Teslenko, V. A. & Zhiltzova, L. A. (2009) Key to the stoneflies (Insecta, Plecoptera) of Russia and adjacent countries. Imagines and nymphs (in Russian). Dalnauka, Vladivostok, 382 pp."]}

Isoperla grammatica (Poda, 1761) sensu lato Ş��ki-Zaqatala region: 20: 1♂ 1♀, 6 larvae (HNHM), 2♂ (LMEE); 22: 1♂ 2♀ (LMEE); 23: 1♂ 8♀ (HNHM), 1♂ (LMEE); 24: 1 larva (HNHM), 1♂ (LMEE); 26: 1 exuviae (HNHM); 27: 1♀ (LMEE); 30: 1♀ (LMEE); 32: 3 exuviae (HNHM); 33: 1 larva (HNHM). Remarks. These Azerbaijan specimens probably belong to Isoperla caucasica. This species, together with I. pulchra, were considered as synonyms of I. bithynica by Zwick (1971). However, the identity of I. bithynica is questionable, since only female paratypes exist in the NHMW. The paratypes were recently studied by DM who found it unlikely they are conspecific with any of the known Caucasian species, being rather big and pale insects. The types of I. caucasica and I. pulchra are also lost (Zwick 1971). Their original description, and the numerous specimens we recently collected in both Georgian and Azerbaijani areas of the Greater Caucasus, suggest that these taxa belong to the I. grammatica species complex sensu Berth��lemy (1979). But the variability of the specimens is rather confusing, as already noted by Balinsky (1950) in the original description of I. caucasica and I. pulchra. None of the Caucasian specimens can be conspecific with I. grammatica sensu stricto, as defined by Rupprecht (1984) on the basis of neotype designation. Further research, especially molecular studies, are needed to clarify the species status of I. caucasica, I. pulchra and I. bithynica. Thus, we enumerate our specimens herein as I. grammatica sensu lato. Isoperla grammatica (probably also in a sensu lato approach) were reported from the whole of Anatolia, northern part of the Greater Caucasus, and the Alborz of Iran (Aubert 1964, Cherchesova & Zhiltzova 2013, Darilmaz et al. 2016). Isoperla caucasica (as I. bithynica) was reported from both the Greater and Lesser Caucasus, while I. pulchra had never been reported outside of the Greater Caucasus (Zhiltzova 1961, 1964b). Azerbaijan specimens were caught at diverse habitats of the Greater Caucasus, but mostly in medium-high elevations. These are the first exact data from Azerbaijan, after Kasymov (1996) noted occurrence of both I. grammatica and I. caucasica, as well as of I. pulchra in the country without locality., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 74, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Poda von Neuhaus, N. (1761) Insecta Musei Graecensis, quae in ordines, genera et species juxta Systema Naturae Linnaei digessit. Graecii, Widmanstad, 127 pp.","Zwick, P. (1971) Plecoptera aus Anatolien und benachbarten Gebieten. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 44 (3 - 4), 233 - 264.","Berthelemy, C. (1979) Mating call and taxonomy in Pyrenean Isoperla. Gewasser und Abwasser, 64, 71 - 72.","Balinsky, B. I. (1950) On the Plecoptera of the Caucasus. Transactions of the Royal Entomological Society of London, 101, 59 - 87. https: // doi. org / 10.1111 / j. 1365 - 2311.1950. tb 00375. x","Rupprecht, R. (1984) Isoperla grammatica Poda, 1761 - Beschreibung eines Neotypus (Plecoptera). Annales de Limnologie, 20 (1 - 2), 81 - 90. https: // doi. org / 10.1051 / limn / 1984026","Aubert, J. (1964) Plecopteres du nord de l'Iran. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 37 (1 - 2), 69 - 80.","Cherchesova, S. K. & Zhiltzova, L. A. (2013) Opredelitel vesnyanok (Plecoptera) Kavkaza. RGAU-MI K. A. T, Moskva-Vladikavkaz, 113 pp.","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1","Zhiltzova, L. A. (1961) On the study of the fauna of Plecoptera of the Caucasus V. Plecoptera of Armenia (in Russian). Entomologicheskoe obozrenie, 40 (4), 872 - 880.","Zhiltzova, L. A. (1964 b) Vesnyanki (Plecoptera) v faune vysokogorya Bolsogo Kavkaza v Gruzii. Fauna vysokogorya Bolsogo Kavkaza v predelah Gruzii, 1964, 35 - 48.","Kasymov, A. G. (1996) Otryad Vesnyanki - Plecoptera. In: Musaev, M. A., Aliev, S. V., Gadziev, D. V., Kasymov, A. G. & Mikailov, T. K. (Eds.), Zivotniy mir Azerbaydzana. Tom II. Til Chlenistonogie. pp. 115 - 116."]}

Leuctra hippopus Kempny, 1899 Ş��ki-Zaqatala region: 20: 5♂ 16♀ (HNHM), 7♂ 9♀ (LMEE); 22: 1♀ (HNHM), 1♂ 2♀ (LMEE); 25: 2♀ (HNHM); 29: 1♂ 4♀ (HNHM), 1♂ (LMEE); 33: 2♀ (HNHM), 2♂ 1♀ (LMEE); 34: 11♂ 7♀ (HNHM), 6♂ 4♀ (LMEE); Nax��ıvan : 36: 1♀ (HNHM). Remarks. A widespread and common species in the whole West Palaearctic, with wide ecological spectrum (Graf et al. 2009). In the Caucasian region, it is known from Armenia, Georgia, Kabardino-Balkaria, North Ossetia and Stavropol Krai (Zhiltzova 2003, Cherchesova & Zhiltzova 2013). A spring and early summer species, in Azerbaijan we found it mostly along forest streams, in moderate numbers. These specimens provide the first confirmed locations for this species in Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 66, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Kempny, P. (1899) Zur kenntniss der Plecopteren. II. Neue und ungenugend bekannte Leuctra- Arten. II. und III. Theile. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 49, 9 - 15, 269 - 278. https: // doi. org / 10.5962 / bhl. part. 24102","Cherchesova, S. K. & Zhiltzova, L. A. (2013) Opredelitel vesnyanok (Plecoptera) Kavkaza. RGAU-MI K. A. T, Moskva-Vladikavkaz, 113 pp."]}

Amphinemura trialetica Zhiltzova, 1957 Ş��ki-Zaqatala region: 34: 1♂ (HNHM). Remarks. This species is distributed in the Greater Caucasus (Georgia, Krasnodar Krai, Karachay-Cherkessia, Kabardino-Balkaria, North Ossetia), Lesser Caucasus (Armenia, Georgia), and the northern half of Anatolia (Cherchesova & Zhiltzova 2013, Darilmaz et al. 2016). It emerges in spring and the whole summer, the only Azerbaijan specimen was found along a brook of the Greater Caucasus in May. New for the fauna of Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 67, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Zhiltzova, L. A. (1957) Contribution a l'etude des Plecopteres du Caucase. 2. Nouvelles especes de la famille Nemouridae dans la faune des Plecopteres des montagnes Trialetzky. Entomologicheskoe obozrenie, 36 (3), 659 - 670. [in Russian]","Cherchesova, S. K. & Zhiltzova, L. A. (2013) Opredelitel vesnyanok (Plecoptera) Kavkaza. RGAU-MI K. A. T, Moskva-Vladikavkaz, 113 pp.","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1"]}

Paragnetina transoxanica (Klap��lek, 1921) Some of the syntypes of its junior synonym, Caucasoperla spinulifera Zhiltzova, 1967 were collected in Azerbaijan: one female in K��lb��c��r-La��ın (Zabux), and one female in Yuxarı-Qarabağ region (H��s��nqaya). Esera caucasica Nav��s, 1931, a nomen dubium, may also be a synonym of this species (Zhiltzova 1967), since it was collected at one of the original localities of C. spinulifera (Georgia, Borjomi). Its single specimen, the female holotype, should be in the Museum of Natural History Paris (MNHP) but seems to be lost (Aubert 1952, 1956). Paragnetina transoxanica was described from Kazakhstan, known from all the Caucasian countries, as well from Eastern Anatolia (Darilmaz et al. 2016)., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 62, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Klapalek, F. (1921) Plecopteres nouveaux. I. Sous-famille des Perlinae et Neoperlinae. Annales de la Societe Entomologique de Belgique, 61, 57 - 67.","Zhiltzova, L. A. (1967) New genus and three new species of stone-flies (Plecoptera) from Caucasus and Crimea (in Russian). Entomologicheskoe obozrenie, 46 (4), 850 - 857.","Navas, L. (1931) Insectos del Museo de Paris. 7. a Serie. Broteria, Serie Zoologica, 27 (3), 101 - 113.","Aubert, J. (1952) Plecopteres decrits par le R. P. L. Navas, S. J. 2. Note sur quelques types des Museums de Barcelone et de Paris. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 25 (3), 239 - 241.","Aubert, J. (1956) Plecopteres decrits par le R. P. L. Navas, S. J. 4. Liste des types actuellement connus. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 29 (4), 437 - 445.","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1"]}

Arcynopteryx dichroa (McLachlan, 1872) It was reported from the ��ki-Zaqatala region (Katex��ay River), under the name A. compacta (McLachlan, 1872) (Kasymov 1972). This is the only mention of the species from the Caucasian region, unreported by the relevant monographs (Cherchesova & Zhiltzova 2013, Teslenko & Zhiltzova 2009). Given its wide distribution in mountainous regions of the Holarctic, the presence of A. dichroa in the Caucasus is possible. However, the Azerbaijan specimen(s) has not been found, so its occurrence must be confirmed by new specimens., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 62, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["McLachlan, M. R. (1872) Materiaux pour une Faune Nevropterologique de l'Asie Septentrionale. Seconde partie. Non-Odonates. Annales de la Societe Entomologique de Belgique, 15, 47 - 77 + Pl. I - II.","Kasymov, A. G. (1972) Presnovadnaya fauna Kavkaza. Elm, Baku, 288 pp.","Cherchesova, S. K. & Zhiltzova, L. A. (2013) Opredelitel vesnyanok (Plecoptera) Kavkaza. RGAU-MI K. A. T, Moskva-Vladikavkaz, 113 pp.","Teslenko, V. A. & Zhiltzova, L. A. (2009) Key to the stoneflies (Insecta, Plecoptera) of Russia and adjacent countries. Imagines and nymphs (in Russian). Dalnauka, Vladivostok, 382 pp."]}

Leuctra fusca ssp. (Linnaeus, 1758) L��nk��ran region: 1: 9♂ 7♀ (HNHM); 2 : 4♂ 1♀ (HNHM); 4: 5♂ 4♀ (HNHM), 1♂ (LMEE); 5: 1♂ 2♀, 1 exuviae (HNHM); 7: 1♂ 4♀ (HNHM); 8: 4♂ 2♀ (HNHM); 9: 3♂ 1♀ (HNHM); 10: 3♂ 4♀ (HNHM), 1♂ 1♀ (LMEE); 12: 3♂, 6 larvae (HNHM); 13: 1♂ (HNHM), 1♀ (LMEE); 14: 6♂ 2♀ (HNHM); 15: 9♂ 6♀ (HNHM), 1♂ 2♀ (LMEE); G��nc��-Qazax region: 37: 1♂ (MM); 38: 2♂ 4♀ (HNHM), 2♂ (LMEE), 1♀ (MM); 40: 2♀ (HNHM), 2♂ 1♀ (MM); 41: 1♂ 1♀ (HNHM), 1♂ (LMEE); 42: 4♂ 4♀ (HNHM), 2♂ 1♀ (LMEE), 2♂ 3♀ (MM); 43: 1♀ (HNHM), 1♂ (SNM); 45: 1♂ 2♀ (HNHM), 1♂ (MM). Remarks. The species has a wide Eurosiberian distribution, with three described subspecies (Mur��nyi & Hwang 2017). It is an autumnal potamal species over most of its range, however, Caucasian populations were noted by their uncommonly wide range of habitats occupied, from rivers to high mountain brooks (Zhiltzova 2003). Winter occurrence in Adjaria, Georgia, was recently recorded (Teslenko et al. 2019). Indeed, Azerbaijani specimens were found in extremely different habitats during our sampling both in the Lesser Caucasus and the Talysh Mts (Figs. 7���10). It is the most common autumnal stonefly in Azerbaijan, it occurs both as the single stonefly species of the given waterflow, or with a diverse community. These are the first exact data from Azerbaijan. The subspecific identity of the Azerbaijan population is problematic. We expect L. fusca latior Berth��lemy & Dia, 1982 since this subspecies inhabits the whole of Anatolia, the Levant and Iran (Darilmaz et al. 2016, Mur��nyi 2005, Vin��on et al. 2013). However, females of L. fusca latior are distinguished by the widely separated posterior lobes of the subgenital plate, while all the Azerbaijani females we have examined have narrowly separated lobes and possess a small, triangular median lobule slightly reminiscent of L. kurui Kazanci, 1983. Coloration of the subgenital plate varies by age but the general shape of the subgenital plate and the spermathecal ring are constant among all of our female specimens (Figs. 5���6). The male paraproct and specillum more or less agree with the description of L. fusca latior (Figs. 3���4) but Azerbaijani specimens exhibit a highly reduced or entirely lacking posteromedial sclerite on tergum VIII and the posteromedial sclerite is entire or only slightly subdivided on tergum IX (Fig. 2). The anterior lobes of tergum X are moderately large, the ventral vesicle small, the epiproct is wider than long with a short, weakly separated rod. Both male and female specimens vary in size and coloration, but we found no correlation of these features with the slight variability of the genitalia., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 66, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Linnaeus, C. (1758) Systema naturae per regna tria natur, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Holmi, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542","Muranyi, D. & Hwang, J. M. (2017) Four new species and further contributions to the Leuctridae (Plecoptera) of the Korean Peninsula. Zootaxa, 4282 (1), 43 - 61. https: // doi. org / 10.11646 / zootaxa. 4282.1.2","Teslenko, V. A., Palatov, D. M. & Semenchenko, A. A. (2019) Description of new apterous winter species of Leuctra (Plecoptera: Leuctridae) based morphology and DNA barcoding and further records to stonefly fauna of the Caucasus, Georgia. Zootaxa, 4585 (3), 546 - 560. https: // doi. org / 10.11646 / zootaxa. 4585.3.9","Berthelemy, C. & Dia, A. (1982) Plecopteres du Liban (Insecta). Annales de Limnologie, 18 (2), 191 - 214. https: // doi. org / 10.1051 / limn / 1982009","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1","Muranyi, D. (2005) Leuctra klapperichi, a new stonefly species (Plecoptera: Leuctridae) from Iran with geographic affinities of related species. Zootaxa, 1078 (1), 59 - 64. https: // doi. org / 10.11646 / zootaxa. 1078.1.5","Vincon, G., Dia, A., Kovacs, T. & Muranyi, D. (2013) A new stonefly from Lebanon, Perlodes thomasi sp. n. (Plecoptera: Perlodidae). Illiesia, 9 (3), 18 - 27.","Kazanci, N. (1983) Plecoptera (Insecta) from Anatolia. Aquatic Insects, 5 (3), 141 - 146. https: // doi. org / 10.1080 / 01650428309361135"]}

Perlodes microcephalus (Pictet, 1833) Nax��ıvan: 35: 2♀ (HNHM). Remarks. A widespread and common species in the whole West Palaearctic; it has been reported from several localities in the Greater Caucasus, Lesser Caucasus, all of Anatolia and the Alborz of Iran (Aubert 1964, Teslenko & Zhiltzova 2009, Cherchesova & Zhiltzova 2013, Darilmaz et al. 2016). It emerges in spring and early summer. Our specimens occurred in high elevations of the Lesser Caucasus. New for the fauna of Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 74, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Pictet, F. J. (1833) Memoire sur les Metamorphoses des Perles. Annales des Sciences Naturelles, 28, 44 - 65. https: // doi. org / 10.5962 / bhl. part. 8008","Aubert, J. (1964) Plecopteres du nord de l'Iran. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 37 (1 - 2), 69 - 80.","Teslenko, V. A. & Zhiltzova, L. A. (2009) Key to the stoneflies (Insecta, Plecoptera) of Russia and adjacent countries. Imagines and nymphs (in Russian). Dalnauka, Vladivostok, 382 pp.","Cherchesova, S. K. & Zhiltzova, L. A. (2013) Opredelitel vesnyanok (Plecoptera) Kavkaza. RGAU-MI K. A. T, Moskva-Vladikavkaz, 113 pp.","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1"]}

Pontoperla teberdinica (Balinsky, 1950) Ş��ki-Zaqatala region: 21: 1♀ (HNHM); 26: 2♂ 9♀, 10 larvae, 6 exuviae (HNHM), 1♀, 4 exuviae (LMEE); 33: 4♂ 5♀, 1 larva (HNHM), 4♂ 3♀, 5 larvae (LMEE); 34: 1♀ (HNHM); Nax��ıvan : 36: 4♀ (HNHM). Remarks. This spring emerging species occurs in the Greater Caucasus (Karachay-Cherkessia), Lesser Caucasus (Georgia), all of Anatolia, and the Alborz of Iran (Zwick 1975, Teslenko & Zhiltzova 2009, Darilmaz et al. 2016). In the Greater and Lesser Caucasus of Azerbaijan, specimens were collected in moderate numbers along waterbodies ranging from open brooks to small rivers, at elevations above 1200 meters. These are the first exact location data from Azerbaijan., Published as part of Mur��nyi, D��vid, Manko, Peter, Kov��cs, Tibor, Vin��on, Gilles, ��iak, Matej, Kerimova, Ilhama G., Snegovaya, Nataly Yurievna & Obo��a, Jozef, 2021, Review and contribution to the stonefly (Insecta: Plecoptera) fauna of Azerbaijan, pp. 58-80 in Zootaxa 4975 (1) on page 75, DOI: 10.11646/zootaxa.4975.1.2, http://zenodo.org/record/4804613, {"references":["Balinsky, B. I. (1950) On the Plecoptera of the Caucasus. Transactions of the Royal Entomological Society of London, 101, 59 - 87. https: // doi. org / 10.1111 / j. 1365 - 2311.1950. tb 00375. x","Zwick, P. (1975) Weitere Plecoptera aus Anatolien. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 48 (3 - 4), 387 - 396.","Teslenko, V. A. & Zhiltzova, L. A. (2009) Key to the stoneflies (Insecta, Plecoptera) of Russia and adjacent countries. Imagines and nymphs (in Russian). Dalnauka, Vladivostok, 382 pp.","Darilmaz, M. C., Salur, A., Muranyi, D. & Vincon, G. (2016) Contribution to the knowledge of Turkish stoneflies with annotated catalogue (Insecta: Plecoptera). Zootaxa, 4074 (1), 1 - 74. https: // doi. org / 10.11646 / zootaxa. 4074.1.1"]}