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Indusiothrips seshadrii Priesner (Fig. 14) Until recently, this species was knOwn Only frOm specimens cOllected in sOuthern India (WilsOn 1975). HOwever, in 2016 it was cOllected On Oahu in the Hawaiian Islands (MOund et al. 2017), and the females listed belOw frOm sOuthern China represent the third published recOrd Of the species. Presumably it is widespread in Asia in assOciatiOn with the frOnds Of One Or mOre species Of ferns. The Only Other member Of the genus, nakaharai, is knOwn frOm members Of the fern family POlypOdiaceae in Japan. These twO species are small pale thrips with the dOrsal surface Of the head reticulate (Fig. 14). New recOrds: China, Jiangxi, Anfu COunty, Taishan TOwn, Mt.WugOngshan, fOur females frOm grass, 12.viii.2016 (ZhaOhOng Wang), in SCAU. Guizhou, Zunyi City, 4 females frOm Mallotus repandus, 16.viii.2016 (Yan XQ); Xingyi City, 2 female frOm Viburnum chinshanense, 10.ix.2017 (Li YJ), in YAU. Yunnan, Yuxi City, TOnghai COunty, Xiushan Park, 15 females, 1 male frOm Dryopteris sp., 22.v.2017 (Zhang HR & Li YJ), in YAU. Chongqing, Xiushan COunty, 7 females frOm fern, 9.viii.2017 (Li YJ), in YAU., Published as part of Zhang, Shimeng, Wang, Zhaohong, Li, Yajin & Mound, Laurence, 2018, One new species, two generic synonyms and eight new records of Thripidae from China (Thysanoptera), pp. 370-378 in Zootaxa 4418 (4) on page 373, DOI: 10.11646/zootaxa.4418.4.3, http://zenodo.org/record/1244922

Baliothrips sunae sp.n. (Figs 5���10) Female macroptera: BOdy, femOra and antennal segments I���II unifOrmly brOwn; tarsi yellOw, alsO distal half Of tibiae; antennal segments III���IV mainly yellOw, V���VII light brOwn; fOre wings banded, basal third including clavus pale, then a shOrt dark band, a slightly lOnger pale band, with wing apex dark. Antennae 7-segmented; segment I withOut paired dOrsO-apical setae; III and IV each with sense cOne shOrt and fOrked, sense cOne Y-shaped On III; apex Of V and base Of VI brOadly transverse (Fig. 10); III���VI with micrOtrichia On bOth surfaces. Head slightly lOnger than width acrOss cheeks with eyes large and swOllen (Fig. 5); lOngitudinal rOw Of tuberculate sculpture anteriOr tO first Ocellus, Ocellar triangle weakly sculptured, pOsteriOr third Of head transversely striate/ reticulate; Ocellar setae pair I absent, pair II wide apart lOng but curved, pair III On margin Of triangle; five pairs Of lOng pOstOcular setae, pair I lOngest, nOt arranged in a straight rOw; maxillary palps 2-segmented. PrOnOtum transverse, with nO discal setae Or sculpture medially; fOur pairs Of pOsterOmarginal setae, S1 lOnger than S2, but S3 and S4 (pOsterOangulars) lOngest (Fig. 8); One pair Of midlateral setae lOng and stOut; anteriOr margin with 4���5 pairs Of fine setae. MesOnOtum with nO sculpture Or campanifOrm sensilla On anteriOr third (Fig. 6); median pair Of setae far frOm pOsteriOr margin; mesOfurca with weak spinula. MetanOtum with irregular lOngitudinal striae, median setae far frOm anteriOr margin; metafurca withOut spinula. PrOsternal basantra membranOus, withOut setae; ferna rObust and cOntinuOus medially; prOspinasternum transverse and slender. Tarsi 2-segmented; hind tibiae with setae On distal half lOnger than width Of tibia. FOre wing first vein with 4���5 widely spaced setae near base, 2 widely spaced setae near apex; secOnd vein with abOut 8 widely spaced setae; clavus with 4 veinal and One discal setae. Tergite I cOvered with transverse irregular reticulate sculpture, median setae minute; II���VIII with nO sculpture medially between setae S1, withOut craspeda (Fig. 7); VI���VIII with paired ctenidia ending at discal setae S3; IV���V with weak ctenidia; pOsteriOr margin Of VIII with a few minute micrOtrichia On lateral thirds; IX with anteriOr campanifOrm sensilla absent (Fig. 7); MD setae stOut and extending beyOnd tergal margin, marginal setae lOng and extending beyOnd bOdy apex; X with median split almOst cOmplete, majOr setae nearly twice length Of segments. Sternites II���VII with transverse reticulatiOn (Fig. 9), II with twO pairs Of marginal setae, III���VII with three pairs; S1 On VII in frOnt Of pOsteriOr margin, but clOse tO Or On margin On II���VI. Measurements (hOlOtype female in micrOns). BOdy length 1600. Head, length 125 (nOt hOrizOntal); width acrOss cheeks 140; width acrOss eyes 175. PrOnOtum, length 130; width 210; pOsterOmarginal S1���S4 setae 35, 25, 60, 50. MetanOtum median setae 25. FOre wing length 900. Tergite IX, length 80; MD setae 55; S1 setae 100. Tergite X length 90; S1 setae 140. Male: NOt knOwn. Comments: This new species is referred tO the genus Baliothrips with sOme hesitatiOn, but the alternative Of placing each Of the three species invOlved intO a separate genus de-emphasises their apparent relatiOnships. In a key tO genera related tO Thrips that are assOciated with grasses, Bhatti and MOund (1980) prOvided fOur character states by which they diagnOsed Baliothrips. TwO Of these are fOund in this new species, but twO are absent. Marginal setae S1 On sternites III���VI are submarginal in bOth dispar (Haliday) and kroli (Schille), and in bOth Of them sternite II bears three pairs Of marginal setae Of which pair S1 is submarginal. Neither Of these character states Occurs in this new species, althOugh all three species share a similar cOnditiOn Of the fOre wing clavus with fOur veinal setae, and chaetOtaxy Of the hind tibiae with twO Or three unusually lOng setae. MOreOver, sunae is particularly similar tO dispar, the type species Of Baliothrips, in the 2-segmented maxillary palps, the shape and chaetOtaxy Of the head, the metanOtal sculpture, and many Other details including the banded fOre wings. The new species differs frOm the previOus twO species in the genus in the unifOrmly pale cOlOur Of the wing base and clavus, the absence Of campanifOrm sensilla On the metanOtum and first abdOminal tergite, and the presence Of a weak mesOsternal spinula. Material studied. HOlOtype female, CHINA, Zhejiang, ShuangXikOu, frOm grasses, 27.v.2017 (Niu MM), in NWAFU. Paratypes, 4 females cOllected with hOlOtype, twO in NWAFU; One in ANIC; One in BMNH., Published as part of Zhang, Shimeng, Wang, Zhaohong, Li, Yajin & Mound, Laurence, 2018, One new species, two generic synonyms and eight new records of Thripidae from China (Thysanoptera), pp. 370-378 in Zootaxa 4418 (4) on pages 371-372, DOI: 10.11646/zootaxa.4418.4.3, http://zenodo.org/record/1244922, {"references":["Bhatti, J. S. & Mound, L. A. (1980) The genera of grass and cereal-feeding Thysanoptera related to the genus Thrips (Thysanoptera: Thripidae). Bulletin of Entomology, 21, 1 - 22."]}

Anaphothrips incertus (Girault) (Figs 1���2) Described frOm Australia in the genus Limothrips, this grass-living species has previOusly been knOwn Only frOm Australia where it is widespread in the warmer eastern parts Of that cOntinent (Pitkin 1978). It is here recOrded frOm China fOr the first time frOm the specimens listed belOw. A full descriptiOn Of the species was prOvided by MOund and MasumOtO (2009). AlthOugh similar in general appearance tO A. populi Zhang & TOng, with which it shares the light brOwn cOlOur Of bOdy and fOre wings, a lOng cOmb On tergite VIII, and similar bOdy sculpture, it is distinguished by the 8-segmented antennae, Ocellar setae pair III mOre widely separated (Fig. 1), tergite IX distinctly lOnger, and antennal segments III and IV variably yellOwish. The male listed belOw frOm Yunnan has the strOngly C-shaped sternal pOre plates (Fig. 2) that are typical Of sO many species in this genus, and tergite IX bears a pair Of shOrt stOut thOrn-like setae. The twO females frOm Tibet are the first knOwn micrOpterae Of this species. The fOre wings are shOrter than the thOrax width, the Ocelli are absent, and the fOur pOstOcular setae are in a transverse rOw, but in Other character states it is similar tO macrOpterae frOm Australia. New records: CHINA, Yunnan, Shi Lin COunty, 4 females 5 males frOm grasses, 10.ix.2015 (Li YJ & Yang CW), in YAU and ANIC; Puer City, Jinggu COunty, 13 females frOm Phragmites communis, 9.viii.2015 (Li YJ), in YAU. Guizhou, Xingyi City, 3 females frOm Themeda caudata, 6.xi.2017 (KOng B & Wu C), in YAU. Guangxi, Langping, 2 females frOm bambOO, 25.vii.2014 (Man Yue), in NWAFU. Tibet, MedOg COunty, 2 micrOpterOus females frOm grass, 15.vii.2016 (Niu MM), in NWAFU., Published as part of Zhang, Shimeng, Wang, Zhaohong, Li, Yajin & Mound, Laurence, 2018, One new species, two generic synonyms and eight new records of Thripidae from China (Thysanoptera), pp. 370-378 in Zootaxa 4418 (4) on page 371, DOI: 10.11646/zootaxa.4418.4.3, http://zenodo.org/record/1244922, {"references":["Mound, L. A. & Masumoto, M. (2009) Australian Thripinae of the Anaphothrips genus-group (Thysanoptera), with three new genera and thirty-three new species. Zootaxa, 2042, 1 - 76."]}

Dictyothrips betae Uzel (Figs 3���4) Described frOm central EurOpe, and recOrded widely frOm nOrthern Italy tO Denmark and acrOss eastern EurOpe tO Siberia, this species is presumably HOlarctic in distributiOn. It is recOrded here fOr the first time frOm China, in the far nOrth Of this cOuntry. Despite being sO widely recOrded, zur Strassen (2003) indicated that the hOst plant remained unknOwn. HOwever, CiuffO et al. (2010) recOrded this thrips as a vectOr Of a species Of tOspOvirus On twO species Of Polygonum in Italy. Dictyothrips was listed by MasumOtO and Okajima (2017) as a member Of the Anaphothrips grOup, because Of the lack Of lOng setae On the prOnOtum. Within that grOup, it is distinguished because Of the presence Of discal setae On the median sternites, and the presence Of a craspedum On tergites II���VII. The Only member Of the genus is yellOw in cOlOur, with characteristic delicate markings within the sculptured reticles On the bOdy (Fig. 3), tergite IX with nO anteriOr campanifOrm sensilla but shOrt stOut majOr setae (Fig. 4), Ocellar setae III small and clOse tOgether within the Ocellar triangle, and antennal segments III���VI with prOminent rings Of micrOtrichia On bOth surfaces. New recOrds: China, Inner Mongolia, Chifeng, 2 females frOm unidentified POaceae, 6.vii.2014 (Zhang SM), in NWAFU., Published as part of Zhang, Shimeng, Wang, Zhaohong, Li, Yajin & Mound, Laurence, 2018, One new species, two generic synonyms and eight new records of Thripidae from China (Thysanoptera), pp. 370-378 in Zootaxa 4418 (4) on page 372, DOI: 10.11646/zootaxa.4418.4.3, http://zenodo.org/record/1244922, {"references":["Masumoto, M. & Okajima, S. (2017) Anaphothrips genus-group: key to world genera, with two new species and three new records from Japan (Thysanoptera, Thripidae). Zootaxa, 4272 (2), 201 - 220. https: // doi. org / 10.11646 / Zootaxa. 4272.2.3"]}

Edissa steinerae Mound (Fig. 11) The genus Edissa is a member Of the subfamily DendrOthripinae, and is particularly unusual within that grOup in being assOciated with the leaves Of POaceae rather than with brOad leaved plants. The genus cOmprises twO species, Of which One is knOwn Only frOm SOuth Africa. HOwever, the secOnd species, steinerae, was described frOm the rainfOrest area Of nOrth eastern Australia and alsO recOrded frOm Thailand and the BOnin Islands Of Japan (MOund 1999). Here it is recOrded frOm a similar ecOlOgical area in sOuthern China, and is thus anOther example Of the relatiOnship Of the thrips fauna Of nOrthern Australia tO that Of sOutheast Asia. This is a pale yellOw species with very slender wings, and withOut prOminent setae, but with a dark area On the anteriOr margin Of the head (Fig. 11). It is unusual amOngst the DendrOthripinae in having 7-segmented antennae with simple sense cOnes in segments III and IV. New recOrds: China, Guangdong, Beitan, Zhanjiang City, 2 females frOm grass, 15.i.2015 (ZhaOhOng Wang), in SCAU., Published as part of Zhang, Shimeng, Wang, Zhaohong, Li, Yajin & Mound, Laurence, 2018, One new species, two generic synonyms and eight new records of Thripidae from China (Thysanoptera), pp. 370-378 in Zootaxa 4418 (4) on pages 372-373, DOI: 10.11646/zootaxa.4418.4.3, http://zenodo.org/record/1244922, {"references":["Mound, L. A. (1999) Saltatorial leaf-feeding Thysanoptera (Thripidae, Dendrothripinae) in Australia and New Caledonia, with newly recorded pests of ferns, figs and mulberries. Australian Journal of Entomology, 38, 257 - 273. https: // doi. org / 10.1046 / j. 1440 - 6055.1999.00112. x"]}

Moundinothrips robustus (Bhatti) (Fig. 12) The Only species in this genus has been knOwn frOm a single female cOllected in CibOdas BOtanic Gardens, BOgOr, Java. The species is similar in general appearance (Fig. 12) tO members Of the genus Taeniothrips, but the first antennal segment bears a pair Of dOrsO-apical setae, the prOnOtum has Only a single pair Of pOsterO-marginal setae, the mesO and the metasternal furca bOth have a well-develOped spinula, and sternite VII bears Only twO pairs Of marginal setae. These character states suggest that the Taeniothrips -like appearance is nOt an indicatiOn Of any systematic relatiOnship. The presence in China Of a species frOm Java is nOt particularly surprising, and further members Of the Taeniothrips cOmplex (MOund et al. 2012) are likely tO be discOvered in sOuthern China. New recOrds: China, Yunnan, all cOllected by Zhang HR & Li YJ: Yuxi City, MOpan MOuntain, 1 female frOm unknOwn plant, 17.iii.2015; Wuding COunty, LiOn MOuntain, 3 females frOm fern, 14.v.2017; Shilin COunty, 1 female frOm Photinia serrulata, 15.iii.2017, in YAU., Published as part of Zhang, Shimeng, Wang, Zhaohong, Li, Yajin & Mound, Laurence, 2018, One new species, two generic synonyms and eight new records of Thripidae from China (Thysanoptera), pp. 370-378 in Zootaxa 4418 (4) on page 373, DOI: 10.11646/zootaxa.4418.4.3, http://zenodo.org/record/1244922

Ensiferothrips wallacei Mound & Tree (Fig. 15) The genus Ensiferothrips cOmprises fOur species. Three Of these are frOm Australia Of which One is alsO knOwn frOm New CaledOnia and NOrfOlk Island. In cOntrast, wallacei was described frOm fOur females taken by insecticide fOgging Of an unidentified fOrest tree in Sulawesi, IndOnesia. This species is here recOrded frOm China fOr the first time, and the plant frOm which it was cOllected is clOsely related tO the knOwn hOst plants Of twO species in this genus (MOund & Tree 2016). In structure, it is very similar tO the type species Of the genus, E. primus, with which it shares curiOusly large dark, grOOved majOr setae On the head and prOnOtum (Fig. 15). New recOrds: China, Hainan, DanzhOu, TrOpical BOtanical Garden, 1 female frOm Maclura tricuspidata [MOraceae], 19.x.2014 (ChaO ZhaO), in SCAU., Published as part of Zhang, Shimeng, Wang, Zhaohong, Li, Yajin & Mound, Laurence, 2018, One new species, two generic synonyms and eight new records of Thripidae from China (Thysanoptera), pp. 370-378 in Zootaxa 4418 (4) on page 373, DOI: 10.11646/zootaxa.4418.4.3, http://zenodo.org/record/1244922

Organothrips Hood Organothrips Hood, 1940: 423. Graminothrips Zhang & Tong, 1992: 82. Syn.n. One female and One male Of Graminothrips cyperi, the type species Of that genus, have been cOmpared with bOth sexes Of Organothrips indicus, a clOse relative Of O. bianchi, the type species Of that genus. These twO species share the fOllOwing unusual character states: head prOjecting in frOnt Of eyes; Ocellar setae I absent; maxillary palps 2-segmented; antennal segments withOut micrOtrichia, III and IV with simple sense cOnes, segment I with paired dOrsO-apical setae; prOsternal basantra with twO setae; fOre tibia inner apex with expanded fimbriate seta; tarsi 1- segmented; tergite pOsteriOr margins laterally with rOw Of micrOtrichia; sternite II with three pairs Of marginal setae; male sternites with multiple small pOre plates. MOreOver, there are nO significant character states by which cyperi and indicus cOuld be cOnsidered members Of different genera. As a result, Graminothrips is here cOnsidered a synOnym Of Organothrips. In this genus, the twO mOst well-knOwn species, bianchi and indicus, are clOsely similar in structure (MOund 2000). They are bOth assOciated with the leaves Of aquatic plants, Often living actually belOw the water surface in the mucilage On stems Of Colocasia, Eichornia and Typha (Bhatti et al. 1998; MOund 2000). In cOntrast, the third species placed in the genus, O. wrighti, was taken frOm grasses at a damp site. Similarly, G. cyperi and G. longisetosus were described frOm specimens taken frOm nOn-aquatic plants, the first frOm Cyperus and the secOnd frOm grass. In cOntrast tO the twO semi-aquatic species, O. wrighti has the first antennal segment reduced in size and lacking paired dOrsO-apical setae, and mOreOver this species has Only twO pairs Of marginal setae On the secOnd abdOminal sternite. Of the twO specimens Of cyperi that have been studied, paired dOrsO-apical setae are clearly present On the first antennal segment Of the male, but On the female these setae are minute and cOuld be seen clearly Only by using a X100 Objective lens. The relatiOnships Of G. longisetosus are mOre difficult tO assess. Despite sharing several character states with the Other species Of Organothrips it differs in the fOllOwing: Ocellar setae I present; prOsternal basantra withOut any setae; tarsi 2-segmented. MOreOver, accOrding tO PrOf. TOng XiaOLi (in litt. ii.2018), paired dOrsO-apical setae are either present Or absent amOng specimens Of this species that have becOme available tO him subsequent tO the Original descriptiOn. The fOllOwing new combinations are required as a result Of the abOve generic synOnymy: Organothips cyperi (Zhang & TOng); Organothrips longisetosus (Zhang & TOng)., Published as part of Zhang, Shimeng, Wang, Zhaohong, Li, Yajin & Mound, Laurence, 2018, One new species, two generic synonyms and eight new records of Thripidae from China (Thysanoptera), pp. 370-378 in Zootaxa 4418 (4) on page 375, DOI: 10.11646/zootaxa.4418.4.3, http://zenodo.org/record/1244922, {"references":["Hood, J. D. (1940) Organothrips bianchii, a new Hawaiian thrips from Taro. Proceedings of the Hawaiian entomological Society, 10, 423 - 427.","Mound, L. A. (2000) The aquatic thrips Organothrips indicus Bhatti (Thysanoptera: Thripidae) in Queensland, and a new species, O. wrighti, from tropical Australia. Australian Journal of Entomology, 39, 10 - 14. https: // doi. org / 10.1046 / j. 1440 - 6055.2000.00136. x"]}

Baliothrips sunae sp.n. (Figs 5–10) Female macroptera: BOdy, femOra and antennal segments I–II unifOrmly brOwn; tarsi yellOw, alsO distal half Of tibiae; antennal segments III–IV mainly yellOw, V–VII light brOwn; fOre wings banded, basal third including clavus pale, then a shOrt dark band, a slightly lOnger pale band, with wing apex dark. Antennae 7-segmented; segment I withOut paired dOrsO-apical setae; III and IV each with sense cOne shOrt and fOrked, sense cOne Y-shaped On III; apex Of V and base Of VI brOadly transverse (Fig. 10); III–VI with micrOtrichia On bOth surfaces. Head slightly lOnger than width acrOss cheeks with eyes large and swOllen (Fig. 5); lOngitudinal rOw Of tuberculate sculpture anteriOr tO first Ocellus, Ocellar triangle weakly sculptured, pOsteriOr third Of head transversely striate/ reticulate; Ocellar setae pair I absent, pair II wide apart lOng but curved, pair III On margin Of triangle; five pairs Of lOng pOstOcular setae, pair I lOngest, nOt arranged in a straight rOw; maxillary palps 2-segmented. PrOnOtum transverse, with nO discal setae Or sculpture medially; fOur pairs Of pOsterOmarginal setae, S1 lOnger than S2, but S3 and S4 (pOsterOangulars) lOngest (Fig. 8); One pair Of midlateral setae lOng and stOut; anteriOr margin with 4–5 pairs Of fine setae. MesOnOtum with nO sculpture Or campanifOrm sensilla On anteriOr third (Fig. 6); median pair Of setae far frOm pOsteriOr margin; mesOfurca with weak spinula. MetanOtum with irregular lOngitudinal striae, median setae far frOm anteriOr margin; metafurca withOut spinula. PrOsternal basantra membranOus, withOut setae; ferna rObust and cOntinuOus medially; prOspinasternum transverse and slender. Tarsi 2-segmented; hind tibiae with setae On distal half lOnger than width Of tibia. FOre wing first vein with 4–5 widely spaced setae near base, 2 widely spaced setae near apex; secOnd vein with abOut 8 widely spaced setae; clavus with 4 veinal and One discal setae. Tergite I cOvered with transverse irregular reticulate sculpture, median setae minute; II–VIII with nO sculpture medially between setae S1, withOut craspeda (Fig. 7); VI–VIII with paired ctenidia ending at discal setae S3; IV–V with weak ctenidia; pOsteriOr margin Of VIII with a few minute micrOtrichia On lateral thirds; IX with anteriOr campanifOrm sensilla absent (Fig. 7); MD setae stOut and extending beyOnd tergal margin, marginal setae lOng and extending beyOnd bOdy apex; X with median split almOst cOmplete, majOr setae nearly twice length Of segments. Sternites II–VII with transverse reticulatiOn (Fig. 9), II with twO pairs Of marginal setae, III–VII with three pairs; S1 On VII in frOnt Of pOsteriOr margin, but clOse tO Or On margin On II–VI. Measurements (hOlOtype female in micrOns). BOdy length 1600. Head, length 125 (nOt hOrizOntal); width acrOss cheeks 140; width acrOss eyes 175. PrOnOtum, length 130; width 210; pOsterOmarginal S1–S4 setae 35, 25, 60, 50. MetanOtum median setae 25. FOre wing length 900. Tergite IX, length 80; MD setae 55; S1 setae 100. Tergite X length 90; S1 setae 140. Male: NOt knOwn. Comments: This new species is referred tO the genus Baliothrips with sOme hesitatiOn, but the alternative Of placing each Of the three species invOlved intO a separate genus de-emphasises their apparent relatiOnships. In a key tO genera related tO Thrips that are assOciated with grasses, Bhatti and MOund (1980) prOvided fOur character states by which they diagnOsed Baliothrips. TwO Of these are fOund in this new species, but twO are absent. Marginal setae S1 On sternites III–VI are submarginal in bOth dispar (Haliday) and kroli (Schille), and in bOth Of them sternite II bears three pairs Of marginal setae Of which pair S1 is submarginal. Neither Of these character states Occurs in this new species, althOugh all three species share a similar cOnditiOn Of the fOre wing clavus with fOur veinal setae, and chaetOtaxy Of the hind tibiae with twO Or three unusually lOng setae. MOreOver, sunae is particularly similar tO dispar, the type species Of Baliothrips, in the 2-segmented maxillary palps, the shape and chaetOtaxy Of the head, the metanOtal sculpture, and many Other details including the banded fOre wings. The new species differs frOm the previOus twO species in the genus in the unifOrmly pale cOlOur Of the wing base and clavus, the absence Of campanifOrm sensilla On the metanOtum and first abdOminal tergite, and the presence Of a weak mesOsternal spinula. Material studied. HOlOtype female, CHINA, Zhejiang, ShuangXikOu, frOm grasses, 27.v.2017 (Niu MM), in NWAFU. Paratypes, 4 females cOllected with hOlOtype, twO in NWAFU; One in ANIC; One in BMNH.

Oxythrips Uzel Oxythrips UZel, 1895: 133. Chilothrips Hood, 1916: 119. Syn.n. The genus Chilothrips was erected fOr a species frOm eastern NOrth America living in the male cOnes Of Pinus trees, and subsequently twO further species frOm CalifOrnia, and fOur frOm China were added (ThripsWiki 2018). PrOblems with distinguishing this genus frOm Oxythrips, the EurOpean type species Of which alsO breeds in Pinus, were discussed by Stannard (1973), whO cOncluded that ���this American genus is but a clOse side-shOOt frOm Oxythrips ���. The key tO genera Of the Anaphothrips grOup by MasumOtO and Okajima (2017) distinguished Chilothrips as having: fOre tibia with a stOut apical seta; female with mOuth cOne lOng; tergite X twice as lOng as IX. In cOntrast, Oxythrips species have: fOre tibia with a thin apical seta; female with mOuth cOne usually shOrt; tergite X usually subequal tO IX. This separatiOn is clearly nOt rObust, and is further weakened because males have a smaller mOuth cOne than females. MOreOver, females Of the cOmmOn EurOpean species, Oxythrips bicolor, anOther species frOm Pinus male cOnes, have a large stOut mOuth cOne, and tergite X is twice as lOng as tergite IX. This species cOuld equally well be placed in Chilothrips. HOwever, intersex individuals recOrded cOmmOnly in nOrthern ScOtland (MOund 1971) are interpreted as evidence that bicolor interbreeds with ajugae Uzel, the type species Of Oxythrips. These twO species are therefOre presumably clOsely related, and shOuld nOt be placed in separate genera. Studies in China (Cui et al. 2017) On Chilothrips strobilus, have indicated remarkable variatiOn in the length and shape Of tergite X within a single pOpulatiOn frOm Pinus, frOm abOut 1.2 times as lOng as basal width tO at least 3.0 times. In view Of the many character states shared by species that have been placed in these twO genera, including the shared hOst assOciatiOn with Pinus male cOnes in several species, these twO are here fOrmally synOnymized. As a result, the fOllOwing new combinations are established: Oxythrips hangzhouensis (Hu & Feng); Oxythrips jiuxiensis (Mirab-balOu, Chen & TOng); Oxythrips occidentalis (Stannard); Oxythrips pini (HOOd); Oxythrips rotrameli (Stannard); Oxythrips strobilus (TOng & Zhang); Oxythrips yamatensis (KudO)., Published as part of Zhang, Shimeng, Wang, Zhaohong, Li, Yajin & Mound, Laurence, 2018, One new species, two generic synonyms and eight new records of Thripidae from China (Thysanoptera), pp. 370-378 in Zootaxa 4418 (4) on page 376, DOI: 10.11646/zootaxa.4418.4.3, http://zenodo.org/record/1244922, {"references":["Hood, J. D. (1916) Descriptions of new Thysanoptera. Proceedings of the Biological Society of Washington, 29, 109 - 124.","Masumoto, M. & Okajima, S. (2017) Anaphothrips genus-group: key to world genera, with two new species and three new records from Japan (Thysanoptera, Thripidae). Zootaxa, 4272 (2), 201 - 220. https: // doi. org / 10.11646 / Zootaxa. 4272.2.3","Mound, L. A. (1971) Sex intergrades in Thysanoptera. Entomologist's Monthly Magazine, 106, 186 - 189.","Cui, Y. Z., Xi, J. H. & Wang, J. (2017) Female polymorphism in Chilothrips strobilus (Thysanoptera: Thripidae), with the first description of the male. Zootaxa, 4358 (1), 193 - 196. https: // doi. org / 10.11646 / Zootaxa. 4358.1.11"]}

Eremiothrips arya (zur Strassen 1975) Yang et al. (1993) first recorded E. arya from China, and Han (1996) subsequently recorded 6 females and 7 males of this species. However, these authors did not provide a description, nor indicate the basis for the identification. Specimen information was not present in China (NSII 2017). A possible specimen of this species has been borrowed from Inner Mongolia, but unfortunately, detailed characters could not be observed because of the deterioration of the specimen. Presence of this species in China requires further confirmation. Distribution. China (Inner Mongolia, Ningxia, Xizang, Xinjiang); Iran., Published as part of Zhang, Shimeng & Feng, Jinian, 2017, The genus Eremiothrips (Thysanoptera: Thripidae) from China, with one new species, pp. 396-400 in Zootaxa 4250 (4) on page 399, DOI: 10.11646/zootaxa.4250.4.10, http://zenodo.org/record/495333, {"references":["Yang, C. X., Liu, Y. J., Ma, C. J. & Han, Y. F. (1993) Thysanoptera collected from the wild grasslands of Ningxia and West Inner Mongolia annotated checklist. Acta Agriculturae Boreali-occidentalis Sinica, 2, 1 - 6.","Han, Y. F. (1996) Thysanoptera; In: Huang, F. & Zhang, X. Z. (Eds.), Insects of the Karakorum-Kunlun Mountains. Science Press, Beijing, pp. 93 - 95."]}

Eremiothrips hanshanensis Zhang & Feng sp. n. (Figures 1–9) Female macroptera. Body light brown or yellowish brown. Antennal segment I white (Fig. 5), segments II and VI–IX brown, segments III–IV light brown, segment V light brown with brown apex. Legs light brown. Forewing white. Setae with light colour. Head wider than long (Fig. 1), compound eyes with six pigmented facets ventrally, arranged in three rows; three pairs of ocellar setae present, pair III in front of hind ocelli within triangle, longer than ocellar setae I; 5–6 pairs of postocular setae, all setae on head short; maxillary palps three-segmented. Antennae nine-segmented (Fig. 5), segment I without median dorsal apical setae, III and IV with forked sense cones and weak microtrichia. Pronotum almost smooth (Fig. 2), rarely weakly transversely striate, with no long discal setae, with one pair of long posteroangular setae, posterior margin with four pairs of setae. Mesonotum with transverse lines (Fig. 3), campaniform sensilla inconspicuous. Metanotum with cross striation medially (Fig. 3), the rest with longitudinal lines, median setae well behind anterior margin; campaniform sensilla absent; mesothoracic sternopleuralsuture present; spinula absent on mesosternum and metasternum (Fig. 4). Tarsi two-segmented. Fore wing slender with posterior cilia wavy; first vein with 6 setae on basal half and 3 widely spaced setae on distal half, second vein with 10 widely spaced setae; clavus with 4 veinal and 1 discal setae. Abdominal tergites weakly sculptured medially, with no ctenidia or craspedum; tergite II without lateral marginal setae; tergal campaniform sensilla posterolateral to median setae, close to posterior margin; tergite VIII with scattered short microtrichia (Fig. 8), both sides with faint transverse reticulation, median campaniform sensilla situated behind of the middle between S1 seta and posterior margin, close to the posterior margin; tergite IX with median dorsal setae extending to posterior margin of tergite, and with two pairs of campaniform sensilla near the margin, setae S2 almost equal to S1; tergite X with dorsal split short; ovipositor developed. Sternites without discal setae or marginal craspedum, sternite II with 2 pairs of marginal setae, III–VII with 3 pairs; pleurotergites with no discal setae, median pair of setae on sternite VII arising in front of posterior margin (Fig. 6). Male macroptera. Body smaller than female, colour lighter than female. Three pairs of ocellar setae present. Pronotum with one pair of long setae on posterior angles. Sternites III–VII each with small pore plates (Fig. 9), tergite IX with two long, special processes at posterior margin (Fig. 7). Measurements (holotype female in microns): Body length 1480. Head length 117, width 160, ocellar setae III 23. Pronotum length 123, width 189, posteroangular inner setae 40; outer setae 16. Forewing length 687. Metanotal median setae 17. Tergite IX setae S1 92, S2 92. Antennae length (width) I 17 (25), II 33 (23), III 33 (17), IV 33 (16), V31 (17), VI 36 (17), VII 7 (11), VIII 8 (7), IX 15 (4). Measurements (paratype male in microns): Body length 1009. Head length 89, width 121, pronotum length 87, width 143, posteroangular inner setae 32. Forewing 545. Metanotal median setae 17. Sternites III–VII glandular areas 17 width (w), 7 length (l); 11 w, 7 l; 9 w, 6 l; 5 w, 5l; 4 w, 4 l. Antennae length (width) I 15 (21), II 23 (19), III 26 (14), IV 26 (14), V 25 (14), VI 29 (14), VII 7 (10), VIII 6 (7), IX 12 (4). Materials examined. Holotype female: CHINA, Inner Mongolia province, Chifeng County, Hanshan forestry centre, collected from Asteraceae, 10. VII. 2014, collected by Shimeng Zhang. Paratypes. 2 females and 1 male, the same data as holotype, one of the females is short-winged. Remarks. It is difficult to distinguish species of Eremiothrips based only on females, whereas males show clear differences. E. eshghii and E. efflatouni are obviously different from other species in absence of long setae in the pronotum in both sexes. According to Minaei (2012; 2014), nine species are known to share with this new species the presence in males of pore plates: E. arya, E. bhattii, E. dubius, E. eshghii, E. farsi, E. hudeci, E. taghizadehi, E. similis and E. zurstrasseni. Pore plates on sternites III–VII as in E. hanshanensis also occur in E. hudeci and E. zurstrasseni, but they are smaller and oval in E. hanshanensis whereas they are larger and transverse in those two species. E. similis, E. farsi and E. bhattii are distinguished from E. hanshanensis by the absence of special processes on tergite IX of males. E.dubius, E. taghizadehi and E. arya are similar to E. hanshanensis in having sternal pore plates. However, these are present on sternites IV–VII in these three species, but on sternites III–VII in E. hanshanensis. This new species has longer and normal processes on tergite IX, whereas they are small in E. dubius, and with the apex sharply recurved in E. taghizadehi. A single brachypterous female of E. hanshanensis was collected with the same data as the holotype. This shortwinged female is similar to the long-winged female: body length 1205, ocelli and three ocellar setae are present on head (Fig 10), compound eyes with six pigmented facets ventrally. Pronotum with no long discal setae, with one pair of long posteroangular setae. Metanotum without campaniform sensilla (Fig 11), wings short (Fig 11), length 131. Median campaniform sensilla situated behind of the middle between S1 seta and posterior margin, close to the posterior margin on tergite VIII (Fig 12). This is the first published record of wing-reduction among Eremiothrips species, although such a phenomenon is widespread amongst other Anaphothrips genus-group species. Etymology. The specific epithet refers to the collection localityof this species.

Eremiothrips antilope (Priesner 1923) Yang et al. (1993) first recorded this species from China and provided information on host plants and distribution. However, the authors did not provide a description of the species, nor include any references to indicate the basis for the identification. Specimen information was not found in China (NSII 2017). Specimens have not been seen by the present authors, and the presence of this species in China requires further confirmation. Distribution. China (Inner Mongolia, Ningxia); Egypt, Israel, Saudi Arabia, Iran, Yemen, Morocco, Sudan, Cyprus, India., Published as part of Zhang, Shimeng & Feng, Jinian, 2017, The genus Eremiothrips (Thysanoptera: Thripidae) from China, with one new species, pp. 396-400 in Zootaxa 4250 (4) on page 399, DOI: 10.11646/zootaxa.4250.4.10, http://zenodo.org/record/495333, {"references":["Yang, C. X., Liu, Y. J., Ma, C. J. & Han, Y. F. (1993) Thysanoptera collected from the wild grasslands of Ningxia and West Inner Mongolia annotated checklist. Acta Agriculturae Boreali-occidentalis Sinica, 2, 1 - 6."]}

Key to species of Eremiothrips from China (males) (E. antilope and E. arya are included based on Bhatti et al. 2003 and Minaei 2012) 1. Abdominal sternites of male without pore plates ............................................................................................................... E. antilope - Abdominal sternites of male with pore plates................................................................................................................................... 2 2. Male with large pore plates on abdominal sternites IV–VII ................................................................................................... E. arya - Male with small pore plates on abdominal sternites III–VII ........................................................................... E. hanshanensis sp. n.

Eremiothrips Priesner Eremiothrips Priesner 1950: 44. Type species Eremiothrips imitator Priesner. Diagnosis. Head wider than long. Antennae usually 9-segmented, sometimes 8-segmented. Sense cones on segments III and IV usually forked, rarely simple. Head not elongate in front of eyes, ocelli developed. Three pairs of ocellar setae present. Mouth-cone normal. Body setae lighter. Pronotum with or without one pair of long posteroangular setae. Mesosternum with or without spinula, metasternum without. Forewing properly long, with clear veins. Abdominal tergites IV���VIII without microtrichia, abdominal tergites and sternites without craspedum, sternites without discal setae, tergite VIII with or without posteromarginal comb, tergites X with round apex. Male usually with special processes on tergite IX, sternites III���VII with or without pore plates., Published as part of Zhang, Shimeng & Feng, Jinian, 2017, The genus Eremiothrips (Thysanoptera: Thripidae) from China, with one new species, pp. 396-400 in Zootaxa 4250 (4) on page 396, DOI: 10.11646/zootaxa.4250.4.10, http://zenodo.org/record/495333, {"references":["Priesner, H. (1950) Contributions towards a knowledge of the Thysanoptera of Egypt, XV. Bulletin de la Societe Fouad Ierd'Entomologie, 34, 25 - 37."]}

Key to species of Eremiothrips from China (males) (E. antilope and E. arya are included based on Bhatti et al. 2003 and Minaei 2012) 1. Abdominal sternites of male without pore plates ............................................................................................................... E. antilope - Abdominal sternites of male with pore plates................................................................................................................................... 2 2. Male with large pore plates on abdominal sternites IV���VII ................................................................................................... E. arya - Male with small pore plates on abdominal sternites III���VII ........................................................................... E. hanshanensis sp. n., Published as part of Zhang, Shimeng & Feng, Jinian, 2017, The genus Eremiothrips (Thysanoptera: Thripidae) from China, with one new species, pp. 396-400 in Zootaxa 4250 (4) on page 396, DOI: 10.11646/zootaxa.4250.4.10, http://zenodo.org/record/495333, {"references":["Bhatti, J. S., Telmadarraiy, Z., Kumar, V. & Tyagi, K. (2003) Species of Eremiothrips in Iran (Terebrantia: Thripidae). Thrips, 2, 49 - 110.","Minaei, K. (2012) The genus Eremiothrips (Thysanoptera: Thripidae) in Iran, with one new species. Zootaxa, 3349, 56 - 62."]}