Search

The article introduces new paleoanthropological materials from the Mayak burial ground near Sidelkino village in the Samara region into scientific discourse. The materials were obtained as a result of excavations in 1995 and only recently was it possible to date them. As a result of AMS analysis fulfilled by the authors, human remains from two burials were dated back to the Early Mesolithic. The analysis was carried out taking into account the influence of the “reservoir effect.Despite the rather poor preservation of individuals from the two described burials, the authors carried out an osteological analysis of an adult male from the second burial. He turned out to be quite tall, with elongated legs, shortened forearms, saber-shaped tibia and relatively massive bones of the lower limbs. In burial 3, the remains of an adult woman and a child of 7–10 years old were found. Comparing the osteological indicators of the man from the second burial with materials from the same burial ground of excavations in 2002, the authors came to the conclusion that the people whose remains were found on the hill Mayak in 1995 and in 2002 probably belong to the same population. Similar morphological characteristics are proof of this.

We present an overview of the beginning and early years of radiocarbon dating in Russia. Achievements of several major scholars in this field from Leningrad (St. Petersburg), Moscow and Novosibirsk are briefly described. The existing and closed Russian laboratories are also mentioned.

This paper reviews significant issues related to the fossil hominins from the Altai Mountains of Siberia (Russia), namely Denisovans, Neanderthals, and early modern humans. Uncritical acceptance of the recovered information by some authors has resulted in unreliable chronologies of the Middle and Upper Paleolithic artifact assemblages and the animal and hominin fossils. We examine the chronostratigraphic contexts and archaeological associations of hominin and animal fossils and the lithics discovered at the Denisova, Okladnikov, Strashnaya, and Chagyrskaya cave sites. Taphonomic, site formation, and geomorphological studies show evidence of disturbance and redeposition caused by carnivore activity and sediment subsidence at these sites, which complicates the dating of the human remains. Our analysis indicates that the Middle Paleolithic is dated to ca. 50,000–130,000 years ago, and the Upper Paleolithic to ca. 12,000–48,000 years ago. The best age estimate for Denisovans is ca. 73,000–130,000 years ago. The ages of Neanderthals can be determined as more than 50,000–59,000 years ago, and of modern humans at roughly 12,000–48,000 years ago. Denisovan and Neanderthal fossils are associated with Middle Paleolithic complexes only.

For more than a half-century, obsidian provenancing has underpinned many archaeological investigations of peoples of the past. The pace of obsidian studies in this regard has gathered significantly since around 2007, and we review the literature to gain a sense of where this momentum has come from, and what it heralds. In part, there is a data revolution underway, arising thanks to the capabilities for rapid survey and analysis enabled by field-portable analytical equipment. Obsidian studies are also gaining a stronger foothold in regions of the world where the approach was previously under-exploited. Our survey spans progress made in obsidian studies in the Mediterranean, Central Europe, the Near East, the Caucasus, Northeast Asia and Tibet, the Eurasian Arctic and Alaska, Southeast Asia, the Americas, Oceania, and Africa and Arabia. We also consider methodological issues related to compatibility of differing geochemical analytical techniques, and the state of the art in obsidian geochemical classification. The proliferation of new observations brings opportunities in terms of development of regional and global databases, as well as challenges of calibration and validation of analyses made by different scientists and laboratories employing diverse instrumentation. Obsidian provenancing demonstrates the astonishing ranges of our ancestors’ interactions and networks, sometimes exceeding 1000 km and involving maritime transport.

The reliability of radiocarbon dates for Palaeolithic human burials is of utmost importance for prehistoric archaeologists. Recently obtained dates for several such burials in central Russia raise important interrelated issues concerning site taphonomy and the precise radiocarbon-dating technique employed, with implications for the ‘true’ age of the burials. A critical review of the dating of the Sungir and Kostenki burials calls into question the reliability of employing ultrafiltration or single amino acids for the radiocarbon dating of Upper Palaeolithic bones.

Since the 1990s, a characteristic obsidian geochemistry has linked widespread archaeological assemblages spanning the Russian Far East, Korean Peninsula, and Northeast China. Referred to as PNK1, the source of this material has yet to be identified. As a contribution to solving this enduring puzzle, we report here analyses of a commercial specimen of obsidian exported from Chongjin in DPR (North) Korea. A combination of Neutron Activation Analysis (NAA) and Potassium Argon dating enable us to compare this piece with a large obsidian database for Northeast Asia. We find that the “Chongjin sample” is identical to PNK1 lithics from archaeological collections. While the exact source of the “Chongjin sample” remains unknown, we can more confidently locate the primary source for PNK1 lithics in DPR Korea. Based on an exhaustive literature review of the geology and geochemistry of volcanic glasses and other volcanic rocks in the northern part of the Korean Peninsula, and drawing on our own unpublished data, we suggest that the PNK1 source is most likely located south of Paektusan Volcano. This corroborates the existing evidence for the long-range transport of the material.

Significance The Siberian Arctic has witnessed numerous societal changes since the first known appearance of dogs in the region ∼10,000 years ago. These changes include the introduction of ironworking ∼2,000 years ago and the emergence of reindeer pastoralism ∼800 years ago. The analysis of 49 ancient dog genomes reveals that the ancestry of Arctic Siberia dogs shifted over the last 2,000 years due to an influx of dogs from the Eurasian Steppe and Europe. Combined with genomic data from humans and archaeological evidence, our results suggest that though the ancestry of human populations in Arctic Siberia did not change over this period, people there participated in trade with distant communities that involved both dogs and material culture., Dogs have been essential to life in the Siberian Arctic for over 9,500 y, and this tight link between people and dogs continues in Siberian communities. Although Arctic Siberian groups such as the Nenets received limited gene flow from neighboring groups, archaeological evidence suggests that metallurgy and new subsistence strategies emerged in Northwest Siberia around 2,000 y ago. It is unclear if the Siberian Arctic dog population was as continuous as the people of the region or if instead admixture occurred, possibly in relation to the influx of material culture from other parts of Eurasia. To address this question, we sequenced and analyzed the genomes of 20 ancient and historical Siberian and Eurasian Steppe dogs. Our analyses indicate that while Siberian dogs were genetically homogenous between 9,500 to 7,000 y ago, later introduction of dogs from the Eurasian Steppe and Europe led to substantial admixture. This is clearly the case in the Iamal-Nenets region (Northwestern Siberia) where dogs from the Iron Age period (∼2,000 y ago) possess substantially less ancestry related to European and Steppe dogs than dogs from the medieval period (∼1,000 y ago). Combined with findings of nonlocal materials recovered from these archaeological sites, including glass beads and metal items, these results indicate that Northwest Siberian communities were connected to a larger trade network through which they acquired genetically distinctive dogs from other regions. These exchanges were part of a series of major societal changes, including the rise of large-scale reindeer pastoralism ∼800 y ago.

We report a new series of radiocarbon (14C) dates on the MIS 3 megafauna for a previously poorly studied region of southeastern West Siberia. Some species, like woolly mammoth and woolly rhinoceros, and Pleistocene bison and horse, existed throughout the MIS 3 (ca. 29–59 ka cal BP); cave hyaena is dated to ca. 46,400 cal BP. The very late 14C dates on Khozarian steppe elephant (Mammuthus trogontherii chosaricus), ca. 45,100–45,400 cal BP, may indicate the survival of this species in Siberia up to MIS 3. More work is needed to confirm or reject this suggestion. Previously, Khozarian steppe elephant was known in Siberia only at the beginning of the Late Pleistocene (MIS 5e).

Zhokhov Island in the Siberian High Arctic has yielded evidence for some of the most remote prehistoric human occupation in the world, as well as the oldest-known dog-sled technology. Obsidian artefacts found on Zhokhov have been provenanced using XRF analysis to allow comparison with known sources of obsidian from north-eastern Siberia. The results indicate that the obsidian was sourced from Lake Krasnoe—approximately 1500km distant—and arrived on Zhokhov Island c. 8000 BP. The archaeological data from Zhokhov therefore indicate a super-long-distance Mesolithic exchange network.

The article was prepared with the support of Tomsk State University grant program; State assignment number 0330-2016-2018; the Russian Science Foundation (RNF), grant number 14-50-00036.

We conducted inter-laboratory AMS 14C dating of bones of the Miesenheim IV elk (Rhineland, Germany), buried under Laacher See tephra dated to ca. 11,060 BP (13,000 cal BP). The weighted mean of A laboratory inter-comparison of AMS 14C dating of bones of the Miesenheim IV elk (Rhineland, Germany) and its implications for the date of the Laacher See eruptionthe new dates, which range from 10,920 to 11,270 BP, is 11,092 ± 19 BP. The consistent results from five AMS laboratories are important in two respects. First, they demonstrate that collagen processed by traditional methods can yield accurate ages; the newly obtained 14C dates are in accord with previous hydroxyproline 14C value generated at the Oxford AMS laboratory within the first round of inter-comparison (Fiedel et al., 2013). The results of the first inter-comparison are clearly flawed, except for hydroxyproline 14C date (see Fiedel et al., 2013), and must be affected by the waxy/dark, presumably humic/organic-based contaminant. Second, they provide a new suite of radiocarbon dates for the Laacher See volcanic eruption, a crucial anchor point for Late Glacial chronology in central Europe.

Geochemical analysis of 102 obsidian artifacts from the lower reaches of the Kolyma River was performed to understand the provenance of the raw material; previously, there were no such studies in this region. Sites under investigation belong to the Arctic Neolithic, generally dated to ca. 6000–1500 BP. Based on the data for potential obsidian sources in Northeastern Siberia and neighboring territories, available to us, it was found that all obsidian artifacts originated from the Lake Krasnoe source in Chukotka, with a straight-line distance of ca. 800–1100 km from archaeological sites of the Kolyma River. This is a remarkable example of long-distance exchange/transport of obsidian in Northeastern Siberia during the Stone Age. The Lake Krasnoe locale was the primary obsidian source for prehistoric populations in this vast region, including Chukotka, the Kolyma River basin, and Okhotsk Sea coast; this obsidian was also identified at some Alaskan sites near the Bering Strait.

Patterns for the emergence of pottery-making in greater East Asia based on radiocarbon dates associated with the earliest pottery assemblages are presented. According to a critical evaluation of the existing evidence, the oldest centers with pottery in East Asia are located in South China (dated to ca. 18,000 cal BP), the Japanese Islands (ca. 16,700 cal BP), and the Russian Far East (ca. 15,900 cal BP). The claim for earlier pottery in South China at the Xianrendong Cave, supposedly dated to ca. 20,000 cal BP, cannot be substantiated. The appearance of pottery in other parts of greater East Asia was a slow process, without clear diffusion from any of these centers toward the periphery. In neighboring Siberia, the oldest pottery dated to ca. 14,000 cal BP is known from the Transbaikal.

During the Late Glacial, hunter-gatherers began using ceramic cooking containers in three separate geographic regions of East Asia: China, Japan and in the Russian Far East. While recent research has clarified the use of early pottery in Japan, very little is known about what led to the emergence of pottery in the other two areas, including the likely environmental, economic or cultural drivers. In this paper we focus on a series of key sites along the Amur River in the Russian Far East, where early pottery has been recovered from securely-dated contexts that span ca. 16,200 to 10,200 years ago (cal BP). Interpreting how these ceramic vessels were used has been difficult because the region’s acidic soils make palaeo-economic reconstructions challenging. To address this gap in knowledge we undertook lipid residue analysis of 28 pot sherds from the sites of Khummi, Gasya, and Goncharka 1 on the Lower Amur River, and the Gromatukha site on the Middle Amur. Our results indicate that pottery was employed to process aquatic oils at sites on the Lower Amur, a pattern of use that aligns closely with studies conducted in Japan, and suggests that fishing – probably of salmonids and freshwater fish – was becoming increasingly important during this period. In contrast, the results from the Middle Amur show a significant contribution of lipids from ruminant animals, indicating that these vessels were being used in different ways. Interestingly, these regional differences in pottery use also map onto contrasting manufacturing techniques, with vessels from the Middle and from the Lower Amur forming distinct pottery-making traditions. These combined insights appear to indicate a greater degree of variability in the development and use of early pottery in East Asia than has hitherto been indicated.

Yue et al. (2021) have proposed a model of two Upper Palaeolithic industries in Northeast China called the “Initial Microblade Industry” and the “Northern Microblade Industry”, with the latter originating from the former, and its spread to the neighbouring regions of Northeast Asia. We have found several inconsistencies in their factual data and methodology, and therefore the conclusions reached by Yue et al. (2021) cannot be accepted at face value. Using the most reliable evidence, the oldest true microblade assemblages are known from the Korean Peninsula. We suggest that this region is the more probable centre of diffusion of microblade technology in mainland Northeast Asia, including China and the Russian Far East.

Basic data on the geology and geochemistry of obsidian from the Lake Krasnoe source in Chukotka (Northeastern Siberia) are reported for the first time. The data are based on 2009 fieldwork and analytical studies of igneous rock samples. The lake shore and surrounding parts of the Rarytkin Range were thoroughly examined. Two geochemical types of rhyolitic obsidian were recognized for the first time: (1) metaluminous obsidian related to the fine-grained crystalline rocks and (2) peralkaline obsidian corresponding to ignimbrite ash-flows or lapilli-tuffs composition. Both types are related to the final phase of acidic volcanism in the Western Kamchatkan-Koryak Volcanic Belt. Based on the results obtained, we conclude that accumulation of obsidian pebbles in the lake’s modern beach deposits is related to silicic melts that erupted during the late Eocene-early Oligocene in the form of extrusive domes or pyroclastic flows, which are now either covered by Quaternary sediments or located below the water level. The Lake Krasnoe obsidian was intensively used by the ancient populations of Chukotka as a raw material for making stone tools. It was also occasionally transported to Alaska across the Bering Strait in later prehistory. The distances between source and utilization sites are up to 700–1100 km. Geochemical data for Lake Krasnoe obsidian, based on neutron activation analysis and X-ray fluorescence that are presented here, can now be used for provenance studies in the Northeastern Siberia and adjacent regions of northern North America.

An Illustrated Companion to Japanese Archaeology. Edited by Werner Steinhaus and Simon Kaner. Archaeopress, Oxford. 2016. ISBN 978-1-78491-425-7 (Paperback £35.00), 344 pp, 210 figures, 248 plates....

For the first time, an obsidian provenance study is performed for artifacts from the Indigirka River basin (Northeast Siberia). The non-destructive ED–XRF analysis of seven obsidian items from the Buolumuna-Taasa site, dated to the Neolithic (ca. 5000–1000 cal BC), shows that all of them originated from the Lake Krasnoe source, situated ca. 1300 km away. This is a remarkable example of super-long-distance transport / exchange of obsidian in prehistory of Siberian Arctic, and it testifies in favor of extensive contacts of ancient population in Northeast Siberia since the Mesolithic, ca. 6900 cal BC, or even earlier.

The current status of research on colonization and early migration in the insular part of the Russian Far East is presented. Based on a critical evaluation of the latest data, it is concluded that the initial colonization of Sakhalin Island occurred at ca. 19,500 BP and possibly earlier, but no solid evidence has yet been found to determine this with more precision. The movement of people from neighboring Hokkaido (northern Japan) to Sakhalin continued throughout the Paleolithic–Neolithic since ca. 10,000 BP with the help of maritime transport. In the Middle–Late Neolithic, ca. 5200–4400 BP, migration from the adjacent mainland (via the lower reaches of the Amur River) to Sakhalin is detected. The southern part of the Kurile Archipelago was initially colonized from Hokkaido in the Early Jomon (i.e., Neolithic) at ca. 7200–7000 BP; the northern part was subsequently settled possibly as early as ca. 5300 BP. Active contacts between Hokkaido and the Kurile Islands continued throughout and after the J...

Shuidonggou localities 1 and 2 provide key evidence for the Initial Upper Palaeolithic of north-east Asia. In a recent article inAntiquity(87 (2013), 368–383), Liet al.proposed a new chronology, building on the earlier results of Madsenet al.(Antiquity75 (2001), 705–716). Here Susan Keates and Yaroslav Kuzmin take issue with the new chronology. The article is followed by a response from Li and Gao.

New paleodietary data were obtained after the discovery and excavation in 2015–2017 of the Cherepakha 13 site in the southern part of Primorye (Maritime) Province in far eastern Russia. The site is located near the coast of Ussuri Bay (Sea of Japan) and belongs to the Yankovsky cultural complex of the Early Iron Age 14C-dated to ca. 3000 BP (ca. 1200 cal BC). The stable isotope composition of the bone collagen for 11 humans and 30 animals was determined. For humans, the following values (with±1 sigma) were yielded: δ13C=–10.2±0.8‰; and δ15N=+12.4±0.3‰. The majority of terrestrial animals show the usual isotopic signals: δ13C=–19.4 ÷ –23.3‰; and δ15N=+4.6÷+6.6‰ (for wolves, up to +10.1‰); dogs, however, have an isotopic composition similar to humans: δ13C= –11.7±1.2‰; and δ15N=+12.4±0.4‰. Marine mammals have common values for pinnipeds: δ13C=–13.7 ÷ –14.6‰; and δ15N=+17.4 ÷ +18.0‰. The main food resources for the population of Cherepakha 13 site were (1) marine mollusks, fish, and mammals; and (2) terrestrial mammals; and possibly C4 plants (domesticated millets).

Obsidian provenance studies, based on geochemical signatures, are important for determining the source regions of obsidian artefacts. Such research depends on the availability of reproducible geochemical data. An inter-laboratory study was conducted to validate analytical methods applied to samples from four obsidian sources in northeast Hokkaido Island (Shirataki, Rubeshibe, and Oketo regions). The methods applied were Wavelength-Dispersive X-ray Fluorescence (WDXRF), Energy-Dispersive X-ray Fluorescence (EDXRF), Inductively Coupled Plasma–Mass Spectrometry (ICP–MS), Neutron Activation Analysis (NAA) and Prompt-Gamma Activation Analysis (PGAA). Eight laboratories in Japan, the Russian Federation, Republic Korea, Hungary, Canada, and the USA took part in the trials. Results indicate discrepancies between laboratories, but compositional data for 53 elements were successfully compiled, and reference compositions for 16 elements in each sample defined. Based on these data, a new chemical discrimination scheme is proposed for obsidian sources in the Shirataki, Rubeshibe, and Oketo regions. This scheme is applicable to the discrimination of obsidian sources using semi-quantitative EDXRF analysis, with this being important in non-destructive provenance studies of artefacts. This study fosters the further establishment of reference materials for obsidian sources in the Hokkaido region, and the sharing of such materials.

An overview of current knowledge on the human paleodiet of the southern Russian Far East is presented. The earliest data are available for the Neolithic complexes of Primorye Province, dated to ~7000–5800 BP. For 10 humans from the coastal site of Boisman 2, a heavy reliance on marine fish and mammals (~70–80% of the total diet) has been established; this is similar to the Jomon and Chulmun complexes of Hokkaido Island and Korea. For two individuals from the inland site of Chertovy Vorota, a mixture of terrestrial and riverine (including anadromous species such as salmon, ~25% of the total diet) food resources is evident. In the Amur River basin, the diet of the Paleometal (i.e. Bronze/Early Iron Ages) population (dated to ~2500–1800 BP) was probably based on a mixture of C4 plants (millet, ~50–60% of the total diet), C3 plants, and terrestrial animals, while in the Middle Ages (~1500–300 BP) the contribution of C4 food was lower at ~20–25%. On Sakhalin Island, the maritime-oriented economy existed for a long time, at least since the Late Neolithic/Paleometal Age (dated to ~2500–1800 BP) and until the ethnographic time period (from the 17th–18th centuries AD onwards). Up to ~80–90% of the diet consisted of marine mammals and fish, and this is in accord with dietary patterns of the Epi-Jomon and Okhotsk cultural complexes on Hokkaido Island. Information on the paleodiet of the coastal populations of the Japan and Okhotsk Seas should be taken into account when calibration of 14C dates run on human bones from these regions is undertaken, because the predominant consumption of marine food caused a distortion of the true 14C age.

Review of the book: E.M. Ineshin & A.V. Tetenkin (translated by P.N. Hommel & N. Reynolds). Humans and the environment in northern Baikal Siberia during the Late Pleistocene. 2017. Newcastle upon Tyne: Cambridge Scholars; 978-1-4438-8277-4 £64.99. This book is about the Terminal Pleistocene archaeology of the Vitim River Basin, located between North Baikalsk and the Patomsk uplands. Hardly known outside of Russia, it is only briefly mentioned even in major summaries on the Upper Palaeolithic of eastern Siberia (Mochanov 2009; Pitul’ko & Pavlova 2016: 113–15). The release of the English translation of this volume, originally published in Russian in 2010, is therefore very welcome. The Foreword by J. Chlachula provides an introduction of this work for an international audience.

The Ekven Settlement: Eskimo Beginnings on the Asian Shore of Bering Strait. Edited by Yvon Csonka. BAR International Series 2624, Archaeopress, Oxford. 2014. ISBN 978-1-4073-1259-0 (Paperback £30....

The updated chronology of the earliest pottery-containing complexes in Siberia and the Russian Far East is presented herein. The appearance of pottery (i.e. the process of Neolithization) in this vast region of Eurasia is discussed based on a model that represents a simple approximation of calendar ages between key sites as isolines. No clear spatiotemporal patterns for the origin and spread of pottery in northern Asia can be observed because pottery-making (unlike agriculture) could have emerged in different parts of the Old World at various times. Before modeling of pottery dispersal is conducted, careful evaluation of typology and technology of ceramics and stone artifacts should be done, in order to avoid the confusing situation when the results of modeling contradict the basic archaeological information. DOI: 10.2458/56.16938

The corpus of radiocarbon dates run directly on Pleistocene-age human remains in Eurasia (∼120 values, with ∼80 of them found to be reliable) is analyzed and interpreted. The latest Neanderthals are dated to ∼34,000–30,500 BP (∼38,800–35,400 cal BP). They probably coexisted with the first modern humans at ∼36,200–30,200 BP (∼42,500–32,800 cal BP) in the western and central parts of Europe. The earliest direct14C dates on modern humans in Eurasia are ∼34,950–33,300 BP (∼40,400–37,800 cal BP). A paucity of14C dates corresponding to the LGM is evident for Europe, but Asia perhaps had larger populations during this timespan. The main criteria for the selection of bone/tooth material for direct14C dating as now widely accepted are (1) the collagen yield (generally, 1% or more) and (2) the C:N ratio (within the 2.9–3.4 range).

A critical evaluation of the existing data corpus on the earliest pottery in East Asia and its chronology as of early 2013 is presented here. Pottery in the Old World emerged in three regions within greater East Asia, namely South China, the Japanese Islands and the Russian Far East, at c. 14,800–13,300 bp (or c. 18,500–15,500 cal. bp). Most probably, pottery-making appeared in these places independently; no solid evidence exists about migrations and/or diffusion of this technology from a supposed single centre in South China. Because the Upper Palaeolithic humans in Eurasia were familiar with clay (as a raw material for making figurines), the most probable driving force for the origin of pottery was the necessity to produce in large amounts durable, light containers for the processing (including boiling) and storing of food.

Intriguing wooden objects, excavated (mostly unstratigraphically) from peat bogs in the Trans-Urals region of Russia, are here dated by AMS and found to belong to the Aeneolithic and Bronze Age. In spite of a long sojourn in museums, and conservation with various chemicals, the dates obtained were consistent and reliable.

The chronology of the Mesolithic, Neolithic, Aeneolithic, and Bronze Age sites in the Trans-Urals (Russia) was studied, and a framework for previously established cultural complexes is suggested based on 153 radiocarbon dates. In this new chronological model, the Mesolithic is dated to ~10,000–6500 cal BC; the Neolithic complexes exist at ~6500–3800 cal BC; the Aeneolithic is dated to ~4300–2800 cal BC; the Early Bronze Age spans ~2500–2100 cal BC; and the Late Bronze Age can be preliminary assigned to ~1600–1100 cal BC.

Obsidian provenance studies, conducted in Northeast Asia over the last 40 years, give us solid evidence about human migrations and contacts in prehistory. Active exchange of raw material began at ca. 33,000–25,000 BP in both the mainland and insular parts of this region, and continued afterward. The scale of interaction between Stone Age people was quite large, with distances sometimes greater than 800–1000 km in later prehistory. Seafaring was most probably practiced since this time, as supported by the movement of obsidian across wide (20–40 km) open sea.

The current state-of-the-art of provenance studies for archaeological obsidian on Kamchatka Peninsula is presented. Prehistoric people widely used obsidian as a raw material for making tools, and acquired it from several primary sources. The exact position of seven sources allowed us to understand the general features of obsidian geochemistry and tectonic position of obsidian-bearing volcanic formations. This also made it possible to suggest the localization of seven still unknown sources for archaeological obsidian on Kamchatka. Verification of our preliminary conclusions can be made by fieldwork in selected areas.

The Khayrgas Cave in Yakutia (eastern Siberia) is one of the most important Upper Paleolithic sites in northern Asia, and has been the subject of extensive 14C dating and study of mammal bones. The upper part of the cave sequence (Layers 2–4) dates to the Holocene (~4100–8200 BP), and the lower part (Layers 5–7) to the Late Pleistocene (~13,100–21,500 BP). In Layers 2–4, only extant animal species are known; ecologically they belong to a forest-type ecosystem. In Layers 5–7, several extinct species were identified, and the environment at that time corresponded to open and semi-open ecosystems. The Khayrgas Cave provides rare but reliable evidence of human occupation in the deep continental region of eastern Siberia at the Last Glacial Maximum, ~20,700–21,500 BP.