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Bdella muscorum Ewing, 1909 Distribution in Iran – Alborz Province: Karaj (Ueckermann et al. 2007), East Azerbaijan Province: Region not mentioned (Abbasi et al. 2016). General distribution – United States (California, Colorado, New Mexico, Kansas, Arkansas, Tennessee, Michigan, Minnesota, Illinois, Maryland), Alaska (Point Barrow, Chandler Lake Region, Umiaat, District of Mackenzie), Germany (Ost-Holstein), Czech Republic and Iceland, Georgia, Japan, Bulgaria, Pakistan, Bohemia, Canada, Ukraine, Iran, Slovakia, China, Hungary (Atyeo 1960; Atyeo and Tuxen 1962; Gomelauri 1963; Sosnina et al. 1965; Shiba and Morikawa 1966; Lelláková-Duškova 1978; Tseng 1978; Kuznetsov and Livshits 1979; Chaudhri et al. 1979; Danks 1980; Lehman 1982; Lee et al. 1997; Lin and Zhang 2000; Ripka et al. 2005; Nakamura et al. 2006; Ueckermann et al. 2007; Kaluz 2008; Bednarskaya 2009, 2010, 2011; Abbasi et al. 2016). Collection place(s) – Grain in storage, silos, and flour (Ueckermann et al. 2007; Abbasi et al. 2016)., Published as part of Ebrahimi, Najmeh & Noei, Javad, 2022, Checklist of mites associated with stored products (Arachnida: Acari) of Iran, pp. 559-631 in Persian Journal of Acarology 11 (4) on page 574, DOI: 10.22073/pja.v11i4.75059, http://zenodo.org/record/7447126, {"references":["Ueckermann, A., Rastegar, J., Saboori, A. & Ostovan, H. (2007) Some mites of the superfamily Bdelloidea (Acari: Prostigmata) of Karaj (Iran), with descriptions of two new species and redescription of Bdelloides Kazeruni. Acarologia, 61: 127 - 138.","Abbasi, H., Ardeshir, F. & Bagheri, M. (2016) Fauna of superfamily Bdelloidea and family Cheyletidae mites in grain stocks of East Azerbaijan province, Iran. Proceedings of the 22 nd Iranian Plant Protection Congress, Karaj, Iran, p. 516.","Atyeo, W. T. (1960) A revision of the family Bdellidae in North and Central America (Acarina: Prostigmata). University of Kansas Science Bulletin, 40: 345 - 499.","Atyeo, W. T. & Tuxen, S. L. (1962) The Icelandic Bdellidae (Acarina). Journal of the Kansas Entomological Society, 35 (3): 281 - 298.","Gomelauri, L. A. (1963) On the study of mites of the family Bdellidae in Georgian S. S. R. Bulletin of the Academy of Sciences of the Georgian SSR, 30 (2): 47 - 51.","Sosnina, E. F., Vysotskaya, S. O., Markov, G. N. & Atanasov, L. K. (1965) Predatory mites of the family Bdellidae (Acarina, Prostigmata) from nests of rodents in Bulgaria. Proceedings of the Zoological Institute, USSR Academy of Sciences, 35: 272 - 287.","Shiba, M. & Morikawa, K. (1966) Prostigmatic mites from Japan. II Bdellidae I (Bdellinae, Cytinae, Spinibdellinae). Reports of Research Matsuyama Shinonome Junior College, 3: 17 - 37.","Lellakova-Duskova, F. (1978) Incidence of mites of the family Bdellidae in moss from a spruce wood in SW-Bohemina. Vestnik Ceskoslovenske spolecnosti zoologicke, 42 (1): 23 - 42.","Tseng, Y. H. (1978) Mites of the family Bdellidae from Taiwan (Acarina: Prostigmata). Journal of the Agricultural Association of China, 104: 25 - 51.","Kuznetsov, N. N. & Livshits, I. Z. (1979) Predatory mites of the Nikita Botanical Garden (Acariformes: Bdellidae, Cunaxidae, Camerobiidae). Proceedings of the State Nikita Botanical Garden, 79: 51 - 104.","Chaudhri, W. M., Akbar, S. & Rasool, A. (1979) Studies on the predatory leaf-inhabiting mites of Pakistan. University of Agriculture, Faisalabad, Pakistan, 234 pp.","Danks, H. V. (1980) Arthropods of Polar Bear Pass, Bathurst Island, Arctic Canada. Syllogeus, Ottawa. National Museum of Natural Sciences, National Museums of Canada, 25: 1 - 69.","Lehman, R. D. (1982) Mites (Acari) of Pennsylvania conifers. Transactions of the America Entomological Society, 180: 181 - 286.","Lee, W. K., Lim, J. W. & Lee, S. Y. (1997) A taxonomic study on the family Bdellidae (Bdelloidea: Prostigmata) in Korea. Korean Journal of Soil Zoology, 2 (2): 65 - 75.","Lin, J. Z. & Zhang, Y. X. (2000) Bdelloidea. In: Huang, B. K. (Ed.), Fauna of insects in Fujian Province of China. Fujian Science and Technology Press, Fuzhou, 9: 109 - 121.","Ripka, G., Fain, A., Kazmierski, A., Kreiter, S. & Magowski, W. L. (2005) New data to the knowledge of the mite fauna of Hungary (Acari: Mesostigmata, Prostigmata and Astigmata). Acta Phytopathologica et Entomologica Hungarica, 40: 159 - 176. DOI: 10.1556 / APhyt. 40.200 5.1 - 2.13","Nakamura, Y., Ishikawa, K., Shiba, M., Fujikawa, T., Ono, H., Tamura, H. & Morikawa, K. (2006) Soil animals of the 88 Buddhist temples in Shikoku Island. Memoirs of the Faculty of Argriculture, Ehime University, 51: 25 - 48.","Kaluz, S. (2008) Soil mites (Acari) of the forests in floodplain areas of the rivers Danube and Morava. Peckiana, 5: 89 - 103.","Bednarskaya, E. V. (2009) Population dynamics of bdellides (Acarina, Prostigmata: Bdellidae) in the caves of central part of Mountain Crimea. Optimization and Protection of Ecosystems, 20: 20 - 24.","Bednarskaya, E. V. (2010) Localization of bdellides (Acarina, Prostigmata: Bdellidae) in the caves of central part of Mountain Crimea. Scientific Notes of Taurida V. I. Vernadsky National University. Series: Biology, chemistry, 23 (62): 41 - 45.","Bednarskaya, E. V. (2011) The main biotic relations and localization in microstations of prostigmatic predatory mites in carst cavities of Mountain Crimea. Scientific Notes of Taurida V. I. Vernadsky National University. Series: Biology, Chemistry, 24 (63): 3 - 9."]}

Spinibdella cronini (Baker & Balock, 1944) Distribution in Iran – Guilan Province: Masal, Astaneh, Fuman, Lahijan, Jafarabad, Roodsar (Noei 2007), Razavi Khorasan Province: Region not mentioned (Khaleghabadian et al. 2015), East Azerbaijan Province: Region not mentioned (Abbasi et al. 2016). General distribution – China, Japan, United States (California, Texas, Utah, Colorado, Washington, Alabama, Maryland), Mexico (Tamaulipas, Guerrero, Nevo León, San Luis Potosí), Australia, Bulgaria, Egypt, Syria (Lattakia), Ukraine, United States, Hawaii, Hungary, Iran, Brazil, Slovakia (Atyeo 1960, 1963; Sosnina et al. 1965; Wallace and Mahon 1972; Soliman 1975; Soliman and Zaher 1975; Kuznetsov and Livshits 1979; Lehman 1982; Swift and Goff 1987; Li et al. 1992; Ostovan and Kamali 1995c; Kamali et al. 2001; Jalaeian et al. 2005; Ripka et al. 2005; Li and Fan 2007; Ueckermann et al. 2007; Noei 2007; Kaluz 2008; Noei et al. 2008b; AbbaszadehRad et al. 2010; Daneshnia and Akrami 2013; Majidi and Akrami 2013; Masoudian and Khanjani 2013; Hagstrum et al. 2013). Collection place(s) – Grain in storage, silos, flour, barley, condiment, dried fruit, dried vegetable, mushroom, processed food, wheat, stored rice, rice dust, and debris (Noei 2007; Noei et al. 2008b; Hagstrum et al. 2013; Khaleghabadian et al. 2015; Abbasi et al. 2016)., Published as part of Ebrahimi, Najmeh & Noei, Javad, 2022, Checklist of mites associated with stored products (Arachnida: Acari) of Iran, pp. 559-631 in Persian Journal of Acarology 11 (4) on page 575, DOI: 10.22073/pja.v11i4.75059, http://zenodo.org/record/7447126, {"references":["Khaleghabadian, Z., Sadeghi Namaghi, H., Ardeshir, F., Nemati, A. & Hatefi, S. (2015) Fauna of predatory mosasticgmatic and prostigmatid mites associated with stored food mites in the Mashhad. Journal of Iranian Plant Protection Research, 28 (4): 555 - 564.","Abbasi, H., Ardeshir, F. & Bagheri, M. (2016) Fauna of superfamily Bdelloidea and family Cheyletidae mites in grain stocks of East Azerbaijan province, Iran. Proceedings of the 22 nd Iranian Plant Protection Congress, Karaj, Iran, p. 516.","Atyeo, W. T. (1960) A revision of the family Bdellidae in North and Central America (Acarina: Prostigmata). University of Kansas Science Bulletin, 40: 345 - 499.","Atyeo, W. T. (1963) The Bdellidae (Acarina) of the Australian Realm. Part I: New Zealand, Campbell Island, and the Auckland Islands. pp. 113 - 166; Part II: Australia and Tasmania. pp. 167 - 210. Bulletin of the University of Nebraska State Museum, 4 (8): 113 - 210.","Sosnina, E. F., Vysotskaya, S. O., Markov, G. N. & Atanasov, L. K. (1965) Predatory mites of the family Bdellidae (Acarina, Prostigmata) from nests of rodents in Bulgaria. Proceedings of the Zoological Institute, USSR Academy of Sciences, 35: 272 - 287.","Wallace, M. M. H. & Mahon, J. A. (1972) The taxonomy and biology of Australian Bdellidae (Acari). I. Subfamilies Bdellinae, Spinibdellinae and Cytinae. Acarologia, 14 (4): 544 - 580.","Soliman, Z. R. & Zaher, M. A. (1975) Bdellid mites of Lattakia, Syria. Bulletin de la Societe Royale Entomologique, 59: 73 - 82.","Kuznetsov, N. N. & Livshits, I. Z. (1979) Predatory mites of the Nikita Botanical Garden (Acariformes: Bdellidae, Cunaxidae, Camerobiidae). Proceedings of the State Nikita Botanical Garden, 79: 51 - 104.","Lehman, R. D. (1982) Mites (Acari) of Pennsylvania conifers. Transactions of the America Entomological Society, 180: 181 - 286.","Swift, S. F. & Goff, M. L. (1987) The family Bdellidae (Acari: Prostigmata) in the Hawaiian Islands. International Journal ofAcarology, 13 (1): 29 - 49. DOI: 10.1080 / 01647958708683478","Li, L. S., Xuan, J. Y. & Fan, Q. H. (1992) Taxonomic investigation of food mites in Sichuan Province. Journal of Southeast Agricultural University, 14 (1): 23 - 34.","Ostovan, H. & Kamali, K. (1995 c) Some snout mites (Acari: Bdellidae) from Iran and a key for their identification. Journal of Agricultural Sciences, 1: 29 - 43.","Kamali, K., Ostovan, H. & Atamehr, A. (2001) A catalog of mites and ticks (Acari) of Iran. Islamic Azad University Scientific Publication Center, Tehran, Iran, 192 pp.","Jalaeian, M., Saboori, A. & Seyedoleslami, H. (2005) Prostigmatid mites (Acari: Prostigmata) associated with fruit trees in the Western area of Isfahan. Journal of Entomological Society of Iran, 25 (1): 67 - 68.","Ripka, G., Fain, A., Kazmierski, A., Kreiter, S. & Magowski, W. L. (2005) New data to the knowledge of the mite fauna of Hungary (Acari: Mesostigmata, Prostigmata and Astigmata). Acta Phytopathologica et Entomologica Hungarica, 40: 159 - 176. DOI: 10.1556 / APhyt. 40.200 5.1 - 2.13","Li, L. S. & Fan, Q. H. (2007) A survey of food mites from four provinces of China. Systematic and Applied Acarology, 2: 247 - 250. DOI: 10.11158 / saa. 2.1.38","Ueckermann, A., Rastegar, J., Saboori, A. & Ostovan, H. (2007) Some mites of the superfamily Bdelloidea (Acari: Prostigmata) of Karaj (Iran), with descriptions of two new species and redescription of Bdelloides Kazeruni. Acarologia, 61: 127 - 138.","Kaluz, S. (2008) Soil mites (Acari) of the forests in floodplain areas of the rivers Danube and Morava. Peckiana, 5: 89 - 103.","Noei, J., Hajizadeh, J., Salehi, L. & Ostovan, H. (2008 b) Prostigmatic stored mites of rice in Guilan province. Abstract Book of 18 th Iranian Plant Protection Congress, Hamedan, Iran, p. 278.","Daneshnia, N. & Akrami, M. A. (2013) Mites (Acari) associated with the fig trees (Ficus carica L.) in Estahban (Fars Province), Iran. Persian Journal of Acarology, 2 (3): 539 - 541.","Majidi, M. & Akrami, M. A. (2013) Mites associated with the date palm (Phoenix dactylifera L.) in Larestan (Fars Province), southern Iran. Persian Journal of Acarology, 2 (2): 335 - 339.","Masoudian, F. & Khanjani, M. (2013) Mites associated with some medicinal plants (Asteraceae) in Hamedan, Iran. Journal of Crop Protection, 2 (2): 209 - 218.","Hagstrum, D. W., Klejdysz, T., Subramanyam, B. & Nawrot, J. (2013) Atlas of stored product insect and mites. AACC International Press, USA, 589 pp. DOI: 10.1016 / B 978 - 1 - 891127 - 75 - 5.50009 - 5"]}

Odontoscirus kazeruni (Ostovan & Kamali, 1995) Distribution in Iran – Fars Province: Kazerun (Ostovan 1993; Ostovan and Kamali 1995 c, Hernandes et al. 2016). General distribution – Iran (Ostovan 1993; Ostovan and Kamali 1995 c, Hernandes et al. 2016). Collection place(s) – Stored barley, hay, and wheat (Ostovan 1993; Ostovan and Kamali 1995c)., Published as part of Ebrahimi, Najmeh & Noei, Javad, 2022, Checklist of mites associated with stored products (Arachnida: Acari) of Iran, pp. 559-631 in Persian Journal of Acarology 11 (4) on page 575, DOI: 10.22073/pja.v11i4.75059, http://zenodo.org/record/7447126, {"references":["Ostovan, H. & Kamali, K. (1995 c) Some snout mites (Acari: Bdellidae) from Iran and a key for their identification. Journal of Agricultural Sciences, 1: 29 - 43.","Castilho, R. C., Silva, E. S., De Moraes, G. J. & Halliday, B. (2016) Catalogue of the family Ologamasidae Ryke (Acari: Mesostigmata). Zootaxa, 4197 (1): 1 - 147. DOI: 10.11646 / zootaxa. 4197.1. 1"]}

Cyta latirostris (Hermann, 1804) Distribution in Iran – East Azerbaijan Province: Region not mentioned (Abbasi et al. 2016). General distribution – Cosmopolitan (Canestrini 1886; Berlese 1888; Voigts and Oudemans 1906; Trägårdh 1910; Halbert 1915; Hull 1918; Schweizer 1922; Summerhayes and Elton 1928; Thor 1930, 1931a, b; Womersley 1933; Willman 1939, 1951, 1952, 1956; Schmölzer 1956; Drummond 1957; Mihelčič 1958; Meyer and Ryke 1959; Atyeo 1960, 1963; Gomelauri 1961; Atyeo and Tuxen 1962; Schweizer and Bader 1963; Sosnina et al. 1965; Shiba and Morikawa 1966; Shiba 1969; Wallace and Mahon 1972; Soliman 1975; Soliman and Zaher 1975; LellákováDuškova 1978; Kuznetsov and Livshits 1979; Den Heyer 1981; Lehman 1982; Zaher 1986; Swift and Goff 1987; Michocka 1987; Lee et al. 1997; Lin and Zhang 2000; Kamali et al. 2001; Halliday 2005; Ripka et al. 2005; Nakamura et al. 2006; Ueckermann et al. 2007; Kaluz 2008; AbbaszadehRad et al. 2010; Bednarskaya 2009, 2010, 2011; Abbasi et al. 2016). Collection place(s) – Grain in storage, silos, flour (Abbasi et al. 2016)., Published as part of Ebrahimi, Najmeh & Noei, Javad, 2022, Checklist of mites associated with stored products (Arachnida: Acari) of Iran, pp. 559-631 in Persian Journal of Acarology 11 (4) on pages 574-575, DOI: 10.22073/pja.v11i4.75059, http://zenodo.org/record/7447126, {"references":["Abbasi, H., Ardeshir, F. & Bagheri, M. (2016) Fauna of superfamily Bdelloidea and family Cheyletidae mites in grain stocks of East Azerbaijan province, Iran. Proceedings of the 22 nd Iranian Plant Protection Congress, Karaj, Iran, p. 516.","Canestrini, G. (1886) Prospetto dellʼ Acarofauna Italiana. Part II, Famiglie: Erythraeini, Cheyletini, Bdellini, Eupodini e Analgesini (Padova). pp. 159 - 311.","Berlese, A. (1888) in Berlese, A. and Balzan, A. (1888) Acari Sud-Americani methodice dispositi, descripti, et iconibus illustrati (con tav.), 20: 171 - 222.","Voigts, H. & Oudemans, A. C. (1906) Zur Kenntnis der Milben-Fauna von Bremen. Bdellidae. In: Leuwer, F. (Ed.), Abhandlungen herausgegeben vom Naturwissenschaftlichen Verein zu Bremen, pp. 241 - 242. DOI: 10.5962 / bhl. part. 27528","Tragardh, I. (1910) Acariden aus dem Sarekgebirge. Naturwissenschaftliche Untersuchungen des Sarekgebirges in Schwedisch-Lappland, 4 (4): 477 - 480.","Halbert, M. R. I. A. (1915) Clare Island Survey Acarinida. Section II. terrestrial and marine Acarina. Proceedings of the Royal Irish Academy, 39: 45 - 136.","Hull, J. E. (1918) Terrestrial Acari of the Tyne province. Transactions of the Natural History Society of Northumberland, Durham, and Newcastle-upon-Tyne, 75: 13 - 88.","Schweizer, J. (1922) Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandlungen der Naturferschenden Gesellschaft in Basel, 33: 23 - 111. DOI: 10.5962 / bhl. part. 3644","Summerhayes, V. S. & Elton, C. S. (1928) Further contributions to the ecology of Spitsbergen. III, VI - VII. Journal of Ecology, 16 (2): 193 - 268.","Thor, S. (1930) Beitrage zur kenntnis der invertebraten fauna von Svalbard. Skrifter Svalbard og Ishavet, 27: 1 - 156.","Thor, S. (1931 a) Acarina. Bdellidae, Nicoletiellidae, Cryptognathidae. Das Tierreich, Berlin, 56: 1 - 87.","Womersley, H. (1933) A preliminary account of the Bdellidae (snout mites) of Australia. Transactions and Proceedings of the Royal Society of South Australia, 57: 97 - 107.","Schmolzer, K. (1956) Landmilben aus dem Dauphine (Acarina terrestria). Osterreichische Zoologische Zeitschrift, 6 (3 - 5): 542 - 565.","Drummond, R. O. (1957) Observations on the fluctuations of acarine populations from nests of Peromyscus leucopus. Ecological Monographs, 27 (2): 137 - 152. DOI: 10.2307 / 1948573","Mihelcic, F. (1958) Prostigmata Sudeuropas (Spanien). EOS-Revista Espanola de Entomologia, 34 (3): 269 - 290.","Meyer, M. K. P. S. & Ryke, P. A. J. (1959) Cunaxoidea (Acarina: Prostigmata) occurring on plants in South Africa. Annals and Magazine of Natural History, 13 (2): 369 - 384. DOI: 10.1080 / 002229 35908655745","Atyeo, W. T. (1960) A revision of the family Bdellidae in North and Central America (Acarina: Prostigmata). University of Kansas Science Bulletin, 40: 345 - 499.","Atyeo, W. T. (1963) The Bdellidae (Acarina) of the Australian Realm. Part I: New Zealand, Campbell Island, and the Auckland Islands. pp. 113 - 166; Part II: Australia and Tasmania. pp. 167 - 210. Bulletin of the University of Nebraska State Museum, 4 (8): 113 - 210.","Gomelauri, L. A. (1961) New species of the family Bdellidae. Bulletin of the Academy of Sciences of the Georgian SSR, 26 (1): 68 - 72.","Atyeo, W. T. & Tuxen, S. L. (1962) The Icelandic Bdellidae (Acarina). Journal of the Kansas Entomological Society, 35 (3): 281 - 298.","Schweizer, J. & Bader, C. (1963) Die Landmilben der Schweiz (Mittelland, Jura and Alpen). Trombidiformes Reuter. Denkschriften Schweizerischen Naturforschenden Gesellschaft, 84: 209 - 378.","Sosnina, E. F., Vysotskaya, S. O., Markov, G. N. & Atanasov, L. K. (1965) Predatory mites of the family Bdellidae (Acarina, Prostigmata) from nests of rodents in Bulgaria. Proceedings of the Zoological Institute, USSR Academy of Sciences, 35: 272 - 287.","Shiba, M. & Morikawa, K. (1966) Prostigmatic mites from Japan. II Bdellidae I (Bdellinae, Cytinae, Spinibdellinae). Reports of Research Matsuyama Shinonome Junior College, 3: 17 - 37.","Shiba, M. (1969) Taxonomic investigations on free-living mites in the subalpine forest on Shiga Heights IBP area II. Prostigmata. Bulletin of Natural Sciences of Tokyo, 12 (1): 65 - 115.","Wallace, M. M. H. & Mahon, J. A. (1972) The taxonomy and biology of Australian Bdellidae (Acari). I. Subfamilies Bdellinae, Spinibdellinae and Cytinae. Acarologia, 14 (4): 544 - 580.","Soliman, Z. R. & Zaher, M. A. (1975) Bdellid mites of Lattakia, Syria. Bulletin de la Societe Royale Entomologique, 59: 73 - 82.","Kuznetsov, N. N. & Livshits, I. Z. (1979) Predatory mites of the Nikita Botanical Garden (Acariformes: Bdellidae, Cunaxidae, Camerobiidae). Proceedings of the State Nikita Botanical Garden, 79: 51 - 104.","Den Heyer, J. (1981) The Afrotropical species of Cyta von Heyden (Bdellidae: Actinedida). Phytophylactica, 13: 31 - 41.","Lehman, R. D. (1982) Mites (Acari) of Pennsylvania conifers. Transactions of the America Entomological Society, 180: 181 - 286.","Zaher, M. A. (1986) Survey and ecological studies on phytophagous mites, predacious and soil mites in Egypt. II A. Predacious and non-phytophagous mites. Faculty of Agriculture, Cairo University, Cairo, 559 pp.","Swift, S. F. & Goff, M. L. (1987) The family Bdellidae (Acari: Prostigmata) in the Hawaiian Islands. International Journal ofAcarology, 13 (1): 29 - 49. DOI: 10.1080 / 01647958708683478","Michocka, S. (1987) Polskie Roztocze (Acari) z rodzin Bdellidae i Cunaxidae. Monografie Fauny Polski, 14: 1 - 128.","Lee, W. K., Lim, J. W. & Lee, S. Y. (1997) A taxonomic study on the family Bdellidae (Bdelloidea: Prostigmata) in Korea. Korean Journal of Soil Zoology, 2 (2): 65 - 75.","Lin, J. Z. & Zhang, Y. X. (2000) Bdelloidea. In: Huang, B. K. (Ed.), Fauna of insects in Fujian Province of China. Fujian Science and Technology Press, Fuzhou, 9: 109 - 121.","Kamali, K., Ostovan, H. & Atamehr, A. (2001) A catalog of mites and ticks (Acari) of Iran. Islamic Azad University Scientific Publication Center, Tehran, Iran, 192 pp.","Halliday, R. B. (2005) Predatory mites from crops and pastures in South Africa: potential natural enemies of redlegged earth mite, Halotydeus destructor (Acari: Penthaleidae). Zootaxa, 1079: 11 - 64. DOI: 10.11646 / zootaxa. 1079.1.2","Ripka, G., Fain, A., Kazmierski, A., Kreiter, S. & Magowski, W. L. (2005) New data to the knowledge of the mite fauna of Hungary (Acari: Mesostigmata, Prostigmata and Astigmata). Acta Phytopathologica et Entomologica Hungarica, 40: 159 - 176. DOI: 10.1556 / APhyt. 40.200 5.1 - 2.13","Nakamura, Y., Ishikawa, K., Shiba, M., Fujikawa, T., Ono, H., Tamura, H. & Morikawa, K. (2006) Soil animals of the 88 Buddhist temples in Shikoku Island. Memoirs of the Faculty of Argriculture, Ehime University, 51: 25 - 48.","Ueckermann, A., Rastegar, J., Saboori, A. & Ostovan, H. (2007) Some mites of the superfamily Bdelloidea (Acari: Prostigmata) of Karaj (Iran), with descriptions of two new species and redescription of Bdelloides Kazeruni. Acarologia, 61: 127 - 138.","Kaluz, S. (2008) Soil mites (Acari) of the forests in floodplain areas of the rivers Danube and Morava. Peckiana, 5: 89 - 103.","Bednarskaya, E. V. (2009) Population dynamics of bdellides (Acarina, Prostigmata: Bdellidae) in the caves of central part of Mountain Crimea. Optimization and Protection of Ecosystems, 20: 20 - 24.","Bednarskaya, E. V. (2010) Localization of bdellides (Acarina, Prostigmata: Bdellidae) in the caves of central part of Mountain Crimea. Scientific Notes of Taurida V. I. Vernadsky National University. Series: Biology, chemistry, 23 (62): 41 - 45.","Bednarskaya, E. V. (2011) The main biotic relations and localization in microstations of prostigmatic predatory mites in carst cavities of Mountain Crimea. Scientific Notes of Taurida V. I. Vernadsky National University. Series: Biology, Chemistry, 24 (63): 3 - 9."]}

Biscirus iranensis Paktinat-Saeej & Bagheri, 2015 Distribution in Iran – East Azerbaijan Province: Region not mentioned (Abbasi et al. 2016), Mazandaran Province: Region not mentioned (Hernandes et al. 2016). General distribution – Iran (Abbasi et al. 2016; Hernandes et al. 2016). Collection place(s) – Grain in storage, silos, flour (Abbasi et al. 2016)., Published as part of Ebrahimi, Najmeh & Noei, Javad, 2022, Checklist of mites associated with stored products (Arachnida: Acari) of Iran, pp. 559-631 in Persian Journal of Acarology 11 (4) on page 574, DOI: 10.22073/pja.v11i4.75059, http://zenodo.org/record/7447126, {"references":["Paktinat-Saeej, S., Bagheri, M., Saboori, A. & Ahaniazad, M. (2015) Two new Bdellidae (Trombidiformes: Bdelloidea) from Iran and the status of Neobiscirus Gomelauri, 1963. Zootaxa, 4013 (4): 519 - 530. DOI: 10.11646 / zootaxa. 4013.4. 3","Abbasi, H., Ardeshir, F. & Bagheri, M. (2016) Fauna of superfamily Bdelloidea and family Cheyletidae mites in grain stocks of East Azerbaijan province, Iran. Proceedings of the 22 nd Iranian Plant Protection Congress, Karaj, Iran, p. 516.","Hernandes, F. A., Skvarla, M. J., Fisher, J. R., Dowling, A. P. G., Ochoa, R., Ueckermann, E. A. & Bauchan, G. R. (2016) Catalogue of snout mites (Acariformes: Bdellidae) of the world. Zootaxa, 4152 (1): 1 - 083. DOI: 10.11646 / zootaxa. 4152.1.1"]}

Odontoscirus longirostris (Hermann, 1804) Distribution in Iran – Fars Province: Region not mentioned (Ostovan and Kamali, 1995c). General distribution – Paraguay, Argentina, Brazil, Italy, Ireland, England, Canada (Herschel Island, Yukon Territory, Bering Island, The commander Islands), Alaska, Australia, Northern Bosnia (Balkan caves), Austria, Germany (Wangerooge Island), United States (Iowa, Missouri, Illinois, Indiana, Ohio, Texas, California, Kansas, Arkansas, Florida, Michigan, Montana), Mexico (Michoacán, Jalisco, Oaxaca, Guanajuato, México, Distrito Federal, Puebla, Nuevo León, Guerrero, Tamaulipas), Cuba, Costa Rica, Jamaica, Argentina, Denmark, Japan, Switzerland, Kure Island, Sainte-Hélène Island, Bohemia, Taiwan, Hawaii, Poland, China (Fujian), Iran, Crimea (Canestrini 1886; Berlese 1888; Hull 1915, 1918; Ewing 1917; Banks 1919, 1923; Womersley 1933; Willmann 1941, 1951, 1952; Atyeo 1960, 1977; Ehara 1961; Shiba 1969; Schweizer and Bader 1963; Butler and Usinger 1963; Garret and Haramoto 1967; Lelláková-Duškova 1978; Tseng 1978; Swift and Goff 1987; Michocka 1987; Ostovan and Kamali 1995c; Lin and Zhang 2000; Kamali et al. 2001; Ueckermann et al. 2007. Bednarskaya 2011). Collection place(s) – Stored products (Ostovan and Kamali, 1995c)., Published as part of Ebrahimi, Najmeh & Noei, Javad, 2022, Checklist of mites associated with stored products (Arachnida: Acari) of Iran, pp. 559-631 in Persian Journal of Acarology 11 (4) on page 575, DOI: 10.22073/pja.v11i4.75059, http://zenodo.org/record/7447126, {"references":["Ostovan, H. & Kamali, K. (1995 c) Some snout mites (Acari: Bdellidae) from Iran and a key for their identification. Journal of Agricultural Sciences, 1: 29 - 43.","Canestrini, G. (1886) Prospetto dellʼ Acarofauna Italiana. Part II, Famiglie: Erythraeini, Cheyletini, Bdellini, Eupodini e Analgesini (Padova). pp. 159 - 311.","Berlese, A. (1888) in Berlese, A. and Balzan, A. (1888) Acari Sud-Americani methodice dispositi, descripti, et iconibus illustrati (con tav.), 20: 171 - 222.","Hull, J. E. (1915) Snout mites: an introduction to the British Bdellidae. The Vasculum, 1: 117 - 123.","Hull, J. E. (1918) Terrestrial Acari of the Tyne province. Transactions of the Natural History Society of Northumberland, Durham, and Newcastle-upon-Tyne, 75: 13 - 88.","Ewing, H. E. (1917) New Acarina, part II-Descriptions of new species and varieties from Iowa, Missouri, Illinois, Indiana and Ohio. Bulletin of the American Museum of Natural History, 37 (2): 149 - 172.","Banks, N. (1919) The Acarina collected by the Canadian Arctic Expedition, 1913 - 1918. Reports of the Canadian Arctic Expedition, 3, Part H, pp. 11 - 13.","Banks, N. (1923) Arachnida. In: Trichoptera, Mecoptera, and Arachnida of the Pribilof Islands, Alaska. North American Fauna, 46: 237 - 239.","Womersley, H. (1933) A preliminary account of the Bdellidae (snout mites) of Australia. Transactions and Proceedings of the Royal Society of South Australia, 57: 97 - 107.","Willmann, C. (1941) Die Acari der Hohlen der Balkanhalbinsel (Nach dem Material der \" Biospeologica balcanica \"). Studien aus dem Gebiete der allgemeinen Karstforschung, der wissenschaftlichen Hohlenkunde, der Eiszeitforschung und den Nachbargebieten; 14, B, Biologische Serie. Nr. 8.","Willmann, С. (1951) Die hochalpine Milbenfauna der mittleren Hohen Tauern insbesondere der Groβlockner-Gebietes (Acari). Bonner Zoologische Beitrage, 2: 141 - 176.","Willmann, C. (1952) Die Milbenfauna der Nordseeinsel Wangerooge. Veroffentlichungen des Instituts fur MeeresforschungBremerhaven, 1: 139 - 186.","Atyeo, W. T. (1960) A revision of the family Bdellidae in North and Central America (Acarina: Prostigmata). University of Kansas Science Bulletin, 40: 345 - 499.","Atyeo, W. T. (1977) Family Bdellidae. La Faune terrestre de I´lle de Sainte-Helene. Annales du Musee Royal de l´Afrique Centrale, 8: 300 - 305.","Ehara, S. (1961) Some snout mites from Japan (Acarina: Bdellidae). Publications of the Seto Marine Biological Laboratory, 9 (2): 247 - 263. DOI: 10.5134 / 175293","Shiba, M. (1969) Taxonomic investigations on free-living mites in the subalpine forest on Shiga Heights IBP area II. Prostigmata. Bulletin of Natural Sciences of Tokyo, 12 (1): 65 - 115.","Schweizer, J. & Bader, C. (1963) Die Landmilben der Schweiz (Mittelland, Jura and Alpen). Trombidiformes Reuter. Denkschriften Schweizerischen Naturforschenden Gesellschaft, 84: 209 - 378.","Butler, G. D. & Usinger, R. L. (1963) Insects and other arthropods from Kure Island. Proceedings of the Hawaiian Entomological Society, 18 (2): 237 - 244.","Garret, L. E. & Haramoto, F. H. (1967) A catalogue of Hawaiian Acarina. Proceedings of the Hawaiian Entomological Society, 19 (3): 381 - 414.","Lellakova-Duskova, F. (1978) Incidence of mites of the family Bdellidae in moss from a spruce wood in SW-Bohemina. Vestnik Ceskoslovenske spolecnosti zoologicke, 42 (1): 23 - 42.","Tseng, Y. H. (1978) Mites of the family Bdellidae from Taiwan (Acarina: Prostigmata). Journal of the Agricultural Association of China, 104: 25 - 51.","Swift, S. F. & Goff, M. L. (1987) The family Bdellidae (Acari: Prostigmata) in the Hawaiian Islands. International Journal ofAcarology, 13 (1): 29 - 49. DOI: 10.1080 / 01647958708683478","Michocka, S. (1987) Polskie Roztocze (Acari) z rodzin Bdellidae i Cunaxidae. Monografie Fauny Polski, 14: 1 - 128.","Lin, J. Z. & Zhang, Y. X. (2000) Bdelloidea. In: Huang, B. K. (Ed.), Fauna of insects in Fujian Province of China. Fujian Science and Technology Press, Fuzhou, 9: 109 - 121.","Kamali, K., Ostovan, H. & Atamehr, A. (2001) A catalog of mites and ticks (Acari) of Iran. Islamic Azad University Scientific Publication Center, Tehran, Iran, 192 pp.","Ueckermann, A., Rastegar, J., Saboori, A. & Ostovan, H. (2007) Some mites of the superfamily Bdelloidea (Acari: Prostigmata) of Karaj (Iran), with descriptions of two new species and redescription of Bdelloides Kazeruni. Acarologia, 61: 127 - 138.","Bednarskaya, E. V. (2011) The main biotic relations and localization in microstations of prostigmatic predatory mites in carst cavities of Mountain Crimea. Scientific Notes of Taurida V. I. Vernadsky National University. Series: Biology, Chemistry, 24 (63): 3 - 9."]}

7. Bdellodes iranensis Ueckermann, Rastegar, Saboori & Ostovan, 2007: 129. Paratype male (ARS-20191021-2b, slide code in the original description: SRB-2001-5), Paratype female (ARS- 20191021-2c, slide code in the original description: SRB-2001-2) and Paratype protonymphs (ARS-20191021-2d, e, slide code in the original description: SRB-2001-4 & 5), IRAN, KARAJ, Coll. Jinoos Rastegar. AVAILABLE., Published as part of Saboori, Alireza & Shirvani, Zeinab, 2021, A checklist of Acari type specimens deposited in the Jalal Afshar Zoological Museum, Karaj, Iran, pp. 289-311 in Zootaxa 4949 (2) on page 291, DOI: 10.11646/zootaxa.4949.2.4, http://zenodo.org/record/4636405, {"references":["Ueckermann, E. A., Rastegar, J., Saboori, A. & Ostovan, H. (2007) Some mites of the superfamily Bdelloidea (Acari: Prostigmata) of Karaj (Iran), with redescription of Bdellodes kazeruni. Acarologia, 47 (3 - 4), 127 - 138."]}

Two new species of Bdellidae, collected from China, are described in this paper. Odontoscirus anzhouensis Wu & Guo n. sp. is described based on adult females, characterized by dorsal hypostomal setae (DHS) shorter than vh3，and different leg chaetotaxy/solenidotaxy. Cyta pseudokreiteri Wu & Guo n. sp. is described based on adult females and larvae, characterized by movable digits smooth and fixed digits with one tooth; chelicera with continuous longitudinal striae; setal formula of telofemora I–IV 6–5(6)–5(6)–4.

Bdella longicornis (Linnaeus, 1758) Description, Published as part of Wu, You-Fang, Jin, Dao-Chao, Yi, Tian-Ci, Chen, Jian-Xin & Guo, Jian-Jun, 2020, First description of immature stages and redescription of female Bdella longicornis (Acari: Prostigmata: Bdellidae), with an ontogeny of chaetotaxy, pp. 102-124 in Zootaxa 4900 (1) on page 103, DOI: 10.11646/zootaxa.4900.1.8, http://zenodo.org/record/4409031, {"references":["Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. 10 th Edition. Impensis Direct. Laurentii Salvii, Holmiae (Salvius), (4) + 824 pp. https: // doi. org / 10.5962 / bhl. title. 542"]}

Bdella longicornis (Linnaeus, 1758) is a widespread species. There has been no description of its immature stages before. In this paper, we redescribe its female adult and describe its four immature stages for the first time based on the specimens from P. R. China. Meanwhile, the ontogeny of chaetotaxy in B. longicornis is discussed.

Paktinat-Saeij, Saeid, Rostami, Aylar, Bagheri, Mohammad, Valizadeh, Saeid (2020): Redescription of Odontoscirus iota Atyeo (Acari: Trombidiformes: Bdellidae) from Iran, with a key to the Iranian species of Odontoscirus. Persian Journal of Acarology 9 (4): 303-310, DOI: 10.22073/pja.v9i4.62736

Made available in DSpace on 2020-12-12T01:47:22Z (GMT). No. of bitstreams: 0 Previous issue date: 2019-11-26 Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (CAPES) Fundação de Amparo à Pesquisa do Estado de São Paulo (FAPESP) Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) A catalog of the type specimens of mites deposited at the Mite Collection of the Zoology and Botany Department of São Paulo State University (São José do Rio Preto, São Paulo state, Brazil) is presented. The collection of type specimens includes 120 species of 64 genera and 16 families, most of which from Brazil, but also from Angola, Australia, Costa Rica, Phillipines, South Africa, Sultanate of Oman and Thailand. For each species the original publication, provenance data, specimens conditions are provided. Programa de Pós-Graduação em Ecologia e Recursos Naturais Universidade Federal de São Carlos, Rodovia Washington, Luís, Km 235, Caixa Postal 676 Laboratório de Estudos Subterrâneos Departamento de Ecologia e Biologia Evolutiva Universidade Federal de São Carlos, Via Washington, Luís, km 235, Caixa Postal 676 Departamento de Zoologia e Botânica Instituto de Biociências Letras e Ciências Exatas (IBILCE) Universidade Estadual Paulista (UNESP), Rua Cristóvão Colombo 2265 Departamento de Zoologia e Botânica Instituto de Biociências Letras e Ciências Exatas (IBILCE) Universidade Estadual Paulista (UNESP), Rua Cristóvão Colombo 2265 CAPES: nº1798353 FAPESP: nº2010/13269-0 FAPESP: nº2012/23824-6 FAPESP: nº2015/05315-5 CNPq: nº303435/2013-5

Eghbalian, Amir Hossein, Khanjani, Mohammad (2018): Re-description of Bdella muscorum Ewing (Acari: Bdellidae) from Western Iran. Persian Journal of Acarology 7 (4): 353-361, DOI: 10.22073/pja.v7i4.41008, URL: https://www.mendeley.com/catalogue/69d47cc1-d84b-3284-b197-4c6a2683a2af/

Rostami, Aylar, Bagheri, Mohammad, Jamshidi, Soleyman, Paktinat-Saeij, Saeid (2017): New records of Bdelloidea (Acari: Trombidiformes: Prostigmata) from Iran with a re-description of Spinibdella tadjikistanica Kuznetzov. Persian Journal of Acarology 6 (4): 245-258, DOI: 10.22073/pja.v6i4.30958

Hexabdella persiaensis Paktinat-Saeej & Bagheri, 2014 Hexabdella persiaensis Paktinat-Saeej and Bagheri, 2014: 3. Distribution: Iran (Paktinat-Saeej et al. 2014b). Previous provincial records from Iran: Mazandaran (Paktinat-Saeej et al. 2014b). Material examined: Two females from rotten leaves under apple tree, Chetab village, 18 December 2015, Miyaneh city, East Azerbaijan province, Iran were collected by Aylar Rostami., Published as part of Rostami, Aylar, Bagheri, Mohammad, Jamshidi, Soleyman & Paktinat-Saeij, Saeid, 2017, New records of Bdelloidea (Acari: Trombidiformes: Prostigmata) from Iran with a re-description of Spinibdella tadjikistanica Kuznetzov, pp. 245-258 in Persian Journal of Acarology 6 (4) on page 251, DOI: 10.22073/pja.v6i4.30958, http://zenodo.org/record/4644776

Spinibdella tabarii Paktinat-Saeej & Bagheri, 2015 Spinibdella tabarii Paktinat-Saeej and Bagheri, 2015b: 696. Distribution: Iran (Paktinat-Saeej et al. 2015b). Previous provincial records from Iran: Mazandaran (Paktinat-Saeej et al. 2015b). Material examined: Two females and one nymph from soil and rotten leaves under apple tree, Chetab village, 18 December 2015; one female from soil under apple tree, Zarnagh village, 26 January 2016, Miyaneh city, East Azerbaijan province, Iran were collected by Aylar Rostami., Published as part of Rostami, Aylar, Bagheri, Mohammad, Jamshidi, Soleyman & Paktinat-Saeij, Saeid, 2017, New records of Bdelloidea (Acari: Trombidiformes: Prostigmata) from Iran with a re-description of Spinibdella tadjikistanica Kuznetzov, pp. 245-258 in Persian Journal of Acarology 6 (4) on page 250, DOI: 10.22073/pja.v6i4.30958, http://zenodo.org/record/4644776

Odontoscirus meridionalis Thor, 1931 Biscirus (Biscirus) meridionalis Thor, 1931: 74. Distribution: Cosmopolitan (Hernandes et al. 2016). Previous provincial records from Iran: Alborz (Ueckermann et al. 2007), East Azerbaijan (Bagheri and Paktinat-Saeij 2016), Mazandaran (Paktinat-Saeij et al. 2016b). Material examined: Two females from soil under apple tree, Chetab village, 11 December 2015, Miyaneh city, East Azerbaijan province, Iran were collected by Aylar Rostami., Published as part of Rostami, Aylar, Bagheri, Mohammad, Jamshidi, Soleyman & Paktinat-Saeij, Saeid, 2017, New records of Bdelloidea (Acari: Trombidiformes: Prostigmata) from Iran with a re-description of Spinibdella tadjikistanica Kuznetzov, pp. 245-258 in Persian Journal of Acarology 6 (4) on page 251, DOI: 10.22073/pja.v6i4.30958, http://zenodo.org/record/4644776, {"references":["Thor, S. (1931) Nordafrikanische Bdellidae und Cunaxidae, von Dr. F. Grandjean (Paris) 1931 gesammelt. Zoologischer Anzeiger, 97 (3 - 4): 62 - 79.","Hernandes, F. A., Skvarla, M. J., Fisher, J. R., Dowling, A. P. G., Ochoa, R., Ueckermann, E. A. & Bauchan, G. R. (2016) Catalogue of snout mites (Acariformes: Bdellidae) of the world. Zootaxa, 4152 (1): 1 - 83. http: // dx. doi. org / 10.11646 / zootaxa. 4152.1.1","Ueckermann, E. A., Rastegar, J., Saboori, A. & Ostovan, H. (2007) Some mites of the superfamily Bdelloidea (Acari: Prostigmata) of Karaj (Iran), with descriptions of two new species and redescription of Bdellodes kazeruni. Acarologia, 47: 127 - 138."]}

Odontoscirus lapidaria (Kramer, 1881) Bdella lapidaria Kramer, 1881: 444. Distribution: Cosmopolitan (Hernandes et al. 2016). Previous provincial records from Iran: Fars (Ostovan and Kamali 1995), Chahar Mahal and Bakhtiari, Khuzestan (Kamali et al. 2001). Material examined: Two females from rotten leaves under apple tree and four nymphs from soil, Balesin village, 26 January 2016; one female from rotten leaves under apple tree, Seyyedlar village, 11 December 2015, Miyaneh city, East Azerbaijan province, Iran were collected by Aylar Rostami., Published as part of Rostami, Aylar, Bagheri, Mohammad, Jamshidi, Soleyman & Paktinat-Saeij, Saeid, 2017, New records of Bdelloidea (Acari: Trombidiformes: Prostigmata) from Iran with a re-description of Spinibdella tadjikistanica Kuznetzov, pp. 245-258 in Persian Journal of Acarology 6 (4) on page 251, DOI: 10.22073/pja.v6i4.30958, http://zenodo.org/record/4644776, {"references":["Kramer, P. (1881) Uber Milben. Zeitschrift fur die gesammten Naturwissenschaft, 54: 417 - 452.","Hernandes, F. A., Skvarla, M. J., Fisher, J. R., Dowling, A. P. G., Ochoa, R., Ueckermann, E. A. & Bauchan, G. R. (2016) Catalogue of snout mites (Acariformes: Bdellidae) of the world. Zootaxa, 4152 (1): 1 - 83. http: // dx. doi. org / 10.11646 / zootaxa. 4152.1.1","Ostovan, H. & Kamali, K. (1995) Some snout mites (Acari: Bdellidae) from Iran and a key for their identification. Journal of Agricultural Sciences, 1: 29 - 43."]}

Bdella muscorum Ewing, 1909 Bdella muscorum Ewing, 1909: 124. Distribution: Cosmopolitan (Hernandes et al. 2016). Previous provincial records from Iran: Alborz (Ueckermann et al. 2007), Razavi Khorasan (Paktinat-Saeej et al. 2012), East Azerbaijan (Navaei-Bonab et al. 2013), Tehran (Cheraghali et al. 2013), Mazandaran (Paktinat-Saeij et al. 2016b). Material examined: 10 females and two males from rotten leaves under apple tree, Hafez village, 1 November 2016; 15 females and three males from soil and rotten leaves under apple tree, Turkman Chay village, 11 December 2015, Miyaneh city, East Azerbaijan province, Iran were collected by AylarRostami., Published as part of Rostami, Aylar, Bagheri, Mohammad, Jamshidi, Soleyman & Paktinat-Saeij, Saeid, 2017, New records of Bdelloidea (Acari: Trombidiformes: Prostigmata) from Iran with a re-description of Spinibdella tadjikistanica Kuznetzov, pp. 245-258 in Persian Journal of Acarology 6 (4) on page 250, DOI: 10.22073/pja.v6i4.30958, http://zenodo.org/record/4644776, {"references":["Ewing, H. E. (1909) Three new species of the genus Bdella (mites). Canadian Entomologist, 41 (4): 124 - 125.","Hernandes, F. A., Skvarla, M. J., Fisher, J. R., Dowling, A. P. G., Ochoa, R., Ueckermann, E. A. & Bauchan, G. R. (2016) Catalogue of snout mites (Acariformes: Bdellidae) of the world. Zootaxa, 4152 (1): 1 - 83. http: // dx. doi. org / 10.11646 / zootaxa. 4152.1.1","Ueckermann, E. A., Rastegar, J., Saboori, A. & Ostovan, H. (2007) Some mites of the superfamily Bdelloidea (Acari: Prostigmata) of Karaj (Iran), with descriptions of two new species and redescription of Bdellodes kazeruni. Acarologia, 47: 127 - 138.","Paktinat-Saeej, S., Sadeghi-Namaghi, S., Hosseini, M., Hatefi, S. & Ueckermann, E. A. (2012) Predatory mites of superfamilies Bdelloidea, Erythraeoidea, Raphignathoidea (Acari: Prostigmata) in pomegranate orchards in Mashhad region, Iran. In: Sarafrazi, A., Asef, M. R., Mozhdehi, M., Mozhdehi, M., Solhjouy Fard, S. & Abdollahi, T. (Eds.), Proceedings of the 20 th Iranian Plant Protection Congress, Shiraz, Iran, p. 474.","Navaei-Bonab, R., Kazazi, M. & Ueckermann, E. A. (2013) Bdelloid mites fauna of Marand (Northwest of Iran) and the first report of the genus Rubroscirus Den Heyer from Iran. In: Joharchi, O. & Saboori, A. (Eds.). Abstract of the 2 nd International Persian Congress of Acarology, Karaj, Iran, p. 27.","Cheraghali, Z., Rastegar, J., Sakenin-Chelav, H., Bagheri, M. & Lin, J. (2013) Fauna of bdelloid and raphignathoid mites (Acari: Trombidiformes) in Rodbar Ghasran region (Shemiranat), Tehran Province, Iran. In: Joharchi, O. & Saboori, A. (Eds.). Abstract book of the 2 nd International Persian Congress of Acarology, Karaj, Iran, p. 8."]}

Fossil BDELLIDAE 279. Bdella vetusta Ewing, 1934: 58; Manitoban amber (Cretaceous, ~ 76–79 Ma), Cedar Lake, Manitoba, Canada. Type deposition. ROME. 280. Bdella lata Koch & Berendt, 1854: 872 —not Bdella lata Ewing (1909b); Baltic amber (~44 Ma). 281. Bdella bicincta Menge in Koch & Berendt, 1854: 108 —Baltic amber (~44 Ma). 282. Bdella bombycina Menge in Koch & Berendt, 1854: 108 —Baltic amber (~44 Ma). 283. Bdella obconica Menge in Koch & Berendt, 1854: 108 —Baltic amber (~44 Ma). 284. Bdellidae sp. (Aoki 1974 apud Dunlop et al. 2012).

Cyta von Heyden Cyta von Heyden, 1826: 608; Thor, 1930a: 89; 1931a: 16; Oudemans, 1937: 1232; Meyer & Ryke, 1959: 377; Atyeo, 1960a: 416; 1963a: 121, 177; Soliman & Zaher 1975: 74; Den Heyer, 1981: 32; Michocka, 1987: 72. Amonia Koch, 1836: 7; type species Amonia cruciata Koch, 1836: 7; Berlese, 1893: 43. Ammonia, Koch, 1842: 75. Cytobdella Mihelčič, 1958a: 271; type species Bdella (Cytobdella) cytoides Mihelčič, 1958a by original designation; synonymy according to Atyeo (1963a: 121). Troglobdella Oudemans, 1937: 1228; type species Troglobdella obisium (Gervais, 1841), Scirus obisium Gervais, 1841 by original designation; synonymy according to Atyeo (1960a: 417). Rigibdella Tseng, 1978; type-species Rigibdella ignea Tseng, 1978: 48 by original designation; new synonym. Type-species: Scirus latirostris Hermann, 1804 by original designation. 65. Cyta americana (Banks, 1902b): 171; Washington D.C., USA. Original designation: Ammonia americana Banks. ��� Cyta americana; Banks, 1907: 596; 1915: 25; Hernandes et al., 2011: 808. Remarks. Although Thor (1931a) treated this species as a synonym of C. latirostris, they have different chaetotaxy on the palpfemur (6���7 in the latter species and 4 in C. americana) (Hernandes 2013); also, the proximal setae of the chelicerae is approximately three times the length of the distal setae (2x in C. latirostris). Type deposition. MCZ. 66. Cyta brevipalpa Ewing, 1909b: 73; Ewing & Webster, 1912: 129; under bark on soft maple tree, Illinois, USA. Type deposition. USNM. 67. Cyta coerulipes (Dug��s, 1834): 21; Europe, unknown locality.��� Thor, 1931a: 20; Willmann, 1939c: 431; 1956: 241; 227; Den Heyer, 1981: 32 (ssp.��� quadrisetosus). Original designation: Bdella coerulipes Dug��s. ��� Walckenaer & Gervais, 1844: 156. a. Amonia chlorophana Koch, 1836: 8; Amonia chloropus, Koch, 1842: 76; Bdella chloropus (Koch), Berlese, 1883: 214; synonymy according to Berlese (1891, 1893). b. Amonia leucocephala Koch, 1839: 1; Walckenaer & Gervais, 1847: 531; Oudemans, 1937: 1238; synonymy according to Thor (1931a: 20). c. Bdella crassirostris Kramer, 1881: 442; synonymy according to Thor (1931a). d. Ammonia coerulipes; Berlese, 1893: 44. e. Cyta lutea George, 1912: 236; synonymy according to Thor (1931a). f. Bdella robustirostris Ewing, 1913: 112; synonymy according to Thor (1931a). g. Cyta coeruleipes; Hull, 1918: 37. Distribution. Morocco (Thor 1931b), Austria (Willmann 1951), Spain (Mihelčič 1958b), Panama, Haiti, Cuba, Mexico (San Luis Potos��, Guerrero, Tamaulipas, Hidalgo), United States (California, Colorado, Texas), Alaska (Umiat) and Sudan Bor (Atyeo 1960a), Georgia (Gomelauri 1963b), Switzerland (Schweizer & Bader 1963), Bulgaria (Sosnina et al. 1965), Egypt (Soliman 1975), Syria (Lattakia) (Soliman & Zaher 1975), Ukraine (Kuznetsov & Livshits 1979a), Afrotropical (Natal) (Den Heyer 1981), Poland (Michocka 1987), Brazil (Hernandes et al. 2011), Iran (Beyzavi et al. 2011). Redescriptions. Atyeo (1960a), Michocka (1987). Remarks. There is confusion between the names Amonia chlorophana Koch and A. chloropus Koch���both names are present in Koch (1836) in the contents page and in text, respectively. Type deposition. Unknown. 68. Cyta cytoides (Mihelčič, 1958a): 271 comb. nov.; in a damp cave in brown clay, Guadarrama Mountains, Cercedilla, Spain. Original designation: Bdella (Cytobdella) cytoides Mihelčič. Remarks. Mihelčič (1958a) illustrates a massive and stout chelicera for this species, a key diagnostic feature for the genus Cyta; therefore this species is herein transferred to the latter genus. Type deposition. Laboratorium Faunistica y Ecolog��a Animal, Instituto de Edafolog��a, Madrid. 69. Cyta grandjeani Gomelauri, 1963a: 169; Tbilisi, Georgia, ex soil, lichens, grass, oak and juniper tree. Kuznetsov & Livshits (1979a: 56). Remarks. Cyta veneta grandjeani Lombardini, 1964: 106 is a junior homonym; although the date printed in Lombardini is 1962, the issue appeared only in 1964. Type deposition. Unknown. 70. Cyta flava Mihelčič, 1958a: 270; between Ciempozuelos and Chinch��n, Spain, on soil. Type deposition. Laboratorium Faunistica y Ecolog��a Animal, Instituto de Edafolog��a, Madrid. 71. Cyta ignea (Tseng, 1978): 48 comb. nov.; Taipu, Chiayi Hsien, Taiwan, on litter. Original designation: Rigibdella ignea Tseng. Remarks. The rationale for moving this species to Cyta was given in the discussion before the taxonomic section. Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.). 72. Cyta kauaiensis Swift & Goff, 1987: 38; Kauai I, Hawaii, ex litter in Metrosideros sp. (Myrtaceae) tree hole. Remarks. Female unknown. Type deposition. BPBM. 73. Cyta latirostris (Hermann, 1804): 62; France, ex moss. Original designation: Scirus latirostris Hermann. ��� Oudemans, 1929: 314. Cyta latirostris, Thor, 1902: 160; Oudemans, 1902: 54; Thor, 1904b: 72; B��bler, 1910: 812. Other names: Leptus latirostris, Lamarck, 1838: 63; Ammonia latirostris, Koch, 1839: 3; Koch, 1842: 76; Walckenaer & Gervais, 1847: 531; Berlese, 1891: 4; Bdella latirostris, Walckenaer & Gervais, 1844: 157; 1847: 531; Canestrini & Fanzago, 1877: 107; Berlese, 1883: 214. a. Ammonia cruciata Koch, 1836: 7; Anders��n, 1863: 185; Berlese, 1893: 44; Oudemans, 1937: 1235; Bdella cruciata; Walckenaer & Gervais, 1844: 156 synonymy according to Berlese (1893). b. Ammonia leucocephala Koch, 1839: 23; 1842: 76; Bdella leucocephala (Koch), Canestrini & Fanzago, 1877: 106; synonymy according to Berlese (1893). c. Ammonia megacephala Koch, 1839: 23; 1842: 76; Walckenaer & Gervais, 1847: 531; Berlese, 1893: 44; Oudemans, 1937: 1239; synonymy according to Berlese (1893). d. Bdella phoenicea Koch, 1839: 7; 1842: 74; Walckenaer & Gervais, 1847: 531; Canestrini & Fanzago, 1877: 103; Thor, 1931a: 36; Oudemans, 1937: 1213; synonym of Bdella vulgaris according to Berlese (1893). f. Bdella cruentata Koch, 1839: 10; 1842: 74; Walckenaer & Gervais, 1847: 532; Anders��n, 1863: 185; Canestrini & Fanzago, 1877: 102; Thor, 1931a: 37; Oudemans, 1937: 1192; synonym of Bdella vulgaris according to Berlese (1893). g. Bdella crassirostris Kramer, 1881: 442; synonymy according to Berlese (1891, 1893). h. Bdella dispar Koch, 1839: 23; 1842: 75; Walckenaer & Gervais, 1847: 532; Canestrini & Fanzago, 1877: 101; Thor, 1931a: 31; Oudemans, 1937: 1194; Willmann, 1951a: 148; 1952: 166; Michocka, 1987: 24; synonymy according to Thor (1931a). h. Bdella amarantina Koch, 1839: 17; 1842: 75; Walckenaer & Gervais, 1847: 532; Anders��n, 1863: 185; Berlese, 1893: 42; Oudemans, 1937: 1191; synonymy according to Thor (1931a: 31). i. Bdella tenuirostris Koch, 1839: 18; 1842: 74; Walckenaer & Gervais, 1847: 532; synonym of Bdella vulgaris according to Berlese (1893). j. Bdella vivida Koch, 1839: 19; 1842: 75; Walckenaer & Gervais, 1847: 532; Berlese, 1893: 43; Thor, 1931a: 37; Oudemans, 1937: 1216; synonym of Bdella vulgaris according to Berlese (1893). k. Bdella vulgaris (Hermann 1804: 61), Koch, 1839: 8; 1842: 74; Walckenaer & Gervais, 1847: 531; Berlese, 1888: 187; Hull, 1918: 38. l. Bdella robustirostris Ewing, 1913: 112; synonymy according to Thor (1931a). m. Cyta novangliae Jacot, 1939: 326; synonymy according to Atyeo (1960a). n. Cyta phaseoli Meyer & Ryke, 1959: 377; Den Heyer, 1981: 32; synonymy according to Wallace & Mahon (1972: 571). o. Amonia latirostris var. fusca Canestrini, 1896: 92; Cyta latirostris var. fusca Thor, 1931a: 19. p. Bdella brevirostris molissima Koch, 1879: 132, Cyta brevirostris; Hull, 1922: 622; Banks, 1923: 237; Ammonia brevirostris; Tr��g��rdh, 1900: 15; 1902a: 8; Cyta latirostris var. brevirostris; Tr��g��rdh, 1904: 49; 1910: 480; 1928: 7; synonymy according to Thor (1904b). q. Bdella arenaria Kramer, 1881: 444; Berlese 1893: 42; Karpelles, 1894: 425; synonymy of Bdella vulgaris according to Berlese (1893). Distribution. Paraguay, Brazil (Berlese 1888), Italy (Canestrini 1886), Sweden (Tr��g��rdh 1910), Germany (Voigts & Oudemans 1906; Willmann 1952), Ireland (Halbert 1915), England (Hull 1918), Switzerland (Schweizer 1922), Canada (Summerhayes & Elton 1928), Germany, Norway (Thor 1930a, 1931a), Northern Africa (Thor 1931b), Australia (Womersley 1933a), Madeira Island (Willman 1939a), Austria (Willmann 1951), Germany (Wangerooge Island) (Willmann 1952), Germany (Willmann 1956), France (Schm��lzer 1956), United States (Maryland) (Drummond 1957), Spain (Mihelčič 1958a), Georgia (Gomelauri 1961), South Africa (Meyer & Ryke 1959; Halliday 2005), Cuba, Jamaica, Mexico (Puebla, M��xico, San Luis Potos��), United States (California, Utah, Idaho, Texas, Arkansas, Kansas, Nebraska, North Dakota, Michigan, Alabama, Tennessee, West Virginia, Maryland, Connecticut), Alaska (Point Barrow), Iceland, Italy, Germany and Australia (Atyeo 1960a), Iceland (Atyeo & Tuxen 1962), Switzerland (Schweizer & Bader 1963), Bulgaria (Sosnina et al. 1965), Japan (Shiba & Morikawa 1966; Shiba 1969a; Nakamura et al. 2006), New Zealand, Australia (Victoria, Tasmania, Western Australia) (Atyeo 1963a; Wallace & Mahon 1972), Egypt (Soliman 1975; Zaher 1986), Syria (Lattakia) (Soliman & Zaher 1975), Bohemia (Lell��kov��-Du��kova 1978), Ukraine (Kuznetsov & Livshits 1979a), Afrotropical (Natal, Cape Province, Pietersburg Dist.) (Den Heyer 1981), United States (Lehman 1982), Hawaii (Swift & Goff 1987), Poland (Michocka 1987), Korea (Lee et al. 1997), China (Fujian) (Lin & Zhang 2000), South Africa (Halliday 2005), Hungary (Ripka et al. 2005), Iran (Ostovan & Kamali 1995; Kamali et al. 2001; Ueckermann et al. 2007; Abbaszadeh et al. 2010), Crimea (Bednarskaya 2009, 2010, 2011), Slovakia (Kaluz 2008). Redescriptions. Atyeo (1960a, 1963a), Atyeo & Tuxen (1962), Sosnina et al. (1965), Shiba (1969a), Lell��kov��-Du��kova (1978), Michocka (1987). Although Berlese (1893: 44) has considered Ammonia leucocephala Koch as a synonym of Cyta latirostris, Koch (1839) illustrated three leg trichobothria (on tibia I, tibia IV and tarsus III), while C. latirostris only has a trichobothrium on tibia IV. Type deposition. Unknown (probably lost). Remarks. Species with world-wide distribution; this species bears trichobothria only on tibia IV; Wallace & Mahon (1972: 579) reports an undescribed species of Cyta with no leg trichobothria at all. 74. Cyta leiliae Eghbalian, Khanjani & Ueckermann, 2014: 568; ex soil and litter under Quercus brantii Lindl (Fagaceae), Kurdistan province, Iran. Type deposition. Holotype and paratypes at the Acarology Laboratory, University of Abu-Ali Sina, Hamedan, Iran, one paratype at NCA. 75. Cyta kurdistanicus Eghbalian, Khanjani & Ueckermann, 2014: 572; ex soil and litter under Amygdalus lycioides Spach (Rosaceae), Kurdistan province, Iran. Type deposition. Holotype and paratypes at the Acarology Laboratory, University of Abu-Ali Sina, Hamedan, Iran, one paratype at NCA. 76. Cyta longiseta Wallace & Mahon, 1972: 575; Mt. Claremont, Perth, Australia, on Banksia sp. (Proteaceae) scrub. Distribution. Iran (Abbaszadeh et al. 2010). Type deposition. Holotype and paratypes at ANIC, paratypes at SAM, BMNH, BPBM, OSAL and USNM. 77. Cyta magdalenae Den Heyer, 1981: 34; Kleinmond, Cape Province, South Africa, in soil under dune vegetation. Type deposition. NCA, IZRPU, ZUNIN. 78. Cyta murrayi Den Heyer, 1981: 37; ex litter of Dombeya rotundifolia (Hochst.) (Malvaceae), UNIN, South Africa. Distribution. Afrotropical (Angola, South Africa, Zimbabwe) (Den Heyer 1981). Type deposition. NCA, ZUNIN, ANIC. 79. Cyta reticulata Soliman & Zaher, 1975: 74; Slinfa, Lattakia, Syria, ex moss. Remarks. Male unknown. Type deposition. CUE. 80. Cyta spuria Atyeo, 1960a: 421; 10 miles east of Xilitla, San Luis Potos��, Mexico, ex bamboo; Distribution. Mexico, USA (Texas) (Atyeo 1960a). Type deposition. Holotype and paratypes at SEMC, paratypes at USNM, BMNH and SAM. 81. Cyta troglodyta Hernandes, 2011: 801; ex feces of Kerodon rupestris Wied (Mammalia, Rodentia), Gruta do Janel��o cave, Minas Gerais, Brazil (Hernandes et al. 2011). Type deposition. DZSJRP. 82. Cyta veneta (Lombardini, 1960): 241; Jesolo, Venice, Italy. Original designation: Neomolgus venetus Lombardini. Cyta veneta; Lombardini, 1964: 105. Type deposition. CRA., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on pages 19-22, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["von Heyden, C. (1826) Versuch einer systematischen eintheiling der Acariden. In: Isis von Oken, 18 (6), 608 - 613.","Thor, S. (1930 a) Beitrage zur kenntnis der invertebraten fauna von Svalbard. Skrifter Svalbard og Ishavet, 27, 1 - 156.","Oudemans, A. C. (1937) Kritisch historisch overzicht der Acarologie. Derede gedeelte, 1805 - 1850, Band C, Tarsonemini Can. & Fanz. 1877, Stomatostigmata Oudms. 1906, Eleutherengona Oudms. 1909. 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Transactions of the America Entomological Society, 180, 181 - 286.","Lee, W. - K., Lim, J. - W. & Lee, S. - Y. (1997) A taxonomic study on the family Bdellidae (Bdelloidea: Prostigmata) in Korea. Korean Journal of Soil Zoology, 2 (2), 65 - 75 [in Korean with English abstract]","Lin, J. Z. & Zhang, Y. X. (2000) Bdelloidea. In: Huang, B. K. (Ed.), Fauna of insects in Fujian Province of China. Vol. 9. Fujian Science and Technology Press, Fuzhou, 109 - 121.","Ripka, G., Fain, A., Kazmierski, A., Kreiter, S. & Magowski, W. L. (2005) New data to the knowledge of the mites fauna of Hungary (Acari: Mesostigmata, Prostigmata and Astigmata). Acta Phytopathologica et Entomologica Hungarica, 40, 159 - 176. http: // dx. doi. org / 10.1556 / APhyt. 40.2005.1 - 2.13","Ostovan, H. & Kamali, K. (1995) Some snout mites (Acari: Bdellidae) from Iran and a key for their identification. Journal of Agricultural Sciences, 1, 29 - 43.","Kamali, K., Ostovan, H. & Atamehr, A. (2001) A catalog of mites & ticks (Acari) of Iran. Islamic Azad University Scientific Publication Center. 204 pp.","Bednarskaya, E. V. (2009) Population dynamics of bdellides (Acarina, Prostigmata: Bdellidae) in the caves of central part of Mountain Crimea. Optimization and Protection of Ecosystems, 20, 20 - 24.","Bednarskaya, E. V. (2010) Localization of bdellides (Acarina, Prostigmata: Bdellidae) in the caves of central part of Mountain Crimea. Scientific Notes of Taurida V. I. Vernadsky National University. - Series: Biology, chemistry, 23 (62), 41 - 45.","Bednarskaya, E. V. (2011) The main biotic relations and localization in microstations of prostigmatic predatory mites in carst cavities of Mountain Crimea. Scientific Notes of Taurida V. I. Vernadsky National University. - Series: Biology, chemistry, 24 (63), 3 - 9.","Kaluz, S. (2008) Soil mites (Acari) of the forests in floodplain areas of the rivers Danube and Morava. Peckiana, 5, 89 - 103.","Eghbalian, A. H., Khanjani, M., Safaralizadeh, M. H. & Ueckermann, E. A. (2014) Two new species of Cyta (Acari: Prostigmata: Bdellidae) from Western Iran. Zootaxa, 3847 (4), 567 - 575. http: // dx. doi. org / 10.11646 / zootaxa. 3847.4.6","Lombardini, G. (1960) Acari nuovi XXXVIII. Redia-Giornale di Entomologia, 45, 241 - 243."]}

Hexabdella quercusi Eghbalian, Khanjani, Safaralizadeh and Ueckermann sp. nov. (Figs. 1���10) Diagnosis. Two pairs of eyes present, fixed cheliceral digit as long as movable digit, chelicerae striated, coxa II with three setae, basifemora II and IV with eight and four setae respectively, genu IV with 5 (+ 1 s) setae, solenidotaxy of tibiae I���IV 3 ��� 2 ��� 1 ���0. Female (n= 4). Total body length (including gnathosoma from apex of hypostome to posterior margin of idiosoma) 602 (608���628), body length (excluding gnathosoma) 446 (455���470); width 270 (230���273). Dorsum (Fig. 1). Prodorsum between setae vi and sci with irregularly broken longitudinal striae; prodorsum with two pairs of eyes, diameters of anterior lateral eye 8 (7���8) and posterior lateral eye 5 (4���5). Striae anterior to internal verticals (vi) broken and transverse; all dorsal setae finely serrate; hysterosomal setae not extending to bases of setae next behind (except seta h 1). Dorsum of hysterosoma with fine, continuous and transverse striae between setae (c 1, d 1, e 1, f 1, h 1 and h 2), but striae obliquely longitudinal between setae c 1���2; seta sce the longest and h 2 and f 2 the shortest; hysterosomal region with three cupules (ia, im and ip) at level of setae d, e and f 1 (Fig. 1). Measurements of dorsal setae as follows: vi 98 (88���93), ve 41 (38���39), sce 69 (65���68), sci 130, c 1 47 (50���51), c 2 50 (40���45), d 1 36 (34���38), e 1 35 (33���38), f 1 41 (35���36), f 2 30 (28���30), h 1 47 (43���45), h 2 30 (27���30). Distance between dorsal setae: vi��� vi 78 (73���75); ve���ve 113 (110���113); vi���ve 33 (30���33); ve���sce 40 (38���40); sci���sci 93 (83��� 87); sce���sci 9 (8���10); sci���c 1 72 (68���69); sci���c 2 102 (103���105); c 1 ���c 1 77 (77���78); c 1 ���c 2 75 (67���70); c 1 ���d 1 60 (58���64); d 1 ���d 1 64 (65���68); d 1 ��� e 1 55 (48���51); e 1 ��� e 1 80 (70���83); e 1 ���f 1 55 (50���54); f 1 ���f 1 48 (43���49); f 1 ���f 2 33 (33���38); h 1 ���h 24 (27���28); h 1 ���h 2 28 (20���27); h 2 ���h 2 80 (70���76). Ratio: vi / ve 2.30 (2.40); vi / sce 0.75 (0.68���0.72); sci / sci���sci 1.39 (1.57 ��� 1.48); vi/vi���vi 0.1. 30 (1.30���1.70), c 1 / c 1 ���c 1 0.61 (0.64���0.65), d 1 / d 1 ���d 1 0.6 (0.50���0.58), e 1 / e 1 ��� e 1 0.44 (0.4���0.54), f 1 / f 1 ���f 1 0.85 (0.73���0.81), h 1 / h 1 ���h 1 1.96 (1.59���1.61), h 2 /h 2 ���h 2 0.34 (0.4���0.43), h 1 /h 2 1.74 (1.43���1.50), c 1 ���c 1: d 1 ���d 1: e 1 ���e 1: f 1 ���f 1: 1.60 (1.60���1.80): 1.33 (1.39���1.51): 1.67 (1.63���1.69): 1.00 (1.00). Gnathosoma (Fig. 4). Subcapitulum 155 (148���165) long, width at base 75 (66���78); base of gnathosoma with transverse striae; palp: five���segmented, tibiotarsus with three setae + one solenidion + two long end setae, DES and VES 83 (83) and 108 (106���108) respectively; genu with four setae; telofemur with one seta; basifemur with five setae; trochanter without setae; measurements of palp segments as follows: trochanter 11 (10���12), basifemur 58 (50���53), telofemur 22 (22���23), genu 12 (13���15), tibiotarsus 35 (35���38). Subcapitulum with six pairs of long ventral subcapitular setae (vh 1���6), distal pair (vh 6) 36 (33���34) about 1.5 x length of proximal pair (vh 1) 21 (19���21); two pairs of short adoral setae near tip of hypostome, avs 10 (11) and ad 9 (8���10). Chelicerae 143 (138���141) long, width 42 (36���38), finely striated and with two dorsal setae (ch 1���2), proximal seta 35 (33���35), and distal seta 50 (46���48), respectively, movable chela with one small tooth and fixed chela smooth and attenuate, equal in length to movable chela; distal seta (ch 1) short, not extending to base of chela (Fig. 4). Venter (Fig. 5). Striae between coxae I���II and III��� IV longitudinal and between coxae II���III finely broken and oblique. One pair of setae between coxae IV. Aggenital region with nine pairs of setae (ag 1���9) by genital valves each with nine pairs of setae (g 1���9) (Fig. 5); anal region surrounded with oblique continuous striae and three pairs of smooth setae (ps 1���3): ps 1 33 (33���38), ps 2 23 (23���25) and ps 3 16 (16���18) long (Fig. 5). Ovipositor (Fig. 6). Ovipositor as indicated in Figure 6; tube like, with 18 smooth setae. Legs (Figs. 7���10). Measurements of legs I-IV: I 255 (238���243), II 231 (215���218), III 272 (250���253), IV 343 (315���338). Setal formulae of leg segments (solenidia in parentheses): coxae I���IV: 5 ��� 3���5 (right leg of holotype with 6)��� 2; trochanters I���IV: 1 ��� 1���2 ��� 1; basifemora I���IV: 8 ��� 8 ��� 7 ��� 4; telofemora I���IV: 5 ��� 5 ��� 5 ��� 5; genua I���IV: 4 ts, 1 s ��� 4 ts, 1 s ��� 4 ts, 1 s ��� 5 ts, 1 s; tibiae I���IV: 6 ts, 3 f, 1 tr��� 7 ts, 1 f, 1 bls��� 7 ts, 1 f ��� 6 ts, 1 tr; tarsi I���IV: 18 ts, 4 w, 1 e ��� 16 ts, 2 w, 1 e ��� 17 ts, 1 tr��� 15 ts, 1 w (Figs. 7���10). Male and immatures: Unknown. Habitat. Litter under oak trees. Distribution. Only known from Western Iran. Remarks. Hexabdella quercusi Eghbalian, Khanjani, Safaralizadeh and Ueckermann sp. nov. is closely related to H. mexicana (Baker & Balock, 1944), redescribed by Atyeo (1960), by having: tibiae I, IV and tarsus III, each with one trichobothrium; opisthosomal setae not branched distally; two pairs of eyes; palp basifemur with five setae; solenidotaxy of tibiae I���IV 3 ��� 2 ��� 1 ���0; it however differs in having: 1) striae between setae vi and sci irregularly broken and longitudinal in the former instead of finely and regularly broken in the latter; 2) fixed digit of equal length as movable digit, instead of two thirds the length of movable digit; 3) genital valve with nine pairs of setae in H. quercusi sp. nov. opposed to eight pairs of setae in H. mexicana; 4) coxa II with three setae in new species while four setae in H. mexicana; 5) subcapitular setae smooth instead of finely serrate; 6) basifemora II and IV with eight and four setae respectively versus seven and five; 7) telofemur IV with five setae instead of four setae; 8) genu IV with 5 (+ 1 ��) setae in H. quercusi whereas 4 (+ 1 ��) in the H. mexicana; 9) tibiae II���IV: 7 ts��� 7 ts��� 6 ts vs. 5 ts��� 5 ts��� 5 ts; 10) tarsi I���IV 18 ts��� 16 ts��� 17 ts��� 15 ts opposed to tarsi I���IV: 14 ts��� 10 ts��� 9 ts��� 8 ts. The new species also resembles H. persiaensis Paktinat Saeej, Bagheri, Saboori & Ueckermann, 2014, in having mostly the same leg chaetotaxy, smooth dorsal idiosomal setae and fixed digit of equal length as the movable digit; however it differs by: 1) palpal basifemur with five setae in the former while six setae in the latter; 2) basifemur III with seven setae in the new species whereas six setae in H. persiaensis; 3) genital valve with nine pairs of setae in H. quercusi sp. nov. while eight pairs of setae in H. persiaensis; 4) aggenital region with nine pairs of setae instead of eight; 5) marginal region between dorsal setae d 1 and e 1 with continuous striae in the former but fine broken striae in the latter; 6) marginal regions between legs II and III with broken striae in new species whereas continuous striae in H. persiaensis; 7) aggenital setae ag 1���5 covered in continuous striae in new species but all aggenital seta, ornamented, on broken striae in H. persiaensis; 8) tarsi I, II and IV with 18, 17 and 15 setae vs 19 (20), 16 and 16 setae. Etymology. The specific epithet is derived from Quercus, due to the oak-dominated forest where the new species was collected. Type material. The holotype and three paratype females were collected from soil and litter under oak trees, Quercus brantii Lindl, (Fagaceae), Baneh, Kurdistan Province, Iran, (35 �� 59 ' N, 46 �� 53 ' E, a.s.l. 1540 m), 17 January 2013, Col. Amir Hossein Eghbalian. Holotype and two paratype females (EHBH 15010 and EPBH 15011 ��� 12, respectively) are deposited in the Mites Collection of the Acarology Laboratory, University of Bu���Ali Sina, Hamedan, Iran, and one paratype female (slide specimen) will be deposited in the Mites Collection of Arachnida, Plant Protection Research Institute, Pretoria, South Africa., Published as part of Eghbalian, Amir Hossein, Khanjani, Mohammad, Safaralizadeh, Mohammad Hassan & Ueckermann, Edward A., 2016, New species of Hexabdella and Neomolgus (Acari: Prostigmata: Bdellidae) from Iran, pp. 291-300 in Zootaxa 4072 (2) on pages 292-295, DOI: 10.11646/zootaxa.4072.2.10, http://zenodo.org/record/260608, {"references":["Baker, E. W. & Balock, J. (1944) Mites of the family Bdellidae. Proceedings of the Entomological Society of Washington, 46 (7), 176 - 184.","Atyeo, W. T. (1960) A revision of the mite family Bdellidae in North and Central America (Acarina, Prostigmata). The University of Kansas Science Bulletin, 40, 345 - 499.","Paktinat Saeej, S., Bagheri, M., Saboori, A. & Ueckermann, E. A. (2014) Hexabdella persiaensis sp. nov. (Acari: Prostigmata: Bdellidae) as a first new species of the genus Hexabdella from Asia. International Journal of Acarology, 40 (5), 384 - 389. http: // dx. doi. org / 10.1080 / 01647954.2014.928366"]}

Tetrabdella Hernandes & Feres Type-species: Tetrabdella neotropica Hernandes & Feres, 2006 by original designation. 158. Tetrabdella neotropica Hernandes & Feres, 2006: 60; Cedral, S��o Paulo, Brazil, ex Hevea brasiliensis M.Arg. (Euphorbiaceae). Type deposition. Holotype and paratypes at DZSJRP, paratype at USNM., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on page 30, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["Hernandes, F. A. & Feres, R. J. F. (2006) Tetrabdella neotropica (Acari: Bdellidae), a new genus and species from Brazil. Zootaxa, 1135, 57 - 68."]}

Genus Hexabdella Van der Schyff et al. 2004 Type species: Hexabdella denheyeri Van der Schyff et al. 2004

Neomolgus Oudemans Molgus Dujardin, 1842: 316 (nomen nudum); Molgus Trouessart, 1894: 118; Thor, 1913: 30; 1931a: 54. Molgus (Hoplomolgus) Berlese, 1923: 237. Neomolgus Oudemans, 1937: 1229; Atyeo, 1960a: 389; Soliman, 1975: 48; Soliman & Zaher, 1975: 78; Wallace & Mahon, 1976: 116; Michocka, 1987: 44. Type-species: Acarus littoralis Linnaeus, 1758 by original designation. 251. Neomolgus aegyptiacus Soliman & Mohamed, 1972b: 90; Faculty of Agriculture farm, Giza, Egypt, from clover field associated with Bryobia cristata (Acari, Tetranychidae). Remarks. Male unknown; although the species was formally described in 1975, three years earlier Soliman & Mohamed (1972b) published the biology of this species along with a morphological description of all stages. The authority and date must be therefore attributed to Soliman & Mohamed, 1972b. Type deposition. Probably at CUE. 252. Neomolgus aequalis (Schweizer & Bader, 1963): 238 comb. nov.; Kreuzlingen, Switzerland, ex moss. Original designation: Hoplomolgus aequalis Schweizer & Bader. Type deposition. NMB (A 211a). 253. Neomolgus berlesei (Tr��g��rdh, 1902b): 17 comb. nov.; in humid areas and under stones, Italy. Original designation: Bdella capillata var. berlesei Tr��g��rdh, 1902b: 17. Other name: Molgus (Hoplomolgus) berlesei, Thor, 1931a: 57. a. Molgus (Hoplomolgus) tuberculatus Berlese, 1923: 238 synonymy according to Thor (1931a: 57); Hoplomolgus tuberculatus, Schweizer & Bader, 1963: 242. Type deposition. Unknown. 254. Neomolgus capillatus (Kramer, 1881): 446; Germany���Willmann, 1939: 431; 1956: 243. Original designation: Bdella capillata Kramer; Canestrini, 1886: 188; Berlese, 1891; Michael, 1896: 514; Kramer, 1896: 447; Tr��g��rdh, 1902b: 17; B��bler, 1910: 812; Halbert, 1915: 112; Hull, 1918: 40. Other names: Molgus capillatus, Trouessart, 1888: 619; Thor, 1904b: 77; 1909: 7; Oudemans, 1914: 123; Schweizer, 1922: 79; Thor, 1926: 139; Thor, 1930a: 95; Hoplomolgus capillata, Tr��g��rdh, 1931: 49; Molgus (Hoplomolgus) capillatus, Vitzthum, 1929: 59; Molgus (Molgus) capillatus, Thor, 1931a: 60. Distribution. Germany (Voigts & Oudemans 1906), Ireland (Hull 1915), Germany, Switzerland, Italy, England, Ireland, Norway, Svalbard Islands (Thor 1931a), France (Schweizer 1922), Austria (Willmann 1951), Iceland (Atyeo & Tuxen 1962), Georgia (Gomelauri 1963b), Switzerland (Schweizer & Bader 1963), Australia (south western W.A.) (Wallace & Mahon 1976), Ukraine (Kuznetsov & Livshits 1979a), Poland (Michocka 1987), Iran (Ostovan & Kamali 1995; Kamali et al. 2001). Redescriptions. Atyeo & Tuxen (1962), Wallace & Mahon (1976), Michocka (1987). Type deposition. CRA. 255. Neomolgus clypeatus (Thor, 1930b): 17; Sjus��rn (Lillehammer), Norway.��� Atyeo & Tuxen, 1962: 290. Original designation: Molgus clypeatus Thor. Other name: Molgus (Hoplomolgus) clypeatus, Thor: 1931a: 55. Distribution. Iceland (Atyeo & Tuxen 1962), Syria (Lattakia) (Soliman & Zaher 1975), Australia (Wallace & Mahon 1976), Ukraine (Kuznetsov & Livshits 1979a), Iran (Ostovan & Kamali 1995; Kamali et al. 2001). Redescriptions. Atyeo & Tuxen (1962), Wallace & Mahon (1976). Type deposition. Lost. 256. Neomolgus egregia (Koch, 1839): 23 comb. nov.; in humid meadows, Europe. Original designation: Bdella egregia Koch; Koch, 1842: 74; Walckenaer & Gervais, 1847: 532; Anders��n, 1863: 184; Canestrini & Fanzago, 1877: 101; Oudemans, 1937: 1196. Other name: Molgus egregious, Thor, 1931a: 63. Bdella vulgaris, Canestrini, 1886: 183; Berlese, 1888: 7; 1893: 43; Halbert, 1915: 113; Schweizer & Bader, 1963: 230; synonymy according to Thor (1931a: 63). Bdella longitarsa Karsch, 1881: 37; Molgus egregius var. longitarsa, Thor, 1931a: 63. Distribution. Middle Europe (Germany, France, Italy, Sweden) (Thor 1931a), Japan (Karsch 1881), Ireland (Hull 1915). Type deposition. Unknown; type of B. longitarisa in ZMB. 257. Neomolgus helveticus (Schweizer & Bader, 1963): 239 comb. nov.; Buffalora mountain, National Park, Switzerland, under stone. Original designation: Hoplomolgus helveticus Schweizer & Bader. Type deposition. NMB (NP 1471). 258. Neomolgus iraniensis Eghbalian, Khanjani, Safaralizadeh & Ueckermann, 2016: 296; ex soil and litter under wild almond trees, Amygdalus scoparia L. (Rosaceae), Palangan village, Kamyaran, Kermanshah Province, Iran. Type deposition. Holotype and paratypes at University of Bu-Ali Sina, Hamedan, Iran; paratype female at NCA. 259. Neomolgus lacustris (Hull, 1915): 121 comb. nov.; on soil, England. Original designation: Bdella lacustris Hull.���Hull, 1918: 40. Other name: Molgus (Molgus) lacustris, Thor, 1931a: 62. a. Molgus (Hoplomolgus) sublimis Berlese, 1923: 238 synonymy according to Thor (1931a: 62). Type deposition. Unknown. 260. Neomolgus littoralis (Linnaeus, 1745): 96; unknown locality.��� Atyeo, 1960a: 390. Original designation: Acarus littoralis Linnaeus, 1745: 96; 1758: 618; Hammer, 1775: 158; Fabricius, 1775: 815; 1780: 225; 1781: 493; 1787: 374; M��ller, 1776: 187; Oudemans, 1926b: 117; Johnston, 1845: 229; Oudemans, 1929: 316. Other names: Bdella littoralis, Neuman, 1875: 104; Hull, 1918: 40; 1922: 622; Oudemans, 1926a: 116; Tr��g��rdh, 1904: 46; 1912: 419; Bdella vulgaris var. littoralis Moniez, 1890a: 30, 1890b: 196. a. Bdella littoralis + capillata var. pallipediformis Tr��g��rdh, 1902b: 18; synonymy according to Thor (1931a: 61); Molgus littoralis, Thor, 1904a: 15; 1904b: 78; 1913: 30; 1928: 216; 1930a: 96; 1931a: 61; King, 1914: 138; Berlese, 1923: 237; Tr��g��rdh, 1928: 8; 1931: 47. Tr��g��rdh, 1928: 8; 1931: 47. b. Bdella groenlandica Tr��g��rdh, 1904: 48; Hull, 1922: 622; synonymy according to Thor (1930a). c. Molgus (Molgus) littoralis var. seurati Berlese, 1923: 237; synonymy according to Thor (1931a). d. Acarus petrarum ruber L.; Str��m, 1762: 196; Olafsen & Povelsen, 1772: 607; synonymy according to Thor (1904). e. Acarus basteri Johnston, 1836: 353; Bdella basteri; Michael, 1896: 478; synonymy according to Thor (1904). f. Acarus longicornis, Johnston, 1845: 227; synonymy according to Thor (1904). g. Bdella podurophila White, 1852: 210; synonymy according to Thor (1931a). h. Bdella arctica Thorell, 1871: 698; Kramer, 1897: 79; Tr��g��rdh, 1900: 9; Banks, 1919: 11; Molgus arcticus (Thorell), Trouessart, 1888: 119; Thor, 1902: 162; synonymy according to Thor (1904). i. Bdella marina Packard, 1873: 544; 1884: 828; Banks, 1894: 220; Michael 1896: 479; Banks 1907: 596; 1908: 5; Thor, 1931a: 39; synonymy according to Thor (1904). j. Molgus longicornis, Murray, 1877: 143; synonymy according to Thor (1931a). k. Bdella grandis Kramer, 1879: 131; synonymy according to Tr��g��rdh (1904: 46). l. Bdella villosa Kramer & Neuman, 1883: 525; Michael 1896: 479; Oudemans & Koenike, 1897: 238; Banks, 1899: 348; 1907: 596; Molgus villosa, Trouessart, 1888: 619; synonymy according to Thor (1904). m. Eupalus sanguineus Trouessart, 1888: 753; Bdella sanguinea, Trouessart, 1894: 119; Michael 1896: 478; Tr��g��rdh, 1900: 9; Molgus sanguineus, Trouessart, 1894: 117; synonymy according to Thor (1904). n. Bdella villosa + frigida Banks, 1899: 348; synonymy according to Thor (1931a). Distribution. England (Hull 1918), Canada (Nunavut) (Summerhayes & Elton 1928), Alaska, Hudson Bay Area, Greenland, Iceland, Spitsbergen, Bering Island, Russian Lapland (Kola-Hafvon), Novaya Zemlya (Matochkin Strait), Finland (Atyeo 1960a), Japan (Ehara 1960, 1961, Shiba 1971), Iceland (Atyeo & Tuxen 1962). Redescriptions. Atyeo (1960a), Atyeo & Tuxen (1962). Remarks. Atyeo (1960a: 391) reported specimens from Matochkin Strait lacking trichobothria on tibia IV, and having a long tactile seta in the position normally occupied by the trichobothria. Type deposition. Unknown. 261. Neomolgus longipalpis (Karpelles, 1893): 106 comb. nov.; Maramures, Hungary, at Pop Ivan mountain. Original designation: Bdella longipalpis Karpelles. Other name: Molgus longipalpis, Thor, 1931a: 63. Type deposition. Unknown. 262. Neomolgus longipalpus Kuznetsov, 1984: 774; park of the Nikita botanical garden, Yalta, Crimea, ex litter. Remarks. Male unknown. According to ICZN (article 57.6), the one-letter difference in spelling in relation to the previous species is enough to prevent the homonymy. Type deposition. NBG. 263. Neomolgus lumarius Atyeo, 1962: 292 in Atyeo & Tuxen; H��itindur, Southeast Iceland. Type deposition. Holotype and paratypes at ZMUC; paratypes at USNM, BMNH and SAM. 264. Neomolgus maculatus (Karpelles, 1893): 26 comb. nov.; Sălaj, Hungary. Original designation: Bdella maculata Karpelles. Other names: Molgus maculatus, Thor, 1931a: 64; Molgus maculatus var. pilosa; Thor, 1931a: 64. Type deposition. Unknown. 265. Neomolgus monticola Willmann, 1951: 162; Hohe Tauern, Switzerland. Other name: Hoplomolgus monticola, Schweizer & Bader, 1963: 240. Type deposition. ZSM. 266. Neomolgus mutabilis Atyeo, 1960a: 392; 2 miles south of Galena, Cherokee Co., Kansas, USA, under board. Distribution. United States (Kansas, Texas), Mexico (Distrito Federal) (Atyeo 1960a). Type deposition. Holotype and paratypes at SEMC, paratypes at USNM, BMNH and SAM. 267. Neomolgus obsoletus (Berlese, 1923): 238 comb. nov.; in Germany and Norway. Original designation: Molgus (Hoplomolgus) obsoletus, Thor, 1931a: 58; Hoplomolgus obsoletus, Schweizer & Bader, 1963: 242. Type deposition. CRA. 268. Neomolgus ontakensis Shiba, 1971: 99; Mt. Ontak��, Nagano pref., Japan. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 269. Neomolgus pallipes (Koch, 1879): 131; Siberia, Russia. Original designation: Bdella pallipes Koch; Hull, 1918: 41; Hull, 1922: 622; Neomolgus pallipes; Mihelčič, 1958a; Willmann, 1952: 166. a. Bdella hirta Kramer, 1896: 447; synonymy according to Thor (1931a: 58). b. Bdella basteri Johnston forma pallipes, Tr��g��rdh, 1901: 61; synonymy according to Thor (1931a: 58). c. Bdella capillata var. pallipes, Tr��g��rdh, 1902b: 16; 1904: 47; synonymy according to Thor (1931a: 59). d. Molgus (Molgus) pallipes, Thor, 1931a: 58. e. Molgus capillatus, Thor, 1927: 139; 1930: 95; synonymy according to Thor (1931a: 59); Molgus capillatus var. pallipes, Tr��g��rdh, 1928: 8; synonymy according to Thor (1931a: 59). Distribution. Canada (Summerhayes & Elton 1928), Northern Europe and Asia (Siberia, Novaja, Semlja, Sweden, Norway, Svalbard, England, Borkum Island) (Thor 1931a), Germany (Wangerooge Island) (Willmann 1952), Spain (Mihelčič 1958a). Type deposition. BMNH. 270. Neomolgus paracapillatus Michocka, 1987: 45; Warsaw, Poland, ex decayed grass. Type deposition. In the author���s collection. 271. Neomolgus pratensis Shiba, 1971: 93; Namekawa, Kawauchi-ch��, Ehime pref., Japan. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 272. Neomolgus pygmaeus Shiba, 1969a: 74; Shiga Heights, Japan. Distribution. Japan (Shiba 1969a), China (Lin et al. 2006). Redescription. Shiba (1971). Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 273. Neomolgus raeticus (Schweizer & Bader, 1963): 238 comb. nov.; NationalPark, Mt. Mezdi, Switzerland. Original designation: Hoplomolgus raeticus Schweizer & Bader. Type deposition. NMB (NP 1845). 274. Neomolgus raptor Kuznetsov & Barilo, 1984: 934; near Samarkand (Tchapan-Atha), Uzbekistan, under stones. Type deposition. NBG. 275. Neomolgus reticulatus (Schweizer & Bader, 1963): 241 comb. nov.; S-chanf, Switzerland. Original designation: Hoplomolgus reticulatus Schweizer & Bader. Type deposition. NMB (NP 283). 276. Neomolgus sabulosus Shiba, 1971: 96; Shigenobu River, Matsuyama, Ehime pref., Japan. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 277. Neomolgus seriatus Shiba, 1971: 101; Otanom��su-no-taira, Shigak��gen, Nagano pref., Japan. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 278. Neomolgus thorianus (Berlese, 1923): 239 comb. nov.; Corfu, Greece. Original designation: Molgus (Hoplomolgus) thorianus Berlese. Other name: Molgus (Molgus) thorianus, Thor, 1931a: 62. Distribution. Greece (Corfu Island) (Thor 1931a), Georgia (Gomelauri 1963b). Type deposition. CRA., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on pages 41-45, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["Dujardin, F. (1842) Zoologie. L'Institut - Journal Universel des Sciences et des Societes Savantes en France et a l'Etranger, 454, 316.","Trouessart, E. L. (1894) Note sur une grande espece de Bdelle maritime originaire d'Island. Journal de l'Anatomie et de la Physiologie, 30, 117 - 125.","Thor, S. (1913) Biscirus genus novum: eine neue Bdelliden-Gattung und zwei neue Untergattungen. Zoologischer Anzeiger, 42, 28 - 30.","Berlese, A. (1923) Centuria sesta di Acari nuovi. Redia-Giornale di Entomologia, 15, 1 - 240.","Oudemans, A. C. (1937) Kritisch historisch overzicht der Acarologie. 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R. & Mohamed, M. I. (1972 b) On the development and biology of the predaceous mites, Neomolgus aegyptiacus Sol. (Acarina: Bdellidae). Zeitschrift fur Angewandte Entomologie, 71, 90 - 95. http: // dx. doi. org / 10.1111 / j. 1439 - 0418.1972. tb 01723. x","Schweizer, J. & Bader, C. (1963) Die Landmilben der Schweiz (Mittelland, Jura and Alpen). Trombidiformes Reuter. Denkschriften Schweizerischen Naturforschenden Gesellschaft, 84, 209 - 378.","Tragardh, I. (1902 b) Zur Kenntnis der litoralen Arten der Gattung Bdella Latr. Bihang Till Kongliga Svenska Vetenskaps- Akademien Handlingar, 27 (4), 1 - 24.","Thor, S. (1931 a) Bdellidae, Nicoletiellidae, Cryptognathidae. Das Tierreich, 56, 87 pp.","Kramer, P. (1881) Uber Milben. Zeitschrift fur die gesammten Naturwissenschaft, 54, 417 - 452.","Canestrini, G. (1886) Prospetto dell'acarofauna italiana. 2: Erythraeini, Cheyletini, Bdellini, Eupodini e Analgesini. pp. 159 - 311, Tav. X - XXII [= 10 - 22]. Padova.","Michael, A. D. 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Odontoscirus Thor Odontoscirus Thor, 1913: 29; Thor, 1931a: 52 (as subgenus of Biscirus); type-species Bdella virgulata Canestrini & Fanzago by original designation; Meyer & Ryke, 1959: 379; Atyeo, 1960a: 386; Michocka, 1987: 50; Wallace & Mahon 1976: 67. Bdellodes Oudemans, 1937: 1217; Meyer & Ryke, 1959: 378; Wallace & Mahon, 1976: 67; Michocka, 1987: 56. Type-species: Scirus longirostris Hermann, 1804 by original designation. Scirus Hermann, 1804: 61; Thor, 1931a: 41 (part.). Hoploscirus Thor, 1937: 43; type-species Scirus dubitatus Womersley, 1933a by original designation; synonymy according to Wallace & Mahon (1976: 67). Thoribdella Grandjean, 1938: 4 type-species Biscirus meridionalis Thor, 1931a by original designation; synonymy according to Wallace & Mahon (1976: 67). Octobdellodes Atyeo, 1960a: 407 type-species O. hurdi Atyeo, 1960a; synonymy according to Atyeo (1963a: 118, 125). 159. Odontoscirus affinis (Atyeo, 1963a): 185 comb. nov.; Rottnest Island, Australia. Original designation: Bdellodes (Hoploscirus) affinis Atyeo. Distribution. Australia (Atyeo 1963a; Wallace & Mahon 1976), Andaman Islands (Gupta & Ghosh 1980). Redescriptions. Wallace & Mahon (1976), Gupta (2002). Type deposition. Holotype and paratypes at SAM, paratypes at BMNH, USNM, ANIC. 160. Odontoscirus agrestis (Atyeo, 1963a): 152 comb. nov.; Auckland, New Zealand, ex moss and lichens. Original designation: Bdellodes (Hoploscirus) agrestis Atyeo. Remarks. Male unknown. Type deposition. Holotype and paratype at MONZ, paratype at NZFRI. 161. Odontoscirus alacris (Atyeo, 1963a): 193 comb. nov.; Glen Osmond, South Australia. Original designation: Bdellodes (Hoploscirusu) alacris Atyeo. a. Biscirus symmetricus (part.), Womersley, 1933a: 106 (not Kramer, 1898); synonymy according to Wallace & Mahon (1976: 103). Distribution. Australia (Atyeo, 1963a, Wallace & Mahon, 1976). Redescription. Wallace & Mahon (1976). Type deposition. SAM. 162. Odontoscirus alpinus Atyeo, 1960a: 388; Piegan Pass, Glacier National Park, Montana, USA. Type deposition. SEMC. 163. Odontoscirus amamiensis Shiba, 1985: 81; Southern Japan (Kyushu). Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 164. Odontoscirus ancalae (Atyeo, 1963a): 148 comb. nov.; Wellington, New Zealand, ex leaf mold. Original designation: Bdellodes (Hoploscirus) ancalae Atyeo. Remarks. Male unknown. Type deposition. MONZ. 165. Odontoscirus angustifolius (Gupta, 1991): 221 comb. nov.; Moreh, Manipur, India, ex Canthium angustifolium (Rubiaceae)��� Gupta, 2002: 31. Original designation: Bdella angustifolius Gupta. Bdellodes angustifolius, Gupta, 2002. Remarks. Male unknown. Redescription. Gupta (2002). Type deposition. NZSI. 166. Odontoscirus annona (Tseng, 1978): 33 comb. nov., new emendation; Kuiyin, Tainan Hsien, Taiwan, ex Annona sp. (Annonaceae). Original designation: Bdellodes anona Tseng. Remarks. The etymology of the species refers to the host plant, originally misspelled as Anona sp. [sic] rather than Annona sp.; therefore this species is herein emendated to Odontoscirus annona. Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.). 167. Odontoscirus asiaticus Kuznetsov & Barilo, 1984: 934; near Samarkand, Agalyk village, Uzbekistan, ex under stones on humid soil; 3 males and 1 female paratypes on low grass and under stones in Samarkand region near Zeravshan mountain ridge. Type deposition. NBG. 168. Odontoscirus atro (Gupta, 1991): 220 comb. nov.; Arunachal Pradesh, Siji, India, ex Viburnum atrocyaneum Clarke (Adoxaceae). Bdellodes atro, Gupta, 2002: 32. Original designation: Bdella atro Gupta. Remarks. Male unknown. Redescription. Gupta (2002). Type deposition. NZSI. 169. Odontoscirus atyeoi Michocka, 1987: 54; Tatra, Poland. Type deposition. In the author���s collection. 170. Odontoscirus augusta (Roy & Saha, 2010): 121 comb. nov.; West Bengal, India. Original designation: Bdellodes augusta Roy & Saha. Type deposition. DZCU. 171. Odontoscirus australicus (Womersley, 1933a): 107 comb. nov.; Waroona, Western Australia. Original designation: Biscirus (Biscirus) australicus Womersley. Other name: Bdellodes (Hoploscirus) australica, Atyeo, 1963a: 188 (neotype designation). Redescriptions. Atyeo (1963a), Wallace & Mahon (1976). Type deposition. Neotype female at SAM. 172. Odontoscirus bidentata (Wallace & Mahon, 1976): 94 comb. nov.; Omeo, Australia, in mountain gully. Original designation: Bdellodes bidentata Wallace & Mahon. Type deposition. Holotype and paratypes at ANIC, paratypes at SAM, BPBM and MONZ. 173. Odontoscirus bifurcata (El-Sherif & Bolland, 1993): 2237 comb. nov., from litter near railway in Amsterdam, The Netherlands. Original designation: Bdellodes bifurcata El-Sherif & Bolland. Type deposition. Unknown. 174. Odontoscirus bisetosa (Atyeo, 1960a): 414 comb. nov.; 10 miles west of Tuxtla Gutierrez, Chiapas, Mexico, under rock. Original designation: Bdellodes bisetosa Atyeo. Type deposition. Holotype and paratypes at SEMC, paratypes at USNM, BMNH and SAM. 175. Odontoscirus brevicornis (Cooremann, 1959): 30 comb. nov.; Muotathal, Switzerland, in cave. Original designation: Thoribdella brevicornis Cooremann. Type deposition. IRSN. 176. Odontoscirus bryi (Atyeo, 1963a): 142 comb. nov.; Otago, New Zealand, ex moss on stones. Original designation: Bdellodes (Hoploscirus) bryi Atyeo. Remarks. Male unknown. Type deposition. MONZ. 177. Odontoscirus californica (Banks, 1904b): 366 comb. nov.; Claremont, LA, USA. Original designation: Bdella californica Banks; Banks, 1907: 596; Thoribdella californica (Banks), Atyeo, 1960a: 396; Bdellodes californica, Hernandes, 2013: 65. Bdella magna Ewing, 1913: 123; synonymy according to Atyeo (1960a). Distribution. United States (Los Angeles, California) (Atyeo 1960a). Redescription. Atyeo (1960a). Thor (1931a) has strangely regarded Bdella californica Banks as both synonyms of Bdellodes longirostris and of Biscirus uncinatus (pages 42 and 50, respectively). Type deposition. MCZ. 178. Odontoscirus camellae (Atyeo, 1963a): 145 comb. nov.; Wellington, New Zealand, ex leaf mold. Original designation: Bdellodes (Hoploscirus) camellae Atyeo. Remarks. Female unknown. Type deposition. MONZ. 179. Odontoscirus communis (Atyeo, 1960a): 399 comb. nov.; west slope of Cortez Pass, Mexico, under rock. Original designation: Thoribdella communis Atyeo. Other name: Bdellodes (Hoploscirus) commnunis, Atyeo, 1963a: 188. Type deposition. Holotype and paratypes at SEMC, paratypes at BMNH, USNM and SAM. 180. Odontoscirus conformis (Atyeo, 1963a): 137 comb. nov.; Otago, New Zealand, beaten off ferns. Original designation: Bdellodes (Hoploscirus) conformis Atyeo. Remarks. Male unknown. Type deposition. MONZ. 181. Odontoscirus consanguinea (Atyeo, 1963a): 194 comb. nov.; Busselton, Western Australia. Original designation: Bdellodes (Hoploscirus) consanguinea Atyeo. a. Biscirus symmetricus (part.), Womersley, 1933a: 106 (not Kramer, 1898); synonymy according to Wallace & Mahon (1976: 84). b. Biscirus intermedius (part.) (Thor, 1928: 213); Womersley, 1933a: 104; synonymy according to Wallace & Mahon (1976: 84). Distribution. Australia (Atyeo 1963a), (WA, SA, Victoria, Tasmania, NSW, south eastern Queensland, Kununurra area WA) (Wallace & Mahon 1976). Redescription. Wallace & Mahon (1976). Type deposition. SAM. 182. Odontoscirus copiosa (Atyeo, 1963a): 136 comb. nov.; Otago, New Zealand, under stone. Original designation: Bdellodes (Hoploscirus) copiosa Atyeo. Type deposition. MONZ. 183. Odontoscirus currax (Atyeo, 1963a): 187 comb. nov.; Moount Toolbrunnup, Western Australia, on mealy litter and moss. Original designation: Bdellodes (Hoploscirus) currax Atyeo. Redescription. Wallace & Mahon (1976). Type deposition. SAM. 184. Odontoscirus curvus (Atyeo, 1963a): 140 comb. nov.; Auckland Islands, New Zealand, under stone at high tide mark. Original designation: Bdellodes (Hoploscirus) curvus Atyeo. Redescription. Wallace (1970). Type deposition. MONZ. 185. Odontoscirus dubitatus (Womersley, 1933a): 102 comb. nov.; Mount Nelson, Tasmania, under stone. Bdellodes dubitata, Wallace & Mahon, 1976: 78. Original designation: Scirus dubitatus Womersley. Other name: Bdellodes (Hoploscirus) dubitata, Atyeo, 1963a: 196. Redescriptions. Atyeo (1963a), Wallace & Mahon (1976). Remarks. Male unknown, species known only from the holotype. Type deposition. SAM. 186. Odontoscirus edentata (Halliday, 2005): 22 comb. nov.; Port Elizabeth, Marine Protea Hotel, South Africa. Original designation: Bdellodes edentata Halliday. Remarks. Male unknown. Type deposition. Holotype and paratypes at NCA, paratype at ANIC. 187. Odontoscirus exilicornis (Berlese, 1910): 347 comb. nov.; Cape Town, South Africa. Bdellodes exilicornis, Meyer & Ryke, 1959: 379. Original designation: Bdella exilicornis Berlese. Other name: Scirus exilicornis, Thor, 1931a: 44. Type deposition. Lost (Castagnoli & Pegazzani, 1985). 188. Odontoscirus flexuosa (Atyeo, 1963a): 139 comb. nov.; Campbell Island, New Zealand, under stone. Original designation: Bdellodes (Hoploscirus) flexuosa Atyeo; Atyeo, 1964: 167. Redescriptions. Wallace (1970). Type deposition. MONZ. 189. Odontoscirus furcatus (Shiba, 1969a): 73 comb. nov.; Shiga Heights, Japan. Original designation: Bdellodes (Hoploscirus) furcatus Shiba; Nakamura et al. (2006). Type deposition. Unknown. 190. Odontoscirus georgianensis (Wallace, 1970): 109 comb. nov.; Royal Bay, Moltke Harbor, South Georgia, Antarctica, under moss and rocks. Original designation: Bdellodes (Bdellodes) georgianensis Wallace. Type deposition. Holotype at BPBM, paratypes at ANIC and BMNH. 191. Odontoscirus gleba Chaudhri & Akbar, 1985: 118; 1 mile from W. Hassanabdal, Pakistan. Type deposition. UAF. 192. Odontoscirus graminis (Wallace & Mahon, 1976): 105 comb. nov.; Pine Creek, Northern Territory, Australia, ex grass, Eucalyptus sp. Original designation: Bdellodes graminis Wallace & Mahon. Type deposition. Holotype and paratypes at ANIC, paratypes at SAM, BPBM and MONZ. 193. Odontoscirus grandiflora (Gupta, 1991): 224 comb. nov.; Arunachal Pradesh, Garu, India, ex Thunbergia grandiflora Roxb. (Acanthaceae). Original designation: Bdellodes grandiflora Gupta. Type deposition. NZSI. Remarks. Male unknown. Redescription. Gupta (2002). 194. Odontoscirus gressitti (Atyeo, 1964): 167 comb. nov.; Beeman Camp, Campbell Island, New Zealand, ex moss. Original designation: Bdellodes (Hoploscirus) gressitti Atyeo. Redescriptions. Wallace (1970). Type deposition. Holotype at MONZ; paratypes at BPBM. 195. Odontoscirus guajavae (Chatterjee & Gupta, 2002): 34 (in Gupta 2002) comb. nov.; Kalyani, West Bengal, India, ex Psidium guajava (Myrtaceae). Original designation: Octobdellodes guajavae Chatterjee & Gupta. Type deposition. ZSI. 196. Odontoscirus hadroseta (Wallace & Mahon, 1976): 110 comb. nov.; Nyabing, Western Australia, Australia, on sand plain scrub on roadside. Original designation: Bdellodes hadroseta Wallace & Mahon. a. Biscirus symmetricus (part.), Womersley, 1933a: 106 (not Kramer, 1898); synonymy according to Wallace & Mahon (1976: 110). b. Biscirus intermedius (Thor 1928: 213) (part.), Womersley, 1933a: 104; synonymy according to Wallace & Mahon (1976: 110). c. Biscirus uncinatus (Kramer 1898: 12), Womersley, 1933a: 106; synonymy according to Wallace & Mahon (1976: 110). Type deposition. Holotype and paratypes at ANIC, paratypes at SAM, BPBM and MONZ. 197. Odontoscirus haramotoi (Swift & Goff, 1987): 35 comb. nov.; Koa, Hawaii, ex litter. Original designation: Bdellodes haramotoi Swift & Goff. Remarks. Female unknown. Type deposition. BPBM. 198. Odontoscirus harpax (Atyeo, 1963a): 128 comb. nov.; Woods Point, South Australia. Original designation: Bdellodes (Bdellodes) harpax Atyeo. a Biscirus symmetricus (part.), Womersley, 1933a: 106 (not Kramer, 1898); synonymy according to Wallace & Mahon (1976: 115). b. Biscirus intermedius (Thor 1928: 213) (part.), Womersley, 1933a: 104; synonymy according to Wallace & Mahon (1976: 115). c. Scirus longirostris Hermann (part.), Womersley, 1933a: 101; synonymy according to Wallace & Mahon (1976: 115). Distribution. Australia, New Zealand, Tasmania (Atyeo, 1963a, Wallace & Mahon, 1976). Redescription. Wallace & Mahon (1976). Type deposition. Holotype and paratypes at SAM, paratypes at NZFRI, MONZ, BMNH, USNM. 199. Odontoscirus hessei (Womersley, 1933c): 111 comb. nov.; Stellenbosch C.P., South Africa.��� Meyer & Ryke, 1959: 379. Original designation: Scirus hessei Womersley. Other name: Bdellodes (Bdellodes) hessei, Atyeo, 1963a: 181. Distribution. South Africa (Meyer & Ryke 1959; Atyeo 1963a; Halliday 2005), Australia (Perth, Nedland) (Atyeo 1963a), Western Australia (Wallace & Mahon 1976). Redescriptions. Meyer & Ryke (1959), Atyeo (1963a), Wallace & Mahon (1976). Type deposition. SAM. 200. Odontoscirus hickmani (Womersley, 1933a): 107 comb. nov.; National Park, Tasmania, Australia, under stones.��� Wallace & Mahon, 1976: 81. Original designation: Biscirus (Biscirus) hickmani Womersley. Redescriptions. Atyeo (1963a), Wallace & Mahon (1976). Remarks. Male unknown. Type deposition. SAM. 201. Odontoscirus hospita (Banks, 1916): 224 comb. nov.; Victoria, Australia, with Polyrhachis hexacantha (Insecta, Formicidae).��� Wallace & Mahon, 1976: 91. Original designation: Bdella (Scirus) hospita Banks. Other name: Bdellodes (Hoploscirus) hospita, Atyeo, 1963a: 19. a. Biscirus symmetricus (part.), Womersley, 1933a: 106 (not Kramer, 1898); synonymy according to Wallace & Mahon (1976: 91). b. Biscirus intermedius (Thor 1928: 213) (part.), Womersley, 1933a: 104; synonymy according to Wallace & Mahon (1976: 91). Redescriptions. Atyeo (1963a), Wallace & Mahon (1976). Type deposition. Cotypes at USNM and SAM. 202. Odontoscirus hurdi (Atyeo, 1960a): 408 comb. nov.; Point Barrow, Alaska, USA. Original designation: Octobdellodes hurdi Atyeo. Other name: Bdellodes (Bdellodes) hurdi; Atyeo, 1963a: 125. Type deposition. Holotype and paratypes at SEMC, paratypes at BMNH, USNM, SAM, CAS. 203. Odontoscirus hygrotes (Swift & Goff, 1987): 32 comb. nov.; Oahu I, Maui I, Hawaii. Original designation: Bdellodes hygrotes Swift & Goff. Type deposition. BPBM. 204. Odontoscirus inflata (Wallace & Mahon, 1976): 78 comb. nov.; Mandurah, Western Australia, coastal sanddune vegetation. Original designation: Bdellodes inflata Wallace & Mahon. Type deposition. Holotype and paratypes at ANIC, paratypes at SAM, BPBM and MONZ. 205. Odontoscirus infrequens (Atyeo, 1960a): 410 comb. nov.; Douglas County, Kansas, USA, on shagbark hickory. Original designation: Octobdellodes infrequens Atyeo. Remarks. According to Atyeo (1960a: 411), this species has 7 to 8 pairs of hypostomal setae rather than the usual 6 pairs. Type deposition. Holotype and paratypes at SEMC, paratypes at USNM. 206. Odontoscirus insolita (Atyeo, 1960a): 398 comb. nov.; 2 miles of west Oakville, Napa Co., California, USA, ��� Laurel association���. Original designation: Thoribdella insolita Atyeo. Type deposition. SEMC. 207. Odontoscirus intermedius (Thor, 1928): 213 comb. nov.; Norway. Original designation: Biscirus (Biscirus) intermedius Thor; Thor, 1931a: 48. Other names: Biscirus intermedius, Schweizer & Bader, 1963: 236; Thoribdella intermedius, Grandjean, 1938: 4; Bdellodes (Hoploscirus) intermedius, Atyeo, 1963a: 128. Distribution. Norway (Thor 1931a), Georgia (Gomelauri 1963b), Switzerland (Schweizer & Bader 1963), Ukraine (Kuznetsov & Livshits 1979a). Type deposition. Lost. 208. Odontoscirus, Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on pages 31-41, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["Thor, S. (1913) Biscirus genus novum: eine neue Bdelliden-Gattung und zwei neue Untergattungen. Zoologischer Anzeiger, 42, 28 - 30.","Thor, S. (1931 a) Bdellidae, Nicoletiellidae, Cryptognathidae. Das Tierreich, 56, 87 pp.","Meyer, M. K. P. & Ryke, P. A. J. (1959) Cunaxoidea (Acarina: Prostigmata) occurring on plants in South Africa. Annals and Magazine of Natural History, 13 (2), 369 - 384. http: // dx. doi. org / 10.1080 / 00222935908655745","Atyeo, W. T. 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Section II-terrestrial and marine Acarina. Proceedings of the Royal Irish Academy, 39, 45 - 136.","Atyeo, W. T. (1977) Family Bdellidae. La Faune terrestre de I´lle de Sainte-Helene. Annales du Musee Royal de l´Afrique Centrale, 8, 300 - 305.","Trouessart, E. L. (1888) Note sur les acariens marins recueillis par M. Giard au laboratoire maritime de Wimereux. Comptes rendus hebdomadaires des seances de l'Academie des Sciences, 107, 753 - 755. http: // dx. doi. org / 10.5962 / bhl. part. 4881","Banks, N. (1894) Some new American Acarina. Transactions of the American Entomological Society, 21, 209 - 222.","Ewing, H. E. (1909 b) A systematic and biological study of the Acarina of Illinois. University of Illinois Bulletin, 7 (14), 1 - 120.","Ewing, H. E. (1917) New Acarina, part II-Descriptions of new species and varieties from Iowa, Missouri, Illinois, Indiana and Ohio. Bulletin of the American Museum of Natural History, 37 (2), 149 - 172.","Hartzell, A. (1918) A preliminary list of the Acarina of Iowa. The Proceedings of the Iowa Academy of Sciences, 25, 205 - 212.","Banks, N. (1899) Reports upon the insects, spiders, mites and myriapods collected by Dr. L. Stejneger and Mr. G. E. H. Barrett- Hamilton on the Commander Islands-Arachnida. Reports of the fur-seal investigations, 347 - 351.","Tragardh, I. (1904) Monographie der arktischen Acariden. Fauna Arctica, 41, 1 - 77.","Koch, C. L. (1835 - 1844) Deutschlands Crustaceen, Myriapoden und Arachniden. Ein Beitrag zur deutschen Fauna. Herausgegeben von Dr. Herrich-Schaffer. Regensburg, Friedrich Pustet. 40 parts.","Koch, C. L. (1842) Ubersicht des Arachnidensystems. Familie II: Schabelmilben Bdellides, pp. 73 - 77.","Walckenaer, C. A. & Gervais, M. P. (1844) Histoire naturelle des insectes apteres III. 1 - 476 pp.","Gervais, H. P. (1841) Note sur quelques especes de l´ordre des Acariens. Annales des Sciences Naturelles. Zoologie et Biologie Animale, 15, 5 - 10.","Murray, A. (1877) Economic Entomology. Aptera. Chapman and Hall. London. 433 pp.","Hull, J. E. (1918) Terrestrial Acari of the Tyne province. Transactions of the Natural History Society of Northumberland, Durham, and Newcastle-upon-Tyne, 75, 13 - 88.","Cambridge, M. A. (1876) On a new order and some new genera of Arachnida from Kerguelen's Land. Proceedings of the Scientific Meetings of the Zoological Society of London, 1, 258 - 264.","Berlese, A. (1888) in Berlese, A. & Balzan, A. (1888) Acari Sud-Americani methodice dispositi, descripti, et iconibus illustrati (con tav.), 20, 171 - 222.","Canestrini, G. (1886) Prospetto dell'acarofauna italiana. 2: Erythraeini, Cheyletini, Bdellini, Eupodini e Analgesini. pp. 159 - 311, Tav. X - XXII [= 10 - 22]. Padova.","Hull, J. E. (1915) Snout mites: an introduction to the British Bdellidae. The Vasculum, 1, 117 - 123.","Banks, N. (1919) The Acarina collected by the Canadian Arctic Expedition, 1913 - 1918. Reports of the Canadian Arctic Expedition, 3, Part H, 11 - 13.","Banks, N. (1923) Arachnida. In: Trichoptera, Mecoptera, and Arachnida of the Pribilof Islands, Alaska. North American Fauna, 46, 237 - 239.","Willmann, С. (1951) Die hochalpine Milbenfauna der mittleren Hohen Tauern insbesondere der Groβlockner-Gebietes (Acari). Bonner Zoologische Beitrage, 2, 141 - 176.","Butler, G. D. & Usinger, R. L. (1963) Insects and other arthropods from Kure Island. Proceedings of the Hawaiian Entomological Society, 18 (2), 237 - 244.","Lellakova-Duskova, F. (1978) Incidence of mites of the family Bdellidae in moss from a spruce wood in SW-Bohemina. Vestnik Ceskoslovenske spolecnosti zoologicke, 42 (1), 23 - 42.","Garret, L. E. & Haramoto, F. H. (1967) A catalogue of Hawaiian Acarina. Proceedings of the Hawaiian Entomological Society, 19 (3), 381 - 414.","Bednarskaya, E. V. (2011) The main biotic relations and localization in microstations of prostigmatic predatory mites in carst cavities of Mountain Crimea. Scientific Notes of Taurida V. I. Vernadsky National University. - Series: Biology, chemistry, 24 (63), 3 - 9.","Atyeo, W. T. (1963 b) New species and records of Bdellidae from Macquarie and the Auckland Islands (Acarina). Pacific Insects, 5 (2), 445 - 450.","Shiba, M. (1978) Taxonomic investigation on free-living Prostigmata from the Malay Peninsula. Nature and Life in Southeast Asia, 7, 83 - 229.","Thor, S. (1931 b) Nordafrikanische Bdellidae und Cunaxidae, von Dr. F. Grandjean (Paris) 1931 gesammelt. Zoologischer Anzeiger, 97 (3 - 4), 62 - 79.","Kaluz, S. (2008) Soil mites (Acari) of the forests in floodplain areas of the rivers Danube and Morava. Peckiana, 5, 89 - 103.","Kaluz, S., Ferencik, J. & Vrabek, M. (2013) Study sites influenced by natural and human impacts in Tanap and their acarofauna. Entomofauna Carpathica, 25 (1), 1 - 12.","Berlese, A. (1923) Centuria sesta di Acari nuovi. Redia-Giornale di Entomologia, 15, 1 - 240.","Atyeo, W. T. (1960 b) A unique species of Thoribdella Grandjean, 1938, from New Zealand (Acarina: Bdellidae). Records of the Dominion Museum, 3, 289 - 291.","Thor, S. (1902) On the systematic representation of the Acarinen familien Bdellidae Koch, 1842, Grube, 1859, Eupodidae Koch, 1842 and Cunaxidae Sig Thor, 1902. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 52, 159 - 165."]}

Monotrichobdella Baker & Balock Monotrichobdella Baker & Balock, 1944: 176. Type-species: Monotrichobdella maxosburni Baker & Balock, 1944 by original designation. 120. Monotrichobdella maxosburni Baker & Balock, 1944: 176; near Tres Cumbres, Mexico-Cuernavaca Highway, Morelos, Mexico, ex lichens. Distribution. Mexico (Morelos, Salazar, Distrito Federal) (Baker & Balock 1944; Atyeo 1960a), Taiwan (Tseng 1978). Redescriptions. Atyeo (1960a), Tseng (1978). Type deposition. USNM.

Genus Neomolgus Oudemans, 1937 Type species: Bdella capillata Berlese 1981, Published as part of Eghbalian, Amir Hossein, Khanjani, Mohammad, Safaralizadeh, Mohammad Hassan & Ueckermann, Edward A., 2016, New species of Hexabdella and Neomolgus (Acari: Prostigmata: Bdellidae) from Iran, pp. 291-300 in Zootaxa 4072 (2) on page 295, DOI: 10.11646/zootaxa.4072.2.10, http://zenodo.org/record/260608, {"references":["Oudemans, A. C. (1937) Kritisch Histrorisch Overzicht der Acarologie. Derde Gedeelte Band C. 3 (part C), 1179 - 1248. [E. J. Brill, Leide]"]}

Polytrichus van Der Schyff, Theron & Ueckermann Type-species: Polytrichus yemenensis van Der Schyff, Theron & Ueckermann, 2003 by original designation. 87. Polytrichus yemenensis van Der Schyff, Theron & Ueckermann, 2003: 21; Djebel An-Nabi Shuaib, Yemen. Type deposition. NCA., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on page 23, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["van Der Schyff, J., Theron, P. D. & Ueckermann, E. A. (2003) Polytrichinae, a new subfamily of Bdellidae (Acari: Prostigmata) from the Afrotropical region. African Plant Protection, 9 (1), 19 - 22."]}

Neomolgus iraniensis Eghbalian, Khanjani, Safaralizadeh and Ueckermann sp. nov. (Figs. 11���19) Diagnosis. Each chelicera with 21 setae, palp basifemur with seven setae, palp tibiotarsus with 14 setae, genital plate with ten pairs of setae, and aggenital region with three pairs of setae. Female (n= 3). Total body length (including gnathosoma from apex of hypostome to posterior margin of idiosoma) 2025 (1750���1813), body length (excluding gnathosoma) 1500 (1275���1350); width 1000 (862���950). Dorsum (Fig. 11). Dorsum with irregularly broken striae; prodorsum with two pairs of eyes, diameters of anterior eye 50 (40���45) and posterolateral eye 40 (33���38). Striae between setae vi transverse; dorsal setae (c 1 -h 2) smooth, prodorsal setae vi, sci and sce broken on three collected specimens, eyes separated by distance ~ 2 (2���2.4) times diameter of anterior pair eyes, interval between eyes with oblique and longitudinal striae; hysterosomal setae not extending to bases of setae next behind. Between hysterosomal setae (c 1, d 1, e 1) fine, longitudinal and transversal broken striae; between setae f 1 longitudinal and setae h 1-2 transverse striae; between setae c 1���2 with oblique striae; hysterosomal region with three cupules (ia, im and ip) at level of setae d 1, e 1 and f 1 (Fig. 11). Measurements of dorsal setae: vi, sce and sci broken, c 1 70 (70���90), c 2 65 (65���88), d 1 55 (55���88), e 1 63 (63���88), f 1 58 (75), f 2 53 (53���75), h 1 63 (63���80), h 2 63 (63���88). Distance between dorsal setae: vi��� vi 84 (63���69); vi���sce 200 (200���240); sci���sci 185 (185���200); sce���sci 35 (38���48); sce���sce 200 (200); sci���c 1 270 (277���288); sci���c 2 338 (200���350); c 1 ���c 1 375 (180���375); c 1 ���c 2 136 (120���140); c 1 ���d 1 195 (175); d 1 ���d 1 312 (185); d 1 ���e 1 200 (185); e 1 ��� e 1 195 (175); e 1 ���f 1 140 (120); f 1 ���f 1 64 (80); f 1 ���f 2 80 (88); f 2 ���f 2 176 (170); f 1 ���h 1 176 (170); h 1 ���h 1 65 (63); h 1 ���h 2 100 (123); h 2 ���h 2 128 (138). Gnathosoma (Fig. 12). Subcapitulum 425 (462���500) long, width at base 225 (200���250); base of subcapitulum with broken and transverse striae, hypostome smooth; palp five���segmented, palp tibiotarsus with 14 setae [DES and VES 203 (215) and 173 (200) respectively]; genu with four setae; telofemur with one seta; basifemur with seven setae; trochanter lacking setae; measurements of palp segments: trochanter 15 (13), basifemur 290 (355), telofemur 45 (55), genu 55 (53), tibiotarsus 305 (310). Subcapitulum with six pairs of setae (vh 1���6), distal pair (vh 6) 75 (88) more than twice length of proximal pair (vh 1) 33 (35); two pairs of short adoral setae near tip of hypostome, avs 28 (30) and ad 19 (21). Cheliceral length 443 (425���433), width 95 (100���105), with 21 smooth setae (ch 1���21), distal setae [ch 1] 98 (75) and proximal setae [ch 21] 38 (50), and; movable digit longer than fixed digit, movable chelae with two teeth and fixed digit smooth; distal seta (ch 1) long extending to base of chela (Fig. 12). Venter (Fig. 13). Aggenital region with three pairs of setae (ag 1���3), each genital plate with ten pairs of setae (g 1���10) (Fig. 13); anal region with three pairs of smooth setae (ps 1���3): ps 1 65, ps 2 63 and ps 3 broken. Ovipositor (Figs. 14-15) Ovipositor with 16 smooth setae and ovipositor gland as depicted in Figs 14-15. Legs (Figs. 16-19). Measurements of leg segments: I 1075 (1050), II 1050 (1115), III 1150 (1125), IV 1350 (1235). Setal formulae of leg segments: coxae I���IV: 5 ��� 3���4 ��� 2; trochanters I���IV: 1 ��� 1 ��� 1 ��� 1; basifemora I���IV: 11 ��� 10 ��� 10 ��� 6; telofemora I���IV: 7 ��� 7 ��� 6 ��� 6; genua I���IV: 7 ts, 11 s ��� 6 ts, 10 s ��� 5 ts��� 7 ts, 3 s; tibiae I��� 16 ts, 16 f, 1 e, 1 tr��� 15 ts, 5 f, 1 tr��� 17 ts, 1 f ��� 15 ts, 1 f, 1 tr; tarsi I���IV: 40 ts, 1 dt, 9 w, 1 e ��� 41 ts, 1 dt, 3 w ��� 41 ts, 1 tr��� 45 ts, 1 tr (Figs. 16-19). Remarks. Neomolgus iraniensis Eghbalian, Khanjani, Safaralizadeh and Ueckermann sp. nov. is closely related to N. pygmaeus Shiba (1969) in having four setae on the papal genu, coxa I with five setae and telofemora II-IV with 7 ��� 6 ��� 6 setae but differs from the latter in having: 1) Dorsal setae smooth in N. iraniensis opposed to plumose in N. pygmaeus; 2) each chelicera with 21 setae in new species while eight setae in N. pygmaeus; 3) papal basifemur with seven setae in the former while four setae in the latter; 4) palp tibiotarsus with 14 setae in the new species opposed to nine setae in the latter; 5) each genital plate with ten setae vs eight setae; 6) Coxae II-IV with 3��� 4 ��� 2 vs 4 ��� 3 ��� 3. Neomolgus iraniensis Eghbalian, Khanjani, Safaralizadeh and Ueckermann sp. nov. is closely related to N. longipalpus Kuznetsov (1984) in having 7 ��� 6 ��� 6 setae on telofemora II-IV and coxa III-IV with 4 ��� 2 setae but differs from the latter in having: 1) each chelicera with 21 setae in the new species instead of 16 (17) setae in N. longipalpus; 2) palp basifemur with seven setae versus 19 setae; 3) palp genu with four setae versus ten setae; 4) palp tibiotarsus with 14 setae versus 27 setae; 5) coxa I-II with 5 ��� 3 setae versus 6 ��� 4 setae; 6) basifemura I-IV with 11 ��� 10 ��� 10 ��� 6 setae versus 22 ��� 17 ��� 15 ��� 8 setae; 7) telofemur I with seven setae instead of eight setae. Etymology. The specific epithet is derived from Iran, due to the country where the new species was collected. Type material. The holotype and two paratype females were collected from soil and litter under wild almond trees, Amygdalus scoparia L. (Rosaceae), "Sirvan river bank", located in Palangan village, Kamyaran, Kermanshah Province, Iran, (35 �� 03.7' N, 46 �� 35.97 ' E, a.s.l. 864 m), 14 IV 2013, by Amir Hossein Eghbalian. The holotype and one paratype females (EHBN 15014 and EPBH 15015, respectively) are deposited in the Mites Collection of the Acarology Laboratory, University of Bu���Ali Sina, Hamedan, Iran, and one paratype female slide will be deposited in the National Collection of Arachnida, Plant Protection Research Institute, Pretoria, South Africa., Published as part of Eghbalian, Amir Hossein, Khanjani, Mohammad, Safaralizadeh, Mohammad Hassan & Ueckermann, Edward A., 2016, New species of Hexabdella and Neomolgus (Acari: Prostigmata: Bdellidae) from Iran, pp. 291-300 in Zootaxa 4072 (2) on pages 296-298, DOI: 10.11646/zootaxa.4072.2.10, http://zenodo.org/record/260608, {"references":["Shiba, M. (1969) Taxonomic investigation on free-living mite in the subalpine forest on Shiga Heights IBP area, II Prostigmata. Bulletin of the National Science Museum, Tokyo, 12 (1), 65 - 115."]}

Bdella Latreille Bdella Latreille, 1795: 18; 1810: 133; Berlese, 1893: 42; Oudemans, 1929: 303; Meyer & Ryke, 1959: 373; Atyeo, 1960a: 372, 1963a: 120, 170; Soliman & Zaher, 1975: 79; Chaudhri et al., 1979: 130; van Der Schyff et al., 2005: 222. nec Bdella Savigny, 1822 (Anellida: Hirudinea) junior homonym. Chelifer Geoffroy, 1762: 617 (part.); type species Acarus cancroides Linnaeus, 1762 (Pseudoscorpiones). Scirus Hermann, 1804: 60 (part.) type species Scirus vulgaris Hermann, 1804: 61. Bdellidium Oudemans, 1929: 449; type species Scirus vulgaris Hermann, 1804: 61; synonymy according to Atyeo (1960a). Caenobdella Oudemans, 1937: 1227; type-species Bdella crassipes Koch, 1839 by original designation; synonymy according to Atyeo (1960a). Type-species: Acarus longicornis Linnaeus, 1758 (= La pince Geoffroy), by subsequent designation (Latreille 1795). 1. Bdella aloios van Der Schyff, Theron & Ueckermann, 2005: 222; Limpopo province, South Africa, ex Canthium mundianum Cham. & Schltdl. (Rubiaceae). Remarks. Male unknown. This species was misspelled as B. alois in van Der Schyff et al. 2005, pages 223, 224. Type deposition. NCA. 2. Bdella bakeri Gupta & Paul, 1985: 14; West Bengal, Midnapur Dist., Patharkumkumi, India; ex nest of dove Streptopelia chinensis (Scopoli) (Aves, Columbidae). Remarks. Male unknown. Type deposition. NZSI. 3. Bdella biroi Supino, 1894: 197; Hungary.��� Thor, 1931a: 37. Type deposition. Unknown. 4. Bdella boskopensis van Der Schyff, Theron & Ueckermann, 2005: 232; North West Province, South Africa; ex Acacia karroo Hayne (Fabaceae). Type deposition. NCA. 5. Bdella calva Hull, 1915: 122; on soil, England.��� Hull, 1918: 39. Remarks. This species probably belongs to a different genus as indicated by the elongated palp tibiotarsus (Hull 1915, figure 6). Type deposition. Unknown. 6. Bdella captiosa Atyeo, 1963a: 170; Groverly, Queensland, Australia; ex unknown host. Distribution. Australia (Wallace & Mahon 1972), Hawaii (Swift & Goff 1987), Iran (Kamali et al. 2001; Ueckermann et al. 2007; Abbaszadeh et al. 2010). Remarks. Although Atyeo (1963a: 171) established a male as the holotype, Wallace & Mahon (1972: 546, footnote) analyzed the types and concluded it was actually a female; the male and other immatures were not reported until 1987 by Swift & Goff (1987: 43). Redescriptions. Swift & Goff (1987), Ueckermann et al. (2007). Type deposition. SAM. 7. Bdella cardinalis Banks, 1894: 219; ex wood under leaves, NY, USA; 1904a: 144; 1907: 596; 1908: 5; Ewing, 1909b: 68; Ewing & Webster, 1912: 129; Hartzell, 1918: 206; Hernandes, 2013: 61. Distribution. United States (Alabama, Florida, Texas, Illinois) and Mexico. Type deposition. MCZ. Remarks. This species has been considered as a junior synonym of Bdella longicornis (Linnaeus) by Thor (1931a), and as a junior synonym of Bdella oblonga by Jacot (1938); however, Hernandes (2013) revalidated the status of this species. 8. Bdella carolae van Der Schyff, Theron & Ueckermann, 2005: 235; Limpopo Province, South Africa; ex Acacia karroo Hayne (Fabaceae). Type deposition. NCA. 9. Bdella consobrinae van Der Schyff, Theron & Ueckermann, 2005: 234; North West Province, South Africa; ex Acacia karroo Hayne (Fabaceae). Type deposition. NCA. 10. Bdella crassipes Koch, 1839: 23; Germany.��� Koch, 1842: 74; Walckenaer & Gervais, 1847: 532; Thor, 1931a: 31; Caenobdella crassipes (Koch), Oudemans, 1937: 1227. Type deposition. Unknown. 11. Bdella distincta Baker & Balock, 1944: 179; Morelos, Mexico, ex lichens.��� Atyeo, 1960a: 381 (lectotype designation). Distribution. United States (Washington D.C., Hawaii, Texas), Puerto Rico (Guayama), Philippines, Indonesia, Japan, China, Indonesia, Thailand (Atyeo 1960a), Taiwan (Tseng 1978), Hawaii (Garret & Haramoto 1967; Swift & Goff 1987), Guadaloupe (Flechtmann & Etienne 2006), Brazil (Lawson-Balagbo et al. 2008). Redescriptions. Atyeo (1960a), Tseng (1978), Swift & Goff (1987). Type deposition. Lectotype female, ex Bambusa parvariabilis, China (detected in Washington DC, USA), at USNM. 12. Bdella distinguenda Berlese, 1905: 157; Bogor (former Buitenzorg, Java). Other names: Scirus distinguendus; Thor, 1931a: 45. Remarks. Suspected synonym of Bdellodes longirostris (Hermann) according to Thor (1931a: 45). Type deposition. Lost (Castagnoli & Pegazano 1985). 13. Bdella dorsata Walckenaer & Gervais, 1844: 157; Paris, France on humid soil with small plants in gardens.��� Berlese, 1893: 43; Thor, 1931a: 37; Oudemans, 1937: 1195. Type deposition. Probably lost. 14. Bdella farabii Paktinat-Saeej & Bagheri, 2015a: 524; Amol, Mazandaran Province, Iran (Paktinat-Saeej et al. 2015a). Type deposition. Holotype at the Acarological Collection, Department of Plant Protection, Faculty of Agriculture, University of Maragheh, Maragheh, Iran; one paratype at ASI. 15. Bdella grandjeani Thor, 1931b: 65; Northern Africa. Type deposition. Lost. 16. Bdella heliophila Mihelčič, 1958a: 273; Cercedilla, Spain, on moss of pinus forest. Type deposition. Laboratorium Faunistica y Ecolog��a Animal, Instituto de Edafolog��a, Madrid. 17. Bdella helvetica Schweizer & Bader, 1963: 232; Val Ftur, National Park, Switzerland, under stone. Original designation: Bdella iconica helvetica Schweizer & Bader. Type deposition. NMB (NP 1931). 18. Bdella horvathi Karpelles, 1888: 275; Bulgaria.��� Thor, 1931a: 37. Type deposition. Unknown. 19. Bdella humida Wallace & Mahon, 1972: 546; Millstream, Fortescue River, West Australia, Australia; ex litter of Acacia sp. (Fabaceae) and Eucalyptus sp. (Myrtacea). Distribution. Australia, Iran (Kamali et al. 2001; Ueckermann et al. 2007). Redescription. Ueckermann et al. (2007). Remarks. Male unknown. Type deposition. Holotype and paratypes at ANIC; paratypes at SAM, BMNH, USNM and OSAL. 20. Bdella iconica Berlese, 1923: 240; on plants, in Florence and Germany���Thor, 1931a: 32. a. Bdella iconica var. veneta Berlese, 1923: 240; Thor, 1931a: 32; Schweizer & Bader, 1963: 231. b. Bdella sardoa Berlese, 1923: 241; B. iconica var. sardoa; Thor, 1931a: 32. Synonymy according to Thor (1931a). Distribution. Italy, Germany and Norway (Thor 1931a), Germany (Willmann 1955), Austria (Willmann 1951), France (Schm��lzer 1956), Spain (Mihelčič 1958a), Iceland (Atyeo & Tuxen 1962), Georgia (Gomelauri 1963b), New Zealand (Atyeo 1963a), Switzerland (Schweizer & Bader 1963), Bulgaria (Sosnina et al. 1965), Bohemia (Lell��kov��-Du��kova 1978), Ukraine (Kuznetsov & Livshits 1979a), Poland (Michocka 1987), China (Lin et al. 2006), Crimea (Bednarskaya 2009, 2010, 2011), Slovakia (Kaluz et al. 2013). Remarks. Michocka (1987: 39) apparently figured trichobothria on tibiae II, a characteristic only found in the subfamily Odontoscirinae; therefore it is likely that the species reported from Poland might be the result of a misidentification. Redescriptions. Atyeo & Tuxen (1962), Atyeo (1963a), Sosnina et al. (1965), Wallace & Mahon (1972), Lell��kov��-Du��kova (1978), Michocka (1987). Type deposition. CRA. 21. Bdella interrupta Evans, 1952: 668; Mid and North Wales, UK, on rocks above tide marks. Type deposition. Unknown. 22. Bdella karajiensis Ueckermann, Rastegar, Saboori & Ostovan, 2007: 133; Karaj, Iran, ex litter and soil beneath ornamental plant. Type deposition. Holotype at SRIAUT, paratypes at ASI. 23. Bdella khasyana Gupta, 1991: 221; Arunachal Pradesh, India; ex Litsea khasyana Meisn. (Lauraceae). Redescription. Gupta (2002). Remarks. Male unknown. Type deposition. NZSI. 24. Bdella kuznetsovi Maslov & Khaustov, 2013: 52; Arabatsky Nature Reserve, Crimea, Ukraine; ex storm detritus on shore of Sivash Gulf. Type deposition. NBG. 25. Bdella lattakia Soliman & Zaher, 1975: 79; Slinfa, Lattakia, Syria, ex moss. Distribution. Syria, Iran (Kamali et al. 2001; Ueckermann et al. 2007). Redescription. Ueckermann et al. (2007). Remarks. Male unknown. Type deposition. CUE. 26. Bdella longicornis (Linnaeus, 1758): 618; in Europe. Original designation: Acarus longicornis Linnaeus (= Acarus petrarum ruber L., 1746: 349).��� M��ller, 1776: 187; Johnston, 1845: 227; Oudemans, 1926b: 116.; Atyeo, 1962: 344 (neotype designation). a. Chelifer totus ruber Geoffroy, 1762: 618; synonymy according to Thor (1902). b. Acarus rupestris Linnaeus, 1758: 618; Hammer, 1775: 157; synonymy according to Thor (1931a). c. Acarus citri Hasselquist, 1757: 431; Oudemans, 1929: 306; synonymy according to Thor (1931a). d. Acarus ruber Linnaeus, 1746; Fabricius, 1775: 815. e. Acarus velox M��ller, 1776: 187; Oudemans, 1929: 312; 1937: 1201. f. Bdella rubra Lamarck, 1801: 179; von Heyden, 1826: 608; synonym of Bdellodes longirostris according to Berlese (1893). g. Scirus vulgaris Hermann, 1804: 61; Oudemans, 1929: 313; Bdella vulgaris, Koch, 1839: 1851; Dug��s, 1834: 21; Voigts & Oudemans, 1906: 242; B��bler, 1910: 812; Oudemans, 1937: 1206; synonymy according to Thor (1904b). h. Bdella decipiens Thorell, 1871: 701; Tr��g��rdh, 1902a: 7; 1904: 47; 1931: 47; Halbert, 1915: 113; Hull, 1918: 39; Banks, 1919: 11; Hull, 1922: 622; Tr��g��rdh, 1931: 47; synonymy according to Thor (1902: 7); Bdella longicornis var. decipiens Thorell; Tr��g��rdh, 1928: 8. i. Bdella borealis Kramer & Neuman, 1883: 525; Banks, 1899: 348; 1907: 596 synonymy according to Thor (1904b). j. Bdella caeca Berlese, 1905: 15; Bdella longicornis var. caeca; synonymy according to Thor (1931a: 28). k. Bdella vestita Koch, 1835: 23; 1842: 74; Walckenaer & Gervais, 1844: 157; Anders��n, 1863: 184; Canestrini & Fanzago, 1877: 105.��� Scirus vestitus, Murray, 1877: 146; synonymy according to Thor (1904b). l. Bdella anguinesetosa Ewing, 1909b: 72; synonymy according to Atyeo (1960a). m. Bdella tessellata Ewing, 1913: 112; Thor, 1931a: 36; synonymy according to Atyeo (1960a). n. Bdella oblonga Say, 1821: 19; Walckenaer & Gervais, 1847: 348; Gervais, 1849: 32; Banks, 1902a: 543; Oudemans, 1937: 1211; Jacot, 1938: 126; Baker & Balock, 1944: 180; Drummond, 1957: 142; synonymy according to Thor (1931a: 25); Bdellodes oblongula, Oudemans, 1937: 1221. o. Bdella egregia Koch, 1839: 23; synonymy according to Thor (1904b). p. Bdella cruentata Koch, 1839: 10; synonymy according to Thor (1904b). q. Bdella tenuirostris Koch, 1839: 18; synonymy according to Thor (1904b). r. B del la vivida Koch, 1839: 19; synonymy according to Thor (1904b). s. Bdella podurophila White, 1852: 210; synonymy according to Thor (1904b). t. Bdella arenaria Kramer, 1881: 444; synonymy according to Thor (1904b). Distribution. Germany (Voigts & Oudemans 1906), Sweden (Tr��g��rdh 1910), England (Hull 1918), Switzerland (Schweizer 1922; Schweizer & Bader 1963), Canada (Summerhayes & Elton 1928), Norway (Thor 1930a), North Africa (Thor 1931b), Austria (Willmann 1951), France (Schm��lzer 1956), Barro Colorado Islands, Panama Canal Zone, Costa Rica, Cuba, Mexico (Oaxaca, Distrito Federal, M��xico, San Luis Potos��), United States (California, Texas, Utah, Arkansas, Illinois, Tennessee, Florida, Missouri, Michigan, New Hampshire, Vermount), Nova Scotia (Atyeo 1960a), Japan (Ehara 1960; Shiba & Morikawa 1966; Shiba 1969a), Canada (Sinha 1963), Taiwan (Tseng 1978), United States (Lehman 1982), Poland (Michocka 1987), Korea (Lee et al. 1997), China (Fujian) (Lin & Zhang 2000), Japan (Nakamura et al. 2006), Iran (Kamali et al. 2001, Ueckermann et al. 2007), Crimea (Bednarskaya 2011). Redescriptions. Walckenaer & Gervais (1844), Thor (1926, 1930a, 1931a), Tr��g��rdh (1928), Oudemans (1929, 1937), Willmann (1956), Atyeo (1960a, 1962), Ehara (1961), Schweizer & Bader (1963), Shiba (1969a), Wallace & Mahon (1972), Tseng (1978), Michocka (1987), Ueckermann et al. (2007). Remarks. Bdella longicornis is the type species of the genus Bdella, and is among the first members of the family ever described. As a consequence, it has appeared under various names in the 18th and 19th century literature. Type deposition. Neotype of B. longicornis at USNM; type of B. caeca at CRA; type of B. cardinalis at MCZ; type of B. anguinesetosa at USNM. 27. Bdella longipalpus Mihelčič, 1958b: 41; Sierra Nevada, Spain. Type deposition. Unknown. 28. Bdella longistriata Atyeo, 1960a: 380; Llera, Tamaulipas, Mexico, ex pineapple epiphyte. Other records: Mexico (Llera, Tamaulipas; Ciudad del Maiz, San Luis Potos��; Antiguo Morelos, Tamaulipas), United States (Texas) (Atyeo 1960a). Type deposition. Holotype and paratypes at SEMC, paratypes at USNM. 29. Bdella malaccensis Shiba, 1978: 99; Malacca, Malaysia; under stones of intertidal zone. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 30. Bdella malawiensis van Der Schyff, Theron & Ueckermann, 2005: 224; Malawi, Bangula, Mozambique; ex Trichilia dregeana Sond. (Meliaceae). Remarks. Male unknown. Type deposition. NCA. 31. Bdella maldahanensis Gupta, 1992: 121; West Bengal, India, ex Mangifera indica Linnaeus (Anacardiaceae). Redescription. Gupta (2002). Type deposition. NZSI. 32. Bdella muscorum Ewing, 1909a: 124; Muncie, Illinois, USA ex moss.��� Thor, 1931a: 38. a. Bdella recens Ewing, 1934: 57 (= Bdella lata Ewing, 1909b: 69); Drummond, 1957: 142; synonymy according to Atyeo (1960a). b. Bdella subnigra Ewing, 1909b: 73; synonymy according to Atyeo (1960a). c. Bdella muscorum var. minnesotensis Ewing, 1913: 113; Thor, 1931a: 38; synonymy according to Atyeo (1960a). Distribution. United States (California, Colorado, New Mexico, Kansas, Arkansas, Tennessee, Michigan, Minnesota, Illinois, Maryland), Alaska (Point Barrow, Chandler Lake Region, Umiaat, District of Mackenzie), Germany (Ost-Holstein), Czeck Republic and Iceland (Atyeo 1960a), Iceland (Atyeo & Tuxen 1962), Georgia (Gomelauri 1963b), Bulgaria (Sosnina et al. 1965), Japan (Shiba & Morikawa 1966), Bohemia (Lell��kov��- Du��kova 1978), Taiwan (Tseng 1978), Ukraine (Kuznetsov & Livshits 1979a), Pakistan (Chaudhri et al. 1979), Canada (Danks 1980), United States (Lehman 1982), Korea (Lee et al. 1997), China (Fujian) (Lin & Zhang 2000), Hungary (Ripka et al. 2005), Japan (Nakamura et al. 2006), Iran (Ueckermann et al. 2007), Crimea (Bednarskaya 2009, 2010, 2011), Slovakia (Kaluz 2008). Redescriptions. Atyeo (1960a), Atyeo & Tuxen (1962), Shiba & Morikawa (1966), Sosnina et al. (1965), Wallace & Mahon (1972), Lell��kov��-Du��kova (1978), Tseng (1978). Type deposition. USNM. 33. Bdella neograndjeani Meyer & Ryke, 1959: 373; Eastern Cape Province, South Africa; ex unidentified shrub. Redescription. van Der Schyff et al. (2005). Type deposition. NCA. 34. Bdella nihoaensis Swift & Goff, 1987: 29; Nihoa Island, Hawaii, ex litter. Remarks. Female unknown. Type deposition. BPBM. 35. Bdella nylsvleyensis van Der Schyff, Theron & Ueckermann, 2005: 226; Limpopo Province, South Africa, ex soil and debris. Type deposition. NCA. 36. Bdella obesa Oudemans, 1937: 1211; unknown locality, Germany. Type deposition. Unknown. 37. Bdella pinicola Cooreman, 1943: 7; on pines, near Brussels, Belgium. Type deposition. IRSN. 38. Bdella piggotti Evans, 1953: 272; Kilimanjaro, Tanzania. Type deposition. Unknown. 39. Bdella pulchella Berlese, 1923: 241; unknown plant and in soil, Italy (Florence and Forli).��� Thor, 1931a: 33. Distribution. Norway and Italy (Thor 1931a), Spain (Mihelčič 1958a). Type deposition. Lost (Castagnoli & Pegazzano 1985). 40. Bdella radhikae Sadanandan et al., 2009: 3; ex Cocos nucifera, Kerala, India. Type deposition. Division of Acarology, Department of Zoology, University of Calicut, Kerala, India. 41. Bdella robusta Banks, 1894: 220; Sea Cliff, New York, USA, on ground.��� Banks, 1907: 596. Other name: Scirus robustus, Thor, 1931a: 45. Distribution. United States (Sea Cliff, New York) (Thor 1931a). Type deposition. Unknown. 42. Bdella semiscutata Thor, 1930a: 92; Svalbard, Norway.��� Thor, 1931a: 29; Willmann, 1939c: 431; 1956: 243. Distribution. Svalbard Island (Norway), Norway, Germany, (Thor 1931a), North Africa (Thor 1931b), Austria (Willmann 1951), Spain (Mihelčič 1958b), Georgia (Gomelauri 1963b), Switzerland (Schweizer &, Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. 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T

Polytrichus van Der Schyff, Theron & Ueckermann Type-species: Polytrichus yemenensis van Der Schyff, Theron & Ueckermann, 2003 by original designation. 87. Polytrichus yemenensis van Der Schyff, Theron & Ueckermann, 2003: 21; Djebel An-Nabi Shuaib, Yemen. Type deposition. NCA.

Spinibdella Thor Spinibdella Thor, 1930b: 22; 1931a: 39; Atyeo, 1960a: 424; Soliman & Zaher, 1975: 80; Tseng, 1978: 38; Chaudhri et al., 1979: 133; Michocka, 1987: 82. Type-species: Spinibdella reducta Thor, 1930b: 23 by original designation. 121. Spinibdella ampulla Wallace & Mahon, 1972: 568; Millstream, Western Australia, ex green herbaceous, couch grass, Eucalyptus sp. litter. Remarks. Male unknown. Type deposition. ANIC. 122. Spinibdella ankylotricha Omukunda, Theron & Ueckermann, 2012: 9; Limpopo Province, South Africa. Type deposition. NCA. 123. Spinibdella antarctica (Tr��g��rdh, 1907): 24; South Georgia, Grytviken Peninsula, Antarctica, under rocks.��� Wallace, 1970: 107. Original designation: Bdella antarctica Tr��g��rdh; Thor, 1931a: 33. Redescription. Wallace (1970). Type deposition. BPBM; USNM, BMNH, ANIC. 124. Spinibdella arenosa Willmann, 1939b: 532; Germany (Wangerooge Island) (Willmann, 1952: 165). Type deposition. ZSM. 125. Spinibdella atyeoi Gupta & Paul, 1985: 14; West Bengal, Midnapur Dist., Patharkumkumi, India, ex nest of Prinia inornata (Sykes 1832) (Aves: Cysticolidae). Remarks. Male unknown. Type deposition. NZSI. 126. Spinibdella bifurcata Atyeo, 1960a: 430; 10 miles west of Tuxtla Gutierrez, Chiapas, Mexico, under rock.��� Soliman, 1975: 48. Distribution. Mexico (Chiapas, Oaxaca, Puebla, Michoac��n, United States (Texas) (Atyeo 1960a), Egypt (Giza) (Soliman 1975; Zaher 1986), Malaysia (Shiba 1978), China (Lin et al. 2006). Redescription. Shiba (1978). Type deposition. Holotype and paratypes at SEMC, paratypes at USNM, BMNH and SAM. 127. Spinibdella bioculata Swift & Goff, 1987: 39; Kahoolawe Island, Hawaii, ex Kiawe duff and grasses. Remarks. Male unknown. Type deposition. BPBM. 128. Spinibdella californica McGregor, 1956: 12; California, USA, ex lemon. Type deposition. Unknown. 129. Spinibdella corticis (Ewing, 1909a): 122; Urbana, Illinois, USA, under bark of cottonwood tree.��� Atyeo, 1960a: 426. Original designation: Bdella corticis Ewing; Thor, 1931a: 36. Other name: Spinibdella cortis [sic]; Rack, 1961: 185. Distribution. United States (Illinois, Texas, Utah, Nebraska), Mexico (Oaxaca), Guatemala (Atyeo 1960a), Japan (Shiba & Morikawa 1966), Australia (Wallace & Mahon 1972). Redescriptions. Atyeo (1960a), Wallace & Mahon (1972). Type deposition. USNM. 130. Spinibdella cronini (Baker & Balock, 1944): 178; Planada, California, USA, on lichens from fig tree.��� Atyeo, 1960a: 432. Original designation: Bdella cronini Baker & Balock. Distribution. United States (California, Texas, Utah, Colorado, Washington, Alabama, Maryland), Mexico (Tamaulipas, Guerrero, Nevo Le��n, San Luis Potos��) (Atyeo 1960a), Australia (Atyeo 1963a; Wallace & Mahon 1972), Bulgaria (Sosnina et al. 1965), Egypt (Soliman 1975), Syria (Lattakia) (Soliman & Zaher 1975), Ukraine (Kuznetsov & Livshits 1979a), United States (Lehman 1982), Hawaii (Swift & Goff 1987), Hungary (Ripka et al. 2005), China (Li et al. 1992; Li & Fan 2007), Iran (Ostovan & Kamali 1995; Kamali et al. 2001, Jalaeian et al. 2005; Ueckermann et al. 2007; Abbaszadeh et al. 2010; Daneshnia & Akrami 2013; Majidi & Akrami 2013; Masoudian & Khanjani 2013), Brazil (Pinto-da-Rocha 1995; Hernandes et al. 2011), Slovakia (Kaluz 2008). Remarks. this species was found in bat guano in USA (Webster & Whitaker 2005). Redescriptions. Atyeo (1960a, 1963a), Sosnina et al. (1965), Wallace & Mahon (1972), Swift & Goff (1987), Ueckermann et al. (2007). Type deposition. USNM. 131. Spinibdella denheyeri Hernandes, Daud & Feres, 2008: 265; ex Coffea arabica leaves (Linnaeus, Rubiaceae) Atibaia, S��o Paulo, Brazil. Type deposition. DZSJRP. 132. Spinibdella depressa (Ewing, 1909a): 125; Arcola, Illinois, USA, under bark.��� Atyeo, 1960a: 428. Original designation: Bdella depressa Ewing; Thor, 1931a: 38. a. Bdella virgata Ewing, 1909b: 70; Hartzell, 1918: 206; Baker & Balock, 1944: 179 synonymy by Atyeo (1960a). b. Bdella chapultepecensis Baker & Balock, 1944: 177 synonymy by Atyeo (1960a). c. Bdella riolermensis Baker & Balock, 1944: 178 synonymy by Atyeo (1960a). Distribution. United States (Maryland) (Drummond 1957), United States (Illinois, Texas, Maryland, Arkansas, Kansas, New Jersey, Connecticut), Mexico (M��xico, Distrito Federal, San Luis Potos��, Morelos) (Atyeo 1960a), Australia (Atyeo 1963a; Wallace & Mahon 1972), Pakistan (Chaudhri et al. 1979), United States (Lehman 1982), Hawaii (Swift & Goff 1987), China (Fujian) (Lin & Zhang 2000), Iran (Kamali et al. 2001; Ueckermann et al. 2007; Abbaszadeh et al. 2010). Remarks. this species has the posterior eye wanting, with circular striation where that eye should normally be. Redescriptions. Baker & Balock (1944), Atyeo (1960a, 1963a), Wallace & Mahon (1972), Swift & Goff (1987), Ueckermann et al. (2007). Type deposition. USNM. 133. Spinibdella dusta Shiba, 1969b: 150; tatami, Kuwabara-ch��, Matsuyama, Japan, ex tatami. Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan. 134. Spinibdella gibberabdomen (Thor, 1931b): 68; Tangier, Morocco; Paktinat-Saeej et al. 2015: 695. Original designation: Bdella gibberabdomen Thor. Remarks. The author mentions only two ventral setae on the hypostome, and illustrates a truncate palptarsus, which is why this species was transferred to the genus Spinibdella. Type deposition. Lost. 135. Spinibdella howarthi Swift & Goff, 1987: 40; Mauna Kea Summit Cone, Hawaii, under stone. Remarks. Male unknown; species known only from the holotype. Type deposition. BPBM. 136. Spinibdella iberica Gomelauri, 1961: 68; close to the Turtle lake near Tbilisi, Georgia, ex lichens. Type deposition. Unknown. Remarks. The choice of the epithet by the author is a mystery, since the type locality was not in the Iberian Peninsula. 137. Spinibdella lignicola (Canestrini, 1886): 184; Italy and Egypt.��� Tseng, 1978: 47. Original designation: Bdella lignicola Canestrini. Distribution. Italy (Canestrini 1886; Thor 1931a), England (Hull 1918), Egypt (Tr��g��rdh 1905; Thor 1931a; Abdel-Shaheed et al. 1971), Spain (Mihelčič 1958b), Switzerland (Schweizer & Bader 1963), Taiwan (Tseng 1978), China (Sichuan) (Li et al. 1992), Czeck Republic (Stejskal & Hubert 2008). Redescription. Tseng (1978). Type deposition. CRA. 138. Spinibdella longistriata Tseng, 1978: 42; Shandimann, Pingtung Hsien, Taiwan, ex litter. Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.). 139. Spinibdella mali Jorgensen, 1967: 98; Spring Lake, Utah Co, Utah, USA, ex bark of apple tree. Remarks. Male unknown. Type deposition. Unknown. 140. Spinibdella namibiensis Omukunda, Theron & Ueckermann, 2012: 15; Namibia and South Africa. Type deposition. NCA. 141. Spinibdella novemsetosa Tseng, 1978: 42; Tainan city, Taiwan, on shallot (Alliaceae). Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.). 142. Spinibdella ornata Atyeo, 1960a: 434; Bear Lake, Rock Mountain National Park, Colorado, USA, ex moss and litter. Distribution. California, (Atyeo 1960a), Wisconsin (Oatman 1963). Remarks. Suspected synonym of S. thori, according to Atyeo (1963a: 174). Type deposition. Holotype and paratypes at SEMC, paratypes at CSUC, USNM. 143. Spinibdella polyattenuata Omukunda, Theron & Ueckermann, 2012: 6; Eastern Cape Province, South Africa. Type deposition. NCA. 144. Spinibdella pongolensis Omukunda, Theron & Ueckermann, 2012: 12; Kwazulu, Natal, South Africa. Type deposition. NCA. 145. Spinibdella quinqueoculata Thor, 1931b: 70; Tangier, Morocco, ex moss. Type deposition. Lost. 146. Spinibdella rapida Kuznetsov & Livshits, 1979b: 608; rocky cliffs in the vicinity of Alupka town, Crimea, Ukraine, ex moss.��� Bednarskaya, 2011: 5. Type deposition. NBG. 147. Spinibdella reducta Thor, 1930b: 23; Norway, in coniferous forest litter.��� Thor, 1931a: 39. Distribution. Norway (Thor 1931a), Poland (Michocka 1987). Redescriptions. Thor (1931a), Michocka (1987). Type deposition. Lost. 148. Spinibdella smileyi Tseng, 1978: 39; Taipei, Chiayi Hsien, Taiwan, ex litter. Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.). 149. Spinibdella subrufa Rack, 1961: 183; Germany. Type deposition. ZMUH. 150. Spinibdella tabarii Paktinat-Saeej & Bagheri, 2015b: 696; Amol city, Mazandaran Province, Iran; also citrus, Noor city, Mazandaran Province, Iran. Type deposition. Holotype and paratypes at the Acarological Collection, Department of Plant Protection, Faculty of Agriculture, University of Maragheh, Maragheh, Iran; paratypes at the Acarological Collection, Jalal Afshar Zoological Museum, Department of Plant Protection, Faculty of Agriculture, University of Tehran, Karaj, Iran; and also at ASI. 151. Spinibdella tadjikistanica Kuznetsov, 1984: 774; Kondara Canyon, Tadjikistan, ex hawthorn (Crataegus sp., Rosaceae) and grape. Type deposition. NBG. 152. Spinibdella tenella (Banks, 1896): 75; Sea Cliffs, Long Island, NY, USA. Original designation: Bdella tenella Banks; Banks, 1904c: 16; 1907: 596; Thor, 1931a: 31; Spinibdella tenella; Hernandes, 2013: 64. Type deposition. MCZ. 153. Spinibdella tenuirostris (Ewing, 1917): 149; Xenia, Ohio, USA, under stones.��� Atyeo, 1960a: 424. Original designation: Bdella tenuirostris Ewing; Berlese, 1893: 43. a. Spinibdella wilsoni Jacot, 1938: 129; synonymy according to Atyeo (1960a: 424). Distribution. Germany (Thor 1931a), United States (Ohio, Florida, Arkansas, Kansas, North Carolina, Vermont, Michigan, California) (Atyeo 1960a), Japan (Shiba & Morikawa 1966), Australia (Atyeo 1963a; Wallace & Mahon 1972), Russia (Wainstein et al. 1978, Ghilarov 1978), Taiwan (Tseng 1978), Korea (Lee et al. 1997), Mexico (Hoffmann & L��pez-Campos 2000), Spain (Domingo-Quero et al. 2003). Redescriptions. Atyeo (1960a, 1963a), Shiba & Morikawa (1966), Wallace & Mahon (1972), Tseng (1978). Remarks. Ewing (1917, not 1914 as mentioned by both Atyeo [1960a] and Wainstein et al. [1978]) described Bdella tenuirostris, without noticing the preoccupied name erected by Koch (1839: 23). Type deposition. USNM. 154. Spinibdella thori (Meyer & Ryke, 1959): 375; Bathurst, South Africa, ex grass and soil.��� Atyeo, 1963a: 174. Original designation: Bdella thori Meyer & Ryke. Distribution. South Africa (Meyer & Ryke 1959; Halliday 2005), Australia (Atyeo 1963a; Wallace & Mahon 1972; Halliday 2005), Hawaii (Swift & Goff 1987; 2001), Mexico (Hoffmann & L��pez-Campos 2000), Iran (Abbaszadeh et al. 2010). Redescriptions. Atyeo (1963a), Wallace & Mahon (1972), Swift & Goff (1987), Omukunda et al. (2012). Remarks. Male unknown. Type deposition. Institute for Zoological Research, Potchefstroom University, South Africa. 155. Spinibdella trinomma Omukunda, Theron & Ueckermann, 2012: 3; Kwazulu, Natal, South Africa. Type deposition. NCA. 156. Spinibdella trisetosa (Jacot, 1938): 128 comb. nov.; Micanope, Florida, USA, ex leaf litter. Original designation: Bdella trisetosa Jacot. Remarks. This species is herein transferred to the genus Spinibdella due to having two ventral setae on the hypostome, tricobothria present on tibiae I, IV, tarsi III and IV, setae lps present, and the palpal tibiotarsus truncate. Type deposition. USNM. 157. Spinibdella yeni Tseng, 1978: 44; Taipu, Chiayi Hsien, Taiwan, ex litter. Type deposition. Supposedly at BSMI, but probably lost (C-C Ho, pers. comm.)., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. 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(2011) Snout mites from caves in Brazil, with description of a new species. Journal of Natural History, 45, 799 - 812. http: // dx. doi. org / 10.1080 / 00222933.2010.535919","Kaluz, S. (2008) Soil mites (Acari) of the forests in floodplain areas of the rivers Danube and Morava. Peckiana, 5, 89 - 103.","Webster, J. M. & Whitaker, J. O. (2005) Study of guano communities of Big Brown bat colonies in Indiana and neighboring Illinois counties. Northeastern Naturalist, 12 (2), 221 - 232. http: // dx. doi. org / 10.1656 / 1092 - 6194 (2005) 012 [0221: SOGCOB] 2.0. CO; 2","Hernandes, F. A., Daud, R. D. & Feres, R. J. F. (2008) Two new species of Bdellidae (Acari: Prostigmata) from Brazil. International Journal of Acarology, 34 (3), 259 - 266. http: // dx. doi. org / 10.1080 / 01647950808684539","Ewing, H. E. (1909 b) A systematic and biological study of the Acarina of Illinois. University of Illinois Bulletin, 7 (14), 1 - 120.","Hartzell, A. (1918) A preliminary list of the Acarina of Iowa. The Proceedings of the Iowa Academy of Sciences, 25, 205 - 212.","Drummond, R. O. (1957) Observations on the fluctuations of acarine populations from nests of Peromyscus leucopus. Ecological Monographs, 27 (2), 137 - 152. http: // dx. doi. org / 10.2307 / 1948573","Lin, J. Z. & Zhang, Y. X. (2000) Bdelloidea. In: Huang, B. K. (Ed.), Fauna of insects in Fujian Province of China. Vol. 9. Fujian Science and Technology Press, Fuzhou, 109 - 121.","Shiba, M. (1969 b) On some mites from \" Tatami \" (Japanese Rice Straw Matting). Reports of Research Matsuyama Shinonome Junior College, 4, 147 - 170.","Thor, S. (1931 b) Nordafrikanische Bdellidae und Cunaxidae, von Dr. F. Grandjean (Paris) 1931 gesammelt. Zoologischer Anzeiger, 97 (3 - 4), 62 - 79.","Gomelauri, L. A. (1961) New species of the family Bdellidae. Bulletin of the Academy of Sciences of the Georgian SSR, 26 (1), 68 - 72.","Canestrini, G. (1886) Prospetto dell'acarofauna italiana. 2: Erythraeini, Cheyletini, Bdellini, Eupodini e Analgesini. pp. 159 - 311, Tav. X - XXII [= 10 - 22]. Padova.","Hull, J. E. (1918) Terrestrial Acari of the Tyne province. Transactions of the Natural History Society of Northumberland, Durham, and Newcastle-upon-Tyne, 75, 13 - 88.","Tragardh, I. (1905) Acariden aus Aegypten und dem Sudan. In: Results of the Swedish zoological expedition to Egypt and the White Nile, 1901. part II. 124 pp.","Abdel-Shaheed, G. A., Hammad, S. M. & El-Sawaf, S. K. (1971) Survey of mites found on some field crops in Alexandria District, U. A. R. Journal of Applied Entomology, 69, 106 - 110. http: // dx. doi. org / 10.1111 / j. 1439 - 0418.1971. tb 03191. x","Mihelcic, F. (1958 b) Zoologisch-systematische Ergebnisse der Studienreise von H. Janetschek und W. Steiner in die spanische Sierra Nevada 1954. IX. Milben (Acarina). Sitzungsberichte der Mathematisch-naturwissenschaftliche Klasse, Abteilung I, 167, 35 - 57.","Schweizer, J. & Bader, C. (1963) Die Landmilben der Schweiz (Mittelland, Jura and Alpen). Trombidiformes Reuter. Denkschriften Schweizerischen Naturforschenden Gesellschaft, 84, 209 - 378.","Stejskal, V. & Hubert, J. (2008) Risk of occupational allergy to stored grain arthropods and false pest-risk perception in Czech grain stores. Annals of Agricultural and Environmental Medicine, 15 (1), 29 - 35.","Jorgensen, C. (1967) A new species of Spinibdella from Utah (Bdellidae: Acarina). The Great Basin Naturalist, 27 (1), 98 - 101.","Oatman, E. R. (1963) Mite species on apple foliage in Wisconsin. In: Advances in Acarology, Naegele, J. A. (Ed.), vol. 1, Comstock Pub. Assoc., 21 - 24.","Kuznetsov, N. N. & Livshits, I. Z. (1979 b) Two new species of Bdellidae (Acariformes) from the Crimea. Zoologicheskii Zhurnal, 58 (4), 607 - 610.","Bednarskaya, E. V. (2011) The main biotic relations and localization in microstations of prostigmatic predatory mites in carst cavities of Mountain Crimea. Scientific Notes of Taurida V. I. Vernadsky National University. - Series: Biology, chemistry, 24 (63), 3 - 9.","Paktinat-Saeej, S., Bagheri, M., Saboori, A., Seilsepour, N. & Ueckermann, E. A. (2015 b) A new snout mite, Spinibdella tabarii sp. nov. (Trombidiformes: Bdellidae) from Iran, with a summary of Spinibdella distributions worldwide. Systematic & Applied Acarology, 20 (6), 693 - 706. http: // dx. doi. org / 10.11158 / saa. 20.6.11","Banks, N. (1896) New North American spiders and mites. Transactions of the American Entomological Society, 23, 57 - 77.","Banks, N. (1904 c) A treatise on the Acarina, or mites. Proceedings of the United States National Museum, 28, 1 - 114.","Hernandes, F. A. (2013) Revision of Nathan Banks' type specimens of Bdellidae Duges (Acari: Trombidiformes) of the Museum of Comparative Zoology, Cambridge. International Journal of Acarology, 39 (1), 1 - 9. http: // dx. doi. org / 10.1080 / 01647954.2012.739642","Ewing, H. E. (1917) New Acarina, part II-Descriptions of new species and varieties from Iowa, Missouri, Illinois, Indiana and Ohio. Bulletin of the American Museum of Natural History, 37 (2), 149 - 172.","Jacot, A. P. (1938) Thomas Say's free-living mites re-discovered. Psyche, 45, 126 - 130. http: // dx. doi. org / 10.1155 / 1938 / 92469","Wainstein, B. A., Kuznetsov, N. N., Livshits, I. Z. & Sosnina, E. F. (1978) Family Bdellidae. In: Gilyarov, M. S. (Ed) Identifcation key to soil-inhabiting mites, Trombidiformes. Nauka, Moscow, 133 - 143.","Lee, W. - K., Lim, J. - W. & Lee, S. - Y. (1997) A taxonomic study on the family Bdellidae (Bdelloidea: Prostigmata) in Korea. Korean Journal of Soil Zoology, 2 (2), 65 - 75 [in Korean with English abstract]","Hoffmann, A. & Lopez-Campos, G. (2000) Biodiversidad de los acaros en Mexico. Universidad Nacional Autonoma de Mexico y Conabio, Mexico, D. F. 230 pp.","Domingo-Quero, T., Alonso-Zarazaga, M. A., Sanchez-Ruiz, A., Araujo Armero, R., Navas Sanchez, A., Sanchez Moreno, S., Garcia Becerra, R., Nebreda, M., Sanchez Ruiz, M., Fontal-Cazalla, F. & Nieves-Aldrey, J. L. (2003) Inventariando la Biodiversidad en el Parque Nacional de la Caldera de Taburiente (La Palma, Islas Canarias, Espana): Novedades cientificas. Graellsia, 59 (2 - 3), 45 - 68. http: // dx. doi. org / 10.3989 / graellsia. 2003. v 59. i 2 - 3.235","Meyer, M. K. P. & Ryke, P. A. J. (1959) Cunaxoidea (Acarina: Prostigmata) occurring on plants in South Africa. Annals and Magazine of Natural History, 13 (2), 369 - 384. http: // dx. doi. org / 10.1080 / 00222935908655745","Halliday, R. B. (2005) Predatory mites from crops and pastures in South Africa: potential natural enemies of redlegged earth mite, Halotydeus destructor (Acari: Penthaleidae). Zootaxa, 1079, 11 - 64.","Swift, S. F. & Goff, M. L. (2001) Mites (Acari) communities associated with ' Ohi'a, Metrosideros polymorpha (Myrtaceae), at Hono O Na Pali and Kui`a Natural area reserves on Kaua`i Island, Hawaiian Islands. Pacific Science, 55 (1), 23 - 40. http: // dx. doi. org / 10.1353 / psc. 2001.0008"]}

Key to adult female Hexabdella of the world (modified from Van der Schyff et al. 2004) and to Neomolgus of Asia (Iran, Japan, Uzbekistan) and the Siberian region of Russia 1. Tibia II without trichobothria, each chelicera with two setae.................. Bdellinae… Hexabdella Van der Schyff … 2 - Tibia II with trichobothria, each chelicera with more than 2 setae........... Odontoscirinae … Neomolgus Oudemans … 11 2. Opisthosomal setae distally branched...................................................................... 3 - Opisthosomal setae smooth or slightly plumose.............................................................. 5 3. Dorsal striae coarsely to sparsely broken; seta ps 1 smooth; coxa IV without macroseta.............................. 4 - Dorsal striae finely broken; seta ps 1 branched; coxa IV with a serrated macroseta.............................................................................................. H. denheyeri Van der Schyff, Theron & Ueckermann 4. Marginal striae of hysterosoma with lobes, visible as dashed lines............... H. cinquaginta Hernandes, Daud & Feres - Marginal striae of hysterosoma smooth............................ H. singula Van der Schyff, Theron & Ueckermann 5. Eyes present......................................................................................... 6 - Eyes absent................................................. H. maraugia Van der Schyff, Theron & Ueckermann 6. Two pairs of eyes present; chelicerae striated............................................................... 7 - One pair of eyes present; chelicerae smooth...................... H. unusoculata Van der Schyff, Theron & Ueckermann 7. Palp basifemur with five or six setae...................................................................... 8 - Palp basifemur with four setae........................................................ H. brevitarsis Hernandes 8. Palp basifemur with five setae........................................................................... 9 - Palp basifemur with six setae.......................... H. persiaensis Paktinat Saeej, Bagheri, Saboori & Ueckermann 9. Solenidotaxy of tibiae I–IV 3 – 2 – 1 –0..................................................................... 10 - Solenidotaxy of tibiae I– IV 1 – 1 – 1 –0............................. H. miranda Van der Schyff, Theron & Ueckermann 10. Coxa II with four setae; basifemura II and IV with seven and fivesetae; genu IV with 4 setae (1 s); fixed digit two thirds the length of movable digit......................................................... H. mexicana (Baker & Balock) - Coxa II with three setae; basifemura II and IV with eight and four setae; genu IV with five setae (1 s); fixed digit as long as movable digit................................ H. quercusi Eghbalian, Khanjani, Safaralizadeh & Ueckermann sp. nov. 11. Palp genu with four setae.............................................................................. 12 - Palp genu with ten setae............................................................ N. longipalpus Kuznetsov 12. Palp tibiotarsus with 8–14 setae......................................................................... 13 - Palp tibiotarsus with more than 20 setae......................................................... N. littoralis (L.) 13. Chelicerae with 7–11 setae............................................................................. 14 - Chelicerae with 21 setae...................... N. iraniensis Eghbalian, Khanjani, Safaralizadeh & Ueckermann sp. nov. 14. Palp basifemur with four setae.......................................................................... 15 - Palp basifemur with five to seven setae................................................................... 16 15. Palp tibiotarsus with eight setae (including end setae); chelicerae with seven setae........... N. raptor Kuznetsov & Barilo - Palp tibiotarsus with nine setae (including end setae); chelicerae with eight setae.................... N. pygmaeus Shiba 16. Prodorsum without distinct dorsal shield.................................................................. 17 - Prodorsum with distinct dorsal shield....................................................... N. clypeatus (Thor) 17. Chelicerae with nine setae; palp basifemur with 5–6 setae........................................... N. pallipes (L.) - Chelicerae with 11 setae; palp basifemur with 7 setae....................................... N. capillatus (Kramer)

Genus Hexabdella Van der Schyff et al. 2004 Type species: Hexabdella denheyeri Van der Schyff et al. 2004, Published as part of Eghbalian, Amir Hossein, Khanjani, Mohammad, Safaralizadeh, Mohammad Hassan & Ueckermann, Edward A., 2016, New species of Hexabdella and Neomolgus (Acari: Prostigmata: Bdellidae) from Iran, pp. 291-300 in Zootaxa 4072 (2) on page 292, DOI: 10.11646/zootaxa.4072.2.10, http://zenodo.org/record/260608

Eghbalian, Amir Hossein, Khanjani, Mohammad, Safaralizadeh, Mohammad Hassan, Ueckermann, Edward A. (2016): New species of Hexabdella and Neomolgus (Acari: Prostigmata: Bdellidae) from Iran. Zootaxa 4072 (2): 291-300, DOI: http://doi.org/10.11646/zootaxa.4072.2.10

Trachymolgus Berlese Trachymolgus Berlese, 1923: 242; Thor, 1931a: 40. Type-species: Bdella nigerrima Canestrini & Fanzago, 1876 by original designation. 83. Trachymolgus jesusi Mej��a-Recamier & Palacios-Vargas, 1999: 165; Jalisco, Chamela, M��xico. Type deposition. Holotype and paratypes at UNAM, paratypes at UGA, CRA and CNC. 84. Trachymolgus nigerrima (Canestrini & Fanzago, 1876): 108; Italy. Original designation: Bdella nigerrima Canestrini & Fanzago.���Canestrini & Fanzago, 1877: 104; Canestrini, 1886: 189; Berlese, 1883: 214; Thor, 1931a: 40; Trachymolgus nigerrimus; Berlese, 1923: 242; Sosnina et al. (1965: 279). a. Acarus citri Hasselquist, 1757: 431; Oudemans, 1926b: 118; synonymy according to Thor, 1931a; Bdella citri, Thor (1931a). Distribution. Italy, Switzerland (Thor 1931a), Georgia (Gomelauri 1963b), Switzerland (Schweizer & Bader 1963), Bulgaria (Sosnina et al. 1965), Ukraine (Kuznetsov & Livshits 1979a). Type deposition. Unknown. 85. Trachymolgus purpureus Fisher & Dowling, 2011: 5 (in Fisher et al. 2011); Ozark Highlands, Arkansas, USA. Type deposition. Holotype and paratypes at ACUA, paratypes at OSAL, FMNH, USNM. 86. Trachymolgus recki Gomelauri, 1961: 69; Sukhumi and Tbilisi, Georgia, ex oak leaves and tree stocks in forest. Type deposition. Unknown., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on pages 22-23, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["Berlese, A. (1923) Centuria sesta di Acari nuovi. Redia-Giornale di Entomologia, 15, 1 - 240.","Thor, S. (1931 a) Bdellidae, Nicoletiellidae, Cryptognathidae. Das Tierreich, 56, 87 pp.","Canestrini, G. & Fanzago, F. (1876) Nuovi Acari Acari Italiani. Atti della Societa Veneto-Trentina di Scienze Naturali, Padova, 5 (1), 99 - 111.","Mejia-Recamier, B. E. & Palacios-Vargas, J. G. (1999) A new species of Trachymolgus (Prostigmata: Bdellidae) from Mexico. Acarologia, 40 (2), 165 - 170.","Canestrini, G. & Fanzago, F. (1877) Intorno agli acari italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, 5 (4), 1 - 140 + plates II - VII.","Canestrini, G. (1886) Prospetto dell'acarofauna italiana. 2: Erythraeini, Cheyletini, Bdellini, Eupodini e Analgesini. pp. 159 - 311, Tav. X - XXII [= 10 - 22]. Padova.","Berlese, A. (1883) Escursione in Sicilia: Acarofauna Sicula. Bolletino della Societa Entomologica Italiana, 15, 212 - 220.","Sosnina, E. F., Vysotskaya, S. O., Markov, G. N. & Atanasov, L. K. (1965) Predatory mites of the family Bdellidae (Acarina, Prostigmata) from nests of rodents in Bulgaria. Proceedings of the Zoological Institute, USSR Academy of Sciences, 35, 272 - 287.","Hasselquist, F. (1757) Iter Palaestinum, Eller, Resa til Heliga Landet, Forrattad Infran ar 1749 til 1752.","Oudemans, A. C. (1926 b) Kritisch Historisch overzicht der Acarologie. Tijdschrift voor Entomologie, 69, 500 pp.","Gomelauri, L. A. (1963 b) On the study of mites of the family Bdellidae in Georgian S. S. R. Bulletin of the Academy of Sciences of the Georgian SSR, 30 (2), 47 - 51.","Schweizer, J. & Bader, C. (1963) Die Landmilben der Schweiz (Mittelland, Jura and Alpen). Trombidiformes Reuter. Denkschriften Schweizerischen Naturforschenden Gesellschaft, 84, 209 - 378.","Kuznetsov, N. N. & Livshits, I. Z. (1979 a) Predatory mites of the Nikita Botanical Garden (Acariformes: Bdellidae, Cunaxidae, Camerobiidae). Proceedings of the State Nikita Botanical Garden, 79, 51 - 104.","Fisher, J. R., Skvarla, M. J., Bauchan, G. R., Ochoa, R. & Dowling, A. P. G. (2011) Trachymolgus purpureus sp. n., an armored snout mite (Acari, Bdellidae) from the Ozark Highlands: morphology, development, and key to Trachymolgus Berlese. Zookeys, 125, 1 - 34. http: // dx. doi. org / 10.3897 / zookeys. 125.1875","Gomelauri, L. A. (1961) New species of the family Bdellidae. Bulletin of the Academy of Sciences of the Georgian SSR, 26 (1), 68 - 72."]}

Bdellidae Dug��s Bdellei Dug��s, 1834: 21; (for exhaustive reference list see Thor, 1931a: 1)., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on page 11, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["Duges, A. (1834) Recherches sur l'ordre des Acariens en generale et la famille des Trombidies en particulier. Annales des Sciences Naturelles, 2, 5 - 46.","Thor, S. (1931 a) Bdellidae, Nicoletiellidae, Cryptognathidae. Das Tierreich, 56, 87 pp."]}

Fossil BDELLIDAE 279. Bdella vetusta Ewing, 1934: 58; Manitoban amber (Cretaceous, ~ 76���79 Ma), Cedar Lake, Manitoba, Canada. Type deposition. ROME. 280. Bdella lata Koch & Berendt, 1854: 872 ���not Bdella lata Ewing (1909b); Baltic amber (~44 Ma). 281. Bdella bicincta Menge in Koch & Berendt, 1854: 108 ���Baltic amber (~44 Ma). 282. Bdella bombycina Menge in Koch & Berendt, 1854: 108 ���Baltic amber (~44 Ma). 283. Bdella obconica Menge in Koch & Berendt, 1854: 108 ���Baltic amber (~44 Ma). 284. Bdellidae sp. (Aoki 1974 apud Dunlop et al. 2012)., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on page 45, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["Ewing, H, E. (1934) Arachnida. Order Acarina. In: Carpenter, F. M., Folsom, J. W., Essig, E. O., Kinsey, A. C., Brues, C. T., Boesel, M. W. & Ewing, H. E. Insects and Arachnids from Canadian amber. Contributions to Canadian Mineralogy. Proceedings of the Geological Society of America, 418, pp. 56 - 62.","Koch, C. L. & Berendt, G. C. (1854) Die im Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt, in Berendt G. C. (Ed.), Die im Bernstein befindlichen organischen Reste der Vorwelt. Nicolaischen Buchhandlung, Berlin. 1 (2), 1 - 124.","Ewing, H. E. (1909 b) A systematic and biological study of the Acarina of Illinois. University of Illinois Bulletin, 7 (14), 1 - 120.","Aoki, J. (1974) On the fossil mites in Mizunami amber from Gifu Prefecture, Central Japan. Bulletin of the Mizunami Fossil Museum, 1, 397 - 399.","Dunlop, J. A., Penney, D. & Jekel, D. (2012) A summary list of fossil spiders and their relatives. In: Platnick, N. I. (Ed.), The world spider catalog, version 12.5 American Museum of Natural History. Avaliable from: http: // research. amnh. org / entomology / spiders / catalog / index. htm (Accessed 15 Aug. 2016)"]}

Tetrabdella Hernandes & Feres Type-species: Tetrabdella neotropica Hernandes & Feres, 2006 by original designation. 158. Tetrabdella neotropica Hernandes & Feres, 2006: 60; Cedral, São Paulo, Brazil, ex Hevea brasiliensis M.Arg. (Euphorbiaceae). Type deposition. Holotype and paratypes at DZSJRP, paratype at USNM.

Monotrichobdella Baker & Balock Monotrichobdella Baker & Balock, 1944: 176. Type-species: Monotrichobdella maxosburni Baker & Balock, 1944 by original designation. 120. Monotrichobdella maxosburni Baker & Balock, 1944: 176; near Tres Cumbres, Mexico-Cuernavaca Highway, Morelos, Mexico, ex lichens. Distribution. Mexico (Morelos, Salazar, Distrito Federal) (Baker & Balock 1944; Atyeo 1960a), Taiwan (Tseng 1978). Redescriptions. Atyeo (1960a), Tseng (1978). Type deposition. USNM., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on page 26, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["Baker, E. W. & Balock, J. W. (1944) Mites of the family Bdellidae. Proceedings of the Entomological Society of Washington, 46 (7), 176 - 184.","Atyeo, W. T. (1960 a) A revision of the family Bdellidae in North and Central America (Acarina: Prostigmata). University of Kansas Science Bulletin, 40, 345 - 499.","Tseng, Y-H. (1978) Mites of the family Bdellidae from Taiwan (Acarina: Prostigmata). Journal of the Agricultural Association of China, 104, 25 - 51."]}

Hexabdella van Der Schyff, Theron & Ueckermann Hexabdella van Der Schyff, Theron & Ueckermann, 2004: 14. Bdella; Baker & Balock, 1944 (part.). Type-species: Hexabdella denheyeri van Der Schyff, Theron & Ueckermann, 2004 by original designation. 55. Hexabdella brevitarsis Hernandes, 2013: 62; Covey Hill, Quebec, Canada. Bdella brevitarsis Banks (in Tothill 1919: 195���196) (nomen nudum). Type deposition. MCZ. 56. Hexabdella cinquaginta Hernandes, Daud & Feres, 2007: 60; Itiquira, Mato Grosso, Brazil, ex Hevea brasiliensis M.Arg. (Euphorbiaceae). Other record: S��o Paulo, Brazil (Demite & Feres 2008). Remarks. Male unknown. Type deposition. DZSJRP. 57. Hexabdella denheyeri van Der Schyff, Theron & Ueckermann, 2004: 15; Angola, ex Eugenia brasiliensis Lam. (Myrtaceae). Type deposition. NCA. 58. Hexabdella maraugia van Der Schyff, Theron & Ueckermann, 2004: 17; Kleinmond, Western Cape Province, South Africa, ex soil under dune vegetation. Remarks. Male unknown. Type deposition. NCA. 59. Hexabdella mexicana (Baker & Balock, 1944): 181; Valle del Bravo, Mexico, ex moss.���van Der Schyff et al., 2004: 13. Original designation: Bdella mexicana Baker & Balock. a. Bdella willisi Baker & Balock, 1944: 182; Drummond, 1957: 142; synonymy according to Atyeo (1960a: 383). Distribution. Mexico (Valle del Bravo, M��xico; Laguna de Zempoala, Morelos; San Luis Potos��, Veracruz), United States (Texas, Kansas, Illinois, Maryland, New York, Connecticut) (Atyeo 1960a) (Wisconsin) (Oatman 1963), Ukraine (Kuznetsov & Livshits 1979a), United States (Lehman 1982), Hawaii (Swift & Goff 1987). Redescriptions. Atyeo (1960a), Swift & Goff (1987), van Der Schyff et al. (2004). Remarks. Atyeo (1960a) set Bdella willisi as a synonym of this species, but mentions that individuals collected from the United States have hysterosomal setae pilose, whereas those of the Mexican specimens are nude. Type deposition. USNM. 60. Hexabdella miranda van Der Schyff, Theron & Ueckermann, 2004: 20; Limpopo Province, South Africa, ex Euphorbia ingens E.Mey (Euphorbiaceae). Remarks. Male unknown. Type deposition. NCA. 61. Hexabdella persiaensis Paktinat-Saeej & Bagheri, 2014: 3; Mazandaran province, Iran, from soil and rotten leaves under Corylus avellana (Betulaceae). Remarks. Male and immatures unknown. Type deposition. Acarological Collection, Department of Plant Protection, University of Maragheh, Iran, and ASI. 62. Hexabdella quercusi Eghbalian, Khanjani, Safaralizadeh & Ueckermann, 2016: 292; ex litter under oak trees, Western Iran. Type deposition. Holotype and paratypes at University of Bu-Ali Sina, Hamedan, Iran; paratype female at NCA. 63. Hexabdella singula van Der Schyff, Theron & Ueckermann, 2004: 14; Cape Town, South Africa, ex Cassine peragua L. (Celastraceae). Remarks. Male and immatures unknown. Type deposition. NCA. 64. Hexabdella unusoculata van Der Schyff, Theron & Ueckermann, 2004: 24; KwaZulu-Natal, South Africa, ex soil. Remarks. Male unknown. Type deposition. NCA., Published as part of Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, pp. 1-83 in Zootaxa 4152 (1) on pages 18-19, DOI: 10.11646/zootaxa.4152.1.1, http://zenodo.org/record/261900, {"references":["van Der Schyff, J., Theron, P. D. & Ueckermann, E. A. (2004) Hexabdella, a new mite genus of Bdellidae (Acari: Prostigmata) from southern Africa, with descriptions of five new species. African Plant Protection, 10 (1), 13 - 25.","Baker, E. W. & Balock, J. W. (1944) Mites of the family Bdellidae. Proceedings of the Entomological Society of Washington, 46 (7), 176 - 184.","Hernandes, F. A. (2013) Revision of Nathan Banks' type specimens of Bdellidae Duges (Acari: Trombidiformes) of the Museum of Comparative Zoology, Cambridge. International Journal of Acarology, 39 (1), 1 - 9. http: // dx. doi. org / 10.1080 / 01647954.2012.739642","Tothill, J. D. (1919) Some notes on the natural control of the oyster-shell scale (Lepidosaphes ulmi, L.). Bulletin of Entomological Research, 9 (3), 183 - 196. http: // dx. doi. org / 10.1017 / S 0007485300037949","Hernandes, F. A., Daud, R. D. & Feres, R. J. F. (2007) A new species of Hexabdella (Acari: Bdellidae) from Brazil. Zootaxa, 1501, 57 - 63.","Demite, P. R. & Feres, R. J. F. (2008) Influence of cerrado fragments in the distribution of mites in rubber tree crop. Neotropical Entomology, 37 (2), 196 - 204. http: // dx. doi. org / 10.1590 / S 1519 - 566 X 2008000200015","Drummond, R. O. (1957) Observations on the fluctuations of acarine populations from nests of Peromyscus leucopus. Ecological Monographs, 27 (2), 137 - 152. http: // dx. doi. org / 10.2307 / 1948573","Atyeo, W. T. (1960 a) A revision of the family Bdellidae in North and Central America (Acarina: Prostigmata). University of Kansas Science Bulletin, 40, 345 - 499.","Oatman, E. R. (1963) Mite species on apple foliage in Wisconsin. In: Advances in Acarology, Naegele, J. A. (Ed.), vol. 1, Comstock Pub. Assoc., 21 - 24.","Kuznetsov, N. N. & Livshits, I. Z. (1979 a) Predatory mites of the Nikita Botanical Garden (Acariformes: Bdellidae, Cunaxidae, Camerobiidae). Proceedings of the State Nikita Botanical Garden, 79, 51 - 104.","Lehman, R. D. (1982) Mites (Acari) of Pennsylvania conifers. Transactions of the America Entomological Society, 180, 181 - 286.","Swift, S. F. & Goff, M. L. (1987) The family Bdellidae (Acari: Prostigmata) in the Hawaiian Islands. International Journal of Acarology, 13 (1), 29 - 49. http: // dx. doi. org / 10.1080 / 01647958708683478","Paktinat-Saeej, S., Bagheri, M., Saboori, A. & Ueckermann, E. A. (2014) Hexabdella persiaensis sp. nov. (Acari: Prostigmata: Bdellidae) as a first new species of the genus Hexabdella from Asia. International Journal of Acarology, 40, 1 - 6. http: // dx. doi. org / 10.1080 / 01647954.2014.928366","Eghbalian, A. H., Khanjani, M., Safaralizadeh, M. H. & Ueckermann, E. A. (2016) New species of Hexabdella and Neomolgus (Acari: Prostigmata: Bdellidae) from Iran. Zootaxa, 4072 (2), 291 - 300. http: // dx. doi. org / 10.11646 / zootaxa. 4072.2.10"]}

Bdellidae (Trombidiformes: Prostigmata) are moderate to large sized predatory mites that inhabit soil, leaves, leaf litter, and intertidal rocks. They are readily recognized by an elongated, snout-like gnathosoma and by elbowed pedipalps bearing two (one in Monotrichobdella Baker & Balock) long terminal setae. Despite being among the first mites ever described, with species described by Carl Linnaeus, the knowledge about bdellids has never been compiled into a taxonomic catalogue. Here we present a catalogue listing 278 valid species; for each species we include distribution information, taxonomic literature, and type depository institutions. The genus Rigibdella Tseng, 1978 is considered a junior synonym of Cyta von Heyden, 1826, and Bdellodes Oudemans, 1937 is considered a junior synonym of Odontoscirus Tohr, 1913. Illustrated keys to subfamilies and genera are presented, as well as keys to species of each genus.

A new species of the genus Hexabdella Van der Schyff, Theron & Ueckermann, Hexabdella cinquaginta sp.n. is herein described, representing the second species of the family Bdellidae from the neotropical region. It can be distinguished from H. singula by its lobed striae on lateral margins of hysterosoma and by the leg chaetotaxy. A key to species of the genus is presented.

Biscirus Thor, 1913 Type species: Bdella silvaticus Kramer, 1881 [original designation] Diagnosis. Hypostome with two pairs of prominent ventral setae; with or without median prodorsal eye; genital tracheae well-developed; chelicerae elongated, movable chela reduced and needle-like; sometimes with unpaired seta between coxae IV; tibiae I, IV and tarsi III, IV with trichobothria; external vertical setae (lps) absent; palp tibiotarsus cylindrical and elongate., Published as part of Paktinat-Saeej, Saeed, Bagheri, Mohammad, Saboori, Alireza & Ahaniazad, Mansoureh, 2015, Two new Bdellidae (Trombidiformes: Bdelloidea) from Iran and the status of Neobiscirus Gomelauri, 1963, pp. 519-530 in Zootaxa 4013 (4) on page 520, DOI: 10.11646/zootaxa.4013.4.3, http://zenodo.org/record/231988, {"references":["Thor, S. (1913) Biscirus genus novum. Eine neue Bdelliden Gattung und zwei neue Untergattugen. Zoologischen Anzeiger, 42, 28 - 30.","Kramer, P. (1881) Ueben Milben; 4. Ueber Scirus taurus n. spec. Zeitschrift fur die gesammten Naturwissenschaft, 54, 418 - 452."]}

Paktinat-Saeej, Saeed, Bagheri, Mohammad, Saboori, Alireza, Ahaniazad, Mansoureh (2015): Two new Bdellidae (Trombidiformes: Bdelloidea) from Iran and the status of Neobiscirus Gomelauri, 1963. Zootaxa 4013 (4): 519-530, DOI: http://dx.doi.org/10.11646/zootaxa.4013.4.3

Biscirus iranensis Paktinat-Saeej & Bagheri sp. nov. (Figs. 1–17) Description. Male (n = 3): Dimensions: Length of body (including gnathosoma) 1240 (1137–1225), width of body 455 (438–495), length of gnathosoma 420 (345–412), length of chelicera 366 (310–368); leg lengths: I 737 (590– 725), II 705 (535–700), III 820 (645–805), IV 962 (788–925); VES 180 (156–190), DES 252 (238–282); palpomeres I–V: 21 (14–18), 233 (190–222), 37 (27–30), 63 (40–52), 162 (117–132); at ? (220), pt ? (218), mps 52 (52), c 1 46 (43–50), c 2 50 (52), d 1 45 (45), e 1 56 (54), f 1 62 (57–63), f 2 58 (55–58), h 1 63 (60), h 2 61 (53); distance: at–at 82 (73), pt–pt 240 (243), c 1 –c 2 138 (140), c 1 –d 1 133 (116–125). Gnathosoma (Figs. 1–3): Two pairs of ventral hypostomal setae longitudinally aligned (vh 1 –vh 2) (Fig. 1); hypostome ending in two lateral lips, bearing two adoral setae ad 1 24 (23), ad 2 25 (24) and with longitudinal striations, which are transverse at base. Chelicera (Figs. 2, 14) with longitudinal striae and bear two setae, proximal seta 80 (58–72) longer than distal seta 58 (46–52); movable digit smooth and with two denticles, fixed digit straight, smooth and slightly shorter than movable digit. Palp (Fig. 3) chaetotaxy: trochanter 0, basifemur 2 sts, telofemur 1 sts, genu 2 sts, tibiotarsus 1 sts, 1 s, 2 long end setae (VES, DES). Setae on telofemur and genu finely pilose. Dorsum (Figs. 4–7, 15– 17): at and pt slender and nude. Striae on propodosoma transverse between setae at but center of propodosoma with roughly broken striations and lateral margins longitudinally striated (Fig. 4); two pairs of posterolateral eyes with oblique striae between each pair and one median eye present (Figs. 4, 15). Dorsal striae of hysterosoma continuous to sparsely broken; all striae ornamented with papillae-like patterns (Figs. 5–6, 16– 17); dorsal setae minutely pilose (Fig. 7). Venter (Figs. 8–9): Ventral setae nude; genital plates each with 12–16 setae arranged in two rows; 12–15 aggenital setae present, first pair between coxae IV; anal valves with three pairs of smooth pseudanal setae (ps 1 – ps 3). Amphioid sclerite with nine pairs of simple setae (Fig. 9). Legs (Figs. 10–13): Leg chaetotaxy: coxae I–IV 4 (5)- 3-6 (7)- 4 sts; trochantera I–IV 1 - 1 - 1 - 1 sts; basifemora I– IV 9 (10)- 9 t- 8 (9)- 4 sts; telofemora I–IV 5 - 5 (4)- 4 - 4 sts; genua I–IV 5 sts, 5 (4)asl- 5 sts, 1 asl- 5 sts, 1 asl- 4 sts, 1 asl; tibiae I–IV 13 (11)sts, 4 asl, 1 bsl, 1 pe, 1 T- 14 (11 / 13)sts, 1 asl, 1 bsl- 14 (10 / 12)sts, 1 asl- 12 sts, 1 T; tarsi I–IV 29 sts, 2 asl, 2 bsl, 1 pe- 30 sts, 2 bsl- 28 sts, 1 T- 25 sts, 1 asl, 1 T. Female (n = 1): Similar to male, but with the following specifications. Dimensions: Length of body (including gnathosoma) 1025, width of body 465, length of gnathosoma 338, length of chelicera 298; leg lengths: I 570, II 537, III 641, IV 745; VES 148, DES 230; palpomeres I–V: 14, 182, 31, 34, 105; at 175, pt 172, mps 48, c 1 42, c 2 46, d 1 43, e 1 44, f 1 62, f 2 57, h 1 60, h 2 51; distance: at–at 60, pt–pt 195, c 1 –c 2 122, c 1 –d 1 115. Ventral side with 15 genital setae on each plate and 15–16 aggenital setae. Leg chaetotaxy: coxae I–IV 4 - 3 -5(7)- 4 sts; trochantera I–IV 1 - 1 - 1 - 1 sts; basifemora I–IV 9 - 9 - 8 (9)- 4 sts; telofemora I–IV 5 - 4 - 4 - 4 sts; genua I–IV 5 sts, 4 asl- 5 sts, 1 asl- 5 sts, 1 asl- 4 sts, 1 asl; tibiae I–IV 11 sts, 4 asl, 1 bsl, 1 pe, 1 T- 10 sts, 1 asl, 1 bsl- 10 sts, 1 asl- 12 (10)sts, 1 T; tarsi I–IV 30 sts, 2 asl, 2 bsl, 1 pe- 29 sts, 2 bsl- 28 sts, 1 T- 25 sts, 1 asl, 1 T. Remarks. Biscirus iranensis Paktinat-Saeej & Bagheri sp. nov. resembles Biscirus silvaticus (Kramer, 1881) and B. arenarius Wallace & Mahon, 1972 in having the same solenidotaxy of tarsi I–IV; and resembles B. kobachidzei comb. nov. in having a median eye. B. iranensis sp. nov. can be distinguished from B. silvaticus by (1) presence of one attenuated solenidion on genua IV (absent in B. silvaticus); (2) presence of one blunt solenidion on tibiae I (absent in B. silvaticus); (3) all eugenital setae on amphioid sclerites are simple (branched and forked in B. silvaticus); (4) coxae II and telofemora IV with 3 and 4 setae, respectively (opposed to 4 and 3 in B. silvaticus). B iranensis sp. nov. differs from B. arenarius by (1) having four and one attenuated solenidion on tibiae I and III, respectively (rather than three attenuated solenidia on tibiae I and one blunt solenidion on tibiae III in B. arenarius); (2) having a median eye on the propodosoma (absent in B. arenarius); (3) coxae II and IV with 3 and 4 setae, respectively (rather than 4 and 3 in B. arenarius). B iranensis sp. nov. differs from Biscirus kobachidzei in having 9 simple eugenital setae on the amphioid sclerites (10 branched amphioid setae B. kobachidzei). Etymology. The specific epithet was derived from the country of origin, Iran. Type material. Holotype and one paratype males were collected from the soil and rotten leaves under citrus trees, 17 June 2013, Amreh village, Sari city, Mazandaran Province, by Saeed Paktinat-Saeej and one paratype male and one paratype female were collected from the soil and humus under apple trees, 25 September 2014, Azarshahr city, East Azerbaijan Province, Iran by Mansoureh Ahaniazad. The holotype male, one paratype male and one paratype female, are deposited in the Acarological Collection, Department of Plant Protection, Faculty of Agriculture, University of Maragheh, Maragheh, Iran and one paratype male is deposited in the Acarological Collection, Acarological Society of Iran (ASI), Department of Plant Protection, Faculty of Agriculture, University of Tehran, Karaj, Iran.

Biscirus iranensis Paktinat-Saeej & Bagheri sp. nov. (Figs. 1���17) Description. Male (n = 3): Dimensions: Length of body (including gnathosoma) 1240 (1137���1225), width of body 455 (438���495), length of gnathosoma 420 (345���412), length of chelicera 366 (310���368); leg lengths: I 737 (590��� 725), II 705 (535���700), III 820 (645���805), IV 962 (788���925); VES 180 (156���190), DES 252 (238���282); palpomeres I���V: 21 (14���18), 233 (190���222), 37 (27���30), 63 (40���52), 162 (117���132); at ? (220), pt ? (218), mps 52 (52), c 1 46 (43���50), c 2 50 (52), d 1 45 (45), e 1 56 (54), f 1 62 (57���63), f 2 58 (55���58), h 1 63 (60), h 2 61 (53); distance: at���at 82 (73), pt���pt 240 (243), c 1 ���c 2 138 (140), c 1 ���d 1 133 (116���125). Gnathosoma (Figs. 1���3): Two pairs of ventral hypostomal setae longitudinally aligned (vh 1 ���vh 2) (Fig. 1); hypostome ending in two lateral lips, bearing two adoral setae ad 1 24 (23), ad 2 25 (24) and with longitudinal striations, which are transverse at base. Chelicera (Figs. 2, 14) with longitudinal striae and bear two setae, proximal seta 80 (58���72) longer than distal seta 58 (46���52); movable digit smooth and with two denticles, fixed digit straight, smooth and slightly shorter than movable digit. Palp (Fig. 3) chaetotaxy: trochanter 0, basifemur 2 sts, telofemur 1 sts, genu 2 sts, tibiotarsus 1 sts, 1 s, 2 long end setae (VES, DES). Setae on telofemur and genu finely pilose. Dorsum (Figs. 4���7, 15��� 17): at and pt slender and nude. Striae on propodosoma transverse between setae at but center of propodosoma with roughly broken striations and lateral margins longitudinally striated (Fig. 4); two pairs of posterolateral eyes with oblique striae between each pair and one median eye present (Figs. 4, 15). Dorsal striae of hysterosoma continuous to sparsely broken; all striae ornamented with papillae-like patterns (Figs. 5���6, 16��� 17); dorsal setae minutely pilose (Fig. 7). Venter (Figs. 8���9): Ventral setae nude; genital plates each with 12���16 setae arranged in two rows; 12���15 aggenital setae present, first pair between coxae IV; anal valves with three pairs of smooth pseudanal setae (ps 1 ��� ps 3). Amphioid sclerite with nine pairs of simple setae (Fig. 9). Legs (Figs. 10���13): Leg chaetotaxy: coxae I���IV 4 (5)- 3-6 (7)- 4 sts; trochantera I���IV 1 - 1 - 1 - 1 sts; basifemora I��� IV 9 (10)- 9 t- 8 (9)- 4 sts; telofemora I���IV 5 - 5 (4)- 4 - 4 sts; genua I���IV 5 sts, 5 (4)asl- 5 sts, 1 asl- 5 sts, 1 asl- 4 sts, 1 asl; tibiae I���IV 13 (11)sts, 4 asl, 1 bsl, 1 pe, 1 T- 14 (11 / 13)sts, 1 asl, 1 bsl- 14 (10 / 12)sts, 1 asl- 12 sts, 1 T; tarsi I���IV 29 sts, 2 asl, 2 bsl, 1 pe- 30 sts, 2 bsl- 28 sts, 1 T- 25 sts, 1 asl, 1 T. Female (n = 1): Similar to male, but with the following specifications. Dimensions: Length of body (including gnathosoma) 1025, width of body 465, length of gnathosoma 338, length of chelicera 298; leg lengths: I 570, II 537, III 641, IV 745; VES 148, DES 230; palpomeres I���V: 14, 182, 31, 34, 105; at 175, pt 172, mps 48, c 1 42, c 2 46, d 1 43, e 1 44, f 1 62, f 2 57, h 1 60, h 2 51; distance: at���at 60, pt���pt 195, c 1 ���c 2 122, c 1 ���d 1 115. Ventral side with 15 genital setae on each plate and 15���16 aggenital setae. Leg chaetotaxy: coxae I���IV 4 - 3 -5(7)- 4 sts; trochantera I���IV 1 - 1 - 1 - 1 sts; basifemora I���IV 9 - 9 - 8 (9)- 4 sts; telofemora I���IV 5 - 4 - 4 - 4 sts; genua I���IV 5 sts, 4 asl- 5 sts, 1 asl- 5 sts, 1 asl- 4 sts, 1 asl; tibiae I���IV 11 sts, 4 asl, 1 bsl, 1 pe, 1 T- 10 sts, 1 asl, 1 bsl- 10 sts, 1 asl- 12 (10)sts, 1 T; tarsi I���IV 30 sts, 2 asl, 2 bsl, 1 pe- 29 sts, 2 bsl- 28 sts, 1 T- 25 sts, 1 asl, 1 T. Remarks. Biscirus iranensis Paktinat-Saeej & Bagheri sp. nov. resembles Biscirus silvaticus (Kramer, 1881) and B. arenarius Wallace & Mahon, 1972 in having the same solenidotaxy of tarsi I���IV; and resembles B. kobachidzei comb. nov. in having a median eye. B. iranensis sp. nov. can be distinguished from B. silvaticus by (1) presence of one attenuated solenidion on genua IV (absent in B. silvaticus); (2) presence of one blunt solenidion on tibiae I (absent in B. silvaticus); (3) all eugenital setae on amphioid sclerites are simple (branched and forked in B. silvaticus); (4) coxae II and telofemora IV with 3 and 4 setae, respectively (opposed to 4 and 3 in B. silvaticus). B iranensis sp. nov. differs from B. arenarius by (1) having four and one attenuated solenidion on tibiae I and III, respectively (rather than three attenuated solenidia on tibiae I and one blunt solenidion on tibiae III in B. arenarius); (2) having a median eye on the propodosoma (absent in B. arenarius); (3) coxae II and IV with 3 and 4 setae, respectively (rather than 4 and 3 in B. arenarius). B iranensis sp. nov. differs from Biscirus kobachidzei in having 9 simple eugenital setae on the amphioid sclerites (10 branched amphioid setae B. kobachidzei). Etymology. The specific epithet was derived from the country of origin, Iran. Type material. Holotype and one paratype males were collected from the soil and rotten leaves under citrus trees, 17 June 2013, Amreh village, Sari city, Mazandaran Province, by Saeed Paktinat-Saeej and one paratype male and one paratype female were collected from the soil and humus under apple trees, 25 September 2014, Azarshahr city, East Azerbaijan Province, Iran by Mansoureh Ahaniazad. The holotype male, one paratype male and one paratype female, are deposited in the Acarological Collection, Department of Plant Protection, Faculty of Agriculture, University of Maragheh, Maragheh, Iran and one paratype male is deposited in the Acarological Collection, Acarological Society of Iran (ASI), Department of Plant Protection, Faculty of Agriculture, University of Tehran, Karaj, Iran., Published as part of Paktinat-Saeej, Saeed, Bagheri, Mohammad, Saboori, Alireza & Ahaniazad, Mansoureh, 2015, Two new Bdellidae (Trombidiformes: Bdelloidea) from Iran and the status of Neobiscirus Gomelauri, 1963, pp. 519-530 in Zootaxa 4013 (4) on pages 520-523, DOI: 10.11646/zootaxa.4013.4.3, http://zenodo.org/record/231988, {"references":["Kramer, P. (1881) Ueben Milben; 4. Ueber Scirus taurus n. spec. Zeitschrift fur die gesammten Naturwissenschaft, 54, 418 - 452.","Wallace, M. M. H. & Mahon, J. A. (1972) The taxonomy and biology of Australian Bdellidae (Acari). I. Subfamilies Bdellinae, Spinibdellinae and Cytinae. Acarologia, 14 (4), 544 - 580."]}

Bdella farabii Paktinat-Saeej & Bagheri sp. nov. (Figs. 18���30) Female (n = 2): Dimensions: Length of body (including gnathosoma) 1150 (1025), width of body 638 (485), length of gnathosoma 263 (250), length of chelicera 245 (237); leg lengths: I 575 (570), II 555 (540), III 637 (612), IV 752 (732); VES 172 (174), DES 210 (202); palpomeres I���V: 18 (20), 140 (145), 35 (35), 22 (25), 75 (76); at 117 (122), lps 80 (78), pt 145 (158), mps 85 (82), c 1 76 (75), c 2 100 (93), d 1 80 (80), e 1 84 (77), f 1 70 (67), f 2 71 (70), h 1 82 (76), h 2 76 (65); distance: at���at 130 (121), pt���pt 136 (121), c 1 ���c 2 180 (115), c 1 ���d 1 125 (95). DHS 37 (34). Gnathosoma (Figs. 18���20): Six pairs of ventral hypostomal setae longitudinally aligned (vh 1 ���vh 6) (Fig. 18); hypostome terminates in two lateral lips, bearing two adoral setae ad 1 9 (9), ad 2 12 (13) and with sparsely longitudinal striations, which are transverse at base. Chelicera (Fig. 19) with longitudinal striae and with two setae, distal seta 80 (76) longer than proximal seta 55 (52); movable and fixed digit smooth, straight and of same length. Palp (Fig. 20) chaetotaxy: trochanter 0, basifemur 7 sts, telofemur 1 sts, genu 4 sts, tibiotarsus 3 sts, 1 s, 2 long end setae (VES, DES). Dorsum (Figs. 21���23): at and pt slender and nude (Fig. 21). Center of prodorsum with continuous to sparsely broken longitudinal striations (Fig. 21); two internal apodemes present (Fig. 21); two pairs of eyes posterolateral to pt with transverse striae between each pair present. Setae lps closer to at than to pt. Dorsal striae of hysterosoma with continuous to sparsely broken striae (Fig. 22); dorsal setae smooth to minutely barbed (Fig. 23). Venter (Figs. 24���26): Ventral setae nude (Fig. 24); genital plates each with 9 setae longitudinally aligned; 10 aggenital setae present (Fig. 25); anal valves with 3 pairs of smooth pseudanal setae (ps 1 ���ps 3), ps 1 68 (62), ps 2 51 (51), ps 3 48 (45); with 1 pair of setae (v 1) between coxae IV. Ovipositor (Fig. 26) present and has 12 subapical and 6 medial setae. Legs (Figs. 27���30): Leg chaetotaxy: coxae I���IV 6 (5)- 5-6 (7)- 4 sts; trochantera I���IV 1 - 1 -2- 1 sts; basifemora I��� IV 8 (7)- 7 (8)- 6 (7)- 3 sts; telofemora I���IV 7 - 7 -8- 7 sts; genua I���IV 7 sts, 1 duplex (dxs), 1 asl- 8 sts, 1 duplex- 8 sts, 1 duplex- 8 sts, 1 duplex; tibiae I���IV 8 sts, 2 asl, 1 pe, 1 T- 9 sts, 1 asl, 1 bsl- 9 sts, 1 asl- 8 sts, 1 T; tarsi I���IV 27 (28)sts, 2 asl, 2 bsl, 1 pe��� 26 (25)sts, 2 bsl��� 24 sts, 1 T��� 23 sts, 1 asl, 1 T. Male and Immature stages. Unknown Remarks. Bdella farabii Paktinat-Saeej & Bagheri sp. nov. is similar to B. muscorum Ewing 1909, B. kuznetsovi Maslov & Khaustov 2013, and B. longistriata Atyeo 1960. B. farabii sp. nov. is similar to B. muscorum in having duplex setae on genua I���III but can be distinguished by (1) tibiae I with two attenuated solenidia instead of four attenuated solenidia in B. muscorum; (2) genua IV with duplex setae which is absent in B. muscorum; (3) prodorsum with longitudinal striations instead of transverse striations in B. muscorum; (4) tarsus II in new species lacks the peg; (5) internal apodemes in new species weakly developed vs. distinctly developed apodemes in B. muscorum. B. farabii sp. nov. resembles B. kuznetsovi in having weakly developed internal apodemes and longitudinal striations on prodorsum but can be distinguished by (1) genua I���IV with duplex setae which is absent in B. kuznetsovi; (2) tibiae I with two attenuated solenidia and one peg vs. four attenuated solenidia and without peg in B. kuznetsovi; (3) tarsus II in new species lacks the peg. B. farabii sp. nov. resembles B. longistriata in having longitudinal striations on the prodorsum and duplex setae on genua I��� IV but it can be separated by (1) different palp chaetotaxy; (2) having 9 genital setae on each genital plate vs. 11 setae in B. longistriata; (3) tarsus II in new species lacks the peg; (4) internal apodemes in the new species weakly developed vs. distinctly developed apodemes in B. longistriata. Etymology. This species is named after���Abu Na���r Mu���ammad Farabi���(872 A.D��� 950 A.D), who was a Persian writer specializing in political philosophy, metaphysics, ethics, and logic. Type material. Holotype female was collected from soil and rotten leaves under citrus trees, 30 October 2013, Saeej-Mahalleh village, Amol city, Mazandaran Province, Iran, by Saeed Paktinat-Saeej and one paratype female was collected from the soil and humus under apple trees, 25 September 2014, Azarshahr city, East Azerbaijan Province, Iran by Mansoureh Ahaniazad. The holotype female is deposited in the Acarological Collection, Department of Plant Protection, Faculty of Agriculture, University of Maragheh, Maragheh, Iran and one paratype female is deposited in the Acarological Collection, Acarological Society of Iran (ASI), Department of Plant Protection, Faculty of Agriculture, University of Tehran, Karaj, Iran., Published as part of Paktinat-Saeej, Saeed, Bagheri, Mohammad, Saboori, Alireza & Ahaniazad, Mansoureh, 2015, Two new Bdellidae (Trombidiformes: Bdelloidea) from Iran and the status of Neobiscirus Gomelauri, 1963, pp. 519-530 in Zootaxa 4013 (4) on pages 524-526, DOI: 10.11646/zootaxa.4013.4.3, http://zenodo.org/record/231988, {"references":["Ewing, H. E. (1909) Three new species of the genus Bdella (mites). Canadian Entomologist, 41 (4), 124 - 125. http: // dx. doi. org / 10.4039 / ent 41122 - 4","Maslov, S. I. & Khaustov, A. A. (2013) A new species of mites of the genus Bdella (Acari, Bdellidae) from Ukraine. Vestnik zoologii, 47 (2), 52 - 56. http: // dx. doi. org / 10.2478 / vzoo- 2013 - 0015","Atyeo, W. T. (1960) A revision of the mite family Bdellidae in North and Central America (Acarina, Prostigmata). The University of Kansas Science Bulletin, 40 (8), 345 - 499."]}

Biscirus kobachidzei (Gomelauri, 1963) comb. nov. Neobiscirus kobachidzei Gomelauri, 1963: 167. Distribution. Georgia S.S.R. Remarks. The monospecific Neobiscirus Gomelauri, 1963 differs from Biscirus by the presence of a median eye and two denticles on the movable digit. These characters are continuous with multiple genera in most subfamilies (Hernandes 2013), including Biscirus, which is often overlooked as possessing a median eye (Alberti 1975). Therefore, we do not consider Neobiscirus a distinct genus, and instead suggest the new combination Biscirus kobachidzei (Gomelauri, 1963)., Published as part of Paktinat-Saeej, Saeed, Bagheri, Mohammad, Saboori, Alireza & Ahaniazad, Mansoureh, 2015, Two new Bdellidae (Trombidiformes: Bdelloidea) from Iran and the status of Neobiscirus Gomelauri, 1963, pp. 519-530 in Zootaxa 4013 (4) on page 520, DOI: 10.11646/zootaxa.4013.4.3, http://zenodo.org/record/231988, {"references":["Gomelauri, L. A. (1963) On the study of mites of the Bdellidae family in Georgia S. S. R. Soobshcheniya Akademii Nauk Gruzinskoy S. S. R., 30 (2), 47 - 51.","Alberti, G. (1975) Pralarven und Phylogenie der Schnabelmilben (Bdellidae, Trombidiformes). Zeitschrift fur zoologische Systematik und Evolutions-Forschung, 13 (1), 44 - 62. http: // dx. doi. org / 10.1111 / j. 1439 - 0469.1975. tb 00227. x"]}