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Genus Microcancilla Dall, 1924 Type species: Admete microscopica (Dall, 1889) [= Cancellaria microscopica Dall, 1889], by original designation. Recent, Caribbean.

Fusceulima watsoni sp. nov. urn:lsid:zoobank.org:act: 2C702913-5836-4DC0-ABA1-1B4ED3E518B0 (Figure 10 A–I) Type material. Holotype NMW 1955.158.11589 (Figure 10A); Paratypes: MNRJ 23054 † [1 dd] (Figure 10C, D), MNRJ 35264 [9 dd] (Figure 10 E–I), Off Rio Grande do Norte, REVIZEE Nordeste (04°51′40″S 35°08′01″W, 384 m), coll. RV Natureza, 24/xi/2001; NMW 1955.158.27216 [1 dd] (Figure 10B), from the type locality. Type locality. Brazil: Pernambuco, Challenger stn. 122. Etymology. After Rev. Robert Boog Watson (1823–1910), who described several eulimids from deep waters off Northeast Brazil collected during the Challenger Expedition. Description (Holotype). Shell subconical, apex broad, obtuse, reaching 2.8 mm long, 1.0 mm wide. Protoconch-teleoconch transition not visible. Whole shell with about 5.5 whorls, slightly sinuous outline, convexity confined to the lower region of each whorl; suture shallow, slightly impressed, sloping; surface glossy and smooth; incremental scar weak, only one appearing on the body whorl. Last whorl about 65% of the shell length; base elongated, slightly rounded outline. Aperture high, pear shaped, acute posteriorly and rounded anteriorly; outer lip thin, slightly sinuous, almost orthocline, projecting gently and gradually from suture to a point below the middle of its height and retracting gently and gradually in the distal region; inner lip sinuous, sloping and well demarcated. Not umbilicate. Shell vitreous or whitish. Measurements. Holotype NMW 1955.158.11589, 5.5 whorls, SL= 2.8 mm; BWL= 1.8 mm; AL= 1.2 mm; SW= 1.0 mm; AW= 0.6 mm. Paratype MNRJ 23054, 6 whorls, SL= 3.0 mm; BWL= 1.8 mm; AL= 1.2 mm; SW= 1.0 mm; AW= 0.6 mm. Remarks. The paratypes MNRJ 23054 (Figure 10C, D) and MNRJ 35264 (Figure 10 E–I) were destroyed during the fire in the MNRJ, but the types in the NMW are safely preserved. The shell of this species is extremely smooth, except for the small wrinkles in the suture of the protoconch (Figure 10J). The protoconch-teleoconch transition is not clearly visible, but by the sculpture and shape of the larval shell this species possibly has non-planktotrophic development, like other species of Fusceulima (Souza & Pimenta 2014). A single incremental scar is present on the body whorl. Fusceulima watsoni sp. nov. resembles F. toffee Souza & Pimenta, 2014, from the continental shelf of Southeast Brazil, by the similar shape of the shell and of the aperture, but has less convex teleoconch whorls and is whitish or completely vitreous (Figure 10 A–D) while F. toffee has a dark brown color (Souza & Pimenta 2014: figure 46–47). Fusceulima watsoni sp. nov. can be distinguished from F. victorhensenae Hoffman & Freiwald, 2017, from western Morocco, by being larger (paratype MNRJ 23054, 6 whorls, SL= 3.0 mm vs. holotype of F. victorhensenae, 6 whorls, SL= 1.8 mm) (Hoffman & Freiwald 2017), with less convex teleoconch whorls and a less protruding outer lip. Fusceulima watsoni sp. nov. differs from other Atlantic congeners mainly by the more conical shell (Figure 10 A–E) and by the higher aperture (Figure 10F). Most of deep-sea Fusceulima from the Atlantic have a more cylindrical shell and a shorter aperture, for example: F. coralensis Hoffmann & Freiwald, 2017, from western Morocco; F. minuta (Jeffreys, 1884) (Souza & Pimenta 2014: figs. 29–31), from the Northeast Atlantic; F. sordida (Watson, 1897) (Figure 10K), from the Madeira archipelago; and F. thalassae Bouchet & Warén, 1986 (Figure 10L), from the Northeast Atlantic. Geographical distribution. Brazil: Rio Grande do Norte and Pernambuco (present study). Bathymetric distribution. From 384 m to 640 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 470-472, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Watson, R. B. (1897) On the Marine Mollusca of Madeira; with Descriptions of Thirty-five new Species, and an Index-List of all the known Sea-dwelling Species of that Island. Journal of the Linnean Society of London, 26 (168), 233 - 329. https: // doi. org / 10.1111 / j. 1096 - 3642.1897. tb 00033. x","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762","Souza, L. S. & Pimenta, A. D. (2014) Fusceulima and Halielloides (Gastropoda: Eulimidae) in the southwestern Atlantic, with descriptions of two new species of Fusceulima. Zoologia, Curitiba, 31 (6), 621 - 633. https: // doi. org / 10.1590 / S 1984 - 46702014000600008","Hoffman, L. & Freiwald, A. (2017) A unique and diverse amalgamated mollusk assemblage from the Coral Patch Seamount, eastern Atlantic. Miscellanea Malacologica, 7 (4), 61 - 79.","Jeffreys, J. G. (1884) On the Mollusca procured during the ' Lightning' and ' Porcupine' expeditions, 1868 - 70. Part VII. Proceedings of the Zoological Society of London, 52 (3), 341 - 372. https: // doi. org / 10.1111 / j. 1096 - 3642.1884. tb 02837. x"]}

Melanella sarissa (Watson, 1883) (Figure 12 A–L) Eulima sarissa Watson, 1883: 119–120. Eulima sarissa: Watson (1886: plate 37, figure 6a–b). Melanella cinca Dall, 1927: 65. syn. nov. Melanella sarissa: Lange-de-Morretes (1949: 83); Rios (1994: 104, plate 34, figure 430 [reproduced from original illustration]); Rios (2009: 192, figure 471 [reproduced from original illustration]). Type material. Holotype NHMUK 1887.2.9.1599 (Figure 12 A–B). Melanella cinca Dall, 1927: Syntype USNM 108028 (Figure 12D), USFC stn. 2668, off Georgia, USA. Type locality. Brazil: off Pernambuco, Challenger stn. 122. Material examined. Type material; USA: USFC stn. 2668: USNM 108036 b [4 dd], USNM 108036 h [2 dd]; USFC stn. 2415 (30°44′N 79°26′W, 805 m), coll. 01/iv/1885: USNM 108379 b [1 dd]. Puerto Rico: North of Culebra Island, off St. Thomas, Challenger stn. 24: NMW 1955.158.26254 [1 dd]; Brazil: Pernambuco: Challenger stn. 120 (08°37′S 34°28′W, 1234 m), coll. 09/ ix/1873: NMW 1955.158.00652 [3 dd]; Rio de Janeiro state, continental slope of Campos Basin: Oceanprof I stn. 44: IBUFRJ 19708 † [1 dd]; (22°22′37″S 40°06′28″W, 650 m), coll. RV Gyre, 22–24/viii/2009: MNRJ 28845 † [1 dd]. Measurements. Holotype NHMUK 1887.2.9.1599, 10 whorls, SL= 4.0 mm; BWL= 1.7 mm; AL= 1.1 mm; SW= 1.2 mm; AW= 0.7 mm. NMW 1955.158.26254, 10 whorls, SL= 4.0 mm; BWL= 1.8 mm; AL= 1.2 mm; SW= 1.2 mm; AW= 0.7 mm. MNRJ 28845, 10 whorls, SL= 3.8 mm; BWL= 1.8 mm; AL= 1.0 mm; SW= 1.2 mm; AW= 0.6 mm. IBUFRJ 19708, 9 whorls, SL= 3.5 mm; BWL= 1.7 mm; AL= 1.1 mm; SW= 1.1 mm; AW= 0.6 mm. Syntype of M. cinca, USNM 108028, 8 whorls, SL= 2.5 mm; BWL= 1.1 mm; AL= 0.7 mm; SW= 0.8 mm; AW= 0.5 mm. Remarks. Watson (1883) only reported material from Challenger stn. 122 in the original description. A survey in the NMW revealed other specimens of M. sarissa collected from Challenger stn. 24 (NMW 1955.158.26254), in Puerto Rico, and stn. 120 (NMW 1955.158.00652), in Pernambuco, but as they are from different localities, we do not include them in the type series. Thus, the shell of NHMUK 1887.2.9.1599 may be inferred as the holotype by monotypy. Shell features of Melanella sarissa fits the wide concept of the genus Melanella, however one specimen from MNRJ 28845 (Figure 12E) with a well-preserved shell shows brownish spiral bands at the suture of the protoconch. The holotype of M. sarissa is also well-preserved but has no color pattern. Nevertheless, the specimen from MNRJ 28845 (Figure 12E) fits perfectly in shape and size with the type of M. sarissa. Species of Melanella are usually glassy and lack brownish bands, but we maintain this classification for now. This species remained for decades without citations in the literature, Lange-de-Morretes (1949) listed Melanella sarissa in his catalogue of Brazilian mollusks based only on the original description. Rios (1970, 1975, 1985) did not list M. sarissa in his catalogues and included the species only in the most recent editions (Rios 1994, 2009). Melanella sarissa is rediscovered after more than a century, with new records from the southeast Brazil. Melanella cinca Dall, 1927 (Figure 12D), only known from the type locality off Fernandina, Florida, is a junior synonym of M. sarissa. The holotype of M. cinca is a young specimen but has a similar outline of the initial whorls and dimensions (Figure 12D), including the protoconch (Figure 12L). Besides these features, the aperture is also similar in shape to the specimens of M. sarissa from Brazil. Thus, the geographic distribution of M. sarissa is greatly enlarged, occurring from Florida, USA to Rio de Janeiro, Brazil. Based on the new material reported here, we can add to the original description that M. sarissa has 4.0 protoconch whorls, with a maximum diameter and height of 340 µm and 470 µm, respectively. Otherwise, the whole shell lacks microscopic sculpture, incremental scars are present and appear at irregular intervals, and the body whorl comprises about 45% of the shell length. Melanella sarissa resembles M. hians (Figure 9J) but differs by the proportionally smaller length (holotype of M. sarissa, ~10 whorls, SL= 4.0 mm vs. syntype of M. hians NHMUK 1887.2.9.1588, 10 whorls, SL= 10.01 mm). Melanella sarissa has similar dimensions to M. ira Dall, 1927, from off Georgia and Fernandina, USA, but differs by being more conical and by the larger lateral expansion of the aperture. Geographical distribution. USA: Georgia (Dall 1927); Puerto Rico (present study); Brazil: Pernambuco (Watson 1883) and Rio de Janeiro (present study). Bathymetric distribution. From 538 m to 806 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 474-476, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Watson, R. B. (1883) Mollusca of H. M. S. \" Challenger \" Expedition. Part XVII. Zoological Journal of the Linnean Society, 17 (99), 112 - 130. https: // doi. org / 10.1111 / j. 1096 - 3642.1883. tb 02043. x","Watson, R. B. (1886) Report on the Scaphopoda and Gasteropoda collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger, Zoology, 15 (2), 1 - 680 + 692 - 756, 50 pls.","Dall, W. H. (1927) Small shells from dredgings off the southeast coast of the United States by the United States Fisheries Steam- er \" Albatross \" in 1885 and 1886. Proceedings of the United States National Museum, 70 (2667), 1 - 134. https: // doi. org / 10.5479 / si. 00963801.70 - 2667.1","Lange-de-Morretes, F. (1949) Ensaio de catalogo dos moluscos do Brasil. Arquivos do Museu Paraense, 7, 1 - 216.","Rios, E. C. (1994) Seashells of Brazil. 2 nd Edition. Editora da Fundac \" o Universidade do Rio Grande, Rio Grande, 368 pp., 113 pls.","Rios, E. C. (2009) Compendium of Brazilian seashells. Evangraf, Rio Grande, viii + 668 pp.","Rios, E. C. (1970) Coastal Brazilian seashells. Museu Oceanografico do Rio Grande, Rio Grande, 250 pp., 50 pls.","Rios, E. C. (1975) Brazilian marine mollusks iconography. Museu Oceanografico do Rio Grande, Rio Grande, 331 pp., 91 pls.","Rios, E. C. (1985) Seashells of Brazil. Editora da Fundac \" o Universidade do Rio Grande, Rio Grande, 328 pp., 102 pls."]}

Melanella doederleini (Brusina, 1886) (Figure 11 A–I) Eulima stalioi auct. non Brusina, 1869: Jeffreys (1884: 368, plate 28, figure 3). Eulima doederleini Brusina, 1886: 214. Melanella doederleini (Brusina, 1886): Bouchet & Warén (1986: 382, figs. 902–903); Utrilla et al. (2020: 15, figure 4A–D). Type material. Lectotype (Bouchet & Warén 1986) NHMUK 1979229 (Figure 11A, B); Paralectotypes NHMUK 1885.11.5.2027–2028 [2 dd], USNM 131144 [1 dd] (not located). Type locality. Southwest of Cadiz, PORCUPINE 1870 stn. 30 (36°15′N 06°52′O, 706 m), coll. H.M.S. Porcupine. Material examined. Lectotype; paralectotypes NHMUK 1885.11.5.2027–2028; Brazil: Off Ceará, RE-VIZEE Nordeste (03°30′51″S 37°59′28″O, 384 m, coll. RV Natureza, 07/xi/2001): MNRJ 35277 † [2 dd]; Off Pernambuco, REVIZEE Nordeste (08°46′00″S 34°44′00″O, 690 m), coll. RV Natureza, 18/xi/2000): MNRJ 35208 † [4 dd]; Espírito Santo: REVIZEE-Central V stn. 25A (19°35′28″S 38°45′28″W, 230 m), coll. Supply Boat Astro Garoupa, 28/vi/2001: IBUFRJ † 19719 [1 dd]. Redescription (Lectotype). Shell conical, slightly curved at the apex, reaching about 3.6 mm long, 1.3 mm wide. Larval shell vitreous or whitish, almost cylindrical, with about 3.0 whorls; smooth, transition to teleoconch marked by an almost indistinct incremental scar. Teleoconch with about 7.0 whorls, flat outline; suture almost indistinct, sloping slowly; subsutural zone about 15% of the whorl height; surface smooth; incremental scars slightly demarcated, intervals close to 1.0 whorl. Last whorl about 50% of the shell length; base rounded. Aperture short, pear shaped, acute posteriorly and rounded anteriorly; outer lip thin, opisthocline, sinuous, slightly retracting near the suture and projecting in the uppermost third; inner lip sinuous, sloping, reflected, well demarcated. Not umbilicate. Shell vitreous, colorless. Measurements. Lectotype NHMUK 1979229, 10 whorls, SL= 3.6 mm; BWL= 1.7 mm; AL= 1.0 mm; SW= 1.3 mm; AW= 0.7 mm. MNRJ 35208†, 10 whorls, SL= 3.6 mm; BWL= 1.8 mm; AL= 1.0 mm; SW= 1.3 mm; AW= 0.7 mm. Remarks. This species has a slightly curved apex and a short and wide aperture; the placement in Melanella is more suitable than the original combination in Eulima. Brusina (1886) and Bouchet & Warén (1986) did not provide a detailed description of the species, therefore we redescribe the shell morphology of the lectotype here. We can add that the specimens from Brazil have a protoconch of 2.8 whorls, 280 μm wide and 250 μm in height. Melanella doederleini is a very conspicuous species. Shells collected in the West Atlantic are indistinguishable from the lectotype of M. doederleini (NHMUK 1979220; Figure 11A, B) and from the shells recorded by Utrilla et al. (2020: figure 4A–D) from the Gulf of Cádiz. A small variation is present in the outline of the base, which is more truncated in the lectotype (Figure 11A), while it is more rounded in shells from Brazil (Figure 11C, D) and in the shells figured by Utrilla et al. (2020). The geographic distribution of M. doederleini is extended by about 7,000 km (from type locality to the coast of Espírito Santo, Southeast Brazil). Hoffman & Freiwald (2020) recorded a few empty shells of “ Melanella cf. doederleini ” from the Azorean seamounts; a confirmed identification can fill part of the wide distributional gap. Bouchet & Warén (1986) noted that M. doederleini is a possible junior synonym of Melanella altavillensis (Seguenza, 1876), but highlighted a small difference in the protoconch of both species. We add that M. doederleini is relatively smaller in length but wider in comparison to the syntype of M. altavillensis figured by Bouchet & Warén (1986: figs. 904–905) and to the shells figured by Landau et al. (2006: plate 19, figure 2A–B). Melanella altavillensis is known only from fossils, while M. doederleini was considered to belong to the biocenosis in the Gulf of Cadiz, with representatives in the taxocoenosis and thanatocoenosis (Utrilla et al. 2020). Melanella doederleini resembles Eulima chyta Watson, 1883 (Figure 11J), only from its type locality in Ascension Island, but it is larger (MNRJ 35208†, 10 whorls, SL= 3.6 mm vs. syntype of E. chyta NHMUK 1887.2.9.1604, 9 whorls, SL= 2.1 mm), has a higher spire angle, thinner outer lip and by the slightly curved spire as a consequence of the position of the incremental scars. In the syntype of E. chyta (Figure 11J) the incremental scars are not clearly visible. Further studies and more samples of both taxa may help clarify the status of these species. Geographical distribution. Eastern Atlantic: Spain: Southwest of Cadiz (Bouchet & Warén, 1986); Azorean seamounts (Hoffman & Freiwald 2020). Western Atlantic: Brazil: Ceará, Pernambuco and Espírito Santo (present study). Bathymetric distribution. From 230 m to 706 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 472-474, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Brusina, S. (1886) Appunti ed osservazioni sull'ultimo lavoro di J. Gwyn Jeffreys. Glasnik Hrvatskoga Naravoslovnoga Drus ˇ tva, Zagreb, 1, 182 - 221.","Brusina, S. (1869) Gasteropodes nouveaux de l'Adriatique. Journal de Conchylologie, 17, 230 - 249.","Jeffreys, J. G. (1884) On the Mollusca procured during the ' Lightning' and ' Porcupine' expeditions, 1868 - 70. Part VII. Proceedings of the Zoological Society of London, 52 (3), 341 - 372. https: // doi. org / 10.1111 / j. 1096 - 3642.1884. tb 02837. x","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762","Utrilla, O., Gofas, S., Urra, J., Marina, P., Mateo-Ramirez, A., Lopez-Gonzalez, N., Gonzalez-Garcia, E., Salas, C. & Rueda, J. L. (2020) Molluscs from benthic habitats of the Gazul mud volcano (Gulf of Cadiz). Scientia Marina, 84 (3). https: // doi. org / 10.3989 / scimar. 05027.17 A","Watson, R. B. (1883) Mollusca of H. M. S. \" Challenger \" Expedition. Part XVII. Zoological Journal of the Linnean Society, 17 (99), 112 - 130. https: // doi. org / 10.1111 / j. 1096 - 3642.1883. tb 02043. x","Hoffman, L. & Freiwald, A. (2020) Bathyal Eulimidae (Gastropoda: Vanikoridoidea) from the Azorean seamounts collected during the R / V Meteor Cruise M 151 Athena. Miscellanea Malacologica, 8 (6), 81 - 99.","Seguenza, G. (1876) Studii stratigrafici sulla formazione pliocenica della'Italia Meridionale. Bollettino del R. Comitato Geologico d'Italia, 7, 7 - 15 + 87 - 103.","Landau, B, La Perna, R. & Marquet, R. (2006) The Early Pliocene Gastropoda (Mollusca) of Estepona, Southern Spain. Part 6: Triphoroidea, Epitonioidea, Eulimoidea. Palaeontos, 10, 1 - 96."]}

Thaleia nisonis (Dall, 1889b) (Figures 22 A–F, 23A–C) Benthonella nisonis Dall, 1889b: 283. Thaleia nisonis: Warén (1979: 284, figure 5); Absal„o et al. (2003: 324, figs. 4–5). Type material. Holotype USNM 93802. Type locality. USA: off Lousiana, UFSC stn. 2384 (28°45′00″N 88°15′30″, 1719 m), coll. 03/iii/1885. Material examined. Holotype; USA: USFC stn. 2668: USNM 108092 [5 dd]. Bahamas: USFC stn. 2654 (27°57′30″N 77°27′30″W, 1207 m), coll. 18/v/1886: USNM 330620 [1 dd]; Cuba: Blake stn. 30 (24°33′N 84°34′W, 1770 m), coll. 01/i/1877: USNM 94289 [2 dd]; USFC stn. 2352 (22°35′N 84°23′W, 847 m), coll. 21/i/1885: USNM 323107 [1 dd]. Brazil: Rio Grande do Norte: off Rio Grande do Norte, REVIZEE Nordeste (06°14′04″S 34°52′33″W, 510 m), 26/xi/2001: MNRJ 35263 † [1 dd]; Bahia: Camamu-Almada Basin (14°30′42″S 38°44′02″W, 1400 m): MNRJ 34498 † [1 dd]; Rio de Janeiro: Oceanprof II stn. A-13: IBUFRJ 16639 † [1 dd]; Oceanprof I stn. 84: MNRJ 33512 † [1 dd]; Oceanprof II stn. 74: IBUFRJ 17317 † [2 dd]; Oceanprof II stn. 68: IBUFRJ 17570 † [1 dd]. Measurements. Holotype USNM 93802, 10 whorls (protoconch partially broken), SL= 9.1 mm; BWL= 4.1 mm; AL= 2.4 mm; SW= 4.0 mm; AW= 1.8 mm. MNRJ 33512†, ~10 whorls, SL= 4.3 mm; BWL= 2.1 mm; AL= 1.3 mm; SW= 2.0 mm; AW= 1.1 mm. MNRJ 34498†, ~8 whorls, SL= 3.6 mm; BWL= 1.8 mm; AL= 1.1 mm; SW= 0.9 mm; AW= 1.7 mm. Remarks. Warén (1979) provided a redescription of the shell and of the soft parts of Thaleia nisonis. Absal„o et al. (2003) recorded T. nisonis for the first time in Northeast and Southeast Brazil. One of the shells recorded by Absal„o et al. (2003) was collected in 50 m (IBUFRJ 12699), an odd shallow water record. This lot was not located and we consider this record doubtful. Thaleia nisonis is a peculiar species and easily distinguished from other deep water eulimids by the brownish protoconch (Figure 23A) and by the sculpture on the protoconch (Figure 23B) and teleoconch (Figure 22C). Other genera occurring in the deep Atlantic have a brown protoconch: Batheulima Nordsieck, 1968, that also presents axial sculpture on the protoconch and teleoconch (Bouchet & Warén 1986: 334); and Fuscapex Warén, 1981a, which usually does not have sculpture on the protoconch, except for F. microcostellata Bouchet & Warén, 1986 (Bouchet & Warén 1986: 325). Despite these similarities, T. nisonis can be distinguished from any of the species in these other genera by the presence of a spiral thread on the upper region of the protoconch whorls and the small denticles present on the thin axial lines of the protoconch (Figure 23B, C). These denticles resemble spiral sculpture (Absal„o et al. 2003). Geographical distribution. USA: Florida (Off Fernandina); Gulf of Mexico; Bahamas; Cuba (Warén 1979); Brazil: Rio Grande do Norte state (present study), Bahia and Rio de Janeiro states (Absal„o et al. 2003, present study). Bathymetric distribution. From 510 m to 1950 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 491-492, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Dall, W. H. (1889 b) Reports on the results of dredgings, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer ' Blake'. Bulletin of the Museum of Comparative Zoology, 18, 1 - 492, pls. 10 - 40.","Waren, A. (1979) On the systematic position of Benthonella nisonis Dall, 1889 (Gastropoda: Prosobranchia). Journal of Molluscan Studies, 45, 284 - 288. https: // doi. org / 10.1093 / oxfordjournals. mollus. a 065501","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762","Waren, A. (1981 a) Eulimid gastropods parasitic on echinoderms in the New Zealand region. New Zealand Journal of Zoology, 8, 313 - 324. https: // doi. org / 10.1080 / 03014223.1981.10430611"]}

Genus Ophieulima Warén & Sibuet, 1981 Type species. Stilifer minima Dall, 1927, by original designation, Recent; North Atlantic. Diagnosis. Shell small, few whorls, colorless, globose; protoconch brownish, cylindrical; teleoconch inflated, strongly convex whorls, surface usually sculptured by microscopic spiral lines; aperture broad, almost half the size of total length, outer lip prosocline (adapted from Warén & Sibuet 1981). Remarks. Ophieulima Warén & Sibuet, 1981 is a genus that parasitizes ophiuroids and comprises three species (MolluscaBase Eds. 2020c): O. minima (Dall, 1927), from the North Atlantic and Mediterranean (Dall 1927; Bouchet & Warén 1986; Romani et al. 2014; Hoffman & Freiwald 2020); O. fuscoapicata Warén, 1981, from New Zealand (Warén 1981a); and O. antecessor Lozouet, 1999, a fossil species from the Miocene of France (Lozouet 1999). The new record of O. minima herein is the first record of the genus in the southwestern Atlantic.

Fusceulima Laseron, 1955 Type species. Fusceulima jacksonensis Laseron, 1955, by original designation (Recent; New South Wales, Australia). Diagnosis. See Souza & Pimenta (2014)., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on page 470, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Laseron, C. F. (1955) Revision of the New South Wales Eulimoid Shells. The Australian Zoologist, 12 (2), 83 - 107.","Souza, L. S. & Pimenta, A. D. (2014) Fusceulima and Halielloides (Gastropoda: Eulimidae) in the southwestern Atlantic, with descriptions of two new species of Fusceulima. Zoologia, Curitiba, 31 (6), 621 - 633. https: // doi. org / 10.1590 / S 1984 - 46702014000600008"]}

Melanella sp. 2 (Figure 18 A–F) Material examined. Brazil: Ceará: (03°30′51″S 37°59′28″W, 384 m), RV Natureza, 07/xi/2001: MNRJ 35267 † [1 dd]; Alagoas: (10°06′35″S 35°46′41″W, 720 m), RV Natureza, 16/xii/2001: MNRJ 35206 † [1 dd]; Rio Grande do Sul: Off Solidão (30°46′S 49°07′W, 176 m), coll. RV Prof. Wladimir Besnard, 27/i/1972: MORG 17624 † [1 dd]. Description. Shell conical, slightly curved, reaching about 8.5 mm long, 3.3 mm wide. Larval shell broken. Teleoconch with about 8.0 whorls, slightly convex outline, convexity accentuated in the lower region of each whorl; suture well demarcated, almost straight; subsutural zone indistinct; surface smooth, incremental scars well demarcated, appearing in intervals of about 1.1 whorl at the last two whorls. Last whorl about 50% of the shell length; base slightly truncated. Aperture high, pear shaped, acute posteriorly and rounded anteriorly; outer lip thin, opisthocline, sinuous, straight near the suture and strongly projecting, reaching the maximum projection at the mid of its height; inner lip thick, sinuous, sloping, reflected and well demarcated. Not umbilicate. Shell vitreous, colorless. Measurements. MNRJ 35206†, 8 whorls (apex broken), SL= 8.5 mm; BWL= 4.5 mm; AL= 2.8 mm; SW= 3.3 mm; AW= 2.0 mm. Remarks. The protoconch is broken in the three shells examined, but the relatively wide and solid shell, with a wide aperture and thick inner lip are characteristic enough to be worth noting (Figure 18 A–F). Melanella sp. 2 resembles M. doederleini (Figure 11 A–I) in shape, but has bigger dimensions and a much more elongated aperture. Even with the apex broken, specimens of Melanella sp. 2 reaches 8 whorls and 8.5 mm long, while M. doederleini reaches 10 whorls and 3.6 mm long. Melanella sp. 2 can also be distinguished from M. anapetes sp. nov. (Figure 16 A–G) by the wider aperture, bigger dimensions, thicker inner lip and the curved spire. Geographical distribution. Brazil: Ceará, Alagoas, Rio Grande do Sul (present study). Bathymetric distribution. From 176 m to 720 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 482-485, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Tryon, G. W. (1886) Manual of Conchology. Vol. 8. Philadelphia, Pennsylvania, 461 pp., 79 pls.","Clarke, A. H. (1974) Molluscs from Baffin Bay and the northern North Atlantic Ocean. Publications in Biological Oceanography, National Museum of Natural Sciences of Canada, 7, 1 - 23."]}

Ophieulima minima (Dall, 1927) (Figure 19 A–G) Stilifer minima Dall, 1927: 73. Ophieulima minima: Warén & Sibuet (1981: 381, figs. 1–5); Hoffman et al. (2011: 34, figs. 45–46); Romani et al. (2014: 1, figure 1A–F); Hoffman & Freiwald (2020: 89–90). Type material. Holotype USNM 108039 (Figure 19A, G). Type locality. USA: Off Georgia, USFC stn. 2668 (30°58′30″N 79°38′30″W, 538 m), 05/v/1886. Material examined. Type material; Brazil: Alagoas: (09°20′S 34°59′W, 452 m), 18/xii/2001: MNRJ 35223 † [1 dd]. Measurements. Holotype USNM 108039, ~6 whorls, SL= 2.3 mm; BWL= 1.6 mm; AL= 1.0 mm; SW= 1.2 mm; AW= 0.8 mm. MNRJ 35223†, ~6 whorls, SL= 1.9 mm; BWL= 1.3 mm; AL= 0.9 mm; SW= 1.0 mm; AW= 0.6 mm. Remarks. Dall (1927) recorded four specimens in the original description. Warén & Sibuet (1981) described two paratypes under the same catalog number as the holotype. No other specimen than the holotype (Figure 19A, G) was found in the USNM and the status of these other shells is unknown or they are presumed lost (E. Strong, personal communication). Warén & Sibuet (1981: 384) redescribed the species and noted that the variation in shape of the type specimens is great and that the identification of the material by them is questionable, because the northeast Atlantic specimens studied are even broader than the types. Bouchet & Warén (1986: 348) also considered the shell morphology of the species quite variable, with an evident sexual dimorphism (males are half the size of females). Beside the variation in size and shape correlated with the sexual dimorphism, a variation in shell sculpture can also be observed. Warén & Sibuet (1981: 384) commented that none of the types shows the spiral striation that occurs in some well-preserved specimens from the Northeast Atlantic examined by them. The latter authors described this spiral sculpture as irregular, extremely fine scratch-like striae present in some small areas of the shell that are less corroded in such specimens. The examination of SEM images of the holotype of Ophieulima minima (USNM 108039) (Figure 19A, G) has not revealed the presence of the spiral scratch-like striae, but only the axial growth lines. Romani et al. (2014) reported O. minima from the Mediterranean and did not observe the spiral sculpture in very fresh shells, with dried specimens. The single shell from Brazil agrees well with the paratypes illustrated by Bouchet & Warén (1986: figs. 827– 830) in the general shape (Figure 19B, C), number of whorls of the larval shell (=3.0 whorls) (Figure 19G) and the axial sculpture of faint growth lines (Figure 19C, D). The holotype of Ophieulima minima presents a protoconch with about 2.6 whorls and 195 µm in diameter (Figure 19G) and the shell from Brazil has 3.0 protoconch whorls, 210 µm wide (Figure 19F). The teleoconch of the holotype of O. minima and the shell from Brazil have about three whorls, but the former reaches 2.8 mm long and the latter reaches 1.9 mm long, which represents about 65% of the shell length of the holotype. The shell from Brazil is fairly corroded, as can be seen from the several pits in the shell surface (Figure 19F) and spiral sculpture was not observed. Even with these small differences in size, shape, and sculpture, the shell from Brazil is identified as Ophieulima minima. This represents the southernmost record of O. minima. It is possible that further material and other techniques will uncover more species of Ophieulima than currently expected in the whole Atlantic. Geographical distribution. Northeastern Atlantic: South Iceland to South of the Bay of Biscay (Bouchet & Warén 1986); Mediterranean (Romani et al. 2014); Azorean seamounts (Hoffman & Freiwald 2020). Western Atlantic: USA: Georgia (Dall 1927); Brazil: Alagoas (present study). Bathymetric distribution. From 452 m to 2100 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 485-487, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Dall, W. H. (1927) Small shells from dredgings off the southeast coast of the United States by the United States Fisheries Steam- er \" Albatross \" in 1885 and 1886. Proceedings of the United States National Museum, 70 (2667), 1 - 134. https: // doi. org / 10.5479 / si. 00963801.70 - 2667.1","Waren, A. & Sibuet, M. (1981) Ophieulima (Mollusca, Prosobranchia), a new genus of ophiuroid parasites. Sarsia, 66, 103 - 107. https: // doi. org / 10.1080 / 00364827.1981.10414528","Hoffman, L., van Heugten, B. & Lavaleye, M. S. S. (2011) Gastropoda (Mollusca) from the Rockall and Hatton Banks, northeastern Atlantic Ocean. 3. Miscellanea Malacologica, 5 (2), 23 - 52.","Romani, L., Bartolini, S., Giusti, F. & Sbrana, C. (2014) Ophieulima (Gastropoda: Caenogastropoda: Eulimidae): a new genus for the Mediterranean Sea. Marine Biodiversity Records, 7, E 131. https: // doi. org / 10.1017 / S 1755267214001304","Hoffman, L. & Freiwald, A. (2020) Bathyal Eulimidae (Gastropoda: Vanikoridoidea) from the Azorean seamounts collected during the R / V Meteor Cruise M 151 Athena. Miscellanea Malacologica, 8 (6), 81 - 99.","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762"]}

Sticteulima cabrali sp. nov. urn:lsid:zoobank.org:act: 84C6DFD0-FD21-4784-A28E-8EF10C922A65 (Figure 20 A–H) Type material. Holotype MNRJ 23195 † (Figure 20A); Paratypes: from type locality, MNRJ 35194 † [2] (Figure 20 B–H). Type locality. Brazil: Off Rio Grande do Norte, REVIZEE Nordeste (04°51′40″S 35°08′01″O, 384 m), coll. RV Natureza, 24/xi/2001. Etymology. named after Enílson Cabral of the Research and Management Center of Fishing Resources of the Northeast Coast (CEPENE/IBAMA) for his personal efforts in the collections of sediment from the continental slope off Northeast Brazil. Description (Holotype). Shell conical, straight, reaching about 2.7 mm long, 1.2 mm wide. Larval shell vitreous, with a brownish spiral band at the suture, conical, 3.4 whorls, 380 µm wide, smooth, transition to teleoconch marked by an almost indistinct incremental scar. Teleoconch with about 4.5 whorls, outline slightly convex; suture almost indistinct, sloping slowly; subsutural zone indistinct; surface slightly sculptured by axial lines; incremental scars demarcated, appearing at irregular intervals. Last whorl about 55% of the shell length; base rounded. Aperture wide, pear shaped, acute posteriorly and rounded anteriorly; outer lip thin, opisthocline, sinuous, slightly retracting near the suture and projecting in the uppermost third; inner lip slightly sinuous, sloping, reflected and well demarcated. Small umbilical chink present. Shell vitreous, with irregular brownish patches. Measurements. Holotype MNRJ 23195†, 8 whorls, SL= 2.7 mm; BWL= 1.5 mm; AL= 1.1 mm; SW= 1.2 mm; AW= 0.7 mm. Paratype MNRJ 35194, 8 whorls, SL= 2.9 mm; BWL= 1.5 mm; AL= 1.1 mm; SW= 1.2 mm; AW= 0.7 mm. Remarks. Sticteulima cabrali sp. nov. (Figure 20 A–H) differs from most of its congeners mainly by the presence of microsculpture of axial lines on the teleoconch (Figure 20D). Furthermore, it differs from S. lata by having more convex teleoconch whorls; from S. richteri and S. wareni by the different color pattern, presenting irregular brownish patches on the shell (Figure 20A) in spite of a well-defined brownish spiral band like in S. richteri or a solid brown shell like in S. wareni; from S. fuscescens by the faster increasing diameter, reaching a greater width and by the smooth surface of the protoconch (Figure 20G, H), while S. fuscescens has strong opisthocline axial lines in the protoconch. Sticteulima cabrali sp. nov. resembles two other species currently in Eulima: E. chaunax Watson, 1883, only known from the type locality in Puerto Rico, and E. fuscozonata, from off Spain and Portugal. Sticteulima cabrali sp. nov. differs from both by the presence of a microsculpture on the teleoconch, while the others are smooth. Furthermore, S. cabrali sp. nov. has a colored shell while E. chaunax is whitish, and is proportionally smaller (holotype: SL= 2.7 mm vs. syntype of E. chaunax NHMUK 1887.2.9.1584: SL= 3.8 mm; both with 8–9 whorls); and E. fuscozonata has no umbilicus. Geographical distribution. Only known from the type locality. Bathymetric distribution. 384 m.

Thaleia Warén, 1979 Type species. Benthonella nisonis Dall, 1889b, by original designation, Recent; western Atlantic. Diagnosis. Shell conical, medium length (5–9 mm), vitreous except protoconch; protoconch dark brown, slightly convex whorls, divided in protoconch I and II, protoconch I smooth, protoconch II sculptured by microscopic sinuous and denticulate axial lines and a straight spiral line close to the suture; teleoconch whorls slightly convex, deep suture, sculptured by microscopic axial lines; aperture short, outer lip very thin, umbilical fissure present (adapted from Warén 1979). Remarks. Warén (1979) erected Thaleia as a monotypic genus with a doubtful classification, but related this genus to the Eulimidae. Since then, no additional data about the systematics of the genus is known and it is still allied to Eulimidae. MolluscaBase Eds. (2020e) include a second species in the genus, Thaleia mucronetincta (Thiele, 1925), from East Africa., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on page 491, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Waren, A. (1979) On the systematic position of Benthonella nisonis Dall, 1889 (Gastropoda: Prosobranchia). Journal of Molluscan Studies, 45, 284 - 288. https: // doi. org / 10.1093 / oxfordjournals. mollus. a 065501","Dall, W. H. (1889 b) Reports on the results of dredgings, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer ' Blake'. Bulletin of the Museum of Comparative Zoology, 18, 1 - 492, pls. 10 - 40.","Thiele, J. (1925) Gastropoda der Deutschen Tiefsee-Expedition. II Teil. Deutsche Tiefsee-Expedition 1898 - 1899, 17 (2), 35 - 382."]}

Genus Umbilibalcis Bouchet & Warén, 1986 Type species. Aclis (supranitida Wood var. ?) lata Dall, 1889b, by original designation, Recent; Caribbean. Diagnosis. See Bouchet & Warén (1986: 444). Remarks. Umbilibalcis Bouchet & Warén, 1986 comprises three species, all from the North Atlantic: Um-bilibalcis crassula Bouchet & Warén, 1986, U. lata (Dall, 1889b) and U. subumbilicata (Jeffreys, 1883) (Bouchet & Warén 1986). The present records of Umbilibalcis lata represents the first record of the the genus in the southwestern Atlantic. The present genus resembles Niso by the presence of microscopic sculpture of axial lines on the protoconch and teleoconch, but has a more rounded base and edge of the umbilicus, and lacks color on the teleoconch. Furthermore, the aperture of Umbilibalcis is more rounded, while in Niso it is usually rhomboid., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 492-493, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762","Dall, W. H. (1889 b) Reports on the results of dredgings, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer ' Blake'. Bulletin of the Museum of Comparative Zoology, 18, 1 - 492, pls. 10 - 40."]}

Melanella paraabida sp. nov. urn:lsid:zoobank.org:act: 8D796B06-259E-445F-8C87-B6C4FF5BE506 (Figure 13 A–H) Type material. Holotype MNRJ 35262 † (Figure 13 A–H). Type locality. Brazil: Off Pernambuco, REVIZEE Nordeste (08°42′06″S 34°44′06″W, 465 m), coll. RV Natureza, 25/iii/2000. Etymology. Named after its similarity to Melanella abida Dall, 1927. Description (Holotype). Shell conical, curved, reaching about 3.4 mm long, 1.1 mm wide. Larval shell whitish, conical, consists of 2.5 whorls, 220 µm in wide, smooth, transition to teleoconch marked by distinct incremental scar. Teleoconch with about 7.5 whorls, flat outline; suture slightly demarcated, almost straight; subsutural zone indistinct; surface sculptured by axial lines, intervals about 25 µm in the penultimate whorl; incremental scars well demarcated, appearing at intervals of about 1.0 whorl in the last three whorls, stepped. Last whorl about 45% of the shell length; base rounded. Aperture short, pear shaped, acute posteriorly and rounded anteriorly; outer lip relatively thick, opisthocline, sinuous, slightly retracting near the suture and slightly projecting in the uppermost third; inner lip sinuous, sloping, reflected and well demarcated. Not umbilicate. Shell vitreous, colorless. Measurements. Holotype MNRJ 35262†, 10 whorls, SL= 3.4 mm; BWL= 1.5 mm; AL= 0.8 mm; SW= 1.1 mm; AW= 0.5 mm. Remarks. Despite having only a single shell, we describe Melanella paraabida based on the combination of consistent shell features that enable us to discriminate this species from the other eulimids known to the Atlantic. The holotype was destroyed during the fire in the MNRJ. Melanella paraabida sp. nov. closely resembles M. abida Dall, 1927 (Figure 14A, B), known only from off Georgia, USA. Dall (1927) cited the type locality of M. abida as “off Fernandina, Florida ” but the label of the holo-type (USNM 108030) clearly indicates “RV Albatross stn. 2668”, thus the correct locality is off Georgia. Melanella corrida Dall, 1927 (Figure 14 C–F), from the same locality, is a synonym of M. abida (MolluscaBase Eds. 2020b). Melanella paraabida sp. nov. can be distinguished from M. abida mainly by the presence of microsculpture on the teleoconch (Figure 13D) and the less protruding outer lip. Additionally, the protoconch of M. paraabida sp. nov. has a smaller number of whorls and diameter. Melanella paraabida sp. nov. has a protoconch 210 µm wide with 2.5 whorls, while the holotype of M. corrida (USNM 108376) (Figure 14F) has a protoconch 260 µm wide with 3.5 whorls. Melanella paraabida sp. nov. resembles M. orphanensis Clarke, 1974 (Figure 17K), from the North Atlantic, and M. scarifata Gofas & Rueda, 2014 (Gofas et al. 2014), from the Alboran and Mediterranean sea, by the teleoconch sculpture. Melanella paraabida sp. nov. differs from M. orphanensis by the narrower apex (Figure 13G), which is dome shaped in the latter (Figure 17K). Melanella paraabida sp. nov. differs from M. scarifata by the smaller dimensions of the protoconch (2.5 whorls, 210 µm wide vs. 3.5 whorls, 350 µm wide in M. scarifata) and the less protruding outer lip (Figure 13F). Melanella paraabida sp. nov. resembles Melanella spiridioni (Dautzenberg & Fischer, 1896), from Northeast Atlantic and Mediterranean, by the curved spire but differs by the presence of microsculpture of axial lines. Geographical distribution. Only known from the type locality. Bathymetric distribution. 465 m.

Eulimacrostoma phorcys sp. nov. urn:lsid:zoobank.org:act: 128029A4-10BA-401F-B39D-9AA3ECEDFC0D (Figure 9 A–I) Melanella hians auct. non Watson, 1883: Lima et al. (2005: 2, figure 1a–d). Type material. Holotype: MNRJ 23191 † (Figure 9A, B). Paratypes: from type locality, MNRJ 35191 † [5 dd] (Figure 9 C–I); Brazil: off Sergipe, REVIZEE Nordeste (10°41′24ʺS 36°18′42ʺW, 365 m), 28/x/2000, MNRJ 35220 † [3 dd]. Type locality. Brazil, Off Rio Grande do Norte, REVIZEE Nordeste (04°51′S 35°06′W, 375 m), coll. RV Natureza, 24/xi/2001. Etymology. A sea god from Greek mythology, Phorcys (Latin spelling) is associated with the hidden dangers of the sea. Epithet as a noun in apposition. Description (Holotype). Shell conical, apex obtuse, reaching about 7.6 mm long, 1.8 mm wide. Larval shell vitreous, with about 1.75 whorls, 300 µm wide, 375 µm in height, smooth, transition to teleoconch marked by a subtle incremental scar. Teleoconch with 9.0 whorls, sinuous outline, convexity more attenuated on the lower region of each whorl; suture deep, well impressed, sloping; subsutural zone about 1/5 of the height of the whorl; surface glossy, presenting micro sculpture of opisthocline axial lines; incremental scars well developed, appearing at irregular intervals. Last whorl about 45% of the shell length; base rounded. Aperture high, wide, pear shaped, slightly expanded laterally, acute posteriorly, rounded and spread anteriorly; outer lip thin, very sinuous, opisthocline, retracted near the suture, after strongly projecting, and retracted in the distal region, maximum projection at the middle of the outer lip height; inner lip thin, sinuous, sloping. Not umbilicate. Teleoconch vitreous, with a wide yellowish spiral band at the lower region of each whorl. Measurements. Holotype MNRJ 23191†, 11 whorls, SL= 7.6 mm; BWL= 3.4 mm; AL= 2.2 mm; SW= 1.8 mm; AW= 1.0 mm. Paratype MNRJ 35191†, 11 whorls, SL= 7.6 mm; BWL= 3.6 mm; AL= 2.5 mm; SW= 2.2 mm; AW= 1.3 mm. Paratype MNRJ 35191†, ~10 whorls, SL= 6.9 mm; BWL= 3.4 mm; AL= 2.3 mm; SW= 1.9 mm; AW= 1.2 mm. Paratype MNRJ 35191†, 9 whorls, SL= 5.8 mm; BWL= 3.0 mm; AL= 2.1 mm; SW= 1.7 mm; AW= 1.1 mm. Remarks. All types of the present species were destroyed; however, it has consistent shell features that allow the description as a new species, and the figures provided here are sufficient for its recognition. The present species fits the recently described genus Eulimacrostoma, because it has a subcylindrical protoconch (Figure 9H), microsculpture of very thin axial lines on the teleoconch (Figure 9E), an aperture gaping anteriorly (Figure 9F) and a strongly protruding outer lip (Figure 9G) (Souza & Pimenta 2019a). The record of Melanella hians (Watson, 1883) from the continental slope of Pernambuco, Brazil, by Lima et al. (2005) is actually a misidentification of Eulimacrostoma phorcys sp. nov. The material studied by these authors could not be found in the LMUFRPE, but the figured specimen (Lima et al. 2005: figure 1a–d) is clearly conspecific with E. phorcys sp. nov. Thus, M. hians is known only from the the type locality in Culebra Islands, Puerto Rico (Challenger stn. 24). Eulimacrostoma phorcys sp. nov. can be distinguished from M. hians by the narrower teleoconch whorls, which results in a smaller spire angle, and by features of the aperture, which is more rounded and gaping in the anterior region but less expanded laterally. Eulimacrostoma phorcys sp. nov. closely resembles E. microsculpturatum Souza & Pimenta, 2019, from the Caribbean and Florida, by the general shape of the shell, color pattern of the teleoconch and the presence of several incremental scars. However, E. phorcys sp. nov. has the first protoconch whorl strongly inflated, with a wider diameter (250 μm vs. 200 μm in E. microsculpturatum) (Figure 9I) and the same maximum diameter (300 μm) despite reaching a smaller number of whorls (2.0 vs. 2.5). Additionally, the protoconch of E. phorcys sp. nov. has no trace of a brownish spiral band while E. microsculpturatum has one (Souza & Pimenta 2019a). Eulimacrostoma phorcys sp. nov. also differs by the larger size; the holotype has about 11 whorls and reaches 7.6 mm long while the paratypes USNM 433081 and USNM 1273894 of E. microsculpturatum with the same number of whorls are 6.5 and 6.6 mm long (Souza & Pimenta 2019a), respectively. No sexual dimorphism is known in the genus. Eulimacrostoma phorcys sp. nov. can be distinguished from E. chascanon (Watson, 1883), known only from the type locality (same as Melanella hians), mainly by the wider apex and anterior region of the aperture, and by the larger size (holotype of E. phorcys, 11 whorls, SL= 7.6 mm vs. syntype NHMUK 1887.2.9.1587, 12–13 whorls, SL= 4.5 mm). Eulimacrostoma phorcys sp. nov. can be distinguished from E. patula (Dall & Simpson, 1901), from the Caribbean and northern Brazil, by the absence of an umbilical chink and by the more rounded base in younger and mature specimens. Eulimacrostoma phorcys sp. nov. can be distinguished from E. lutescens (Simone, 2002), from Southeast Brazil, by being proportionally smaller in length (holotype, 11 whorls, SL= 7.6 mm vs. holotype of E. lutescens, SL = 8.9 mm) and by the shorter and narrower aperture (AL= 2.2 mm, AW= 1.0 mm, AW/AL= 0.4 vs. AL= 3.1, AW= 1.6, AW/AL= 0.5, holotypes of both species). Geographical distribution. Brazil: Rio Grande do Norte (present study), Pernambuco (Lima et al. 2005) and Sergipe (present study). Bathymetric distribution. From 365 m to 690 m (Lima et al. 2005; present study)., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 468-470, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Watson, R. B. (1883) Mollusca of H. M. S. \" Challenger \" Expedition. Part XVII. Zoological Journal of the Linnean Society, 17 (99), 112 - 130. https: // doi. org / 10.1111 / j. 1096 - 3642.1883. tb 02043. x","Lima, S. F. B., Barros, J. C. N., Silva, S. V., Santos, M. C. F. & Cabral, E. (2005) Ocorrencia da especie Melanella hians (Watson, 1883) em aguas profundas no estado de Pernambuco, Brasil. Boletim Tecnico Cientifico do CEPENE, 13 (2), 151 - 155.","Souza, L. S. & Pimenta, A. D. (2019 a) Eulimacrostoma gen. nov., a new genus of Eulimidae (Gastropoda, Caenogastropoda) with description of a new species and reevaluation of other western Atlantic species. Zoosystematics and Evolution, 95 (2), 403 - 415. https: // doi. org / 10.3897 / zse. 95.33880","Dall, W. H. & Simpson, C. T. (1901) The Mollusca of Porto Rico. United States Fisheries Commission Bulletin, 20, 351 - 524, pls. 53 - 58.","Simone, L. R. L. (2002) Three new deepwater species of Eulimidae (Caenogastropoda) from Brazil. Novapex, 3 (2 - 3), 55 - 60."]}

Eulimacrostoma Souza & Pimenta, 2019 Type species. Eulimacrostoma microsculpturatum Souza & Pimenta, 2019, by original designation; Recent, Caribbean. Diagnosis. See Souza & Pimenta (2019a)., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on page 468, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Souza, L. S. & Pimenta, A. D. (2019 a) Eulimacrostoma gen. nov., a new genus of Eulimidae (Gastropoda, Caenogastropoda) with description of a new species and reevaluation of other western Atlantic species. Zoosystematics and Evolution, 95 (2), 403 - 415. https: // doi. org / 10.3897 / zse. 95.33880"]}

Eulima psila Watson, 1883 (Figure 6 A–M) Eulima psila Watson, 1883: 112. Eulima psila: Watson (1886: 508, plate 35, figure 1a–c). Type material. Lectotype (herein designated): NHMUK 1887.2.9.1585 (Figure 6A). Type locality. Puerto Rico: North Culebra Island, off St. Thomas, Challenger stn. 24 (18°38′30″N 65°05′30″W, 713 m), coll. 25/iii/1873. Material examined. Type material; USA: Georgia: USFC stn. 2668: USNM 108031 [1 dd]; Florida: Eolis stn. 193: USNM 433355 [1 dd]; Eolis stn. 305: USNM 433360 [1 dd]; Eolis stn. 351: USNM 417633 [3 dd]; Eolis stn. 372: USNM 417532 [1 dd]; Eolis stn. 376: USNM 417541 [1 dd]; Eolis stn. 379: USNM 433085 [1 dd]; USFC stn. 2400 (28°41′N 86°07′W, 309 m), coll. 14/iii/1885: USNM 323841 [1 dd]. Barbados: St. James, Holetown (13°10′52″N 59°38′30″W), coll. F. Sander: ANSP 353545 [1 dd]. Brazil: Pernambuco: off Archipelago Fernando de Noronha, REVIZEE Nordeste (03°54′54″S 32°37′48″W), 12/x/2001: MNRJ 35203† [1 dd]; Espírito Santo: AMBES11 stn. B5: MNRJ 34490† [1 lv, destroyed for dissection]; Rio de Janeiro: REVIZEE-Central V stn. 52f (21°46′01″S 40°04′58″W, 450 m), coll. Supply Boat Astro Garoupa, 21/vii/2001: IBUFRJ 13301† [2 dd]; Campos Basin (23°16′43″S 40°59′51″W, 600 m), coll. 19/viii/2009: MNRJ 25613† [1 dd]; HAB16 stn. E5: MNRJ 16188† [1 dd]; MD55 stn. CB102 (23°07′S 42°04′W, 100 m), coll. RV Marion Dufresne, v/1987: MNRJ 25901† [2 dd]; Campos Basin (23°17′16″S 41°00′36″W, 1800 m), coll. RV Ocean Stalwart, 05/ii/2016: MNRJ 27968† [1 lv]; Cabo Frio VII stn. 6194 (24°03′36″S 44°07′36″W, 134 m), coll. RV Almirante Saldanha, 01/iv/1983: IBUFRJ 15233† [1 dd]; Campos Basin (800 m), coll. iv/2008: MNRJ 35855† [1 dd]; Santos Basin (24°11′52″S 43°15′18″W, 600 m), coll. 28/x/2002: MNRJ 35925† [3 dd]; S„o Paulo: REVIZEE Sul stn. 6678: MNRJ 28245† [4 lv]; REVIZEE Sul stn. 6657: MNRJ 28243† [1 lv]; REVIZEE Sul stn. 6656: MNRJ 28242† [2 lv]; Rio Grande do Sul: REVIZEE Sul stn. 6858: MORG 42540† [1 dd]; REVIZEE Sul stn. 6843: MORG 42448† [1 dd]; Off Rio Grande do Sul (153 m), coll. RV Atlântico Sul, v/1986: MORG 24716† [1 dd]; Off Rio Grande do Sul, stn. 1656 (33°17ʹS 50°34ʹW, 173 m), coll. RV Prof. Wladimir Besnard, 20/i/1972: MORG 17586† [1 dd]. Measurements. Lectotype NHMUK 1887.2.9.1585, 8 whorls, SL= 4.3 mm; BWL= 2.0 mm; AL= 1.4 mm; SW= 1.1 mm; AW= 0.6 mm. USNM 433360, 14 whorls, SL= 11.4 mm; BWL= 5.1 mm; AL= 3.4 mm; SW= 2.4 mm; AW= 1.5 mm. IBUFRJ 15233†, 12 whorls, SL= 8.0 mm; BWL= 3.6 mm; AL= 2.3 mm; SW= 1.6 mm; AW= 1.0 mm. MNRJ 27968†, 12 whorls, SL= 7.2 mm; BWL= 3.5 mm; AL= 2.3 mm; SW= 1.6 mm; AW= 1.0 mm. MNRJ 35203†, 11 whorls, SL= 6.6 mm; BWL= 3.3 mm; AL= 2.0 mm; SW= 1.3 mm; AW= 0.8 mm. Remarks. Two syntypes of the present species were found in the NHMUK and NMW; the shell from the former (NHMUK 1887.2.9.1585) is the figured specimen by Watson (1886: plate 35, figure 1a–c) and is here designated as the lectotype (Figure 6A) to clarify the name of the species. The shell from the NMW, also collected at Challenger stn. 24, is not conspecific and is probably an undescribed species of Eulima known only from this locality. The latter species will be treated latter with additional material from the Caribbean. This species has slightly convex teleoconch whorls, a relatively higher aperture occupying 70% of the body whorl (vs. ~60% in E. psila), a slightly protruding outer lip and a colorless larval shell (Figure 6N, O). The description of Watson (1883) is appropriate for the lectotype and there is no need for a redescription of the specimen. However, since the lectotype is a young specimen it is important to highlight some features of the shell based on the material examined here. Despite the differences in size of other specimens studied here, they fit perfectly with the dimensions of the protoconch (± 420 µm long, 320 µm wide) and the initial teleoconch whorls of the lectotype. They also share a brownish spiral band at the suture of the protoconch. Eulima psila reaches about 14 whorls, 11.4 mm long and 2.4 mm wide (Figure 6E); the body whorl occupies 45% of total length and the outer lip is consistently straight and orthocline in specimens at different ontogenetic stages (Figure 6G, I). The false suture occupies a small height of the whorl (1/10) and the incremental scars are weak and present at regular intervals (0.7 whorl apart). Fresh specimens usually present a faint, wide, brownish or yellowish spiral band in the last whorls (Figure 6B, C, E). The operculum is slender with faint growth lines. This species is rediscovered after more than a century. The species was found in several localities of the western Atlantic, from Florida, USA, to Rio Grande do Sul, South of Brazil. On the Northeast coast of Brazil only a single shell was examined (Figure 6D). The bathymetric range of Eulima psila is extensive; live specimens were collected at 60 m (MNRJ 28243†) and at 1800 m (MNRJ 27968†), but most of the empty shells were collected in depths of more than 100 m. All live collected specimens have eyes. Other species of Eulima present a wide bathymetric range, for example E. bilineata Alder, 1848, from the Northeast Atlantic and Mediterranean, which occurs from 50 to 900 m (Bouchet & Warén 1986: 320). Other deep water Eulima from the Atlantic with a similar shell morphology are Eulima incolor (Figure 5F, G) and E. grimaldii Bouchet & Warén, 1986 (Figure 6P, Q), both from the Northeast Atlantic. Eulima psila can be distinguished from E. incolor by the presence of a brownish spiral band at the protoconch (Figure 6K) and by reaching a higher number of whorls in the protoconch (3.5 vs. 2.0 in E. incolor) (Bouchet & Warén 1986). Eulima psila differs from E. grimaldii by the straighter outer lip (Figure 6I), which is more protruding in E. grimaldii (Figure 6Q) and by the relatively less convex whorls of the protoconch (see Bouchet & Warén 1986: figure 793 for comparison). Geographical distribution. USA: Florida (present study); Puerto Rico (Watson 1883); Brazil: Archipelago of Fernando de Noronha, Espírito Santo, Rio de Janeiro, S„o Paulo, Rio Grande do Sul (present study). Bathymetric distribution. From 60 m to 1800 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 462-464, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Watson, R. B. (1883) Mollusca of H. M. S. \" Challenger \" Expedition. Part XVII. Zoological Journal of the Linnean Society, 17 (99), 112 - 130. https: // doi. org / 10.1111 / j. 1096 - 3642.1883. tb 02043. x","Watson, R. B. (1886) Report on the Scaphopoda and Gasteropoda collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger, Zoology, 15 (2), 1 - 680 + 692 - 756, 50 pls.","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762","Alder, J. (1848) Catalogue of the Mollusca of Northumberland and Durham. Transactions of Tyneside Naturalists Field Club, 1848, 97 - 209. https: // doi. org / 10.5962 / bhl. title. 11190"]}

Melanella sp. 1 (Figure 17 A–G) Material examined. Brazil: Off Pernambuco, REVIZEE Nordeste (08°46′00″S 34°44′00″O, 690 m), coll. RV Natureza, 18/xi/2000: MNRJ 35251 † [1 dd]. Description. Shell subcylindrical, straight, reaching about 3.6 mm long, 1.0 mm wide. Larval shell vitreous, cylindrical, with about 2.0 whorls, 370 µm wide; smooth, except for microscopic striations close to suture, transition to teleoconch marked by an almost indistinct incremental scar. Teleoconch with about 7.0 whorls, flat outline; suture almost indistinct, sloping; subsutural zone about 20% of the whorl height; surface polished, sculptured by straight axial lines spaced every 10–15 μm (Figure 17D); incremental scars slightly demarcated, appearing at irregu-lar intervals. Last whorl about 40% of the shell length; base rounded. Aperture short, pear shaped, acute posteriorly and rounded anteriorly; outer lip thin, opisthocline, sinuous, slightly retracting near the suture and after projecting, maximum projection in the lowermost third of the height, retracted in distal region; inner lip sinuous, sloping, reflected, well demarcated. Not umbilicate. Shell colourless. Measurements. MNRJ 35251†, 9 whorls, SL= 3.6 mm; BWL= 1.4 mm; AL= 0.9 mm; SW= 1.0 mm; AW= 0.5 mm. Remarks. The species that most resembles Melanella sp. 1 (Figure 17 A–G) is Melanella jeffreysi (Tryon, 1886) (Figure 17 H–J), with the subcylindrical shape, dome shaped apex, similar outline of the outer lip and similar dimensions of the protoconch (~ 370–380 µm wide) and teleoconch. According to Jeffreys (1884), the shell surface of M. jeffreysi (as “ Eulima solida ”) is covered by longitudinal, extremely thin axial lines, but Bouchet & Warén (1986) disagree with Jeffrey’s description. The latter authors commented that there are “only a few scattered, indistinct, very thin axial lines”, and that Jeffrey’s description is possibly related to a specimen collected in the PORCUPINE 1869 Expedition stn. 23. Bouchet & Warén (1986) identified the latter specimen as M. orphanensis despite uncertainties. We examined the lectotype of E. solida (USNM 133126) under SEM and verified that the teleoconch surface is almost entirely smooth, marked only by strong incremental scars (Figure 17J). Melanella sp. 1 has thin axial lines, distributed over the whole teleoconch in extremely small intervals (Figure 17D). Melanella sp. 1 differs from M. orphanensis (Figure 17K) by the straight spire, with shallower sutures and a slower increasing diameter. Despite the differences mentioned above, we refrain to describe Melanella sp. 1 as new because there are several Melanella s.l. with a subcylindrical shape currently known or awaiting description (Bouchet & Warén 1986; Personal observation). We prefer to wait for new material to become available. Geographical distribution. Brazil: Pernambuco. Bathymetric distribution. 690 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on page 482, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Tryon, G. W. (1886) Manual of Conchology. Vol. 8. Philadelphia, Pennsylvania, 461 pp., 79 pls.","Jeffreys, J. G. (1884) On the Mollusca procured during the ' Lightning' and ' Porcupine' expeditions, 1868 - 70. Part VII. Proceedings of the Zoological Society of London, 52 (3), 341 - 372. https: // doi. org / 10.1111 / j. 1096 - 3642.1884. tb 02837. x","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762"]}

Costaclis Bartsch, 1947 Type species. Aclis nucleata Dall, 1889b [= Costaclis hyalina (Watson, 1881a)], by original designation; Recent, Caribbean and Northeast Brazil. Diagnosis. Shell elongated, large (> 10 mm), vitreous; protoconch smooth, subcylindrical, dome shaped; teleoconch with strongly convex whorls, sculptured by numerous axial ribs usually stronger in early whorls; aperture commonly short, umbilicus fissure commonly present (adapted from Bartsch 1947)., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on page 455, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Bartsch, P. (1947) A monograph of the West Atlantic mollusks of the family Aclididae. Smithsonian Miscellaneous Collections, 106 (20), 1 - 29, pls. 1 - 6.","Dall, W. H. (1889 b) Reports on the results of dredgings, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer ' Blake'. Bulletin of the Museum of Comparative Zoology, 18, 1 - 492, pls. 10 - 40.","Watson, R. B. (1881 a) Mollusca of H. M. S. \" Challenger \" Expedition. Part VII. Zoological Journal of the Linnean Society, 15 (85), 245 - 274. https: // doi. org / 10.1111 / j. 1096 - 3642.1881. tb 00360. x"]}

Eulima ephamilla Watson, 1883 (Figure 5 A–E) Eulima ephamilla Watson, 1883: 116–117. Watson (1886: plate 35, figure 6a–c). Eulima (“ Liostraca ”) stenostoma auct. non Jeffreys, 1858: Dall (1889a: 126). Eulima (Leiostraca) rectiuscula Dall, 1890: 160 (in part). Melanella ephamilla (Watson, 1883): Lange-de-Morretes (1949: 82); Rios (1994: 103, plate 34, figure 426 [reproduced from original illustration]); Rios (2009: 191, figure 468 [reproduced from original illustration]). Balcis ephamilla (Watson, 1883): Rios (1975: 61, plate 16, figure 239 [reproduced from original illustration]); Rios (1985: 54, plate 20, figure 242 [reproduced from original illustration]). Type material. Lectotype (herein designated): NHMUK 1887.2.9.1589 (Figure 5A, B). Paralectotypes (herein designated): NMW 1955.158.11088 [2 dd] (Figure 5C), from type locality. Holotype of Eulima rectiuscula Dall, 1890: USNM 87343 (Figure 5D, E), USFC stn. 2668 (30°58′30″N 79°38′30″W, 538 m), coll. 05/v/1886. Type locality. Brazil: off Pernambuco, Challenger stn. 122. Material examined. Type material. Measurements. Lectotype NHMUK 1887.2.9.1589, ~10 whorls (protoconch broken), SL= 8.5 mm; BWL= 3.4 mm; AL= 2.1 mm; SW= 1.8 mm; AW= 1.2 mm. Paralectotype NMW 1955.158.11088, ~10 whorls (protoconch broken), SL= 6.3 mm; BWL= 2.7 mm; AL= 1.6 mm; SW= 1.4 mm; AW= 0.9 mm. Paralectotype NMW 1955.158.11088, ~10 whorls (apex and aperture broken), SL= 8.0; BWL= 3.3 mm; AL= 1.6 mm; SW= 1.7 mm; AW= 0.9 mm. Holotype of Eulima rectiuscula, USNM 87343, ~8 whorls (apex broken), SL= 9.6; BWL= 4.0 mm; AL= 2.4 mm; SW= 2.0 mm; AW= 1.3 mm. Remarks. The shell numbered NHMUK 1887.2.9.1589 (Figure 5A, B) is the figured specimen by Watson (1886: plate 35, figure 6a–c) and is here designated as the lectotype. There are two other shells from the type locality stored in the NMW (NMW 1955.158.11088), which are here considered as paralectotypes. Watson (1883: 116) cited that “the specimen is dead and discoloured”, which could indicate that he referred to a single shell. However, Watson (1883) did not designate holotypes in his descriptions, and as other shells from the same locality were found, we prefer to consider that all were originally syntypes. See discussion for the status of types in the NMW. Watson’s (1883) description is appropriate to the lectotype and there is no need for a redescription. We only add that there is no presence of an umbilical fissure as stated by him in describing a “very minute furrow” behind the inner lip. Eulima ephamilla presents the general shape typical of Eulima sensu stricto, with an elongated, conical shell of slowly increasing diameter, flat whorls, narrow and elongated aperture as the type species of the genus, E. glabra (da Costa, 1778) (Warén 1989). MolluscaBase Eds. (2020a) cited Eulima rectiuscula Dall, 1890, from Georgia, USA, as a junior synonym of E. ephamilla without discussion. The holotype of E. rectiuscula (Figure 5D, E), collected from the USFC stn. 2668 (vide original label), also has a broken protoconch, but the general outline and dimensions are indistinguishable from the holotype of E. ephamilla. When Dall (1890) described E. rectiuscula, he also referred to samples of this species from the Caloosahatchee, a Plio-Pleistocene formation in Florida (Allmon et al. 1996). These fossil specimens were later recognized as a distinct species described by Gardner & Aldrich (1919: 39, plate 2, figure 5), currently known as Eulima dalli. In addition to the differences pointed by Gardner & Aldrich (1919), we add that the outer lip of E. ephamilla is more retracted at the proximal area than that of E. dalli. Thus, we consider E. rectiuscula as a junior synonym of E. ephamilla and that E. dalli is a valid species known only from the Miocene and Pliocene (Olsson & Harbison 1953). Eulima ephamilla is known only from two localities distant from each other by more than 6,000 km. The general shape of Eulima ephamilla resembles E. incolor Bouchet & Warén, 1986 (Figure 5F, G), from deep waters off Spain, and E. psila Watson, 1883 (Figure 6 A–M), from the western Atlantic. However, E. ephamilla can be distinguished from E. incolor by the relatively bigger shell (holotype of E. ephamilla, 10 whorls, SL= 8.5 mm vs. holotype MNHN-IM-2000-5660 of E. incolor, 11 whorls, SL= 6.6 mm) and by the more elongated shape of the aperture. The aperture of both species is oblong but the ratio AW/AL in E. ephamilla and E. incolor is very different (0.69 and 0.48, respectively). Eulima ephamilla differs from E. psila by the outline of the outer lip, which protrudes in the former (Figure 5B, E) and is straight in the later (Figure 6I). Geographical distribution. USA: Florida (Dall 1890); Brazil: Pernambuco (Watson 1883). Bathymetric distribution. From 538 m to 640 m (Watson 1883; Dall 1890)., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 460-462, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Watson, R. B. (1883) Mollusca of H. M. S. \" Challenger \" Expedition. Part XVII. Zoological Journal of the Linnean Society, 17 (99), 112 - 130. https: // doi. org / 10.1111 / j. 1096 - 3642.1883. tb 02043. x","Watson, R. B. (1886) Report on the Scaphopoda and Gasteropoda collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger, Zoology, 15 (2), 1 - 680 + 692 - 756, 50 pls.","Jeffreys, J. G. (1858) XIV-Gleanings in British Conchology. Annals and Magazine of Natural History, 2, 117 - 133. https: // doi. org / 10.1080 / 00222935808696993","Dall, W. H. (1889 a) A preliminary catalogue of the shell-bearing marine mollusks and brachiopods of the southeastern coast of the United States, with illustrations of many of the species. Bulletin of the United States National Museum, 37, 1 - 221, pls. 1 - 74.","Dall, W. H. (1890) Contributions to the Tertiary fauna of Florida, with special reference to the Miocene silex-beds of Tampa and the Pliocene beds of the Caloosahatchie River. Part I. Pulmonate, opisthobranchiate and orthodont gastropods. Transactions of the Wagner Free Institute of Science of Philadelphia, 3, 1 - 200, pls. 1 - 12.","Lange-de-Morretes, F. (1949) Ensaio de catalogo dos moluscos do Brasil. Arquivos do Museu Paraense, 7, 1 - 216.","Rios, E. C. (1994) Seashells of Brazil. 2 nd Edition. Editora da Fundac \" o Universidade do Rio Grande, Rio Grande, 368 pp., 113 pls.","Rios, E. C. (2009) Compendium of Brazilian seashells. Evangraf, Rio Grande, viii + 668 pp.","Rios, E. C. (1975) Brazilian marine mollusks iconography. Museu Oceanografico do Rio Grande, Rio Grande, 331 pp., 91 pls.","Rios, E. C. (1985) Seashells of Brazil. Editora da Fundac \" o Universidade do Rio Grande, Rio Grande, 328 pp., 102 pls.","Da Costa, E. M. (1778) Historia Naturalis Testaceorum Britanniae or The British Conchology. Printed for the author, London, 254 pp.","Waren, A. (1989) Designation of neotypes of ' Melanella alba (da Costa, 1778) ' and ' Eulima glabra (da Costa, 1778) ' (Proso- branchia). Journal of Conchology, 33, 219 - 224.","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762","Allmon, W. D., Rosenberg, G., Portell, R. W. & Schindler, K. (1996) Diversity of Pliocene-Recent mollusks in the Western Atlantic: Extinction, Origination, and Environmental change. In: Jackson, J. B., Budd, A. F. & Coates, A. G. (Eds.), Evolution and Environment in Tropical America. The University of Chicago Press, Chicago, pp. 271 - 302.","Gardner, J. A. & Aldrich, T. H. (1919) Mollusca from the Upper Miocene of South Carolina with descriptions of new species. Proceedings of the Academy of Natural Sciences of Philadelphia, 71, 17 - 53, pls. 1 - 4.","Olsson, A. A. & Harbison, A. (1953) Pliocene Mollusca of southern Florida with special reference to those from North Saint Petersburg. Academy of the Natural Sciences of Philadelphia, Monograph, 8, 1 - 361."]}

Eulima cracentis sp. nov. urn:lsid:zoobank.org:act: 90A622EA-9131-4BB6-8AAC-6187B4071FD8 (Figure 7 A–I) Type material. Holotype MNRJ 35202 † (Figure 7 A–C); Paratypes: MNRJ 23225 † [1 dd], from the type locality; MNRJ 35216 † [2 dd] (Figure 7 D–I), off Ceará, REVIZEE Nordeste (04°15′02″S 37°12′37″W, 206 m), 10/xi/2001; MNRJ 35265 † [1 dd], off Alagoas, REVIZEE Nordeste (09°20′00″S 34°59′00″W, 452 m), 18/xii/2001. Type locality. Brasil: Off Pernambuco, REVIZEE Nordeste (08°42′06″S 34°44′06″W, 465 m), coll. RV Natureza, 25/iii/2000. Etymology. From the Latin cracens, meaning neat, graceful; relative to the appearance of the shell. Description (Holotype). Shell conical, apex small and obtuse, reaching 5.9 mm long, 1.6 mm wide. Larval shell with about 2.25 whorls, 380 µm wide, 430 µm in height; smooth, transition to teleoconch marked by an almost indistinct incremental scar. Teleoconch with about 5 whorls, slightly sinuous outline, convexity confined to the lower region of each whorl; suture shallow, slightly impressed, sloping; subsutural zone about 1/4 of the height of the whorl; surface glossy and smooth; incremental scars weak, appearing at irregular intervals, frequent. Last whorl about 60% of the shell length; base elongated with slightly rounded outline. Aperture high, narrow, pear shaped, acute posteriorly and rounded anteriorly; outer lip thin, almost straight, opisthocline, slightly projecting near the suture and slightly retracting in the distal region; inner lip sinuous, sloping and well demarcated. Not umbilicate. Shell vitreous, or whitish, with a brownish spiral band at the suprasutural region of the teleoconch, last whorl with a duplicate spiral band in dorsal view. Measurements. Holotype MNRJ 35202†, 7.25 whorls, SL= 5.9 mm; BWL= 3.6 mm; AL= 2.4 mm; SW= 1.6 mm; AW= 1.0 mm. Paratype MNRJ 35216†, 8 whorls, SL= 6.8 mm; BWL= 4.2 mm; AL= 2.7 mm; SW= 1.9 mm; AW= 1.3 mm. Remarks. All types of the present species were destroyed; however, it has consistent shell features that allow the description as a new species, and the figures provided here are sufficient for its recognition. Eulima cracentis sp. nov. presents brownish spiral bands on the teleoconch (Figure 7 A–C), which is a common feature in the genus. Eulima cracentis sp. nov. can be distinguished from E. bifasciata, a shallow water species of the western Atlantic, by the presence of a single brownish spiral band on the teleoconch, except on the last whorl (Figure 7C), while E. bifasciata shows two brownish spiral bands per whorl (see Lamy & Pointier 2017: plate 88, figure 1a, b). Furthermore, E. cracentis sp. nov. has more convex teleoconch whorls and a more inflated body whorl. Eulima cracentis sp. nov. can also be distinguished from E. ephamilla (Figure 5 A–E) and E. psila (Figure 6 A–M) by the presence of the brownish spiral band at the teleoconch, more convex teleoconch whorls and the relatively higher aperture in relation to the body whorl length (ratio AL/BWL= 0.67, 0.60 and 0.63, respectively). Furthermore, E. cracentis sp. nov. has a smaller number of protoconch whorls (2.25 vs. 3.5) and a wider protoconch (380 µm vs. 320 µm) than E. psila. The shell figured by Lamy & Pointier (2017: plate 88, figure 4a–b) of an unnamed species of Eulima collected in Guadeloupe at 200 m resembles E. cracentis sp. nov., but has a narrower protoconch and teleoconch, and seems to have just a single spiral band on the body whorl. Geographical distribution. Brazil: Ceará, Pernambuco, Alagoas. Bathymetric distribution. From 206 m to 465 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 464-466, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Lamy, D. & Pointier, J-P. (2017) Marine and freshwater molluscs of the French Caribbean. Vols. 1 - 2. PLB Editions, Beta, 785 pp."]}

Eulima sp. 1 (Figure 8 A–I) Material examined. Brazil: Rio Grande do Norte: REVIZEE Nordeste (04°51′00″S 35°06′00″W, 375 m), coll. RV Natureza, 24/xi/2001: MNRJ 29631 † [1 dd], MNRJ 35215 † [1 dd]. Description. Shell conical, apex rounded, reaching about 6.4 mm long, 1.6 mm wide. Larval shell with about 2.5 whorls, 290 µm wide, 370 µm in height; smooth, transition to teleoconch marked by a subtle incremental scar. Teleoconch with about 7.5 whorls, almost flat; suture shallow, slightly impressed, sloping; surface glossy and smooth; incremental scars weak, appearing at irregular intervals. Last whorl about 50% of the shell length; base elongated, slightly rounded outline, connected close to the most anterior region of the aperture. Aperture high, narrow, pear shaped, acute posteriorly and rounded anteriorly; outer lip thin, slightly sinuous, opisthocline, slightly projecting near the suture and slightly retracting in the distal region; inner lip sinuous, sloping and well demarcated. Not umbilicate. Shell vitreous or whitish, with a brownish spiral band at the suture of the larval shell. Measurements. MNRJ 29631, 10 whorls, SL= 6.4 mm; BWL= 3.2 mm; AL= 2.3 mm; SW= 1.6 mm; AW= 1.0 mm. Remarks. The shell surface of the two specimens examined is corroded (Figure 8E) and at first, it seems that it presents a sculpture of axial lines, like some species in the genera Melanella Bowdich, 1822 and Niso Risso, 1826. However, this is an artifact as examined in SEM and there are only incremental scars (Figure 8E). The general shape and the aperture are more similar to Eulima s.s. Other Eulima species that resemble Eulima sp. 1 are E. psila (Figure 6 A–M) and E. grimaldii (Figure 6P, Q), both with a brownish spiral band at the protoconch. However, Eulima sp. 1 differs from E. psila by features of the protoconch, which has a smaller number of whorls (2.5 vs. 3.5 in E. psila) and is shorter (370 µm vs. 420 µm), and by the outline of the outer lip, which is slightly protruding in E. sp. 1 (Figure 8G) and straight in E. psila (Figure 6I). Eulima sp. 1 also differs from E. grimaldii by features of the protoconch, which has less convex whorls in E. sp. 1 (see Figure 8H and Bouchet & Warén 1986: figure 93) and a smaller number of whorls (2.5 vs. 3.3 in E. grimaldii). Furthermore, the body whorl comprises 50% of the shell length in E. sp. 1 and 40% of the shell length in E. grimaldii, and the aperture is relatively wider and higher in E. sp. 1 comparing shells with a similar number of whorls (AL= 2.3 mm, AW= 1.0 mm vs. AL= 1.6 mm, AW= 0.7 mm, holotype of E. grimaldii). Eulima sp. 1 can also be distinguished from Eulima incolor (Figure 5F, G) by the brownish spiral band on the protoconch, which is not present in E. incolor, and by the presence of several incremental scars on the teleoconch, while Bouchet & Warén (1986) described that E. incolor does not present any incremental scar on the teleoconch. Despite the differences mentioned above, we refrain to name Eulima sp. 1 as a new species because of the almost inconspicuous features of the species. Most species of Eulima s.s. are very similar to each other, especially those that do not have a colored teleoconch and we prefer to leave this species undescribed until more samples become available. Geographical distribution. Brazil: Rio Grande do Norte. Bathymetric distribution. 375 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 466-468, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Bowdich, T. E. (1822) Elements of Conchology. I. J. Smith, Paris, 79 pp.","Risso, A. (1826) Histoire naturelle de l'Europe meridionale. 4. Levrault Libraire, Paris, 439 pp.","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762"]}

Costaclis hyalina (Watson, 1881) (Figure 2 A–G) Aclis hyalina Watson, 1881a: 246–247. Watson (1886: plate 34, figure 2a–d); Lange-de-Morretes (1949: 82); Rios (1975: 62, plate 16, figure 245 [reproduced from original illustration]); Rios (1985: 52, plate 19, figure 234 [reproduced from original illustration]); Rios (1994: 106, plate 34, figure 438 [reproduced from original illustration]); Aclis nucleata Dall, 1889b: 325–326, plate 18, figure 7. Synonymized by Bouchet & Warén (1986). Aclis lata auct. non. Dall, 1889b: Dall (1927: 70). Hemiaclis benedicti Bartsch, 1947: 22–23, plate 5, figure 5. Synonymized by Bouchet & Warén (1986). Hemiaclis hyalina auct. non (Watson, 1881a): Bartsch (1947: 21, plate 6, figure 5). Costaclis hyalina (Watson, 1881a): Bouchet & Warén (1986: 450). Type material. Lectotype (herein designated): NHMUK 1887.2.9.1575 (Figure 2A, B); paralectotypes (herein designated): NMW 1955.158.26258 [3 dd], from type locality. Aclis nucleata Dall, 1889b: holotype USNM 126830 (Figure 2C, D); Hemiaclis benedicti Bartsch, 1947: holotype USNM 87356 (Figure 2E). Type locality. Brazil: Off Pernambuco, Challenger stn. 122 (09°04′48″S 34°49′48″W, 640 m), coll. 10/ ix/1873. Material examined. Type material. Measurements. Lectotype NHMUK 1887.2.9.1575, ~12 whorls, SL= 10.4 mm; BWL= 3.8 mm; AL= 2.0 mm; SW= 3.2 mm; AW= 1.7 mm. Paralectotype NMW 1955.158.26258, 9 whorls, SL= 5.4 mm; BWL= 2.4 mm; AL= 1.5 mm; SW= 2. 0 mm; AW= 1.0 mm. Paralectotype NMW 1955.158.26258, 8 whorls, SL= 4.0 mm; BWL= 1.9 mm; AL= 1.1 mm; SW= 1.5 mm; AW= 0.8 mm Remarks. Watson (1881a) did not fix a holotype and did not mention the number of specimens upon which the description was based. Two lots of Aclis hyalina collected from Challenger stn. 122 were found in the NHMUK and NMW, which are considered syntypes. Watson (1881a) recorded the species only from the Challenger stn. 122. The original label of the NHMUK syntype states “f’ type” [figured type], and comparing this syntype to the illustration of Watson (1886: plate 34, figure 2a–d), it is clear that he figured and measured the shell from NHMUK 1887.2.9.1575 (Figure 2A, B). This shell is selected as the lectotype (Figure 2A, B). Barros et al. (2003) wrongly cited holotype and paratypes of “ H. hyalina ” under “BMNH [=NHMUK] 2346”. As discussed above, the material originally consisted of syntypes and the catalog number is incorrect. The original description of the species is appropriate to the lectotype and there is no need for a redescription. We just emphasize that the species does not present spiral sculpture as described by Watson (1881). The spiral lines cited by him (“visible in a changing light”) are probably a refractive phenomenon commonly seem in eulimid shells (Warén 1983: 14). Costaclis hyalina was formerly described in Aclis (Watson 1881a). Bartsch (1947) erected the genus Costaclis Bartsch, 1947 for species of Aclididae with well or poorly developed axial ribs, but classified A. hyalina in the genus Hemiaclis G.O. Sars, 1878. Bouchet & Warén (1986: 300) transferred the genera Costaclis and Hemiaclis to Eulimidae and proposed the combination C. hyalina, which we follow in the present study. The lectotype of Costaclis hyalina (Figure 2A, B) presents poorly developed axial ribs, which are more evident in the incremental scars, appearing as gentle varices. The holotypes of A. nucleata (USNM 126830) (Figure 2C, D, F, G) and H. benedicti (USNM 87356) (Figure 2E), junior synonyms of C. hyalina (Bouchet & Warén 1986), present the ribs more pronounced, especially in the initial teleoconch whorls of A. nucleata (Figure 2G). We consider these small differences in the development of the sculpture as a variation or a condition of the worn status of these shells. Bouchet & Warén (1986: 450) mentioned that Costaclis mizon (Watson, 1881) (Figure 3E), from the eastern Atlantic (Watson 1881a) and Gulf of Mexico (Rosenberg et al. 2009: 582), may be a synonym of C. hyalina. Indeed, the spire angle and width of the whorls are very similar in the lectotype of C. hyalina and in the holotype of C. mizon (NHMUK 1887.2.9.1574) (Figure 3E), however, C. mizon has a much more well-developed sculpture than any other specimen of C. hyalina from the western Atlantic. For now, we consider C. mizon as a valid species based on the teleoconch sculpture. Costaclis hyalina was known only from the type locality for a century, until some new records in the last decades. However, subsequent records of C. hyalina in Brazil are very dubious or based on misidentifications. Mello & Perrier (1986: 121) recorded “ Aclis hyalina ” based on material collected in the intertidal zone, thus this is possibly a misidentification. There is no illustration of the species and the material could not be traced in the LMUFRPE. Barros et al. (2001: figure 2d) recorded “ Hemiaclis hyalina ”, but this is a misidentification and the shell figured by these authors is possibly a new species of Aclis. The shell illustrated has more convex whorls and the aperture is further expanded laterally. The record by Barros et al. (2002) presents no illustrations and could not be traced in LMUFRPE, but it seems dubious by the shallow depth of the collecting stations (around 50 m). The record of “ H. hyalina ” by Barros et al. (2003: figure 1) is also dubious, because the figured specimen is very young and does not permit a precise identification. Thus, based on the reevaluation of previous records of C. hyalina in the literature, we consider them invalid and the species is known in Brazil by the type material only. Geographical distribution. USA: Georgia, Florida (Dall 1927; Bartsch 1947); St. Vincent and the Grenadines: St. Vincent (Dall 1889b); Brazil: Pernambuco (Watson 1881a). Bathymetric distribution. From 538 m to 848 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 456-457, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Watson, R. B. (1881 a) Mollusca of H. M. S. \" Challenger \" Expedition. Part VII. Zoological Journal of the Linnean Society, 15 (85), 245 - 274. https: // doi. org / 10.1111 / j. 1096 - 3642.1881. tb 00360. x","Watson, R. B. (1886) Report on the Scaphopoda and Gasteropoda collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger, Zoology, 15 (2), 1 - 680 + 692 - 756, 50 pls.","Lange-de-Morretes, F. (1949) Ensaio de catalogo dos moluscos do Brasil. Arquivos do Museu Paraense, 7, 1 - 216.","Rios, E. C. (1975) Brazilian marine mollusks iconography. Museu Oceanografico do Rio Grande, Rio Grande, 331 pp., 91 pls.","Rios, E. C. (1985) Seashells of Brazil. Editora da Fundac \" o Universidade do Rio Grande, Rio Grande, 328 pp., 102 pls.","Rios, E. C. (1994) Seashells of Brazil. 2 nd Edition. Editora da Fundac \" o Universidade do Rio Grande, Rio Grande, 368 pp., 113 pls.","Dall, W. H. (1889 b) Reports on the results of dredgings, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer ' Blake'. Bulletin of the Museum of Comparative Zoology, 18, 1 - 492, pls. 10 - 40.","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762","Dall, W. H. (1927) Small shells from dredgings off the southeast coast of the United States by the United States Fisheries Steam- er \" Albatross \" in 1885 and 1886. Proceedings of the United States National Museum, 70 (2667), 1 - 134. https: // doi. org / 10.5479 / si. 00963801.70 - 2667.1","Bartsch, P. (1947) A monograph of the West Atlantic mollusks of the family Aclididae. Smithsonian Miscellaneous Collections, 106 (20), 1 - 29, pls. 1 - 6.","Barros, J. C. N., Mello, R. L. S., Barros, F. N., Lima, S. F. B., Santos, M. C. F., Cabral, E. & Padovan, I. P. (2003) Sistematica dos gastropodes Aclididae G. O. Sars, 1878 da plataforma continental e em aguas profundas do Nordeste do Brasil. Boletim Tecnico Cientifico do CEPENE, 11 (1), 63 - 90.","Waren, A. (1983) A generic revision of the family Eulimidae (Gastropoda: Prosobranchia). Journal of Molluscan Studies, Supplement 13, 1 - 96. https: // doi. org / 10.1093 / mollus / 49. Supplement _ 13.1","Sars, G. O. (1878) Mollusca regionais arcticae Norvegiae. Br ˆ gger, Christiania, 466 pp.","Rosenberg, G. F., Moretzsohn, F., Garcia, E. F. (2009) Gastropoda (Mollusca) of the Gulf of Mexico. In: Felder, D. L. & Camp, D. K. (Eds.), Gulf of Mexico-Origins, Waters and Biota. Biodiversity. Texas A & M University Press, College Station, Texas, pp. 579 - 699.","Mello, R. L. S. & Perrier, L. L. (1986) Polyplacophora e Gastropoda do litral Sul de Pernambuco, Brasil. Caderno Omega, Universidade Federal Rural de Pernambuco. Serie Ciencias Aquaticas, 2, 107 - 144.","Barros, J. C. N., Santos, F. N., Santos, M. C. F., Cabral, E. & Acioli, F. D. (2001) Redescoberta de moluscos obtidos durante a \" Challenger Expedition \" (1873 - 1876): micromoluscos de aguas profundas. Boletim Tecnico Cientifico do CEPENE, 9 (1), 9 - 24.","Barros, J. C. N., Santos, F. N., Cabral, E. & Santos, M. C. F. (2002) Moluscos dragados pelo Navio Oceanografico \" Victor Hensen \", ao largo do Arquipelago de Fernando de Noronha, Nordeste do Brasil. Boletim Tecnico Cientifico do CEPENE, 10, 27 - 37."]}

Umbilibalcis lata (Dall, 1889b) (Figure 24 A–I) Aclis (supranitida var. ?) lata Dall, 1889b: 324–325, plate 18, figure 8. Aclis lata: Dall (1889a: 126, plate 18, figure 8) [reproduced from original illustration]. Hemiaclis lata: Bartsch (1947: 18, plate 4, figure 4) Umbilibalcis lata: Bouchet & Warén (1986: 444–445, figure 1056–1057). Type material. Holotype USNM 126790 (Figure 24A). Type locality. Barbados, 183 m. Material examined. Type material; USA: USFC stn. 2404 (28°44′N 85°16′W, 110 m), coll. 15/iii/1885: USNM 323963 [1 dd]; USBF stn. 20030 (~24°25′N 82°58′, 164 m): USNM 407033 [1 dd]. Antigua and Barbuda: English Harbor, S.U.I. stn. 496 (219 m): USNM 459689 [1 dd]. Barbados: Off Pelican Islands, S.U.I. Expedition stn. 454 (182 m): USNM 459677 [1 dd]; Off Lazareto, S.U.I. Expedition stn. 497 (171 m): USNM 459708 [1 dd]. Brazil: Rio Grande do Norte: (04°51′S 35°06′W, 375 m), 24/xi/2001: MNRJ 35200 † [1 dd]; Bahia: (11°35′31″S 37°12′18″W, 510 m), 30/x/2000: MNRJ 35268 † [3 dd]; REVIZEE-Central V stn. 504 (14°28′55″S 38°54′03″W, 278 m), coll. Supply Boat Astro Garoupa, 02/vii/2001: IBUFRJ 14399 † [5 dd]; Espírito Santo: REVIZEE-Central V stn. 25a (19°33′28″S 38°45′28″W, 230 m), coll. Supply Boat Astro Garoupa, 28/vi/2001: IBUFRJ 19717 † [3 dd]; Rio de Janeiro: Campos Basin (22°18′53″S 40°11′59″W, 200 m), coll. Supply Boat Astro Garoupa, 18/xii/2004: MNRJ 31867 † [2 dd]; Campos Basin (22°42′33″S 40°41′24″W, 110 m), coll. 19/ix/2003: MNRJ 31881 † [2 dd], MNRJ 31895 † [1 dd]; PADCT stn. 6541: MNRJ 28285 † [3 dd]; PADCT stn. 6627: MNRJ 28287 † [1 dd]; Cabo Frio VII stn. 6194 (24°03′36″S 44°07′36″W, 134 m), coll. RV Almirante Saldanha, 01/iv/1983: IBUFRJ 19727 † [1 dd]; S„o Paulo: REVIZEE Sul stn. 6672: MNRJ 28289 † [1 dd]; PADCT stn. 6573: MNRJ 23226 † [1 dd]; Paraná: REVIZEE Sul stn. 6653: MNRJ 28288 † [1 dd]; Santa Catarina: REVIZEE Sul stn. 6695: MNRJ 28290 † [1 dd]; PADCT stn. 6595: MNRJ 28283 † [7 dd]; PADCT stn. 6606: MNRJ 28286 † [1 dd]; Rio Grande do Sul: off Bojuru (220 m), coll. RV Almirante Saldanha, xi/1979: MORG 21416 † [1 dd]. Redescription (Holotype). Shell conical, straight, reaching about 5.4 mm long, 2.4 mm wide. Larval shell brownish, partially broken. Teleoconch with about 8.0 whorls, slightly convex outline; suture distinct, sloping; subsutural region not visible; surface polished, presenting slightly opisthocline axial lines; incremental scars slightly demarcated, appearing at irregular intervals. Last whorl about 45% of the shell length; base rounded.Aperture wide, pear shaped, acute posteriorly and rounded anteriorly; outer lip thin, slightly prosocline, sinuous, slightly retracted near the suture, reaching a maximum projection in the middle of its height, retracted at distal region; inner lip sinuous, sloping, reflected and well demarcated. Umbilicus present. Shell whitish. Measurements. Holotype USNM 126790, 10 whorls (protoconch partially broken), SL= 5.4 mm; BWL= 2.5 mm; AL= 1.6 mm; SW= 2.4 mm; AW= 1.2 mm. IBUFRJ 14399, 10 whorls, SL= 5.0 mm; BWL= 2.3 mm; AL= 1.4 mm; SW= 2.2 mm; AW= 1.1 mm. Remarks. The protoconch is partially broken in the holotype of Umbilibalcis lata (Figure 24A), but the general shape, dimensions and sculpture fits perfectly with specimens from Brazil (Figure 24 B–I). The protoconch of the shells from Brazil have about 3.0 whorls, the first (embryonic) whorl with a smooth surface, the remaining whorls with strongly opisthocline axial lines (Figure 24 H–I). The protoconch is about 270 μm wide and 390 μm in height. A weak brownish band close to suture is present on the protoconch of fresh specimens. The outer lip of the holotype is also partially broken, but it seems uniform in all specimens that the proximal area is slightly advanced, giving a prosocline outline. Umbilibalcis lata differs from U. crassula Bouchet & Warén, 1986 and U. subumbilicata (Jeffreys, 1883) by the higher number of protoconch whorls (3.0 vs. 1.5 in U. crassula and U. subumbilicata) and by the straighter outer lip (Bouchet & Warén 1986). Geographical distribution. USA: Florida (present study); Antigua and Barbuda (present study); Barbados (Dall 1889b); Brazil: Rio Grande do Norte, Bahia, Espírito Santo, Rio de Janeiro, S„o Paulo, Paraná, Santa Catarina, Rio Grande do Sul (present study). Bathymetric distribution. From 60 m to 375 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 493-495, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Dall, W. H. (1889 b) Reports on the results of dredgings, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer ' Blake'. Bulletin of the Museum of Comparative Zoology, 18, 1 - 492, pls. 10 - 40.","Dall, W. H. (1889 a) A preliminary catalogue of the shell-bearing marine mollusks and brachiopods of the southeastern coast of the United States, with illustrations of many of the species. Bulletin of the United States National Museum, 37, 1 - 221, pls. 1 - 74.","Bartsch, P. (1947) A monograph of the West Atlantic mollusks of the family Aclididae. Smithsonian Miscellaneous Collections, 106 (20), 1 - 29, pls. 1 - 6.","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762"]}

Sticteulima Laseron, 1955 Type species. Sticteulima cameroni Laseron, 1955, by original designation, Recent; New South Wales, Australia. Diagnosis. Shell small (1.5–4.0 mm), conical, straight or curved, vitreous and commonly with brownish/yellowish spots, spiral and/or axial bands; protoconch paucispiral or multispiral, whorls usually flat; teleoconch whorls slightly convex; umbilicus absent or an umbilical fissure present (adapted from Laseron 1955; Warén 1983; Mifsud & Ovalis 2019). Remarks. MolluscaBase Eds. (2020d) included 16 species in Sticteulima. In the Atlantic Ocean, the genus is known from a few species that occur in the East Atlantic: Sticteulima badia (Watson, 1897), S. fuscopunctata (E.A. Smith, 1890), Sticteulima lata Bouchet & Warén, 1986, Sticteulima richteri Engl, 1997, Sticteulima wareni Engl, 1997 (Smith 1890; Watson 1897; Bouchet & Warén 1986; Engl 1997a, b). Only S. lata occurs in deep waters, but Bouchet & Warén (1986: 322) commented that this species “may ultimately prove to belong to more shallow water”, but they did not find material to corroborate this assumption. In the western Atlantic, several species of Sticteulima occur, but were never described or treated under this genus (personal observation). Sticteulima badia is possibly a junior synonym of S. fuscescens.The holotype of S. badia (NMW 1955.158.00653) is severely damaged by Byne’s disease, which makes comparison difficult, but the shape, dimensions and color pat-tern is the same as the syntypes of S. fuscescens (NHMUK 1889.10.1.803–5)., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on page 487, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Laseron, C. F. (1955) Revision of the New South Wales Eulimoid Shells. The Australian Zoologist, 12 (2), 83 - 107.","Waren, A. (1983) A generic revision of the family Eulimidae (Gastropoda: Prosobranchia). Journal of Molluscan Studies, Supplement 13, 1 - 96. https: // doi. org / 10.1093 / mollus / 49. Supplement _ 13.1","Mifsud, C. & Ovalis, P. (2019) Two new species of Sticteulima Laseron, 1955 (Gastropoda: Eulimidae) from Turkey, eastern Mediterranean. Bollettino Malacologico, 55, 68 - 71.","Watson, R. B. (1897) On the Marine Mollusca of Madeira; with Descriptions of Thirty-five new Species, and an Index-List of all the known Sea-dwelling Species of that Island. Journal of the Linnean Society of London, 26 (168), 233 - 329. https: // doi. org / 10.1111 / j. 1096 - 3642.1897. tb 00033. x","Smith, E. A. (1890) Report of the marine molluscan fauna of the Island of St. Helena. Proceedings of the Zoological Society of London, 1890, 247 - 317, pls. 21 - 24.","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762"]}

Melanella adiastalta sp. nov. urn:lsid:zoobank.org:act: 3A3EAB3A-11EB-4CD3-8D7C-C50738650225 (Figure 15 A–G) Type material. Holotype MNRJ 35250 † (Figure 15 A–G). Type locality. Brazil: Off Rio Grande do Norte, REVIZEE Nordeste (06°14′04″S 34°52′33″W, 510 m), coll. RV Natureza, 26/xi/2001. Etymology. From adiastaltos Gr., meaning not clearly defined, indefinite, referring to the inconspicuous features of the species. Description (Holotype). Shell conical, straight, reaching about 3.6 mm long, 1.3 mm wide. Larval shell whitish, conical, with 2.75 convex whorls, 330 µm wide, 380 µm in height; smooth, transition to teleoconch marked by distinct incremental scar. Teleoconch with about 5.5 whorls, convex outline; suture well demarcated, sloping; subsutural zone indistinct; surface smooth, incremental scars well demarcated, appearing at intervals of about 0.7 whorl. Last whorl about 50% of the shell length; base rounded. Aperture short, pear shaped, acute posteriorly and rounded anteriorly; outer lip thin, opisthocline, sinuous, slightly retracting near the suture and slightly projecting in the uppermost third; inner lip sinuous, sloping, reflected and well demarcated. Not umbilicate. Shell vitreous, colorless. Measurements. Holotype MNRJ 35250†, 8 whorls, SL= 3.6 mm; BWL= 1.9 mm; AL= 1.2 mm; SW= 1.3 mm; AW= 0.7 mm. Remarks. Melanella adiastalta sp. nov. at first has inconspicuous features but can be distinguished from other similar species occurring in deep waters of the Atlantic. Despite the presence of only a single shell, we describe M. adiastalta sp. nov. based on the combination of consistent shell features that enable us to discriminate this species from remaining eulimids. Melanella adiastalta sp. nov. can be distinguished from M. sarissa (Figure 12 A–K) because it has more convex teleoconch whorls, a wider spire angle (26° vs. 22° in M. sarissa) and a smaller number of protoconch whorls (2.75 vs. 4.0). Melanella adiastalta sp. nov. can be distinguished from M. insculpta Bouchet & Warén, 1986, from the North Atlantic, by the faster increase in diameter of the spire. Both species reach about 8.5 whorls, but M. adiastalta sp. nov. has 1.3 mm wide while M. insculpta has 0.9 mm wide (Bouchet & Warén 1986: 376). Additionally, the outer lip of M. adiastalta sp. nov. protrudes gradually (Figure 15E) while it is more accentuated in M. insculpta (see Bouchet & Warén 1986: figs. 885, 887). Melanella adiastalta sp. nov. differs from M. martynjordani (Jordan, 1895), M. obtusoapicata Bouchet & Warén, 1986, M. pseudoglabra (Dautzenberg & Fischer, 1912) and M. turrita Bouchet & Warén, 1986, all from the Northeast Atlantic area, mainly by the more convex teleoconch whorls (Figure 15A, B). Furthermore, M. adiastalta sp. nov. has a more protruding outer lip (Figure 15E) than M. martynjordani and a higher number of whorls in the protoconch (Figure 15G) than M. turrita (2.75 vs. 1.5). Melanella adiastalta sp. nov. resembles Eulima anonyma Bouchet & Warén, 1986, from off Portugal, in the number and shape of protoconch whorls, but the protoconch is shorter (380 μm high vs. 500 μm high in E. anonyma) (Bouchet & Warén 1986: 467). Furthermore, M. adiastalta sp. nov. has a wider shell, despite reaching a smaller number of whorls (8 whorls, SW= 1.3 mm vs. 10 whorls, SW= 1.2 mm). Geographical distribution. Only known from the type locality. Bathymetric distribution. 510 m.

Costaclis egregia (Dall, 1889b) (Figures 3 A–D, 4A–G) Aclis egregia Dall, 1889b: 325, plate 18, figure 12. Aclis (Costaclis) egregia Dall, 1889b: Abbott (1974: 132, figure 1456). Costaclis egregia (Dall, 1889b): Absal„o et al. (2003: 324, figure 1–3); Rios (2009: 194, figure 480 [reproduced from Absal„o et al. 2003]); Absal„o (2010: 92, not figured). Type material. Lectotype (herein designated): USNM 126791 (Figure 3A). Paralectotype (herein designated): USNM 126799 (Figure 3B), “Blake” stn. 163 (16°03′10″N 61°52′20″W, 1406 m), coll. 20/i/1879, Guadeloupe. Type locality. Off St. Vincent and the Grenadines, “Blake” stn. 228 (13°11′20″N 61°19′48″W, 1436 m), 19/ ii/1879. Material examined. Type material; REVIZEE-Central V stn. 4f (14°48′50″S 38°43′58″W, 1700 m), coll. Sup-ply Boat Astro Garoupa, 02/vii/2001: IBUFRJ 12360 † [2 dd]; Oceanprof I stn. 85: IBUFRJ 15247 † [1 dd]. Redescription (Lectotype). Shell whitish, conical, apex small and obtuse, reaching 13.3 mm long, 5.4 mm wide. Larval shell subcylindrical, with about 3.0 convex whorls; smooth; transition to teleoconch unclear. Teleoconch with about 7 whorls, strongly convex; suture deep, well impressed, sloping; false suture not visible; presence of axial ribs, more developed at mid whorls; growth lines visible through all whorls; incremental scars strong, appearing at irregular intervals. Last whorl about 50% of the shell length; base strongly rounded. Aperture broad, pear shaped, acute posteriorly and rounded and slightly advanced anteriorly; outer lip thin, opisthocline, rectilinear, advancing uniformly, maximum projection at distal area; inner lip sinuous, sloping and well demarcated. Umbilical fissure present. Measurements. Lectotype USNM 126791, ~10 whorls, SL= 13.3 mm; BWL= 6.7 mm; AL= 4.4 mm; SW= 5.4 mm; AW= 3.1 mm. Paralectotype USNM 126799 (protoconch partially broken), ~9 whorls, SL= 11.0 mm; BWL= 5.8 mm; AL= 3.7 mm; SW= 4.6 mm; AW= 2.4 mm. IBUFRJ 15247†, ~11 whorls (protoconch partially broken), SL= 19.8 mm; BWL= 9.4 mm; AL= 5.6 mm; SW= 7.5 mm; AW= 4.2 mm. IBUFRJ 12360, ~8 whorls, SL= 6.9 mm; BWL= 3.9 mm; AL= 2.5 mm; SW= 3.1 mm; AW= 1.6 mm. Remarks. The shell from USNM 126791 is selected as the lectotype (Figure 3A) since it is the best preserved and the figured specimen by Dall (1889b). A redescription is provided here to be more specific about the features present in the lectotype and to add features not cited in the original description (i.e., protoconch, incremental scars, base, outer lip). Some shells of C. egregia (e.g., paralectotype USNM 126799 [Figure 3B] and IBUFRJ 12360 [Figure 4 A–G]) are covered by a brownish layer of possible minerals, which can be related to the type of environment they inhabit in the deep sea. Costaclis egregia was formerly introduced in Aclis but later transferred to Costaclis by Bartsch (1947). Bouchet & Warén (1986) did not comment about the status of this species in Costaclis, but their statement that “ … Costaclis was erected by Bartsch for a mixture of slender species with axial ribs, but only the type species can be kept in the genus”, indicates that they disregard this species as a member of Costaclis. However, the presence of numerous and well-developed axial ribs fits the concept of the genus established by Bartsch (1947). Dall (1889b) described the species without an umbilicus, but an umbilical fissure is usually present in adult specimens (Figure 3C). We can also add to the original description that the protoconch is subcylindrical. Absal „o et al. (2003) were the first to record C. egregia in Brazil based on material from the continental slope off Northeast Brazil. These authors cited this species in a depth of 250 m, which would be the shallowest record of C. egregia. However, it is an incorrect indication of coordinate and depth. Actually, REVIZEE-Central V stn. 4f corresponds to the coordinate 14°48′50″S 38°43′58″W and depth of 1700 m (Lavrado 2006), which becomes the deepest record of the species. We reassessed the record by Absal„o (2010: annotated list) and this represents the southernmost record of the species (~ 22°S). Costaclis egregia can be distinguished from C. hyalina, a congener that also occurs in Brazil, by the teleoconch sculpture with more numerous and well-developed axial ribs, and by the more convex teleoconch whorls. Costaclis egregia can also be distinguished from C. mizon by the less pronounced axial ribs, wider spire angle and by the greater length. One shell of C. egregia studied here (Figure 3C) with 11 whorls reaches 19.8 mm long while the lectotype of C. mizon (Figure 3E) has 15 whorls and is 15.4 mm long. Geographical distribution. Guadeloupe, St. Vincent and Grenadines (Dall 1889b); Brazil: Bahia, Rio de Janeiro (Absal„o et al. 2003; Absal„o 2010; present study). Bathymetric distribution. From 1350 m to 1700 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 458-460, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Dall, W. H. (1889 b) Reports on the results of dredgings, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer ' Blake'. Bulletin of the Museum of Comparative Zoology, 18, 1 - 492, pls. 10 - 40.","Abbott, R. T. (1974) American seashells. The marine Mollusca of the Atlantic and Pacific coasts of North America. Van Nostrand Reinhold Company, New York, 663 pp.","Rios, E. C. (2009) Compendium of Brazilian seashells. Evangraf, Rio Grande, viii + 668 pp.","Bartsch, P. (1947) A monograph of the West Atlantic mollusks of the family Aclididae. Smithsonian Miscellaneous Collections, 106 (20), 1 - 29, pls. 1 - 6.","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762","Lavrado, H. P. (2006) Caracterizac \" o do ambiente e da comunidade bentonica. In: Lavrado, H. P. & Ignacio, B. L. (Eds.), Biodiversidade bentonica da regiao central da Zona Economica Exclusiva brasileira. Museu Nacional, UFRJ, Rio de Janeiro, pp. 19 - 64."]}

Melanella Bowdich, 1822 Type species. Melanella dufresnii Bowdich, 1822, by monotypy (Recent / Mauritius Islands, Indian Ocean). Diagnosis. See Souza & Pimenta (2019b). Remarks. Souza & Pimenta (2019b) discussed the shell morphology of the type species in detail. A concise diagnosis of the genus is doubtful since Melanella comprises a wide variety of species (Bouchet & Warén 1986; Souza & Pimenta 2019b). Here, we follow previous authors in considering a great diversity in shell morphology in the present genus., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on page 472, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Bowdich, T. E. (1822) Elements of Conchology. I. J. Smith, Paris, 79 pp.","Souza, L. S. & Pimenta, A. D. (2019 b) Taxonomy of littoral Melanella (Gastropoda: Eulimidae) from Brazil, with comments on the Eulima species described by Verrill and Bush (1900). Marine Biodiversity, 49 (1), 425 - 442. https: // doi. org / 10.1007 / s 12526 - 017 - 0825 - x","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762"]}

Eulima Risso, 1826 Type species. Turbo subulatus Donovan, 1804 [= Eulima glabra (da Costa, 1778)], by subsequent designation; Recent, Northeast Atlantic and Mediterranean. Diagnosis. Shell elongated, slender, small to large (5–15 mm), commonly presents brownish/yellowish spiral and/or axial bands close or at the suture; protoconch smooth, usually conical, whorls slightly convex; teleoconch smooth, with flat (more frequently) or slightly convex whorls; sutures almost indistinct; aperture elongated, slender, outer lip usually straight or slightly sinuous, distinct callus on the parietal wall and the columella; umbilicus absent (adapted from Bouchet & Warén 1986; Warén 1989)., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on page 460, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Risso, A. (1826) Histoire naturelle de l'Europe meridionale. 4. Levrault Libraire, Paris, 439 pp.","Da Costa, E. M. (1778) Historia Naturalis Testaceorum Britanniae or The British Conchology. Printed for the author, London, 254 pp.","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762","Waren, A. (1989) Designation of neotypes of ' Melanella alba (da Costa, 1778) ' and ' Eulima glabra (da Costa, 1778) ' (Proso- branchia). Journal of Conchology, 33, 219 - 224."]}

Eulima? hebes Watson, 1883 (Figure 25 A–K) Eulima hebes Watson, 1883: 127–128. Watson (1886: 521, plate 37, figure 6a–b). Melanella hebes (Watson, 1883): Lange-de-Morretes (1949: 82); Rios (1994: 104, plate 34, figure 428 [reproduced from original illustration]); Barros et al. (2001: figure 5a); Rios (2009: 192, figure 471 [reproduced from original illustration]). Balcis (?) hebes (Watson, 1883): Rios (1975: 61, plate 16, figure 240 [reproduced from original illustration]); Rios (1955: 54, plate 20, figure 243 [reproduced from original illustration]). Zebina hebes (Watson, 1883): MolluscaBase Eds. (2020f). Type material. Lectotype (herein designated) NHMUK 1887.2.9.1618 (Figure 25A); paralectotypes (herein designated) NHMUK 1887.2.9.1619–1621 [3 dd] (Figure 25 B–D), from the type locality; paralectotype NMW 1955.158.26256 (Figure 25E), from the type locality. Type locality. Brazil: off Pernambuco, Challenger stn. 122. Material examined. Type material; Brazil: Pernambuco: (08°42′06″S 34°44′06″W, 465 m) coll. RV Natureza, 25/iii/2000: MNRJ 35196 † [6 dd]; (08°46′00″S 34°44′00″W, 690 m) coll. RV Natureza, 18/xi/2000: MNRJ 35214 † [1 dd], MORG 49210 † [6 dd], MORG 50046 † [20 dd]. Measurements. Lectotype NHMUK 1887.2.9.1618, 5 whorls (aperture partially broken), SL= 3.7 mm; BWL= 2.2 mm; AL= 1.4 mm; SW= 1.8 mm; AW= 0.9 mm. MNRJ 33556†, 6 whorls, SL= 5.1 mm; BWL= 3.0 mm; AL= 1.9 mm; SW= 2.3 mm; AW= 1.3 mm. Remarks. The shell of NHMUK 1887.2.9.1618 is the figured specimen by Watson (1886: plate 37, figure 6) and is here selected as the lectotype (Figure 25A). There are paralectotypes in the NHMUK (Figure 25 B–D) and possibly in the NMW (Figure 25E). Watson (1883) classified this species in Eulima, but it clearly differs from Eulima sensu stricto by its conical shape with a subtle increase in diameter, well-demarcated sutures and by the wide, obtuse apex. The present species was classified in Melanella by some authors (e.g., Lange-de-Morretes 1949; Rios 1994). The well-demarcated suture and the outline of the outer lip of “ E. hebes ” are not similar to other species of Melanella despite the great conchological variation considered in the latter genus (Bouchet & Warén 1986). MolluscaBase Eds. (2020f) classified the species in Zebina H. Adams & A. Adams, 1854 (Zebinidae Coan, 1964), but treated the species as a taxon inquirendum, mentioning that “the description strongly suggests that it belongs neither to Eulima, nor to Zebina ”. MolluscaBase Eds. (2020f) argued that based on features described by Watson (1883), as “unequal, close-set, hair-like microscopic lines of growth and faint broader spirals”, and especially the faint broader spirals indicate that it is not a Zebina, but possibly a species of Caelatura s.l. (Barleeiidae Gray, 1857). However, there is no publication reviewing the taxonomy of “ E. hebes ”. Based only on conchological features the systematic position of this species remains unclear, but in fact the species resembles some members of Rissooidea. No soft parts are available to aid and elucidate the systematic position of “ E. hebes ” and a final decision is postponed, but we figured as many details as possible of the shell morphology to help in further investigations. We keep the original classification of the species here, but we draw attention to the uncertainties of the classification of this species. Geographical distribution. Brazil: Pernambuco. Bathymetric distribution. From 465 m to 690 m., Published as part of Souza, Leonardo Santos De, Pimenta, Alexandre Dias & Barros, José Carlos Nascimento De, 2021, Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil, pp. 451-504 in Zootaxa 4927 (4) on pages 495-497, DOI: 10.11646/zootaxa.4927.4.1, http://zenodo.org/record/4542923, {"references":["Watson, R. B. (1883) Mollusca of H. M. S. \" Challenger \" Expedition. Part XVII. Zoological Journal of the Linnean Society, 17 (99), 112 - 130. https: // doi. org / 10.1111 / j. 1096 - 3642.1883. tb 02043. x","Watson, R. B. (1886) Report on the Scaphopoda and Gasteropoda collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger, Zoology, 15 (2), 1 - 680 + 692 - 756, 50 pls.","Lange-de-Morretes, F. (1949) Ensaio de catalogo dos moluscos do Brasil. Arquivos do Museu Paraense, 7, 1 - 216.","Rios, E. C. (1994) Seashells of Brazil. 2 nd Edition. Editora da Fundac \" o Universidade do Rio Grande, Rio Grande, 368 pp., 113 pls.","Barros, J. C. N., Santos, F. N., Santos, M. C. F., Cabral, E. & Acioli, F. D. (2001) Redescoberta de moluscos obtidos durante a \" Challenger Expedition \" (1873 - 1876): micromoluscos de aguas profundas. Boletim Tecnico Cientifico do CEPENE, 9 (1), 9 - 24.","Rios, E. C. (2009) Compendium of Brazilian seashells. Evangraf, Rio Grande, viii + 668 pp.","Rios, E. C. (1975) Brazilian marine mollusks iconography. Museu Oceanografico do Rio Grande, Rio Grande, 331 pp., 91 pls.","Bouchet, P. & Waren, A. (1986) Revision of the northeast Atlantic bathyal and abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762"]}