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Cabras, Analyn A., Lam, Athena W., Van Dam, Matthew H. (2021): Metapocyrtus um sp. nov., a new weevil species (Coleoptera, Curculionidae, Entiminae, Pachyrhynchini) from Davao City, Mindanao Island, Philippines. Zootaxa 5068 (4): 597-600, DOI: https://doi.org/10.11646/zootaxa.5068.4.10

The genusMetapocyrtusHeller, 1912 is the most speciose and complex amongst the tribe Pachyrhynchini with seven subgenera and more than 200 described species. The genus is endemic to the Philippines and remains largely unknown particularly in the less explored areas or mountains.Four new species ofMetapocyrtusHeller, 1912 (Coleoptera, Curculionidae, Entiminae, Pachyrhynchini), are described from Mindanao Island, Philippines. Brief bionomical notes and mimicry with their sympatric beetles and other insect counterparts are also reported.

Undarobius gen. n., a new genus of cavernicolous weevils with two new species, U. howarthi sp. n. and U. irvini sp. n., is described from the Undara Lava Cave system in north-eastern Queensland, Australia. These are the first cavernicolous weevils to be described from Australia, and U. howarthi is a new addition to the rich arthropod fauna of Bayliss Cave. Undarobius weevils are relatively large in size (4.0–5.5 mm long), anophthalmic and apterous with a robust, flattened body and long legs. The genus has affinities with Leptopiini, but its placement in the tribe is uncertain. We also provide a list of the known anophthalmic and microphthalmic weevils in Australia, spanning 65 species classified in 20 genera, eight tribes and about seven subfamilies and found in diverse hypogean habitats, mainly leaf litter but also soil, beach sand, subterranean aquifers and mosses.

Abstract Lectotypes of Cneorhinus viridimetallicus Motschulsky, 1860, C. cuprescens Motschulsky, 1866, C. nodosus Motschulsky, 1860, and Dermatodes carinulatus Motschulsky, 1866 are designated. A new synonymy Catapionus viridimetallicus (Motschulsky, 1860) (= Cneorhinus cuprescens Motschulsky, 1866, syn. n.) and a new combination Scepticus carinulatus (Motschulsky, 1866), comb. n. are established. New data on the morphology of some genera of the tribes Cneorhinini and Tanymecini are presented. The record of Amystax fasciatus Roelofs, 1873 for Kunashir Island is shown to be based on misidentification of Scepticus carinulatus.

Two new species of Metapocyrtus Heller, 1912, subgenus Orthocyrtus Heller, 1912 ( Coleoptera , Curculionidae , Entiminae , Pachyrhynchini ) are described and illustrated from Mindanao Island, Philippines. The species are Metapocyrtus ( Orthocyrtus ) davaoensis sp. nov. and Metapocyrtus ( Orthocyrtus ) hirakui sp. nov. from Davao City and Bukidnon, respectively. Brief bionomical notes and phenotypic characters compared to their sympatric Entiminae counterparts are also reported. The discovery of M. ( O. ) davaoensis sp. nov. in Davao City confirms how understudied Coleoptera are in Mindanao and underlines the potential for the discovery of new species even in highly urbanized areas.

Based on a phylogenetic analysis of a large number of mainly undescribed edaphic Entiminae from South Africa, a new tribe of entimine weevils is described, which includes six new genera. Taxa included in Namaini trib. nov. are clustered into seven clades that are used to delimit the following genera: Nama, type genus of the tribe, plus the new genera Cederbergia gen. nov., Cervellaea gen. nov., Namaquania gen. nov., Pentamerica gen. nov., Springbokia gen. nov. and Yamalaka gen. nov. A key to the genera is given and four new species are described.

Lectotype of the southern Siberian Otiorhynchus (Pseudocryphiphorus) pullus Gyllenhal is designated and a closely related new species Otiorhynchus (Pseudocryphiphorus) matisi sp. n. from the left bank of the Irtysh River in Northeastern Kazakhstan or, less probably, in Omsk Province of Russia, is described.

Broad-nosed weevils in the subfamily Entiminae (Coleoptera: Curculionidae) are highly diverse, not only in terms of number of species, but also in their sizes, forms and colours. There are eight tribes, 50 genera and 224 entimine species recorded from Colombia: seven genera and 142 species are considered endemic and only a handful of species, which are recognised as pests ofCitrusor potatoes, are broadly known. The large diversity of this subfamily in the country is only superficially known and even though genus level identifications are generally achievable, species identification remains quite challenging, due in part to limited access to broadly-scattered basic information. Summaries of available information and bibliographic resources for each of the tribes represented in Colombia are offered, along with a checklist of the species of Entiminae recorded from the country, obtained from literature and a pictorial key for tribal recognition. New combinations are proposed for eight species of the genusLanteriusAlonso-Zarazaga & Lyal. Information on the distribution of entimine species in Colombia is compiled for the first time, including complete references to each original description and available taxonomic revisions. About a third of the species of Entiminae remain as recorded from the country without specific locality information. In addition, genus level distributional maps are presented, generated from data obtained from four Colombian entomological collections. Lastly, some challenges for entimine identification in Colombia, which likely extend throughout the Neotropical region, are briefly discussed. This contribution aims, in part, to facilitate and promote entimine research in northern South America.

A new species, Otiorhynchus (Stupamacus) yakovlevi Legalov, sp. nov. from Western Siberia is described and illustrated. This new species is most similar to O. scintillus but differs from it in a slender body, sparsely punctation of pronotum, less rounded, and finer punctate elytral interstriae. A map with localities of newly described species is added. A key for identifying species from the subgenus Stupamacus from Altai is also included.

Brief morphological characteristics of the genera Blosyrus Schoenherr, 1823, Blosyrodes Jekel, 1875, and Blosyridius Fairmaire, 1891 are given. The new genus Stiltoblosyrus gen. n. (type species S. belousovi sp. n.) with 17 new species is described from Yunnan and Sichuan provinces of China. Stiltoblosyrus gen. n. is closely related to Blosyrodes but differs in the tuberculate pronotum and elytra, in the presence of an elongate process of the apex of the male hind tibia, in the armament of the endophallus consisting of tiny spicules, and in the usually hair-like vestiture of the body. The new genus includes S. andreii sp. n., S. asyae sp. n., S. belousovi sp. n. (type species), S. brevicornis sp. n., S. curvimanus sp. n., S. delavayi sp. n., S. fairmairei sp. n., S. gracilis sp. n., S. laticollis sp. n., S. kabaki sp. n., S. khalidi sp. n., S. lisu sp. n., S. longipes sp. n., S. pulcher sp. n., S. titanicus sp. n., S. vulgaris sp. n., and S. zhangi sp. n. A key to the 17 species is given. Most of the species were collected from Rhododendron at night time.

Cydianerus latruncularius (Perty, 1832) is a broad-nosed weevil (Entiminae) with South American distribution. In the past few years, some cases were observed where morphological modifications can occur on different populations of widely distributed species in various animal groups, including insects. Currently, geometric morphometrics is one of the most reliable and substantial ways to analyze shape variation among organisms. This technique is based on analysis of landmarks and their deviation on different individuals, identifying or not, conspicuous changes among them. It is a very useful tool to test shape variation among groups of organisms under different environmental pressures, whatever they might be. Knowing about the wide distribution and environmental diversity (Rainforests, Cerrado, Caatinga) occupied by C. latruncularius, the following research aims to discover if there are any shape patterns among sexes and individuals from different Brazilian ecosystems. Images of males and females’ pronotum, elytra and ventrite were landmarked in order to obtain morphological data which was then analyzed through MorphoJ software. Variations were found in all structures, which presented different results throughout the analysis. Pronotum proved to be a determinant structure to separate Amazonian populations from the others. The elytra formed groups only among males from Cerrado and Rainforests, while all females were overlapping. Also, the ventrites proved to be the best structures to describe sexual dimorphism. Geometric morphometrics of C. latruncularius allowed us to verify that the shape of different body parts varies in an independent way among them and are possibly influenced by the environment.

Rhinodontodes alashanensissp. nov., Trachyphloeosoma honzasp. nov., T. jirkasp. nov., and T. martinsp. nov. are described from China, illustrated and compared with similar species. The genus Rhinodontodes and the species Rhinodontodes subsignatus Voss, 1967 and Rhinodontus mongolicus Borovec, 2003 are recorded from China for the first time. Keys to all Chinese genera of Trachyphloeini, and to the Chinese species of Rhinodontodes and Trachyphloeosoma, are provided.

Lectotype of Alophusmarginatus Faust, 1887 is designated. Trichalophuscaudiculatus Fairmaire, 1886 and T. compressicauda Fairmaire, 1887 are transferred to the genus Pseudalophus Suvorov, 1915, where new combinations are formed: Pseudalophus caudiculatus (Fairmaire, 1886), comb. n., and Pseudalophus compressicauda (Fairmaire, 1887), comb. n. New taxa Pseudalophusangulatussp. n.,Ps. brevirostrissp. n.,Ps. korolevisp. n.,Ps. muliensissp. n.,Ps. muliensis meridionalissubsp. n., and Ps. scylloidessp. n. are described from the Sino-Tibetan (Hengduan) Mountains in China. A species closely related to Potentillapeduncularis David Don, 1825 is reported as a host plant of Ps. sp. pr. caudiculatus.

A new species Stiltoblosyrus serratulus Davidian, sp. nov. is described from Sichuan Province of China. It is the northernmost species of the genus Stiltoblosyrus (Davidian, 2020). From all of the congeners it easily differs in row of large and stout spines at the inner side of hind tibia of the male. New species closely related to S. laticollis Davidian, 2020 and S. brevicornis Davidian, 2020 in size of the process at the inner apical angle of hind tibia of the male, which is distinctly shorter than 2nd segment of hind tarsus. Colour illustrations of habitus and important morphological characters are presented.

The genusMetapocyrtusHeller, 1912 is the most speciose and taxonomically complex genus in the tribe Pachyrhynchini. It is known to be endemic in the Philippines, with most species having a very narrow range of distribution. There are already more than 230 species ofMetapocyrtusdocumented in the Philippines.Metapocyrtus dagtumsp. nov., a new species of the genusMetapocyrtusHeller, 1912 from Davao de Oro, Mindanao Island, Philippines, is described with brief notes about its ecology. Its specific epithet is from the Cebuano word “dagtum” which means pitch black referring to the colour of the integument of the species.

A new genus Rovnoslonik Legalov, Nazarenko and Perkovsky gen. nov. (type species Rovnoslonik damzeni Legalov, Nazarenko and Perkovsky sp. nov.; tribe Stromboscerini, subfamily Dryophthorinae), new species Caulophilus martynovae Legalov, Nazarenko and Perkovsky sp. nov. (Cossoninae: Dryotribini), Dorytomus vlaskini Legalov, Nazarenko and Perkovsky sp. nov. (Curculioninae: Ellescini: Dorytomina), Paonaupactus katyae Legalov, Nazarenko and Perkovsky sp. nov., and Paonaupactus gracilis Legalov, Nazarenko and Perkovsky sp. nov. (Entiminae: Anypotactini) are described from the Late Eocene Rovno amber. These are the first records of the genera Dorytomus Germar, 1817 and Paonaupactus Voss, 1953 from Rovno amber.

Several new tribes, genera and species of Curculionoidea are described from the earlymost Eocene Oise amber (France), viz., the new tribes Oiserhinini trib. nov. (type genus: Oiserhinus gen. nov.) (in Anthribidae: Anthribinae), Palaeotanaini trib. nov. (type genus: Palaeotanaos Kirejtshuk, Legalov et Nel, 2015) (in Brentidae: Apioninae), Sciabregmini trib. nov. (type genus: Sciabregma Scudder 1893) (in Curculionidae: Molytinae); the new genera Oiserhinus gen. nov. (type species: Oiserhinusinsolitus sp. nov.), Oisecalles gen. nov. (type species: Oisecalles latosquamosus sp. nov.) in Cryptorhynchini: Cryptorhynchina), Rhinoporkus gen. nov. (type species: Rhinoporkus gratiosus sp. nov.) (in Dryophthorinae: Dryophthorini), Jantarhinus gen. nov. (type species: Jantarhinus compressus sp. nov.) (in Conoderinae: Conoderini), Jantaronosik gen. nov. (type species: Jantaronosik nebulosus sp. nov.) (in Curculioninae: Acalyptini), Tolstonosik gen. nov. (type species: Tolstonosikoisensis sp. nov.) (in Entiminae: Eudiagogini); and the new species Oiserhinusinsolitus sp. nov., Sciabregmasquamosa sp. nov., Oisecalles latosquamosus sp. nov., Rhinoporkus gratiosus sp. nov., Jantarhinus compressus sp. nov., Jantaronosik nebulosus sp. nov., and Tolstonosikoisensis sp. nov. The generic name Protoceletes Rheinheimer, 2007, syn. nov. is synonymized under Electrotribus Hustache, 1942. New combination of name (Electrotribuswolfschwenningerae (Rheinheimer, 2007), comb. nov. is established. Synopsis of the weevil findings from described groups are presented.

Four new weevil species, Mesocordylusdominicus sp. nov. (Dryophthorinae: Orthognathini), Derelomus miocaenicus sp. nov. (Curculioninae: Acalyptini), Anthonomus (Anthonomus) clarki sp. nov. (Curculioninae: Anthonomini) and Compsus (Compsus) poinari sp. nov. (Entiminae: Eustylini) are described from Early Miocene Dominican amber.

El gorgojo de la raíz de la alfalfa, Sitona discoideus Gyllenhal 1834 (Curculionidae: Entiminae), es reportado por primera vez en la Argentina. Este gorgojo es nativo del sur de Europa y noreste de África, y fue introducido accidentalmente en Australia, Nueva Zelanda, Estados Unidos y Chile. En Argentina fue colectado en el Alto valle de Río Negro, asociado con plantas de alfalfa (Medicago sativa L.). The alfalfa root weevil Sitona discoideus Gyllenhal 1834 (Curculionidae:Entiminae) is herein reported for the first time for Argentina. This weevil is native to southern Europe and northern Africa and was accidentally introduced in Australia, New Zealand, the United States of America, and Chile. In Argentina, it was collected in the Upper valley of Río Negro, associated with alfalfa plants (Medicago sativa L.). Fil: del Rio, Maria Guadalupe. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. División Entomología; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - La Plata; Argentina Fil: Lanteri, Analia Alicia. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. División Entomología; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - La Plata; Argentina Fil: Gittins López, Cecilia Gabriela. Instituto Nacional de Tecnología Agropecuaria. Centro Regional Patagonia Norte. Instituto de Investigación y Desarrollo Tecnológico para la Agricultura Familiar Región Patagonia; Argentina Fil: Rivero, Victoria Ileana. Universidad Nacional del Comahue. Facultad de Ciencias Agrarias; Argentina Fil: González, Diego A.. Universidad Nacional del Comahue. Facultad de Ciencias Agrarias; Argentina Fil: Lopez Armengol, Maria Fernanda. Instituto Nacional de Tecnología Agropecuaria. Centro Regional Patagonia Norte. Instituto de Investigación y Desarrollo Tecnológico para la Agricultura Familiar Región Patagonia; Argentina. Universidad Nacional del Comahue. Facultad de Ciencias Agrarias; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Patagonia Confluencia; Argentina

Species of the genus Pseudocneorhinus occurring in or near China are reviewed, with description of five new species, Pseudocneorhinusangustussp. nov., P.glabersp. nov., P.hlavacisp. nov., P.obliquehumeralissp. nov., and P.setosicallussp. nov. from the provinces of Beijing, Gansu, Shaanxi, Sichuan, and Zhejiang. They are illustrated and compared with similar species, and a key is provided to all presently known species of the genus. Lectotypes of the following species are designated: Callirhopalussubcallosus Voss, 1956 [current name Pseudocneorhinussubcallosus (Voss, 1956)] and P.squamosus Marshall, 1934. Pseudocneorhinussquameus Morimoto, 2015 is confirmed for the fauna of China.

A new genus, Coniophloeus gen. n., is described for five species of South African weevils known from the Western and Eastern Cape and assigned to the tribe Embrithini Marshall, 1942: Coniophloeus squalidus (Boheman, 1842) comb. n. (type species, transferred from the genus Trachyphloeus Germar, 1817 and tribe Trachyphloeini Lacordaire, 1863), C. alternans sp. n., C. oberprieleri sp. n., C. robustus sp. n. and C. obrieni sp. n. All five species are illustrated and keyed.

A new weevil, Arostropsis perkovskyi Bukejs and Legalov, sp. nov. (urn:lsid:zoobank.org:act:408ECE71-B191-4381-AA39-C27A3A1DA082; Coleoptera: Curculionidae: Entiminae), is described from Late Eocene Rovno amber. The new species differs from Arostropsis groehni Yunakov et Kirejtshuk, 2011 from Baltic amber in the pronotum lacking a distinct lateral ridge, it has a shorter rostrum, antennomere 2 is subequal to antennomere 3, it has wider elytra with weaker convex humeral calli and the pronotum is wider than it is long. It is the first record of the tribe Naupactini in Rovno amber and the second finding of Arostropsis in Eocene ambers.

The North American species of the broad-nosed weevil genus Pachyrhinus Schönherr 1823 (Coleoptera: Curculionidae, Entiminae) are revised. Three species of Pachyrhinus are here recognized in North America: P. elegans (Couper 1865), P. californicus (Horn 1876), and P. cinereus (Casey 1888). Pachyrhinus lateralis (Casey 1888) and P. miscix (Fall 1901) are here designated as synonyms of P. elegans. Pachyrhinus crassicornis (Casey 1888) and P. albidus (Fall 1901) are here designated synonyms of P. cinereus (Casey 1888) The previously proposed synonymy of P. ferrugineus (Casey 1888) with P. californicus was confirmed. This revision includes detailed images of diagnostic characters as well as scanning electron micrographs of scale morphology for all species. A key to the Nearctic species of Pachyrhinus is provided. All Nearctic species of Pachyrhinus are considered minor pests of Pinus spp. [Pinaceae].

International audience; Polyphagous broad nosed weevils (Curculionidae: Entiminae) constitute a large and taxonomically challenging subfamily that contains economically significant agricultural pests worldwide. South Africa is a hot-spot for biodiversity and several species of indigenous and endemic genera of Entiminae have shifted on to cultivated plants, with some being phytosanitary pests. The sporadic pest status of many species (where the species has an occasional economic impact on the agricultural industry, but is not encountered often enough that is is readily recognisable by researchers and agricultural extension workers) and the presence of pest complexes and cryptic species represent an identification challenge to non-specialists. Furthermore, no comprehensive identification tools exist to identify immature stages that may be found in crops/soil. In this paper, a curated barcoding database with 70 COI sequences from 41 species (39 Entiminae, 2 Cyclominae) is initiated, to assist with the complexity of identification of species in this group.

Two new species of Curculionidae, Pachyrhynchini, Metapocyrtus waoensis sp. nov., and M. lanaoensis sp. nov., both from western Mindanao are described. Images of habitus and fully everted endophallus and illustrations of male genitalia are provided for each species.

Saotomia tuberculata, a new genus and species of Peritelini (Curculionidae; Entiminae) from São Tomé, West Africa is described. The new genus is most similar to the other African genera Dysommatus Marshal, 1933 and Fernandius Marshall, 1954. Important taxonomic characters are illustrated.

A new genus of Entiminae, an endogean weevil of the tribe Otiorhynchini, Giavarhynchus Bellò, Osella & Ruzzier, gen. n., and its type species Giavarhynchus amicorum Bellò, Osella & Ruzzier, sp. n. are described. The new taxon is readily distinguished from all other members of the tribe due to the unique combination of lack of eyes, elongate rostrum with a ventral transverse furrow and excised lateral margins located at apical third, punctation of pronotum of two distinct sizes arranged in a distinctive pattern, interval 7 of elytra protruding from base of pronotum and crenulate basally, metafemora bearing a spine-like tooth much larger than that of pro- and mesofemora, female tibiae granulate on inner margin, bisinuous mesotibiae. The following synonymy is proposed: Nematocerus Reiche, 1849 (= Cyrtozemia Pascoe, 1872, syn. n.; = Holcorhinosoma Voss, 1939, syn. n.). New combinations are: Nematocerus cognatus (Marshall, 1916), comb. n.; Nematocerus dispar (Pascoe, 1872), comb. n.; Nematocerus pilipes (Morimoto, 2015), comb. n., all from Cyrtozemia; Nematocerus subtuberculatus (Voss, 1939), comb. n. from Holcorhinosoma. New tribal placement is: Pseudocratopus Hustache, 1921 from Otiorhynchini to Peritelini. New subgeneric placement is that of Otiorhynchus deceptorius Białooki, Germann & Pelletier, 2017 and of Otiorhynchus incisirostris Białooki, Germann & Pelletier, 2017 from Otiorhynchus (Lixorrhynchus) Reitter, 1914 to Otiorhynchus (Aranihus) Reitter, 1912.

The genus Rhyncogonus Sharp, 1885, is represented in French Polynesia by 65 species found in the Austral, Marquesas, Society and Tuamotu archipelagos. Hitherto unknown from the fifth archipelago of French Polynesia, the Gambier Islands, a new species of Rhyncogonus has been discovered by the botanist Jean-François Butaud on Motu Teiku, a small island never previously investigated for plants and insects. Rhyncogonus duhameli sp. nov. is here described and illustrated.

Two new species belonging to the genus Pachyrhynchus Germar, 1824 (Curculionidae: Entiminae: Pachyrhynchini Schoenherr, 1826) from Northern Mindanao are described: Pachyrhynchus yoshitakei sp. nov. and Pachyrhynchus imitans sp. nov. The endophallus of Pachyrhynchus zamboanganus Yoshitake, 2012 is illustrated as well.

Four new Tapinomorphus species are described and illustrated: Tapinomorphus angolanus sp. nov. from Angola, T. kudrnai sp. nov. from Zambia, T. latipennis sp. nov. and T. verunkae sp. nov. from Zimbabwe. This brings the total number of species in the genus to 20. Habitus images and illustrations of important characters are provided.

Pantomorus albosignatus Boheman, 1840 (Entiminae: Naupactini), type species of the genus Pantomorus Schoenherr, 1840, is broadly distributed in Mexico, from Oaxaca and Veracruz to Chihuahua and Coahuila, and is probably related to P. parvulus Sharp 1891 (México: Oaxaca and Veracruz), P. andersoni sp. nov. (México: Guerrero), and P. crinitus (Boheman, 1840) (southern Mexico, El Salvador and Guatemala). The four species are small (5-8 mm), apterous, usually show a characteristic maculation and erect setae on the elytra, short antennae, vestigial to absent humeri, and well-developed, squamose corbels of the metatibiae, and are here referred to as the Pantomorus albosignatus species group. Our study was based on the examination of type material and about 500 specimens from different collections, and provides a dichotomous key, descriptions or redescriptions of the species, habitus photographs, line drawings of female and male genitalia, a map of distribution, new locality records and new plant associations. We propose that Pantomorus nobilis (Boheman 1840) is a new junior subjective synonym of P. crinitus (Boheman, 1840), and we designate lectotypes for the species Pantomorus albosignatus Boheman, P. parvulus Sharp, P. crinitus (Boheman), P. nobilis (Boheman) and P. affinis Sharp, 1891 (the latter is also a junior synonym of P. crinitus). Pantomorus albosignatus usually inhabits Acacia grasslands and cactus deserts of the Mexican Plateau, at higher elevations than remaining species; P. parvulus occurs in cloud forests and tropical deciduous forests of southern Mexico; P. crinitus in open oak pine forests, cloud forests and tropical deciduous forests of southern Mexico, Guatemala and El Salvador; and P. andersoni is endemic to the state of Guerrero, in Acacia-cactus woodlands.

The golden-headed weevil Compsus auricephalus is a native and fairly widespread species across the southern U.S.A. extending through Central America south to Panama. There are two recognised morphotypes of the species: the typical green form, with pink to cupreous head and part of the legs and the uniformly white to pale brown form. There are other Central and South American species of Compsus and related genera of similar appearance that make it challenging to provide accurate identifications of introduced species at ports of entry. Here, we re-describe the species, provide images of the habitus, miscellaneous morphological structures and male and female genitalia. We discuss the morphological variation of Compsus auricephalus across its distributional range, by revising and updating its distributional range, based on data from entomological collections in the U.S.A. and Canada. The revised distribution of C. auricephalus extends as far south as Zacapa in Guatemala. Records south from there correspond to a different species, with affinities to C. auricephalus that we discuss and illustrate. We also discuss morphological affinities and differences with other similar species. Furthermore, we summarise information regarding the biology, host plants and natural enemies of C. auricephalus.

A new species of Naupactini (Curculionidae: Entiminae) endemic to Brazil, Espírito Santo and Minas Gerais states, is herein described. It resembles the monotypic genus Hadropus Schoenherr in its general appearance, particularly in the shape of the elytra, and the color of the vestiture, but the results of a cladistics analysis herein conducted suggest that it belongs to the genus Stenocyphus Marshall. This genus ranges in São Paulo, Rio de Janeiro and Espírito Santo, mainly in the Atlantic forest of Brazil, and includes three other species. Stenocyphus costae sp. nov., distinguishes from the remaining species of Stenocyphus by the green iridescent scaly vestiture, having long stiff setae on the two pairs of elytral tubercles; the more slender rostrum; the shorter antennae; the convex disc of the pronotum; and the shorter and broader elytra. This paper includes a cladogram of the Naupactini genera showing the phylogenetic position of the new species, its complete description, photographs of male and female habitus, line drawings of genitalia of both sexes, and a key of the Stenocyphus species., Facultad de Ciencias Naturales y Museo

Polydrusus (s. str.) akreanus sp. n. is a new record of Curculionidae found in Iraqi Kurdistan. The new species is described, illustrated and compared with Polydrusus (s. str.) kadleci Borovec & Ge...

International audience; Phlyctinus callosus (Schoenherr, 1826) (Curculionidae, Entiminae, Oosomini) is a species native to the Western Cape province of the Republic of South Africa. It is regarded as a key pest of orchards and vineyards. Detailed observations of its morphology and ecology have shown this species to be a complex of closely related species. In this study we describe the species of this complex, provide morphological and molecular diagnostic tools to distinguish between species, and review their distribution and host plant associations. In all, six species are recognized in the P. callosus complex and five are described as new (P. grootbosensis Haran sp. nov., P. xerophilus Haran sp. nov., P. planithorax Haran sp. nov., P. littoralis Haran sp. nov., P. aloevorus Haran sp. nov.). Sequences of the mitochondrial gene used (cytochrome oxidase I, COI) indicated that P. xerophilus Haran sp. nov. might comprise several cryptic species for which stable morphological features could not be identified at this stage. The species of the genus Phlyctinus appear to be primarily associated with various Asteraceae found in humid, sandy and disturbed habitats.

Two new species of the subgenus Pseudotiorhynchus Magnano, 2004 of the genus Otiorhynchus Germar, 1822 are described from Iran: O. demavendussp. n. from Mount Demavend and O. binaludussp. n. from the Binalud Range.

A new genus, Afromuelleria gen. n., assigned to the tribe Trachyphloeini Lacordaire, 1863, is described for four South African species of weevils: A. awelani sp. n., A. baobab sp. n., A. limpopo sp. n. and A. venda sp. n. All species are illustrated and keyed. Taxonomic status of the new genus is discussed and compared with similar genera of Trachyphloeini and Embrithini Marshall, 1942.

Pachyrhinuslethierryi (Desbrochers des Loges, 1875) is a Mediterranean weevil species that has become remarkably well known as a result of a series of recent introductions across Western and Central Europe. This species has recently reached Asia Minor and the Crimean Peninsula, as confirmed by several new records. The vectors of invasion in Crimea and possible further expansion are suggested.

In the summer and autumn of 2017, two mostly Southeastern European weevils Otiorhynchus albidus Stierlin, 1861 and O. asphaltinus Germar, 1824, and also O. rotundus Marseul, 1872, previously unknown from Russia, were found in numbers on Syringa vulgaris L. and S. josikaea Jack. in St. Petersburg. New data on the distribution of O. smreczynskii and a key for differentiation of O. rotundus, O. smreczynskii and closely allied to the latter O. ukrainicus Korotyaev, 1984 from southeastern Ukraine and eastern Crimea are reported.

A complete and updated list of 1,066 species and subspecies of the superfamily Curculionoidea recorded from Poland until the end of 2017 is provided, including over 290 comments on all taxonomic, nomenclatural and faunistic changes made since the first edition of the checklist published in 2005. An additional 37 species in the list are considered as erroneously recorded and excluded from the fauna of Poland. An update of the index of genera and subgenera, together with their type species, is provided separately. The following two new synonyms are recognised: Bruchela rufipes nigritarsis (Reitter, 1916) = B. rufipes rufipes (Olivier, 1790) (Anthribidae: Urodontinae), Otiorhynchus proximus iteratus Magnano, 2001 = O. proximus proximus Stierlin, 1861 (Curculionidae: Entiminae). Six species are resurrected from synonymy with the species given in parentheses: Squamapion serpyllicola (Wencker) (S. minutissimum (Rosenhauer)), Polydrusus thalassinus Gyllenhal (P. formosus (Mayer)), Phyllobius vespertinus (Fabricius) (Ph. pyri (L.)), Microplontus melanostigma (Marsham) (M. rugulosus (Herbst)), Otiorhynchus smreczynskii Cmoluch (O. rotundus Marseul), Romualdius angustisetulus (Hansen) (R. scaber (Linnaeus)). Phyllobius fessus Boheman is replaced in the list with Ph. jacobsoni Smirnov due to the former misidentification of Polish specimens. Brachysomus strawinskii Cmoluch is transferred from subgenus Hippomias to Brachysomus s. str. Eleven new weevil species have been discovered in Poland since 2005, including the still unpublished records of Pachycerus madidus (Olivier). The occurrence in Poland of four species, having hitherto only uncertain old records, has been confirmed with recent findings. The number of dubious species in Poland increased to 27 after the inflow of uncertain data in the literature and probable cases of extinction.

A new genus, Lyalinus gen. nov. (Coleoptera: Curculionidae: Entiminae: Cneorhinini) with one new species, Lyalinus bimaculatus sp. nov. from Burkina Faso, is described, illustrated and compared with other related genera.

Nine species of the genus Blosyrus (Schoenherr, 1826) (Curculionidae: Entiminae) occurring in India, Sri Lanka and Myanmar are revised. The earlier known seven species are redescribed, whil...

The subgenera Eunihoides subgen. n. (type species O. echinopterus sp. n.), and Nubidanops subgen. n. (type species Otiorhynchus tumidicollis Stierlin, 1861) in the genus Otiorhynchus Germar, 1822 are newly described. The following species of Otiorhynchus Germar, 1822 are described: echinopterus sp. n., rhizophilus sp. n. and agilis sp. n. from south-western Turkey (subgenus Eunihoides subgen. n.); brevipilosus sp. n. from Turkey (subgenus Pocodalemes); nigrosetosus sp. n., flavosetosus sp. n., lacunosus sp. n., adopertus sp. n., implicatus sp. n. and sculptipterus sp. n. from Turkey (subgenus Tecutinus Reitter, 1912); forficulatus sp. n. and amplicornis sp. n. from south-western Turkey; crocotillus sp. n. from Greece (subgenus Fondajenus Reitter, 1912); crassiphallicus sp. n. from northern Turkey (subgenus Otismotilus Reitter, 1912); agmentosus sp. n., armentalis sp. n., cappadocicus sp. n., crepuscularis sp. n., hebetes sp. n., malinkai sp. n., porculus sp. n., vavrai sp. n. and wanati sp. n. (all from Turkey, subgenus Odelengus); spectativus sp. n. from Turkey (subgenus Aequipennis); anabolicus sp. n. and pulcher Białooki & Fremuth sp. n. from Turkey (subgenus Pterygodontus); rhinorostris sp. n. and curiosus sp. n. from Greece (subgenus Podonebistus); microcornis sp. n. from Turkey (subgenus Arnoldinus Davidian, Gültekin & Korotyaev, 2017). Otiorhynchus (Eunihus) pisidicus (Magnano, 2003) is hereby transferred to Otiorhynchus (Eunihoides subgen. n.), Otiorhynchus tumidicollis Stierlin, 1861 from Otiorhynchus incertae sedis and Otiorhynchus (Nubidanus) torosicus Davidian & Gültekin, 2016 to the subgenus O. (Nubidanops subgen. n.); all are new subgeneric placements.

Austromonticola gen. n. is proposed for a group of eight New Zealand alpine broad-nosed weevil species, all of which are here described: A. atriarius sp. n. (type locality: Umbrella Mountains, Central Otago), A. caelibatus sp. n. (type locality: Ohau Range, Mackenzie), A. furcatus sp. n. (type locality: Old Man Range, Central Otago), A. inflatus sp. n. (type locality: Hawkdun Range, Central Otago), A. planulatus sp. n. (type locality: St Marys Range, Central Otago), A. postinventus sp. n. (type locality: Kirkliston Range, South Canterbury), A. mataura sp. n. (type locality: Mt Dick, Otago Lakes) and A. rotundus sp. n. (type locality: Old Man Range, Central Otago). All species occur exclusively above 1000 m elevation in the mountains of Central Otago and South Canterbury in the South Island. A phylogeny of the genus, including six outgroups, was inferred from 33 morphological characters. It resolved the genus as monophyletic, and revealed two strongly supported clades within Austromonticola. DNA sequences of four gene regions were obtained from five species. Of these, the 3' end of COI proved to be the most suitable for the identification of specimens. Females of all species have diagnostic secondary sexual structures on the elytra and ventrites. These structures are hypothesised to have evolved to assist with oviposition in and beside cushion plants or by selection for structures to mitigate the costs to females of prolonged mating.

Morphological description of the pistachio weevil, Polydrusus davatchii Hoffmann, 1956, is supplemented and an additional distinctive character of the subgenus Orodrusus to which this species belongs, the strongly reduced hair brushes on the tarsal segments, is reported. Polydrusus davatchii was known as a minor pest for pistachio trees in Iran until the first decade of the XXI century, but is considered now as an important herbivore in pistachio orchards in early spring. The adults were not found on other plants in the pistachio growing areas. Only females of this species are known. Polydrusus davatchii produces one generation a year and is distributed now almost throughout the pistachio cultivation areas in central and southern parts of Iran.

The genus Apodrosus Marshall is newly recorded for, and revised for Cuba. Nine new species are recognized as follows: Apodrosus alberti (type locality, Granma, Parque Nacional Pico Turquino), A. alternatus (type locality, Guantánamo, El Yunque), A. franklyni (type locality, Cienfuegos, Parque Nacional Pico San Juan), A. griseus (type locality, Santiago de Cuba, Siboney-Jutici Ecological Reserve), A. mensurensis (type locality, Holguin, Parque Nacional La Mensura-Piloto), A. pseudoalternatus (type locality, Matanzas, Varahicacos), A. beckeli (type locality, Guantánamo, 8 km W. Imias), A. sandersoni (type locality, Guantánamo, Loma Lafarola), and A. zayasi (type locality, Cienfuegos, Parque Nacional Pico San Juan). A key for their identification, descriptions, summaries of natural history information and data on distributions are presented. A molecular phylogeny based on 11 species of Apodrosus from Cuba, Hispaniola and Puerto Rico is reconstructed. A sister group relationship between Polydrusus and Apodrosus is recovered with a limited sampling of the former genus. The monophyly of Apodrosus is recovered with strong support. Cuban Apodrosus are not monophyletic. Five of the six sampled Cuban species form a clade, sister to an undescribed Apodrosus species from the Dominican Republic; and, Apodrosus alternatus is sister to A. quisqueyanus Girón & Franz, 2010, a species from the Dominican Republic. Biogeographic implications for Cuban species are discussed in light of the phylogeny.

An account is given of the twelve species of Leptomias Faust, 1886 occurring in the Sichuan Province of China, including the description of a new species, Leptomias verticalis Ren, Zhang & Song, sp. n. from Jiulong County, Southwest Sichuan. New locality data and remarks for the other eleven species, a key to and distribution map of all twelve Sichuan species are provided. Leptomias chenae Alonso-Zarazaga & Ren is transferred to Geotragus Schoenherr, 1845, where its valid name is G. granulatus (Chao, 1980), comb. n. in application of Art. 59.4. Structural details of Leptomias verticalis and Geotragus granulatus are illustrated.

Statistical historical biogeographical methods rely on the use of models that assume various biogeographic processes. Until recently model selection remains an explored topic and the impacts of using different models on inferring biogeographic history are poorly understood. Focusing on the Neotropical weevils in theExophthalmusgenus complex (Insecta: Curculionidae: Entiminae), we compare three commonly used biogeographic models – DIVA (Dispersal-Vicariance Analysis), DEC (Dispersal-Extinction-Cladogenesis) and BayArea (Bayesian Analysis of Biogeography), and examine the impact of modeling founder-event jump dispersal on biogeographic history estimation. We also investigate the biogeographic events that have shaped patterns of distributions, diversification, and endemism in this group of weevils. We sample representatives of 65 species of theExophthalmusgenus complex and 26 outgroup terminals from the Neotropics including Caribbean islands and mainland. We reconstruct a molecular phylogeny based on six genes and performed molecular dating using a relaxed clock with three fossil calibration points. We conduct biogeographic history estimations and compare alternative biogeographic models with the R package BioGeoBEARS. Model selection strongly favors biogeographic models that include founder-event jump dispersal. Without modeling jump dispersal, estimations based on the three biogeographic models are dramatically different, especially at early diverging nodes. When jump dispersal is modeled, however, the three biogeographic models perform similarly. Accordingly, we show that the Neotropical mainland was colonized by Caribbean species in the early Miocene, and thatin situdiversification accounts for a majority (~75%) of the biogeographic events in theExophthalmusgenus complex. Our study highlights the need for testing for wide-ranging historical biogeographic processes in the study of Caribbean biogeography and the importance of comparing and selecting the best-fitting model in statistical biogeographic inferences. We demonstrate that modeling founder-event jump dispersal significantly improves the fit of the biogeographic history estimation of Caribbean and Neotropical mainland weevils. We establish thatin situdiversification acts as a dominant biogeographic force in the evolution of theExophthalmusgenus complex. The colonization of the Neotropical mainland from Caribbean islands reinforces the notion that islands can be an important source of continental diversity.