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The typification of the name Atriplex mollis, validly published by R. L. Desfontaines in Flora Atlantica, is discussed. A specimen deposited at G (barcode G00386803) is designated as the lectotype of the name. The distribution in Europe of the species is also discussed. It was indicated for Sardinia only (Italy, Southern Europe) in a single locality (San Paolo pond, Cagliari Province). No specimen that proves the occurrence of the species in Sardinia is in extant. Moreover, the overlapping of a map published in 1912 and the current satellite images, allowed to verify that the area in which A. mollis grew up, including wetlands at the beginning of the XX century, is currently almost totally covered by buildings and commercial/industrial sites. In other words, the habitat where the species occurred was destroyed. As a consequence, at the current state of knowledge, the species is here assessed as extinct (EX) at national level according to the IUCN categories.

Amaranthus emarginatus belongs to the Amaranthus blitum complex, a critical taxonomic group. Within the variability of A. emarginatus, two taxa are often accepted, ‘emarginatus’ and ‘pseudogracilis’, which differ by habitus, synflorescence structure, and leaf size. However, their taxonomic value is debated, and A. emarginatus itself is sometimes included in A. blitum as subspecies. In this paper, molecular, morphological, phytogeographical, and ecological data are discussed to assess the most correct rank of these taxa. We conclude that A. emarginatus deserves specific distinctness from A. blitum, and A. pseudogracilis can be regarded as a subspecies of the former. This choice reflects both the molecular data and geographical distributions of A. blitum and A. emarginatus, i.e., Mediterranean area, Europe, and North Africa for the former vs. neotropics for the latter). Within A. emarginatus, in addition, to the different morphology, A. emarginatus var. pseudogracilis differs also on account of its ecological preferences, because it occurs in warm and humid areas, whereas A. emarginatus var. emarginatus is better adapted to cold and dry habitats. However, since molecular analyses cannot clearly separate the two taxa, we consider the subspecific rank as the most appropriate. We also found a further morphological difference in seed coat ornamentation, hitherto not pointed out: the shape of the peripheral cells is tetragonal in A. emarginatus subsp. pseudogracilis, but polygonal (cells with ≥ 5 sides) in subsp. emarginatus. Distributional maps and status assessment at regional level for the two taxa are provided. According to this revision, A. emarginatus subsp. emarginatus is to be regared as naturalized in central and southern Italy (Latium and Campania, respectively).

In this contribution, new data concerning the distribution of native vascular flora in Italy are presented. It includes new records, confirmations, and exclusions to the Italian administrative regions. Nomenclatural and distribution updates, published elsewhere, and corrigenda are provided as supplementary material.

The names Arenaria mattfeldii, A. pallens, A. peruviana, A. pintaudii, and A. stuebelii (Caryophyllaceae, Arenarieae) from Peru and Bolivia were studied and neotypified based on specimens preserved at B and P.

The nomenclatural status and typification of the names Arthrocaulon macrostachyum, Salicornia fruticosa, S. fruticosa var. deflexa, S. fruticosa var. glaucescens, S. fruticosa var. intermedia, S. fruticosa var. humilis, S. fruticosa var. pachystachya, S. fruticulosa, S. glauca, S. lignosa, S. macrostachya var. virescens, S. macrostachya var. glaucescens, S. perennis, S. radicans, S. radicans var. caespitosa, S. sarmentosa, S. sempervirens, and S. virginica, as well as an unnamed β-variety of S. fruticosa proposed by A. Bertoloni, are investigated. Concerning A. macrostachyum, we document that the type indicated in literature (G00177362) is not a holotype, and that lectotypification is necessary. A specimen from G (G00687638) is here designated as a lectotype. On the level of variety, Arthrocnemum fruticosum var. macrostachyum is an earlier legitimate name for Salicornia fruticosa var. pachystachya. Furthermore, Piirainen et al. are incorrect when citing Forsskål’s “Salicornia” from Alexandria as “S. virginica Forssk.”; it is not a new name and should be cited as S. virginica auct. non L., as published in Forsskål’s Flora Aegyptiaco-Arabica. Like with numerous other parallel cases in Flora Aegyptiaco-Arabica, Forsskål’s designation of “Salicornia virginica” for an Arabian plant is to be considered a misapplication of the earlier Linnaean name for an American plant. Arthrocnemum glaucum (a nomen illegitimum of Ungern-Sternberg), was listed as type species of Arthrocnemum by the Names in Current Use project; the basionym, Salicornia glauca Delile, is here lectotypified and identified as Arthrocaulon meridionale, published by Ramirez et al. Updated synonymies of Arthrocaulon macrostachyum, A. meridionale, Salicornia fruticosa, and S. perennis are proposed. Salicornia sempervirens is an invalid name according to Art. 36.1a of ICN. No original material was found for S. radicans var. caespitosa. This paper also refer to lecto- or neotypifications on specimens deposited at BM, G, LINN-HS, LY, MPU, NAP, and PAL, and their current taxonomic positions are suggested in a taxonomic part of the paper.

In this contribution, new data concerning the distribution of vascular flora alien to Italy are presented. It includes new records, confirmations, and status changes for Italy or for Italian administrative regions. Nomenclatural and distribution updates, published elsewhere, and corrections are provided as Suppl. material 1.

Como parte de los estudios en curso del género Polycarpon (Caryophyllaceae) y la preparación de su tratamiento para la Flora Argentina, presentamos aquí una nota taxonómica sobre la identidad del nombre P. apurense. Un espécimen preservado en P es designado como el lectotipo de este nombre. Se discute su identidad y se descarta la sinonimia con P. prostratum, recurrente en la literatura, sobre la base de algunos caracteres morfológicos como el hábito, forma y tamaño de las hojas. Se propone un cambio nomenclatural (P. tetraphyllum subsp. apurense), sobre la base del concepto actual en el género Polycarpon. Sobre la distribución de este taxon, se excluye de las floras de Brasil y Uruguay y, se considera un registro dudoso para Perú.

A contribution to the nomenclature of the genus Sagina is presented. The following 10 taxa are recognized as being part of the Italian flora: S. alexandrae, S. apetala, S. glabra, S. maritima, S. micropetala, S. nodosa, S. pilifera, S. procumbens, S. revelierei, and S. saginoides subsp. saginoides. The names S. apetala var. decumbens (=S. apetala subsp. apetala), S. bryoides (=S. procumbens), S. patula (=S. apetala subsp. apetala), S. revelierei, Spergula glabra (=S. glabra), Spergula pilifera (=S. pilifera), and Spergella subulata var. macrocarpa (=S. saginoides subsp. saginoides) are here typified. Specimens deposited at B-W, C, E, and LY, and illustrations by Reichenbach were considered for the typifications. Specifically, two Reichenbach’s illustrations are chosen for S. bryoides and S. saginoides var. macrocarpa. A specimen at B-W is designated as the lectotype of S. glabra. Two specimens at C and G are designated as the lectotypes of S. apetala var. decumbens and S. revelierei, respectively. A specimen at LY is designated for S. patula. As we did not find original material, a neotype at G is designated for S. pilifera.

Nomenclatural and taxonomic issues concerning Amaranthus caudatus and the related taxa are presented. Types are designated for names A. caudatus var. albiflorus (neotype at RO), A. caudatus var. atropurpureus (neotype at GH), A. caudatus var. gibbosus (neotype at RO), A. dussi (neotype at NAP), and A. edulis (lectotype at LP). Holotypes are indicated for the names A. caudatus var. pseudopaniculatus f. oblongipetalus (EA), A. caudatus var. pseudopaniculatus f. pseudopaniculatus (EA), A. caudatus subsp. saueri (PR), and Amaranthus edulis var. spadiceus (CORD). The names A. caudatus var. albiflorus, A. caudatus var. atropurpureus, A. caudatus subsp. saueri, A. dussi, and Amaranthus edulis var. spadiceus are considered as hererotypic synonyms of A. caudatus. On the basis of morphological, cytological, and molecular data, the taxa caudatus, mantegazzianus, and gibbosus are here proposed to be treated as different species. A new name—Amaranthus baileyanus—is proposed for A. caudatus var. gibbosus because of a previous and validly published Amaranthus gibbosus.

In this contribution, new data concerning the distribution of vascular flora alien to Italy are presented. It includes new records, confirmations, exclusions, and status changes for Italy or for Italian administrative regions. Nomenclatural and distribution updates published elsewhere are provided as Suppl. material 1.

In this contribution, new data concerning the distribution of native vascular flora in Italy are presented. It includes new records, confirmations, exclusions, and status changes to the Italian administrative regions. A new combination in the genus Pilosella is proposed. Nomenclatural and distribution updates, published elsewhere, and corrigenda are provided as Suppl. material 1.

Amaranthus is a genus taxonomically complex because of its high morphological variability, which led to nomenclatural disorders, misapplication of names, and misidentifications. Floristic and taxonomic studies on this genus are still incomplete, and many questions remain open. Seed micromorphology has been shown to play an important role in the taxonomy of plants. Regarding Amaranthaceae and Amaranthus, investigations are rare, and they refer to one or a few species. With the primary aim to test if seed features are helpful in the taxonomy of Amaranthus, we here present a detailed SEM study on seed micromorphology in 25 Amaranthus taxa using morphometric methods. Seeds were collected from field surveys and herbarium specimens; 14 seed coat features (7 qualitative and 7 quantitative) were measured on 111 samples (up to 5 seeds per sample). The results obtained revealed that seeds micromorphology provides interesting new taxonomic data concerning some taxa (species and below ranks). In fact, we were able to distinguish a few seed types, including one or more taxa, i.e., blitum-type, crassipes-type, deflexus-type, tuberculatus-type, and viridis-type. On the other hand, seed features are not useful for other species, for example, those included in the deflexus-type (A. deflexus, A. vulgatissimus, A. cacciatoi, A. spinosus, A. dubius, and A. stadleyanus). A diagnostic key of the studied taxa is proposed. Subgenera cannot be distinguished using seed features, thus confirming the published molecular data. All these facts reveal, once again, the taxonomic complexity of the genus Amaranthus since, e.g., just a few seed types can be defined.

All the names in Paronychia described from South America are investigated. Five names (P. arbuscula, P. brasiliana subsp. brasiliana var. pubescens, P. coquimbensis, P. hieronymi, and P. mandoniana) are lecto- or neotypified on specimens preserved at GOET, K, LP, and P. The typification of nine names, first proposed by Chaudhri in 1968 as the “holotype” are corrected according to Art. 9.10 of ICN. Three second-step typifications (Art. 9.17 of ICN) are proposed for P. camphorosmoides, P. communis, and P. hartwegiana. The following nomenclatural changes are proposed: P. arequipensis comb. et stat. nov. (basionym: P. microphylla subsp. microphylla var. arequepensis), P. compacta nom. nov. pro P. andina (Philippi non Gray; Art. 53.1 of ICN), P. jujuyensis comb. et stat. nov. (basionym: P. hieronymi subsp. hieronymi var. jujuyensis), P. compacta subsp. boliviana comb. nov. (basionym: P. andina subsp. boliviana), and P. compacta subsp. purpurea comb. nov. (basionym: P. andina subsp. purpurea). A new species (P. glabra sp. nov.) is proposed based on our examination of live plants and herbarium specimens. P. johnstonii subsp. johnstonii var. scabrida is synonymized (syn. nov.) with P. johnstonii. Finally, P. argyrocoma subsp. argyrocoma is excluded from South America since it was based on misidentified specimens (deposited at MO) of P. andina subsp. andina. A total of 30 species (43 taxa including subspecies, varieties, subvarieties, and forms) are recognized, highlighting that for some (Paronychia chilensis, P. communis, P. setigera) we provisionally accept Chaudhri’s infraspecific classification, since the high phenotypic variability of these taxa is quite complicated and further investigations need to solve their taxonomy.

Salicornia is a highly taxonomically problematic genus due to the reduced morphological observable characters. Ten Eurasian species are currently recognized: S. alpini, S. europaea, S. fruticosa, S. hispanica, S. lagascae, S. perennans, S. perennis, S. persica, S. procumbens, and S. pruinosa. In addition, eleven subspecies are accepted, mainly based on their distribution areas. Along the Venetian coasts and in Sardinia, in the past, an endemic species called S. veneta was recognized, but this name was later synonymized with S. procumbens subsp. procumbens. The aim of the present research is investigating different Italian Salicornia populations by a molecular point of view, using the nuclear ribosomal external transcribed spacer ETS and the plastid psbA-trnH intergenic spacer. A particular focus is on the comparison between Venetian (including those occurring in locus classicus of S. veneta) and Sardinian S. procumbens and other Italian populations of this species. The molecular analyses based on the plastid marker highlight that the Italian S. procumbens populations form two well distinct groups. In particular, some of the Venetian (Locus classicus of S. veneta) and all the Sardinian specimens are genetically distinct (=plastid haplotype 1) from the other investigated populations (=plastid haplotype 2). This indicates that the psbA-trnH haplotype 1 glassworts represent a distinct entity, which we suppose to coincide with the former S. veneta. Therefore, we suggest to recognize this taxonomic entity at the subspecies rank, as S. procumbens subsp. veneta comb. and stat. nov. However, contrary to the results found with the plastid psbA-trnH intergenic spacer, the ETS locus does not show a separation into two distinct clades for S. procumbens, probably due to a different evolution of the two loci. Nevertheless, in the ETS phylogenetic reconstruction, the Sardinian specimens (=ribotypes 2 and 3) are placed, together with a Moroccan sample, in a subclade separated from all the other S. procumbens. These results suggest that the Sardinian populations can represent a subspecies/incipient speciation process, probably due to geographic isolation. In the light of this, morphometric analyses (k-means, MANOVA, PCA, DA, and Box-Plot) have been carried out on the Sardinian and Venetian populations to verify if this distinction is detectable also by a morphological point of view. The morphometric analyses highlight the existence of two groups, concerning both the nuclear and plastid trees. Six characters were found to be diagnostic.

Sixteen new Asteraceae are added to the non-indigenous flora of Tunisia. Six of them (Dimorphotheca ecklonis, Gaillardia pulchella, Gazania linearis var. linearis, Guizotia abyssinica, Rudbeckia triloba and Tithonia diversifolia) are also new for the flora of North Africa. Chrysanthemum morifolium is here first reported for the Mediterranean Basin. Seven genera (Argyranthemum, Bidens, Dimorphotheca, Gaillardia, Guizotia, Rudbeckia and Tithonia) are recorded for the first time for the vascular flora of Tunisia. Most of these species escaped from gardens and pots and became casual along roadsides and/or disturbed areas after cypsela dispersion (except for Chrysanthemum morifolium). Some species, even though appearing to be well established locally and are potentially invasive at a worldwide level, still do not behave so in Tunisia. In addition to the floristic notes, the typification of the names Dimorphotheca ecklonis, Gorteria linearis and Mirasolia diversifolia are discussed.

In this contribution, new data concerning the distribution of vascular flora alien to Italy are presented. It includes new records, confirmations, exclusions, and status changes for Italy or for Italian administrative regions. Furthermore, three new combinations are proposed. Nomenclatural and distribution updates published elsewhere are provided as Suppl. material 1.

Hertia cheirifolia y H. maroccana (Asteraceae) son dos endemismos norteafricanos que crecen, respectivamente, en Argelia y Túnez, y en Marruecos. Presentamos en este trabajo descripciones morfológicas, distribución y categorización en la lista roja de la UICN. Además, se lleva a cabo un estudio nomenclatural de Hertia maroccana de Battandier and Othonna cheirifolia de Linneo y se tipifican ambos nombres sobre especímenes conservados, respectivamente, en MPU y LINN.

The delimitation of Limonium taxa is highly complicated due to hybridization, polyploidy, and apomixis. Many “microspecies” were described and aggregated into groups, most of which are still poorly known from both molecular and morphological points of view. The aim of this study is to investigate four endemic species from the Tyrrhenian coast of central Italy and the Ponziane Archipelago belonging to the L. multiforme group (L. amynclaeum, L. circaei, L. pandatariae, and L. pontium) by means of molecular and morphometric analyses. Molecular data by sequencing ITS and three plastid markers and morphometric data highlight new information about the taxonomy of these taxa so as to reduce them into a single specific entity. In fact, the better taxonomic choice is to consider the populations studied as part of a single species, i.e., Limonium pontium. Three subspecies are recognized, i.e., subsp. pontium [= L. circaei = L. amynclaeum; from Circeo to Gianola localities (excluding Terracina) and from islands Ponza, Palmarola, Zannone, and Santo Stefano], subsp. pandatariae comb. et stat. nov. (from island of Ventotene), and subsp. terracinense subsp. nov. (from Terracina).

The genus Mcneillia has been recently segregated from Minuartia L. based on molecular results, also supported by morphology. However, to date, a comprehensive study on the phylogenetic relationships within this genus is lacking. In this paper, we provide a multigene phylogeny of all the species and subspecies of Mcneillia employing two nuclear and six chloroplast markers. We documented extensive gene flow between taxa, sometimes separated at specific rank. In addition, Mcneillia as currently circumscribed, is not monophyletic. In fact, Mcneillia graminifolia subsp. brachypetala, strictly endemic to Greece, truly belongs to Minuartiella, a genus otherwise limited to South-West Asia. Moreover, even after removal of this taxon, our results do not support the monophyly of the taxa included in M. graminifolia s.l., the most variable and widespread species of the genus. Further controversial subspecies of Mcneillia graminifolia, i.e., subsp. hungarica and subsp. rosanoi, are shown to deserve taxonomic recognition as separate species, whereas Mc. moraldoi is not distinct at specific rank. In addition, Mc. saxifraga subsp. tmolea is here regarded as a further distinct species. A consistent taxonomic treatment is therefore proposed with six new combinations and nomenclatural notes, providing the necessary typifications.

The first inventory of the flora of Appia Antica Regional Park (Italy), one of the largest protected urban areas in Europe (4580 ha), its biological, ecological and biogeographical composition, and notes of the vegetation physiognomies and landscape are presented; physical characteristics of the territory (geomorphology, lithotypes, and phytoclimate) are also given. The landscape is defined by an agricultural matrix with natural and seminatural areas as patches, and riparian vegetation communities as corridors. The vegetation physiognomies are represented by types linked to the Mediterranean climate (mixed, Mediterranean, and riparian forests; scrubby, rocky, aquatic, and helophytic vegetation; anthropogenic communities). The floristic list includes 714 taxa (104 families and 403 genera). Therophytes prevail over hemicryptophytes; woody flora comprises about 30% of alien species. As regards chorotypes, together with a considerable number of Mediterranean species, there are many exotic species with wide distribution areas testifying to a long-lasting anthropic impact. Floristic novelties (european, national, and regional levels) for 21 taxa are reported. The extraordinary species diversity discovered (43% of flora of Rome and 20% of regional flora) is linked to the landscape heterogeneity, the characteristics of which are: (1) persistence of residual natural patches, (2) occurrence of quite well-preserved aquatic habitats and humid meadows, (3) a rich anthropogenic flora, (4) an interesting flora of archeological sites, (5) occurrence of species not common in Latium, (6) occurrence of populations of aliens in crops (which cause economic impact), (7) presence of aliens on archeological ruins (which cause economic-social impacts). The extensive set of data provided represents a general base framework for guiding future research efforts and landscape action plans consistent with environmental sustainability.

In this contribution, new data concerning the distribution of vascular flora alien to Italy are presented. It includes new records, confirmations, exclusions, and status changes for Italy or for Italian administrative regions of taxa in the genera Acalypha, Acer, Canna, Cardamine, Cedrus, Chlorophytum, Citrus, Cyperus, Epilobium, Eucalyptus, Euphorbia, Gamochaeta, Hesperocyparis, Heteranthera, Lemna, Ligustrum, Lycium, Nassella, Nothoscordum, Oenothera, Osteospermum, Paspalum, Pontederia, Romulea, Rudbeckia, Salvia, Sesbania, Setaria, Sicyos, Styphnolobium, Symphyotrichum, and Tradescantia. Nomenclature and distribution updates, published elsewhere, and corrigenda are provided as supplementary material.

A morphometric analysis has been carried out for the first time in order to investigate the variability of the infraspecific taxa included in the steno-Mediterranean Malva cretica s.l., i.e. M. Cretica subsp. althaeoides, M. cretica subsp. cretica, and the poorly-known M. Cretica var. montana, described from southern Italy. According to our results, M. cretica subsp. althaeoides and subsp. cretica are worth of taxonomic recognition at subspecies rank and occupy respectively the western and the central-eastern sectors of the species range. Concerning M. Cretica var. montana, characterized by some intermediate features, it should be included in M. Cretica subsp. cretica.

In this contribution, new data concerning the Italian distribution of alien vascular flora are presented. It includes new records and exclusions for Italy or for Italian administrative regions of taxa in the genera Cedrus, Cenchrus, Citrus, Cyrtomium, Diospyros, Elaeagnus, Erigeron, Iris, Oenothera, Pavonia, Phytolacca, Styphnolobium, and Verbena. Furthermore, a new combination in the genus Amaranthus is proposed.

In this contribution, new data concerning the Italian distribution of alien vascular flora are presented. It includes new records, exclusions, confirmations, and status changes for Italy or for Italian administrative regions for taxa in the genera Acer, Amaranthus, Araujia, Aubrieta, Avena, Bidens, Calycanthus, Celtis, Elaeagnus, Eragrostis, Euonymus, Fallopia, Ficus, Hedera, Lantana, Ligustrum, Ludwigia, Morus, Oenothera, Opuntia, Oxalis, Parkinsonia, Paspalum, Paulownia, Platycladus, Pleuropterus, Rumex, Salvia, Senecio, Setaria, Syagrus, Tradescantia, Trifolium and Yucca. Furthermore, a new combination in the genus Vicia is proposed.

The names of the Italian taxa in Cirsium sect. Eriolepis are discussed. The accepted names are: Cirsium echinatum, C. eriophorum subsp. eriophorum, C. eriophorum subsp. spathulatum, C. ferox, C. italicum, C. lacaitae, C. lobelii, C. morisianum, C. scabrum, C. tenoreanum, C. vallis-demonii subsp. vallis-demonii, C. vallis-demonii subsp. calabrum comb. nov., and C. vulgare (= C. crinitum, C. sylvaticum). Four accepted names are typified by specimens preserved at FI (one lectotype), G (one lectotype and one neotype), P (one lectotype), and by illustrations (two lectotypes). Several other heterotypic synonyms of taxa described from Italy are discussed and six of them are typified. A new combination and status are proposed: C. vallis-demonii subsp. calabrum, based on C. eriophorum var. vallis-demonii f. calabrum.

The typification of the name Bosea yervamora L. (Amaranthaceae) is discussed under the light of historical knowledge of Canarian plants in Linnean herbaria. An illustration from Sloane is designated as the lectotype. A specimen from BM-SL (Sloane collection) is designated as typotype.