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Two new and very similar species of the genus Xynobius Foerster, 1863 are described and illustrated, X. subparallelus Han & van Achterberg, sp. nov. from Japan (Honshu) and X. setosiscutum van Achterberg, sp. nov. from Norway. Three species are newly reported from Norway: Xynobius aciculatus (Thomson, 1895), X. comatus (Wesmael, 1835), and X. polyzonius (Wesmael, 1835). X. polyzonius (Wesmael, 1835) and X. sapporanus (Fischer, 1963) are new combinations. Identification keys to the Xynobius species known from Norway and Japan are added.

Gupta, Ankita, Achterberg, Cornelis Van, Pattar, Rohit, Minz, Kriti Arpana (2022): On the occurrence of a Palaearctic species of the genus Aphaereta Foerster (Braconidae, Alysiinae) and description of a new species of the genus Leiophron Nees (Braconidae, Euphorinae) from central India. Zootaxa 5209 (4): 455-462, DOI: https://doi.org/10.11646/zootaxa.5209.4.4

Centistidea yamu Ranjith & Priyadarsanan sp. nov. (Figures 13–14) urn:lsid:zoobank.org:act: 2938672F-EA05-47CF-9D33-1AB9CD2EC936 Material examined. Holotype. ♀ INDIA: Arunachal Pradesh, Ramsing, 28°65.538’ N, 94°97.543’ E, 2087 m.a.s.l., secondary evergreen forest, Malaise trap, 9–12.v.2022, coll. Ranjith, A.P. (AIMB). Description. Female Body measurements. Length of body 2.2 mm, length of fore wing 2.5 mm. Head. Antenna with 14 antennomeres, all antennomeres longer than wide (Fig. 13A). First flagellomere 1.1 × as long as second flagellomere. First, second and penultimate flagellomeres 4.8, 4.6 and 2.4 × as long as wide. Maxillary palp with four segments (Figs 13A, B), fourth segment 1.2 × as long as third segment. Face distinctly convex medially and laterally (Fig. 13B), 1.5 × as wide as long, smooth, setose. Clypeus 2.4 × wider ventrally than medially high with convex posterior margin (Fig. 13B). Malar suture distinct (Fig. 13B). Length of malar space 1.2 × basal width of mandible. Mandible distinctly twisted (Fig. 13B), inner tooth distinctly shorter than outer tooth. Frons smooth, convex medially, setose (Fig. 13C). Vertex smooth, setose (Fig. 13C). OOL: transverse diameter of posterior ocellus: POL= 2.8: 1.0: 1.2. Length of eye 1.4 × as long as temple in dorsal view. Mesosoma. Mesosoma 1.4 × as long as high. Pronotum and propleuron smooth laterally (Fig. 13D). Mesoscutum punctate in anterior half smooth posteriorly, setose, carinate postero-laterally (Fig. 13E). Notauli only impressed anteriorly, faintly crenulated (Fig. 13E). Scutellar sulcus represented only by a transverse groove, not crenulated (Fig. 13E). Scutellum evenly setose, smooth (Fig. 13E), with a pair of large pits medio-posteriorly separated by a carina, sides of scutellum irregularly sculptured (Fig. 13F). Mesopleuron smooth, glabrous, setose ventrally (Fig. 13D). Metapleuron with a deep small pit antero-dorsally, with deep shallow longitudinal depressed line, glabrous (Fig. 13D). Propodeum with a distinct and complete medio-longitudinal carina absent behind costula, dorsal part of midlongitudinal carina associated with irregular sculpture, sparsely setose, antero-lateral part with shallow declivity, postero-lateral part faintly crenulated, posterior part well differentiated from dorsal part, posterior transverse carina distinct (Fig. 13F). Wings. Fore wing: Vein r distinct, 0.3 × as long as 2RS (Fig. 14C). R1 distinct, wide (Fig. 14C). Vein (RS+M)b 0.6 × as long as 1m-cu. Vein 1-M 1.5 × as long as m-cu. Vein (RS+M)a 2.6 × (RS+M)b. Vein 1CUa thickened, 0.6 × as long as 1CUb (Fig. 14C). Hind wing: Vein cu-a straight. Veins M+CU: 1M: 1r-m= 2.3: 2.7: 1.0. Legs. Hind coxa smooth, setose (Figs 13A, 14A). Tarsal claws without basal lobe. Hind femur, tibia and basitarsus 4.2, 10.4 and 4.6 × as long as wide respectively. Metasoma. First tergite smooth, widening medially, weakly narrowing apically, distinctly narrowing basally, 3.4 × as long as its maximum width (Fig. 14B). Second tergite longitudinally striate, setose, medial length 0.7 × as long as third tergite (Fig. 14B). Third tergite longitudinally striate, setose (Fig. 14B). Ovipositor straight. Ovipositor sheath 0.1 × as long as fore wing, slightly narrowing apically with long setae apically (Figs 14A, B). Colour. Body reddish brown except face, clypeus, mandible, maxillary and labial palps, vertex laterally, tegula, legs except hind tarsus yellow. Male. Unknown. Etymology. The species is named after its brown body; “ yamu ” is brown the language of Adi indigenous people of Arunachal Pradesh. Biology. Unknown. Distribution. India (Arunachal Pradesh). Comparative diagnosis. The new species is coming close to C. sii (Maetô, 1995) by the following characters, pterostigma brown, first flagellomere 1.1 × as long as second flagellomere and mesoscutum brownish. However, the new species differs from the latter in having the following differences; length of eye 1.4 × as long as temple in dorsal view (1.1 × in C. sii), first flagellomere 4.8 × as long as wide (5.0–6.0 × in C. sii), OOL 2.3 × POL (1.6 in C. sii) and hind femur 4.2 × as long as wide (3.6–3.8 × in C. sii).

Centistidea furca Ranjith & Priyadarsanan, sp. nov. (Figures 7–8) urn:lsid:zoobank.org:act: A6BBDC10-1474-43B1-8A4A-41FF5763A2FF Material examined. Holotype. ♀ INDIA: Karnataka, Chamarajanagar, Biligiri Ranganathaswamy Temple Tiger Reserve, 12°00.345′ N, 77°07.526′ E, 975 m. a.s.l., dry deciduous forest, Malaise trap, 14.x–17.xi.2005, coll. Priyadarsanan, D. R. (AIMB). Description. Female Body measurements. Length of body 2.1 mm, length of fore wing 2.0 mm. Head. Antenna with 14 antennomeres, all antennomeres longer than wide (Fig. 7A). First flagellomere 1.0 × as long as second flagellomere. First, second and penultimate flagellomeres 4.4, 4.2 and 2.0 × as long as wide. Maxillary palp with four segments (Figs 7B, D), fourth segment 1.3 × as long as third segment. Face distinctly convex medially (Fig. 7B), 1.7× as wide as long, granulate, setose. Clypeus 1.8 × wider ventrally than medially high with straight posterior margin (Fig. 7B). Malar suture distinct (Figs 7B, D). Length of malar space 1.4 × basal width of mandible. Mandible distinctly twisted (Fig. 7B), inner tooth distinctly shorter than outer tooth. Frons granulate, moderately convex medially, sparsely setose laterally (Fig. 7C). Vertex granulate, sparsely setose, setose laterally (Fig. 7C). OOL: transverse diameter of posterior ocellus: POL= 2.4: 1.0: 1.0. Length of eye 1.5 × as long as temple in dorsal view. Mesosoma. Mesosoma 1.3 × as long as high. Pronotum and propleuron smooth laterally, lateral pronotum shallowly impressed medially (Fig. 7D). Mesoscutum punctate, setose, without carinae laterally (Fig. 7E). Notauli only impressed anteriorly (Fig. 7E). Scutellar sulcus represented only by a transverse groove, faintly crenulated medially (Fig. 7E). Scutellum evenly setose, smooth (Fig. 7E), with a pair of large pits medio-posteriorly separated by a 0.5 × diameter of the pit, sides of scutellum irregularly sculptured (Fig. 7F). Mesopleuron smooth, glabrous, setose ventrally (Fig. 7D). Metapleuron with a deep small pit antero-dorsally, with deep shallow longitudinal depressed line, glabrous (Fig. 7D). Propodeum with a distinct and complete anteriorly forked medio-longitudinal carina extended behind costulae, costulae arched, incomplete, dorsal part of midlongitudinal carina associated with transverse carinae posteriorly, sparsely setose, antero-lateral part with shallow declivity, postero-lateral part faintly crenulated, posterior part indistinctly differentiated from dorsal part, transverse carina present between costulae and posterior transverse carina, posterior transverse carina distinct (Fig. 7F). Wings. Fore wing: Vein r distinct, 0.1 × as long as 2RS (Fig. 8C). R1 indistinct (Fig. 8C). Vein (RS+M)b 0.8 × as long as 1m-cu. Vein 1-M 1.7 × as long as m-cu. Vein (RS+M)a 2.1 × (RS+M)b. Vein 1CUa thickened, 0.7 × as long as 1CUb (Fig. 8C). Hind wing: Vein cu-a straight. Veins M+CU: 1M: 1r-m= 2.2: 2.3: 1.0. Legs. Hind coxa smooth, setose (Figs 7A, 8A). Tarsal claws without basal lobe. Hind femur, tibia and basitarsus 3.0, 5.9 and 3.3 × as long as wide respectively. Metasoma. First tergite smooth, narrow medially, widening subapically, parallel sided apically, distinctly narrowing basally, 4.5 × as long as its maximum width (Fig. 8B). Second tergite smooth, sparsely punctate, setose, medial length 0.6 × as long as third tergite (Fig. 8B). Third tergite longitudinally striate, setose (Fig. 8B). Ovipositor straight. Ovipositor sheath 0.1 × as long as fore wing, slightly narrowing apically with long sparse setae apically (Fig. 8A). Colour. Body yellow except antennae, apex of mandibles, propodeal carinae, first and second metasomal tergites, apical metasomal tergites, apex of ovipositor brown. Male. Unknown. Etymology. The new species is named after the presence of anteriorly bifurcate midlongitudinal carina of propodeum. Biology. Unknown. Distribution. India (Karnataka). Comparative diagnosis. Centistidea furca sp. nov. can be differentiated from the rest of Centistidea species by the combination of the following characters, anteriorly forked propodeal carina, incomplete and curved costula and presence of transverse carina between costula and posterior transverse carina. This new species comes close to C. kumatai (Maetô) in having arched costula and dorsal length of eye 1.5 × as long as temple. Apart from the forked midlongitudinal propodeal carina C. furca sp. nov. differs from C. kumatai by the following characters, scape and pedicel yellow (brown in C. kumatai), medio-posterior pits of scutellum separated by ½ diameter of the pit (separated by a thin longitudinal carina in C. kumatai), length of malar space 1.4 × as long as basal width of mandible (1.0 × in C. kumatai), hind femur 3.0 × as long as wide (3.5–3.8 × as long as wide in C. kumatai) and first metasomal tergite 4.5 × aslong as its maximum width (3.0–3.5 × in C. kumatai).

Key to the Oriental species of the genus Centistidea Rohwer 1. Median carina of propodeum absent behind level of costulae (Figs 3F, 5F, 9F, 13F)................................. 2 - Median carina of propodeum present behind level of costulae and reaching posterior transverse carina of propodeum (Figs 7F, 11F)............................................................................................... 15 2. Pterostigma short triangular and without distinct slender triangular apex or vein R1; vein 1RS of fore wing distinctly developed and narrow, up to 0.4 × as long as vein 1M; first flagellomere 1.2–1.6 × as long as second flagellomere [China, Indonesia, Japan, Korea, Russia]............................................................. Centistidea irruptor (Papp, 1987) - Pterostigma elongate triangular and with distinct slender triangular apex or with distinct vein R1 (Figs 4C, 6D, 8C, 10C, 14C); vein 1RS of fore wing less developed and wider, 0.2–0.3 × as long as vein 1M (Figs 4C, 6D, 8C, 10C, 14C); first flagellomere 1.0–1.2 × as long as second flagellomere................................................................... 3 3. Dorsal length of eye 1.4 × temple; mesopleuron dark brown (Fig. 13D); propodeum crenulated near posterior transverse carina (Fig. 13F); vein R1 of fore wing distinctly vein-like and narrow (Fig. 14C) [India]........................................................................................... Centistidea yamu Ranjith & Priyadarsanan, sp. nov. - Dorsal length of eye 1.6–3.1 × temple; mesopleuron often yellow (Figs 3D, 5D, 7D, 9D); propodeum not crenulated near posterior transverse carina (Fig. 5F); vein R1 of fore wing triangular and not vein-like (Fig. 12C) or if vein-like then rather wide (Fig. 14C)........................................................................................... 4 4. Vein 1CUa of fore wing 0.4–0.7 × as long as vein 1Cub....................................................... 5 - Vein 1CUa of fore wing 0.8–1.2 × as long as vein 1Cub....................................................... 9 5. Dorsal length of eye more than 2.0 × temple................................................................ 6 - Dorsal length of eye less than 2.0 × temple................................................................. 7 6. Apex of antenna pale yellowish; scutellum with pits medio-posteriorly separated by 0.50 × diameter of pit; scutellar sulcus shallow without crenulations [India]........................ Centistidea mellapicalis Ranjith & van Achterberg, 2018 - Apex of antenna pale brown; scutellum with pits medio-posteriorly separated by carina; scutellar sulcus distinctly crenulated [India]................................................. Centistidea crenulator Ranjith & van Achterberg, 2018 7. OOL 4.0 × POL; face 1.2 × as wide as long; length of malar space 0.8 × basal width of mandible; first metasomal tergite 3.5 × as long as its maximum width [China]......................................... Centistidea zhaoi Chen & He, 1997 - OOL 1.9–2.4 × POL; face 1.6–1.8 × as wide as long; length of malar space 0.9–1.1 × basal width of mandible; first metasomal tergite 3.2–3.4 × as long as its maximum width.............................................................. 8 8. Scutellar pits separated by width of pit; propodeum yellow and smooth near midlongitudinal carina; dorsal part of metasomal tergite 1 & 2 yellow; hind basitarsus 3.4 × as long as wide [India].... Centistidea lineator Ranjith & van Achterberg, 2018 - Scutellar pits separated by a carina; propodeum brown and with striae near midlongitudinal carina; dorsal part of metasomal tergite 1 & 2 brown; hind basitarsus 4.1 × as long as wide [India]...................................................................................................... Centistidea scutellator Ranjith & van Achterberg, 2018 9. Scutellar pits separated by a thin carina (Figs 3F, 13F)....................................................... 10 - Scutellar pits separated by at least 0.25 × diameter of pit (Figs 5F, 9F).......................................... 12 10. Scutellum with oval pits medio-posteriorly; propodeum smooth near midlongitudinal carina; first flagellomere as long as second [China, Japan, Korea, Russia, Vietnam]..................................... Centistidea mogra (Papp, 1987) - Scutellum with subrectangular pits medio-posteriorly (Fig. 3F); propodeum sculptured near midlongitudinal carina (Fig. 3F); first flagellomere longer than second..................................................................... 11 11. OOL 4.2 × POL; first flagellomere 5.8 × as long as wide; mesoscutum and propodeum yellow; hind tibia 8.8 × as long as wide; first metasomal tergite 3.0 × as long as its maximum width [India]... Centistidea rugator Ranjith & van Achterberg, 2018 - OOL 2.5 × POL; first flagellomere 5.3 × as long as wide; mesoscutum and propodeum mostly brown (Figs 3E, F); hind tibia 6.8 × as long as wide; first metasomal tergite 3.5 × as long as its maximum width [Sri Lanka]................................................................................ Centistidea burwelli Ranjith & Priyadarsanan, sp. nov. 12. First metasomal tergite 2.4 × as long as its maximum width................................................... 13 - First metasomal tergite 2.7–3.2 × as long as its maximum width............................................... 14 13. Pterostigma brown; OOL 2.2 × POL; length of malar space 1.5 × basal width of mandible; clypeus 2.2 × wider ventrally than medially high; hind basitarsus 4.5 × as long as wide........... Centistidea procarinator Ranjith & van Achterberg, 2018 - Pterostigma yellow (Fig. 6D); OOL 1.8 × POL; length of malar space 1.1 × basal width of mandible; clypeus 1.5 × wider ventrally than medially high; hind basitarsus 4.0 × as long as wide [India].......................................................................................... Centistidea flavostigmata Ranjith & Priyadarsanan, sp. nov. 14. Mesoscutum and propodeum brown; fore wing vein 1CUa 0.9 × as long as 1Cub; OOL 2.0 × POL; hind basitarsus 4.5 × as long as wide [India]............................................... Centistidea cosmopteryxi Ahmad & Pandey, 2019 - Mesoscutum and propodeum yellow (Figs 9E, F); fore wing vein 1CUa 1.2 × as long as 1Cub; OOL 2.5 × POL; hind basitarsus 3.6 × as long as wide [Sri Lanka].............................. Centistidea lanka Ranjith & Priyadarsanan, sp. nov. 15. Junction of propodeal costula pointed posteriorly (Fig. 7F)................................................... 16 - Junction of propodeal costula pointed anteriorly (Fig. 11F)................................................... 17 16. Midlongitudinal carina of propodeum not forked anteriorly; pterostigma brown; vein R1 of fore wing indistinct; first metasomal tergite 4.0 × as long as its maximum width; OOL 4.0 × POL [India]..... Centistidea acrocercopsi Ahmad & Pandey, 2019 - Midlongitudinal carina of propodeum forked anteriorly (Fig. 7F); pterostigma yellow (Fig. 8C); vein R1 of fore wing distinctly vein-like and wide (Fig. 8C); first metasomal tergite 4.5 × as long as its maximum width; OOL 2.4 × POL [India].................................................................. Centistidea furca Ranjith & Priyadarsanan, sp. nov. 17. Antero-dorsal part of propodeum about 1.5 × as long as metanotum medially; length of eye 1.2 × as long as temple in dorsal view; fore wing vein 1CUa of fore wing slender; first flagellomere as long as second [China]............................................................................................... Centistidea immitis Wu & Chen, 2000 - Antero-dorsal part of propodeum about 2.5–3.0 × as long as metanotum medially; length of eye 1.8–2.0 × as long as temple in dorsal view; fore wing vein 1CUa of fore wing thickened (Fig. 12C); first flagellomere 1.3–1.6 × as long as second...... 18 18. Medio-posterior pits on scutellum circular and without crenulations; clypeus brown, 2.0 × as wide as medially long; metasomal tergite 3 brown; first flagellomere 6.5 × as long as wide [China-Taiwan]....... Centistidea taiwanica (Papp & Chou, 1996) - Medio-posterior pits on scutellum oval and crenulated (Fig. 11F); clypeus yellow (Fig. 11B), 2.6 × as wide as medially long; metasomal tergite 3 yellow (Fig. 12B); first flagellomere 5.5 × as long as wide [India].................................................................................. Centistidea nyeringensis Ranjith & Priyadarsanan, sp. nov.

Centistidea nyeringensis Ranjith & Priyadarsanan sp. nov. (Figures 11–12) urn:lsid:zoobank.org:act: 336BAD51-A3E8-472C-8AD1-B6B58783FF9D Material examined. Holotype. ♀ INDIA: Arunachal Pradesh, Nyering, 28°95.784′ N, 94°98.847′ E, 777 m. a.s.l., 20.x.2022, yellow pan trap, coll. A.P. Ranjith (AIMB). Paratype, 1 female same data as holotype. Description. Female Body measurements. Length of body 1.9 mm, length of fore wing 1.9 mm. Head. Antenna with 14 antennomeres, all antennomeres longer than wide (Fig. 11A). First flagellomere 1.3 × as long as second flagellomere. First, second and penultimate flagellomeres 5.5, 4.0 and 2.8 × as long as wide. Maxillary palp with four segments (Figs 11B, D), fourth segment 1.3 × as long as third segment. Face distinctly convex medially (Fig. 11B), 1.8 × as wide as long, smooth, setose. Clypeus 2.6 × wider ventrally than medially high with straight posterior margin (Fig. 11B). Malar suture distinct (Figs 11B, D). Length of malar space 0.8 × basal width of mandible. Mandible distinctly twisted (Fig. 11B), inner tooth distinctly shorter than outer tooth. Frons smooth, moderately convex medially, sparsely setose laterally (Fig. 11C). Vertex smooth, sparsely setose, setose laterally (Fig. 11C). OOL: transverse diameter of posterior ocellus: POL= 3.3: 1.0: 1.6. Length of eye 1.9 × as long as temple in dorsal view. Mesosoma. Mesosoma 1.4 × as long as high. Pronotum and propleuron smooth laterally, lateral pronotum indistinctly crenulate medially (Fig. 11D). Mesoscutum punctate in anterior half smooth posteriorly, setose, without carinae laterally (Fig. 11E). Notauli only impressed anteriorly (Fig. 11E). Scutellar sulcus represented only by a transverse groove, not crenulated (Fig. 11E). Scutellum evenly setose, smooth (Fig. 11E), with a pair of large crenulated pits medio-posteriorly separated by a carina, sides of scutellum irregularly sculptured (Fig. 11F). Mesopleuron smooth, glabrous, setose ventrally (Fig. 11D). Metapleuron with a deep small pit antero-dorsally, with deep shallow longitudinal depressed line, glabrous (Fig. 11D). Propodeum with a distinct and complete medio-longitudinal carina extended behind costula, dorsal part of midlongitudinal carina associated with irregular sculpture, sparsely setose, antero-lateral part with shallow declivity, postero-lateral part faintly crenulated, posterior part well differentiated from dorsal part, posterior transverse carina distinct (Fig. 11F). Wings. Fore wing: Vein r distinct, 0.2 × as long as 2RS (Fig. 12C). R1 indistinct (Fig. 12C). Vein (RS+M)b 0.6 × as long as 1m-cu. Vein 1-M 1.7 × as long as m-cu. Vein (RS+M)a 2.7 × (RS+M)b. Vein 1CUa thickened, 0.6 × as long as 1CUb (Fig. 12C). Hind wing: Vein cu-a straight. Veins M+CU: 1M: 1r-m= 2.1: 2.7: 1.0. Legs. Hind coxa smooth, setose (Figs 11A, 12A). Tarsal claws without basal lobe. Hind femur, tibia and basitarsus 3.6, 8.1 and 5.2 × as long as wide respectively. Metasoma. First tergite smooth, narrow medially, widening subapically, weakly narrowing apically, distinctly narrowing basally, 3.1 × as long as its maximum width (Fig. 12B). Second tergite longitudinally striate, smooth medially, setose, medial length 0.5 × as long as third tergite (Fig. 12B). Third tergite longitudinally striate, setose (Fig. 12B). Ovipositor straight. Ovipositor sheath 0.09 × as long as fore wing, slightly narrowing apically with long setae apically (Figs 12A, B). Colour. Body reddish brown except head, maxillary and labial palps, propleuron, legs, pterostigma, metasomal tergites 1–3, metasomal sternites, ovipositor sheath except apically yellow. Male. Unknown. Etymology. The species is named after the village, Nyering in which the specimens were collected. Biology. Unknown. Distribution. India (Arunachal Pradesh). Comparative diagnosis. The new species comes close to C. taiwana (Papp & Chou, 1996) in having scape and pedicel yellow, first flagellomere 1.3 × as long as second flagellomere, dorsal length of eye 1.8 × as long as temple. However, the new species can be distinguished from C. taiwana in having the following characters; face and clypeus yellow (brown in C. taiwana), clypeus 2.6 × as wide as long medially (2.0 × in C. taiwana), first flagellomere 5.5 × as long as wide (6.5 × in C. taiwana) and medioposterior pits on scutellum separated by a thin carina (separated by 0.25 × diameter of the pit in C. taiwana).

Centistidea flavostigmata Ranjith & Priyadarsanan, sp. nov. (Figures 5–6) urn:lsid:zoobank.org:act: 1C0C03B8-81F0-429F-BFFA-D3E83AE2A58E Material examined. Holotype. ♀ INDIA: Karnataka, Chamarajanagar, Biligiri Ranganathaswamy Temple Tiger Reserve, Gombekallu, 11°54.363′ N, 77°11.235′ E, evergreen forest, Malaise trap, 14.ii–26.iii.2006, coll. Priyadarsanan, D. R. (AIMB). Paratypes: 3 males with same label data (AIMB). Description. Female Body measurements. Length of body 2.8 mm, length of fore wing 2.7 mm. Head. Antenna with 14 antennomeres, all antennomeres longer than wide (Figs 5A, 6E). First flagellomere 1.2 × as long as second flagellomere. First, second and penultimate flagellomeres 4.8, 4.2 and 2.6 × as long as wide. Maxillary palp with four segments (Figs 5B, D), fourth segment 1.4 × as long as third segment. Face distinctly convex medially (Fig. 5B), 1.8× as wide as long, smooth, setose. Clypeus 1.5 × wider ventrally than medially high with convex posterior margin (Fig. 5B). Malar suture distinct (Figs 5B, D). Length of malar space 1.1 × basal width of mandible. Mandible distinctly twisted (Fig. 5B), inner tooth distinctly shorter than outer tooth. Frons smooth, convex medially, setose laterally, glabrous medially (Fig. 5C). Vertex smooth, glabrous medially, setose laterally (Fig. 5C). OOL: transverse diameter of posterior ocellus: POL= 1.8: 1.0: 1.0. Length of eye 2.0 × as long as temple in dorsal view. Mesosoma. Mesosoma 1.6 × as long as high. Pronotum and propleuron smooth laterally (Fig. 5D). Mesoscutum punctate in anterior half smooth posteriorly, setose, without carinae laterally (Fig. 5E). Notauli only impressed anteriorly, faintly crenulated (Fig. 5E). Scutellar sulcus represented only by a transverse groove, not crenulated (Fig. 5E). Scutellum evenly setose, smooth (Fig. 5E), with a pair of large pits medio-posteriorly separated by less than diameter of a pit and rugose between large pits, sides of scutellum irregularly sculptured (Fig. 5F). Mesopleuron smooth, glabrous, setose ventrally (Fig. 5D). Metapleuron with a deep small pit antero-dorsally, with deep shallow longitudinal depressed line, glabrous (Fig. 5D). Propodeum with a distinct and complete medio-longitudinal carina absent behind costula, dorsal part of midlongitudinal carina associated with irregular sculpture, sparsely setose, antero-lateral part with shallow declivity, postero-lateral part faintly crenulated, posterior part well differentiated from dorsal part, posterior transverse carina distinct (Fig. 5F). Wings. Fore wing: Vein r distinct, 0.2 × as long as 2RS (Fig. 6D). R1 indistinct (Fig. 6D). Vein (RS+M)b 0.7 × as long as 1m-cu. Vein 1-M 1.5 × as long as m-cu. Vein (RS+M)a 2.4 × (RS+M)b. Vein 1CUa thickened, 0.8 × as long as 1CUb (Fig. 6D). Hind wing: Vein cu-a straight. Veins M+CU: 1M: 1r-m= 2.3: 3.2: 1.0. Legs. Hind coxa smooth, setose (Figs 5A, 6A). Tarsal claws without basal lobe. Hind femur, tibia and basitarsus 3.2, 5.7 and 4.0 × as long as wide respectively. Metasoma. First tergite smooth, widening medially, weakly narrowing apically, distinctly narrowing basally, 2.4 × as long as its maximum width (Figs 6B, C). Second tergite longitudinally striate, setose, medial length 0.65 × as long as third tergite (Figs 6B, C). Third tergite longitudinally striate, setose (Fig. 6C). Ovipositor straight. Ovipositor sheath 0.04 × as long as fore wing, slightly narrowing apically with long setae apically (Fig. 6A). Colour. Body reddish brown except head, maxillary and labial palps, mesoscutum medially, scutellum, pterostigma, legs, metasoma except second tergite dorsally yellow. Male. Unknown. Etymology. The species is named after its yellow pterostigma; “ flavus ” is yellow in Latin. Biology. Unknown. Distribution. India (Karnataka). Comparative diagnosis. The new species comes close to C. procarinator Ranjith & van Achterberg, 2018, sharing the posterior pits of scutellum separated by 0.5 × diameter of pit. But the new species can be separated from C. procarinator by the following differences: transverse posterior carina of propodeum arched and close to an additional transverse carina (transverse posterior carina rounded medially and without additional transverse carina in C. procarinator), shorter malar space than in C. procarinator; lateral depressions of metanotum smooth (crenulate in C. procarinator), first flagellomere 4.8 × as long as wide (4.1 × in C. procarinator), hind basitarsus 4.0 × as long as wide (4.5 × in C. procarinator) and OOL 1.8 × POL (2.2 × in C. procarinator). Based on the character of midlongitudinal carina of propodeum bifurcated posteriorly, the new species comes close to C. tihamica Ghramh & Ahmad, 2019 described from Saudi Arabia. However, the new species can be distinguished from the latter in having mesoscutum yellowish (blackish brown in C. tihamica), dorsal length of eye 2.0 × as long as temple (1.5 × in C. tihamica), scutellar sulcus without crenulae (with crenulae in C. tihamica), hind tibia 5.7 × as long as wide (7.0 × in C. tihamica) and first metasomal tergite 2.4 × as long as maximum width (4.0 × in C. tihamica).

Key to the genera of Miracinae 1 Ovipositor sheath with tuft of setae apically (Fig. 2C); pair of posterior depressions of scutellum circular and distinctly separated (Fig. 2A); apical antennomeres of ♀ shortened (Fig. 1D); propodeum longitudinally or obliquely carinate and no medio-longitudinal carina or transverse carina (Figs 2A–B), if with irregular medio-longitudinal carina antero-medially then fourth flagellomere with acute spine apico-ventrally (Fig. 1D); vein 2RS of fore wing less than 2.0 × as long as vein (RS+M)b (Fig. 2F); notauli long and crenulated (Fig. 1F); vein R1 of fore wing longer than pterostigma (except in R. fereta) (Fig. 2F)................................................................... Rugosimirax Ranjith & van Achterberg gen. nov. - Ovipositor sheath with scattered setae apically (Fig. 4A); pair of posterior depressions of scutellum elliptical and closer to each other (Fig. 3F), obsolescent or absent; apical antennomeres of ♀ elongate (Figs 3A, 4E); propodeum smooth or with medio-longitudinal and transverse carinae (Fig. 3F); fourth flagellomere without acute spine apico-ventrally (Fig. 4E); vein 2RS of fore wing 2.0–3.0 × as long as vein (RS +M)b or veins obsolescent (Fig. 4D); notauli either short (Fig. 3E) and smooth or absent; vein R1 of fore wing either absent or present and distinctly shorter than pterostigma (Fig. 4D)........................ 2 2. Propodeum smooth medially and without transverse carinae; notauli largely absent...................... Mirax Haliday - Propodeum with a longitudinal carina medially and connected to more or less developed transverse carinae (Figs 3F, 4C); notauli short and shallow or obsolescent (Fig. 3E)............................................ Centistidea Rohwer

Ranjith, A.P., Achterberg, Cornelis Van, Priyadarsanan, Dharma Rajan (2023): A new genus in the braconid subfamily Miracinae from the Oriental region, with descriptions of seven new species from India and Sri Lanka. Zootaxa 5318 (4): 451-473, DOI: 10.11646/zootaxa.5318.4.1, URL: http://dx.doi.org/10.11646/zootaxa.5318.4.1

Rugosimirax expectata Ranjith & van Achterberg, sp. nov. (Figures 1–2) urn:lsid:zoobank.org:act: 8DC61CB9-A48A-4BC0-84FE-61AFA198870D Material examined. Holotype. ♀ INDIA: Karnataka, Chamarajanagar, Biligiri Ranganathaswamy Temple Tiger Reserve, Malaki Betta, 11°57.230′ N, 77°08.241′ E, 1306 m a.s.l., scrub jungle forest, Malaise trap, 31.v.2005, coll. Priyadarsanan, D. R. (AIMB). Paratypes: 4 females with same label data, one female INDIA: Tamil Nadu, Kalakad Mundanthurai Tiger Reserve (KMTR), primary forest, understorey, Malaise trap, 3.xii.2008, coll. Priyadarsanan, D. R. (AIMB). Description. Female Body measurements. Length of body 2.1 mm, length of fore wing 2.1 mm. Head. Head 1.3 and 1.9 × as wide as long in anterior and dorsal views respectively. Scape 1.6 × as long as wide. Pedicel 1.3 × as long as wide. First flagellomere 1.2 × as long as second flagellomere. First, second and penultimate flagellomeres 6.0, 5.3 and 1.3 × as long as wide. Medial flagellomere as long as wide (Fig. 1D). Fourth segment of maxillary palp 1.5 × as long as third segment. Face 1.3 × as wide as long, smooth, setose. Clypeus 2.1 × wider ventrally than medially high with convex posterior margin (Fig. 1B). Length of malar space 1.7 × basal width of mandible. OOL: transverse diameter of posterior ocellus: POL= 2.5: 1.3: 1.0. Length of eye 2.0 × as long as temple in dorsal view. Mesosoma. Mesosoma 1.5 × as long as high. Scutellar sulcus narrow, slightly wavy (Fig. 1F). Scutellum glabrous medially (Fig. 1F). Propodeum smooth medially and laterally (Fig. 2A). Wings. Fore wing: Pterostigma 2.0 × as long as wide. Vein r 0.3 × as long as 2RS. Vein (RS+M)b 0.8 × as long as 1m-cu. Vein 1-M 1.5 × as long as m-cu. Vein (RS+M)a 1.9 × (RS+M)b. Vein 1CUa 0.6 × as long as 1CUb. Hind wing: Veins M+CU: 1M: 1r-m= 2.1: 2.4: 1.0. Legs. Ratio of fore femur: fore tibia: fore tarsus = 1.0: 1.1: 1.1. Fore femur and tibia 3.5 and 6.3 × as long as wide, respectively. Hind femur, tibia and basitarsus 3.5, 5.4 and 4.0 × as long as wide, respectively. Metasoma. First tergite 5.4 × as long as its maximum width, 4.5 × as long as basal width, 1.8 × as long as second tergite. Second tergite 0.7 × as long as third tergite. Ovipositor straight (Fig. 2C). Ovipositor sheath 0.1 × as long as fore wing. Colour. Body brown, dark brown or reddish brown but face, clypeus, mandible except apically, malar space, maxillary and labial palps, scape and pedicel, flagellomeres 10–12, temples posteriorly, ocelli, pronotum, propleuron, tegula, pterostigma, wing veins except 1CUa, legs (except ivory coxae), metasomal tergites except medial sclerotised part of T1 and T2, ovipositor and sheath basally yellow. Male. Unknown. Etymology. The species is named after the high chances for finding more new species of Miracinae considering the very limited study of this subfamily. Biology. Unknown. Distribution. India (Karnataka).

Centistidea burwelli Ranjith & Priyadarsanan, sp. nov. (Figures 3–4) urn:lsid:zoobank.org:act: 4FD49EDF-4241-4858-B23A-88E98D92E3DD Material examined. Holotype. ♀ SRI LANKA: Sigiriya, 7°57’ N, 80°46’ E, 20.iii.1999, coll. C.J. Burwell. Paratype, 1 male same data as holotype. Description. Female Body measurements. Length of body 2.2 mm, length of fore wing 2.0mm. Head. Antenna with 14 antennomeres, all antennomeres longer than wide (Fig. 3A). First flagellomere 1.1 × as long as second flagellomere. First, second and penultimate flagellomeres 5.3, 5.0 and 2.0 × as long as wide. Maxillary palp with four segments (Fig. 3D), fourth segment 1.2 × as long as third segment. Face distinctly convex medially (Fig. 3B), 1.6 × as wide as long, smooth, setose. Clypeus 2.0 × wider ventrally than medially high with convex posterior margin (Fig. 3B). Malar suture distinct (Figs 3B, D). Length of malar space 1.2 × basal width of mandible. Mandible distinctly twisted (Fig. 3B), inner tooth distinctly shorter than outer tooth. Frons smooth, moderately convex medially, sparsely setose laterally (Fig. 3C). Vertex smooth, sparsely setose, setose laterally (Fig. 3C). OOL: transverse diameter of posterior ocellus: POL= 2.5: 1.25: 1.0. Length of eye 1.6 × as long as temple in dorsal view. Mesosoma. Mesosoma 1.5 × as long as high. Pronotum and propleuron smooth laterally, lateral pronotum shallowly impressed medially (Fig. 3D). Mesoscutum punctate, setose, with carinae laterally (Fig. 3E). Notauli only impressed anteriorly (Fig. 5E). Scutellar sulcus represented only by a transverse groove, not crenulated (Fig. 3E). Scutellum evenly setose, smooth (Fig. 3E), with a pair of large pits medio-posteriorly separated by a carina, sides of scutellum irregularly sculptured (Fig. 3F). Mesopleuron smooth, glabrous, sparsely setose ventrally (Fig. 3D). Metapleuron with a deep small pit antero-dorsally, with deep shallow longitudinal depressed line, glabrous (Fig. 3D). Propodeum with a distinct and complete medio-longitudinal carina absent behind costula, dorsal part of midlongitudinal carina associated with irregular sculpture, faintly crenulated near costula, sparsely setose, antero-lateral part with shallow declivity, postero-lateral part faintly crenulated, posterior part well differentiated from dorsal part, posterior transverse carina distinct (Fig. 3F). Wings. Fore wing: Vein r distinct, 0.2 × as long as 2RS (Fig. 4C). R1 indistinct (Fig. 4C). Vein (RS+M)b 0.6 × as long as 1m-cu. Vein 1-M 1.5 × as long as m-cu. Vein (RS+M)a 2.9 × (RS+M)b. Vein 1CUa thickened, 0.9 × as long as 1CUb (Fig. 4C). Hind wing: Vein cu-a straight. Veins M+CU: 1M: 1r-m= 2.3: 2.4: 1.0. Legs. Hind coxa smooth, setose (Fig. 4A). Tarsal claws without basal lobe. Hind femur, tibia and basitarsus 3.5, 6.8 and 3.8 × as long as wide respectively. Metasoma. First tergite smooth, narrow medially, widening subapically, weakly narrowing apically, distinctly narrowing basally, 3.5 × as long as its maximum width (Fig. 4B). Second tergite longitudinally striate, smooth medially, setose, medial length 0.7 × as long as third tergite (Fig. 4B). Third tergite longitudinally striate, setose (Fig. 4B). Ovipositor straight. Ovipositor sheath 0.09 × as long as fore wing, slightly narrowing apically with long, sparse setae apically (Fig. 4A). Colour. Body reddish brown except head except ocellar region and apex of mandibles, maxillary and labial palps, propleuron, mesoscutum postero-laterally, scutellum medially, mesopleuron, metapleuron, legs, pterostigma, latero-tergites of first metasomal segment, metasomal sternites yellow. Male. Same as female. Etymology. The species is named after the collector Dr Chris J. Burwell, Senior Curator of Insects, Queensland Museum, Australia. Biology. Unknown. Distribution. Sri Lanka (Central Province, Sigiriya). Comparative diagnosis. The new species comes close to C. cosmopteryxi Ahmad & Pandey in having colouration of the body, first flagellomere 1.1 × as long as second flagellomere, yellow head, fore wing vein R1 distinctly vein like. However, the new species differs from the latter in having the following characters, propodeum sculptured medially near midlongitudinal carina (smooth in C. cosmopteryxi), penultimate flagellomere 2.0 × as long as wide (2.5–3.0 × in C. cosmopteryxi), hind basitarsus 3.8 × as long as wide (4.5 × in C. cosmopteryxi) and mesopleuron metapleuron yellow (brown in C. cosmopteryxi).

Centistidea lanka Ranjith & Priyadarsanan, sp. nov. (Figures 9–10) urn:lsid:zoobank.org:act: B3E6CA1E-AC20-45E2-8A9B-C78C2822D22B Material examined. Holotype. ♀ SRI LANKA: Sigiriya, 7°57’ N, 80°46’ E, 21.iii.1999, coll. C. J. Burwell (AIMB). Description. Female Body measurements. Length of body 1.7 mm, length of fore wing 1.6 mm. Head. Antenna with 14 antennomeres, all antennomeres longer than wide (Fig. 9A). First flagellomere 1.1 × as long as second flagellomere. First, second and penultimate flagellomeres 4.0, 3.8 and 2.8 × as long as wide. Maxillary palp with four segments (Figs 9B, D), fourth segment 1.4 × as long as third segment. Face distinctly convex medially (Fig. 9B), 1.6 × as wide as long, smooth, setose. Clypeus 2.3 × wider ventrally than medially high with convex posterior margin (Fig. 9B). Malar suture distinct (Figs 9B, D). Length of malar space 1.4 × basal width of mandible. Mandible distinctly twisted (Fig. 9B), inner tooth distinctly shorter than outer tooth. Frons smooth, moderately convex medially, sparsely setose laterally (Fig. 9C). Vertex smooth, sparsely setose, setose laterally (Fig. 9C). OOL: transverse diameter of posterior ocellus: POL= 2.5: 1.0: 1.0. Length of eye 1.9 × as long as temple in dorsal view. Mesosoma. Mesosoma 1.4 × as long as high. Pronotum and propleuron smooth laterally, lateral pronotum shallowly impressed medially (Fig. 9D). Mesoscutum granulate, setose, without carinae laterally (Fig. 9E). Notauli only impressed anteriorly (Fig. 9E). Scutellar sulcus represented only by a transverse groove, not crenulated (Fig. 9E). Scutellum evenly setose, smooth (Fig. 9E), with a pair of large pits medio-posteriorly separated by 0.25 × diameter of pit, sides of scutellum irregularly sculptured (Fig. 9F). Mesopleuron smooth, glabrous, sparsely setose ventrally (Fig. 9D). Metapleuron with a deep small pit antero-dorsally, with deep shallow longitudinal depressed line, glabrous (Fig. 9D). Propodeum with a distinct and complete medio-longitudinal carina absent behind costula, dorsal part of midlongitudinal carina associated with irregular sculpture, sparsely setose, antero-lateral part with shallow declivity, postero-lateral part faintly crenulated, posterior part well differentiated from dorsal part, posterior transverse carina distinct (Fig. 9F). Wings. Fore wing: Vein r distinct, 0.3 × as long as 2RS (Fig. 10C). R1 indistinct (Fig. 10C). Vein (RS+M)b 0.8 × as long as 1m-cu. Vein 1-M 1.6 × as long as m-cu. Vein (RS+M)a 1.9 × (RS+M)b. Vein 1CUa thickened, 1.2 × as long as 1CUb (Fig. 10C). Hind wing: Vein cu-a straight. Veins M+CU: 1M: 1r-m= 1.7: 1.8: 1.0. Legs. Hind coxa smooth, setose (Figs 9A, 10A). Tarsal claws without basal lobe. Hind femur, tibia and basitarsus 3.1, 6.7 and 3.6 × as long as wide respectively. Metasoma. First tergite smooth, narrow medially, widening subapically, weakly narrowing apically, distinctly narrowing basally, 2.7 × as long as its maximum width (Fig. 10B). Second tergite smooth, setose, medial length 0.3 × as long as third tergite (Fig. 10B). Third tergite longitudinally striate, setose (Fig. 10B). Ovipositor straight. Ovipositor sheath 0.1 × as long as fore wing, slightly narrowing apically with long sparse setae apically (Fig. 10A). Colour. Body yellow except antenna, apex of mandibles, propodeal carina, dorsal part of metasomal segments 1–2, apical half of ovipositor sheath brown. Male. Unknown. Etymology. The species is named after the country, Sri Lanka where the specimens were collected. Biology. Unknown. Distribution. Sri Lanka (Central Province, Sigiriya). Comparative diagnosis. This new species is similar to the other species having hind basitarsus less than 4.0 × as long as wide. Centistidea lanka sp. nov. can be differentiated from the group of species by 1CUa is longer than 1Cub. Based on the additional characters the new species comes close to C. procarinator. Centistidea lanka sp. nov. differs from C. procarinator by the following characters, hind basitarsus 3.6 × as long as wide (4.5 × in C. procarinator), propodeum yellow (brown in C. procarinator), penultimate flagellomere 2.8 × as long as wide (2.3 × in C. procarinator), dorsal length of eye 1.9 × as long as temple (3.1 × in C. procarinator), metasomal tergite yellow (brown in C. procarinator) and second tergite shorter than third tergite (longer in C. procarinator).

Two new species of the genus Cratospila Foerster, 1863 (Braconidae: Alysiinae), Cratospila albosignatasp. nov. and C. longivenasp. nov., are described and illustrated. In addition, the DNA barcode region of the mitochondrial cytochrome c oxidase subunit I (COI) of both species has been sequenced with three previously described species (C. albifera, C. luteocephala and C. syntoma). Alysia ponerola Papp, 2009 which was recorded from North Korea is transfered in Cratospila (C. ponerola (Papp, 2009) comb. nov.). All species validly recorded from Korea are included in a revised key.

Blacus (Contochorus) mellitarsis van Achterberg, 1988 (Figs 3–4) Blacus (Contochorus) mellitarsis van Achterberg, 1988: 77. Material examined. 1 ♀, China, Hainan, Diaoluo Mtn., 16–17. XII.2006, Weng Liqiong, No. 200802539 (ZJUH). Distribution. China (Hainan); Indonesia. New record for China., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, pp. 277-287 in Zootaxa 5306 (2) on page 281, DOI: 10.11646/zootaxa.5306.2.7, http://zenodo.org/record/8058845, {"references":["van Achterberg, C. (1988) Revision of the subfamily Blacinae Foerster (Hymenoptera: Braconidae). Zoologische Verhandelingen Leiden, 249, 1 - 324."]}

Blacus (Contochorus) capillosus sp. nov. (Figs 1–2) Material examined. Holotype: ♀, China, Shaanxi, Qinling, Tiantai Mtn, 4.IX.1999, Chen Xuexin, No. 991398 (ZJUH). Paratype: 1♀, China, Shaanxi, Qinling, Tiantai Mtn, 3.IX.1999, He Junhua, No. 990176 (ZJUH). Description. Female. Length of body 2.4 mm, length of fore wing 2.35 mm. Body setosity. Body with dense setae, although sparser on mesopleuron. Head. Width of head in dorsal view 1.4 × its length (Fig. 2A); antennomeres 16 (Fig. 2C); length of first flagellomere 1.3 × length of second flagellomere; length of first, second and penultimate flagellomeres 4.6, 4.2 and 1.6 × their width, respectively; length of maxillary palp 1.0 × height of head; OOL: diameter of posterior ocellus: POL = 22:9:11; eyes with short dense setae (Fig. 2A); length of eye in dorsal view 0.85 × length of temple; occiput almost level (Fig. 2A); face with dense setae; temple with dense setae; malar suture present, length of malar space 1.75 × basal width of mandible. Mesosoma. Length of mesosoma 1.45 × its height; pronotal sides coarsely reticulate ventrally, dorsally smooth; area behind prepectal carina less conspicuously crenulate (Fig. 2E); precoxal sulcus narrow, with a few very short crenulae (Fig. 2E); notauli absent (Fig. 2D), without transverse carinae anteriorly; scutellar sulcus with medio-longitudinal carina (Fig. 2D); scutellum smooth, its lateral carina present, not protruding apically; surface of propodeum smooth between carinae, its medial area rectangular (Fig. 2F). Wings. Fore wing: first discal cell widely truncate anteriorly (Fig. 2H); vein r 1.2 × longer than maximum width of pterostigma; 1-CU1:2-CU1 = 13:21 (Fig. 2H). Legs. Hind coxa smooth, but with a distinct carina dorsally (Fig. 2F); length of femur, tibia and basitarsus of hind leg 6.2, 9.9 and 11.0 × their width, respectively; fore and mid claws with bristles (Figs 2I, 2J). Metasoma. Length of first tergite 2.6 × its apical width, almost parallel-sided, its surface rugulose, dorsal carinae complete (Fig. 2F); second tergite rugose; length of ovipositor sheath 0.10 × as long as fore wing. Colour. Body reddish-brown; palpi, mandible, tegulae, fore and middle legs, hind coxa and trochanter yellow; pterostigma, parastigma, veins, ovipositor, ovipositor sheath and all basitarsus brown; bristles of fore and mid claws blackish; wing membrane slightly infuscate. Variation. Length of first tergite 2.4–2.6 × its apical width. Male. Unknown. Biology. Unknown. Distribution. China (Shaanxi). Etymology. The specific name capillosus refers to the setose body and is derived from “ capillosus ” (Latin for hairy). Notes. The new species is similar to B. (C). turbidus Papp, 1985. Both species have medial area of propodeum rectangular, first discal cell of fore wing widely truncate anteriorly, and area behind prepectal carina less conspicuously crenulate than B. (C). mellitarsis and B. (C). epomidus but B. (C). capillosus differ by having 16 antennomeres of female (21–23 in B. (C). turbidus), lateral carina of scutellum not protruding apically (strongly protruding apically in B. (C). turbidus), mid claws of female with blackish bristles (only setose in B. (C). turbidus), length of eye of ♀ in dorsal view 0.85 × temple (1.3–1.6 × in B. (C). turbidus) and notauli absent (complete in B. (C). turbidus)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, pp. 277-287 in Zootaxa 5306 (2) on pages 278-279, DOI: 10.11646/zootaxa.5306.2.7, http://zenodo.org/record/8058845, {"references":["Papp, J. (1985) Three new Blacus Nees, 1818 species from India (Hym., Braconidae: Blacinae). Annales historico-naturales Musei nationalis hungarici, Zoology, 77, 227 - 232."]}

Blacus (Contochorus) planus sp. nov. (Figs 5–6) Material examined. Holotype: ♀, China, Hunan, Shimen, Huping Mtn, Xiangbizigou, 8.VII.2009, Ma Li, No. 200903023 (ZJUH). Paratype: 1♀, China, Zhejiang, Anji, Longwang Mtn, 21.VII.1995, Wu Hong, No. 971319 (ZJUH). Description. Female. Length of body 2.1 mm, length of fore wing 1.8 mm. Body setosity. Body with long pubescence, pubescence on head, side of pronotum and mesopleuron sparser than on remainder of body. Head. Head longitudinal and flat in dorsal view, width of head in dorsal view 1.1 × its length (Fig. 6A); antennomeres 16 (Fig. 6C); length of first flagellomere 1.2 × length of second flagellomere; length of first, second and penultimate flagellomeres 4.0, 3.3 and 1.4 × their width, respectively; length of maxillary palp 0.7 × height of head; OOL: diameter of posterior ocellus: POL = 16:6:7; eyes with short sparse setae (Fig. 6A); length of eye in dorsal view 0.8 × length of temple; occiput concave (Fig. 6A); face with dense long pubescence; temple smooth; malar suture weakly developed, length of malar space 1.5 × basal width of mandible. Mesosoma. Length of mesosoma 1.5 × its height; pronotal sides coarsely reticulate ventrally, dorsally smooth; area behind prepectal carina less conspicuously crenulate (Fig. 6E); precoxal sulcus narrow, with short crenulae (Fig. 6E); notauli absent posteriorly (Fig. 6D), impressed and crenulate anteriorly (Fig. 6D), without transverse carinae anteriorly; scutellar sulcus with medio-longitudinal carina (Fig. 6D); scutellum smooth, its lateral carina lamelliform, not protruding apically; surface of propodeum smooth between carinae, its medial area almost rectangular (Fig. 6F). Wings. Fore wing: first discal cell widely truncate anteriorly (Fig. 6I); vein r 0.6 × as long as maximum width of pterostigma; 1-CU1:2-CU1 = 1:3. Legs. Hind coxa smooth, but with a distinct carina dorsally (Figs 6G, 6H); length of femur, tibia and basitarsus of hind leg 5.0, 9.1 and 8.4 × their width, respectively; fore and mid claws with bristles (Figs 6J, 6L). Metasoma. Length of first tergite 2.9 × its apical width, parallel-sided, its surface rugulose, dorsal carinae complete (Fig. 6G); second tergite smooth; length of ovipositor sheath 0.13 × fore wing. Colour. Reddish-brown; palpi, mandible, tegulae, three basal segments of antenna, fore and middle legs, hind coxa and trochanter yellow; remainder of antenna, pterostigma, parastigma, veins, ovipositor, ovipositor sheath and all basitarsi brown; bristles of fore and mid claws blackish; wing membrane slightly infuscate. Variation. Precoxal sulcus with short to medium-sized striae. Male. Unknown. Biology. Unknown. Distribution. China (Hunan, Zhejiang). Etymology. The specific name planus refers to the rather flattened head and is derived from “ planus ” (Latin for flat). Notes. The new species is similar to B. (C). glaber van Achterberg, 1976.In both species the middle lobe of the mesoscutum lacks the strong carinae anteriorly, lateral carina of scutellum not strongly protruding apically, and middle claws with blackish bristles, but the two species differ by having 16 antennomeres of ♀ (17 in B. (C). glaber), notauli incomplete (complete in B. (C). glaber), medial area of propodeum rectangular and somewhat longer than wide (quadrangular, its height subequal to its width in B. (C). glaber), width of head in dorsal view 1.1 × its length (1.4 × in B. (C). glaber), length of eye in dorsal view 0.8 × length of temple (1.1 × in B. (C). glaber) and length of hind femur 5.0 × its width (7.0 × in B. (C). glaber).

Blacus (Contochorus) turbidus Papp, 1985 (Figs 7–8) Blacus (Ganychorus) turbidus Papp, 1985: 229. Blacus (Contochorus) turbidus: van Achterberg, 1988: 78; Papp, 1993: 159; Ahmad & Ahmed, 2008: 84. Material examined. 3♀♀, China, Hainan, Jianfengling, Tianchi, 25. XI.2008, Wang Manman, No. 200903082; Tan Jiangli, No. 200903085; Bawangling, 26.XI.2008, Wang Manman, No. 200805647; 1♀, China, Yunnan, Xishuangbanna, 28.XI.1993, Fan Xiang, No. 940417 (ZJUH). Distribution. China (Hainan, Yunnan); India; Nepal; Thailand. New record for China., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, pp. 277-287 in Zootaxa 5306 (2) on pages 285-287, DOI: 10.11646/zootaxa.5306.2.7, http://zenodo.org/record/8058845, {"references":["Papp, J. (1985) Three new Blacus Nees, 1818 species from India (Hym., Braconidae: Blacinae). Annales historico-naturales Musei nationalis hungarici, Zoology, 77, 227 - 232.","van Achterberg, C. (1988) Revision of the subfamily Blacinae Foerster (Hymenoptera: Braconidae). Zoologische Verhandelingen Leiden, 249, 1 - 324.","Papp, J. (1993) New braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 4. Annales historico-naturales Musei nationalis hungarici, 84, 155 - 180.","Ahmad, Z. & Ahmed, Z. (2008) Description of a new species of the genus Blacus Nees (Hymenoptera: Braconidae), along with a key to Indian species. Journal of the Bombay Natural History Society, 105 (1), 84 - 85."]}

Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu, Chen, Xue-Xin (2023): The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China. Zootaxa 5306 (2): 277-287, DOI: 10.11646/zootaxa.5306.2.7, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.7

Key to world species of subgenus Contochorus van Achterberg 1 Posterior half of notauli absent (Figs 2D, 6D); antennomeres of ♀ 16; length of eye of ♀ in dorsal view 0.8 × length of temple.............................................................................................. 2 - Posterior half of notauli complete (Figs 4D, 8D); antennomeres of ♀ 17–23; length of eye of ♀ in dorsal view 1.1–2.1 × length of temple............................................................................................ 3 2 Vein r of fore wing much shorter than maximum width of pterostigma (Fig. 6I); third antennomere yellow; head longitudinal and flat in dorsal view, width of head in dorsal view 1.1 × its length; notauli present anteriorly; vein 2-CU1 of fore wing 3 × longer than vein 1-CU1 (Fig. 6I); second tergite smooth...................................... B. ( C.) planus sp. nov. - Vein r of fore wing about as long as maximum width of pterostigma (Fig. 2H); third antennomere dark brown; width of head in dorsal view 1.4 × its length; notauli absent anteriorly; vein 2-CU1 of fore wing 1.6 × longer than vein 1-CU1 (Fig. 2H); second tergite rugose.................................................................... B. ( C.) capillosus sp. nov. 3 Antennomeres of ♀ 17; notauli anteriorly smooth, without transverse carinae; middle claw of ♀ with blackish bristles; medial area of propodeum quadrangular, its height subequal to its width; width of head in dorsal view 1.4 × its length; lateral carina of scutellum slightly protruding apically........................................ B. ( C.) glaber van Achterberg, 1976 - Antennomeres of ♀ 21–23; notauli with two transverse carinae anteriorly; middle claw of ♀ without blackish bristles; medial area of propodeum rectangular somewhat longer than wide; head transverse and flat in dorsal view, width of head in dorsal view 1.6–1.8 × its length; lateral carina of scutellum strongly protruding apically....................................... 4 4 First discal cell of fore wing widely truncate anteriorly; length of eye of ♀ in dorsal view 1.3–1.6 × length of temple; notauli comparatively narrow (Fig. 8D); area behind prepectal carina less conspicuously crenulate..... B. ( C.) turbidus Papp, 1985 - First discal cell of fore wing narrowly truncate anteriorly; length of eye of ♀ in dorsal view 1.7–2.1 × length of temple; notauli comparatively wide (Fig. 4D); conspicuously crenulate behind prepectal carina.................................... 5 5 Vein M+CU of hind wing shorter than vein 1-M; notauli widened anteriorly; eye of ♀ in dorsal view 2.0–2.1 × length of temple; hind tarsus of ♀ somewhat shorter than hind tibia; tarsi (pale) brownish-yellow....................................................................................................... B. ( C.) mellitarsis van Achterberg, 1988 - Vein M+CU of hind wing subequal to vein 1-M; notauli evenly wide anteriorly and posteriorly; eye of ♀ in dorsal view approx. 1.7 × length of temple; hind tarsus of ♀ as long as hind tibia; tarsi whitish-yellow...................................................................................................... B. ( C.) epomidus van Achterberg, 1988, Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, pp. 277-287 in Zootaxa 5306 (2) on page 278, DOI: 10.11646/zootaxa.5306.2.7, http://zenodo.org/record/8058845, {"references":["van Achterberg, C. (1976) A revision of the tribus Blacini (Hym., Braconidae, Helconinae). Tijdschrift voor Entomologie, 118, 159 - 322.","Papp, J. (1985) Three new Blacus Nees, 1818 species from India (Hym., Braconidae: Blacinae). Annales historico-naturales Musei nationalis hungarici, Zoology, 77, 227 - 232.","van Achterberg, C. (1988) Revision of the subfamily Blacinae Foerster (Hymenoptera: Braconidae). Zoologische Verhandelingen Leiden, 249, 1 - 324."]}

Blacus (Hysterobolus) xui sp. nov. (Figs. 15–16) Material examined. Holotype: ♀, Guangdong, Fuyuan, Nanling, 4.VIII.2004, Xu Zaifu, No. 20049864 (ZJUH). Paratypes: 3 ♀♀, Guangdong, Fuyuan, Nanling, 4.VIII.2004, Xu Zaifu, No. 20049813, 20049819, 20049936 (ZJUH). Description. Female. Length of body 2.9 mm, length of fore wing 2.4 mm. Body setosity. Body with long pubescence, pubescence on propodeum denser than the remainder. Head. Antennomeres 19; length of first flagellomere 1.7 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.75, 2.5, 1.5 and 2.7 × their width, respectively; length of maxillary palp 1.0 × height of head; OOL: diameter of posterior ocellus: POL = 11:4:5; frons smooth, with a longitudinal groove in upper second; eyes with short setose, sparse; length of eye in dorsal view 1.0 × temple; occiput concave; face transversely rugose; temple smooth; clypeus convex, slightly punctate, smooth; width of clypeus 2.6 × its medial height; tentorial pits large, distance between pits 2.1 × distance from pit to eye; malar suture absent, length of malar space 2.4 × basal width of mandible, mandible with long setae. Mesosoma. Length of mesosoma 1.4 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with long striae, reaching the anterior edge of mesopleuron; notauli complete, wide and crenulate; mesoscutal lobes rather convex, with long and soft pubescence; scutellar sulcus with distinct median carina and four carinae on both sides; scutellum reticulate, its lateral carina lamelliform; propodeal tubercles small, obtuse apically, lateral apical angle slightly less than 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 3.5 × its maximum width; length of vein r 1.1 × width of pterostigma; vein r arising from apical of pterostigma; 1-CU1:2-CU1 = 4:5. Hind wing: 1r-m:1-M = 19:25; M+CU subequal to1-M; 2-1A present and short, slightly branched down. Legs. Hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 4.8, 8.9 and 8.0 × their width, respectively; fore tarsal claws with brownish bristles. Metasoma. Length of first tergite 1.8 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.6; remainder of tergites smooth; length of ovipositor sheath 0.21 × length of fore wing. Colour. Brown; dorsal head (stemmaticum darker), scapus and dark brown; palpi, labrum, tegulae, fore and middle legs yellow; several basal flagellomeres brown, gradually paler towards apex, but several apical flagellomeres dark brown; parastigma and pterostigma basally white; scutellum reddish-brown; most mesoscutum black, partly yellowish-red; mesopleuron, propodeum and metasomal first tergite black; remainder of metasomal, ovipositor and sheath, veins and pterostigma brown; hind leg blackish-brown; all basitarsi apically brown; wing membrane slightly infuscate. Variation. Length of first tergite 1.7–1.8 × its apical width; length of apical flagellomeres 2.7–2.9 × their width, dorsal head dark brown to blackish; most mesoscutum dark brown to black, partly yellowish-red to reddish-brown; hind leg dark brown to blackish-brown. Male. Unknown. Biology. Unknown. Distribution. China (Guangdong). Etymology. The specific name refers to the collector of the species, the late Prof. Zai-fu Xu, who was a wellknown specialist of Chrysidoidea. Notes. The new species is similar to B. (H.) fuscitibialis van Achterberg,1988 because the scutellum is coarsely rugose, the hind tibia infuscated medially, the first metasomal tergite widened apically, the face transversely rugose, length of eye in dorsal view approx. 1.1 times temple, and fore claw with brownish bristles, but differs by having the precoxal sulcus with long striae (with a few medium-sized striae in B. fuscitibialis), dorsal carinae distinct in basal 0.6 of first tergite (basal 0.7), length of ovipositor sheath 0.21 × length of fore wing (0.18 ×), vein 2-SR of the fore wing 1.9 × longer than vein r (1.6 x), vein 1-CU1 oblique and nearly as long as vein 2-CU1 (horizontal and shorter than vein 2-CU1), and most of mesoscutum dark brown to black (mesoscutum reddish-brown)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 491-495, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330

Blacus (Hysterobolus) niger sp. nov. (Figs. 11–12) Material examined. Holotype: ♀, China, Yunnan, Baoshan, Gaoligongshan Nature park (24º50′N, 98º46′E), 11.V.2009, Wang Manman, No. 200904572 (ZJUH). Description. Female. Length of body 2.35 mm, length of fore wing 2.4 mm. Body setosity. Body with long pubescence, pubescence on mesopleuron sparse, propodeum and side of pronotum ventrally densely setose. Head. Antennomeres 19; length of first flagellomere 1.6 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.75, 3.0, 1.5 and 3.0 × their width, respectively; length of maxillary palp 1.1 × height of head; OOL: diameter of posterior ocellus: POL = 20:7:9; frons smooth, with a shallow longitudinal groove in upper second; eyes with short setose, sparse; length of eye in dorsal view 1.0 × temple; occiput slightly concave; face with shallow transversely rugae; temple smooth; clypeus convex, slightly punctate, smooth; width of clypeus 2.7 × its medial height; tentorial pits large, distance between pits 1.7 × distance from pit to eye; malar suture absent, length of malar space 1.8 ×basal width of mandible, mandible with long setae. Mesosoma. Length of mesosoma 1.35 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some medium-sized striae, only one stria reaching the anterior edge of mesopleuron; notauli complete and crenulate; mesoscutal lobes rather convex, with long and soft pubescence; scutellar sulcus with distinct median carina and two weakly developed carinae (Fig. 12D); scutellum reticulate, its lateral carina lamelliform; propodeal tubercles small, slightly acute apically, lateral apical angle slightly less than 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 3.5 × its maximum width; length of vein r 1.1 × width of pterostigma; vein r arising from apical of pterostigma; 1-CU1:2-CU1 = 2:3. Hind wing: 1r-m:1-M = 4:7; M+CU slightly less than 1-M; 2-1A absent. Legs. Hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.55, 9.6 and 7.1 × their width, respectively; fore tarsal claws with blackish bristles. Metasoma. Length of first tergite 1.8 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.5; remainder of tergites smooth; length of ovipositor sheath 0.23 × length of fore wing. Colour. Dark reddish-brown; dorsal head black; palpi, tegulae, ovipositor, fore and middle legs yellow; scape, clypeus reddish-brown; several basal flagellomeres yellowish-brown, gradually paler towards apex, but several subapical flagellomeres brown; mandible yellow; parastigma and base of pterostigma white; most of mesonotum black; metasomal tergite (except first) brown; ovipositor sheath, veins and pterostigma reddish-brown; hind legs coxa brown, femur and tibia blackish-brown; all apical basitarsus (basal yellow) and claws brown; wing membrane slightly infuscate. Male. Unknown. Biology. Unknown. Distribution. China (Yunnan) Etymology. The specific name refers to the black mesonotum and is derived from “ niger ” (Latin for black). Notes. The new species is similar to B. (H.) fuscitibialis van Achterberg,1988 because the scutellum is coarsely rugose, the malar suture absent, the hind tibia infuscated medially, the first metasomal tergite widened apically, the face transversely rugose, and the precoxal sulcus with medium-sized striae, but differs by having the fore claw with blackish bristles (brownish), dorsal carinae distinct in basal 0.5 (0.7), length of ovipositor sheath 0.23 × length of fore wing (0.18 ×), and the mesonotum black (mesoscutum reddish-brown)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 486-488, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330

Blacus (Hysterobolus) variegatus sp. nov. (Figs. 13-14) Material examined. Holotype: ♀, China, Chongqing, Jinfo Mtn, (29º3′N, 107º7′E), 18.VIII.2012, Huang Xinlei, No. 201207984 (ZJUH). Paratypes: 5 ♀ ♀, China, Zhejiang, Tianmu Mtn, 3-5.X.1983, Ma Yun, No. 834329; 25-29.VII.2011, yellow pan trap, No. 201501198; Chanyuan Temple, 9. V.1998, Zhao Mingshui, No. 20003401; Zhejiang, Fengyang Mtn, 1. VI.2005, Liu Jingxian, No. 200809039; Zhejiang, Longtang Mtn, 29. V.2012, Tang Pu, No. 201204960; 1 ♀, China, Hunan, Shimen, Huping Mtn, 12.VII.2009, Zeng Jie, No. 200903015; 1 ♀, China, Guizhou, Leigong Mtn, Tree Farm, 12.VII.2009, Zhang Hongying, No. 200606873 (ZJUH). Description. Female. Length of body 2.6 mm, length of fore wing 2.65 mm. Body setosity. Body with dense pubescence, pubescence on head and mesopleuron sparser, propodeum denser. Head. Antennomeres 18; length of first flagellomere 1.7 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.0, 2.4, 1.2 and 2.8 × their width, respectively; length of maxillary palp 1.1 × height of head; OOL: diameter of posterior ocellus: POL = 24:8:9; frons smooth, with a shallow longitudinal groove in upper second and five groove between antennal sockets; eyes with short setose, sparse; length of eye in dorsal view 0.9 × temple; occiput concave; face transversely rugose; temple smooth; clypeus convex, smooth; width of clypeus 2.4 × its medial height; tentorial pits large, distance between pits 1.55 × distance from pit to eye; malar suture absent, length of malar space 2.0 × basal width of mandible, mandible with long setae. Mesosoma. Pubescence on mesonotum sparser than remainder. Length of mesosoma 1.25 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some long striae, reaching the anterior edge of mesopleuron; notauli complete, wide and crenulate, with short transversely carina in anterior third; mesoscutal lobes rather convex, densely pubescent; scutellar sulcus with distinct median carina, and distinctly crenulate laterally; scutellum sculpture reduced and smooth medially, its lateral carina lamelliform, distinctly protruding apically; propodeal tubercles small, obtuse apically, lateral apical angle almost 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 5.0 × its maximum width; length of vein r 1.4 × width of pterostigma; vein r arising from apical of pterostigma; 1-CU1:2-CU1 = 13:15. Hind wing: 1r-m:1-M = 9:17; M+CU about subequal to 1-M; 2-1A absent. Legs. Fore leg tibial spur large and long; hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 4.8, 9.3 and 8.0 × their width, respectively; fore tarsal claws with blackish bristles. Metasoma. Length of first tergite 2.0 × its apical width, widened apically, medio-dorsally rugose, both sides with longitudinally striae; dorsal carinae distinct in basal 0.4; remainder of tergites smooth; length of ovipositor sheath 0.16 × length of fore wing. Colour. Dark reddish-brown; dorsal head, propodeum and metasomal first tergite black; palpi, tegulae, fore and middle legs yellow; several basal flagellomeres reddish-brown, gradually paler towards apex, but two apical flagellomeres dark brown; clypeus and mandible yellowish-red; parastigma and bottom margin of pterostigma white; side of pronotum yellowish-red, remainder reddish-brown; most mesonotum and scutellum yellowish-red, mesoscutum partly brown; metasomal second tergite yellowish-red, remainder of metasomal tergites and sternum, ovipositor and sheath, veins and pterostigma brown; hind leg (except second, third and fourth tarsus) reddish-brown; all basitarsi apically brown; wing membrane slightly infuscate. Variation. Length of first tergite 1.8–2.1 × its apical width; penultimate flagellomere 1.25–1.3 (occasionally 1.4) × its width; length of fore wing 2.4–2.65 mm; length of ovipositor sheath 0.16–0.18 × length of fore wing; propodeal tubercles medium size to large; scutellum sculpture reduced, at most slightly rugose; head and propodeum dark reddish-brown; two (or at most three) apical flagellomeres dark brown. Male. Unknown. Biology. Unknown. Distribution. China (Chongqing, Hunan, Zhejiang, Guizhou) Etymology. The specific name refers to the particular different difference in colour of head and mesonotum and is derived from “ varius” (Latin for different). Notes. The new species is similar to B. (H.) fuscitibialis van Achterberg, 1988 because the malar suture is absent, the face transversely rugose, and the first metasomal tergite widened apically, but differs by having the precoxal sulcus with long striae (with a few medium-sized striae in B. fuscitibialis), hind leg (except second, third and fourth tarsal flagellomeres) reddish-brown (only hind tibia infuscated medially), fore claw with blackish bristles (brownish bristles), length of penultimate flagellomere 1.2 × its width (1.5 ×), dorsal carinae distinct in basal 0.4 of tergite (basal 0.7), and length of ovipositor sheath 0.16 × length of fore wing (0.18 ×). This new species is also similar to B. (H.) melanoapicalis sp. nov., but differs by having 18 antennomeres (19 antennomeres in B. melanoapicalis), occiput comparatively slightly concave (distinctly concave), and the colour of head and mesonotum particularly different (similar)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 488-491, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330

Blacus (Hysterobolus) longiapicalis sp. nov. (Figs. 7–8) Material examined. Holotype: ♀, China, Liaoning, Laotudingzi National Nature Reserve (41º22′N, 124º57′E), 16-19.VII.2011, Chen Huayan, NO. 202027906 (ZJUH). Description. Female. Length of body 3.2 mm, length of fore wing 2.7 mm. Body setosity. Body with dense setae, setae on head sparser than on mesosoma. Head. Antennomeres 19; length of first flagellomere 1.6 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.8, 3.0, 1.2 and 3.0 × their width, respectively; length of maxillary palp 1.1 × height of head; OOL: diameter of posterior ocellus: POL = 12:4:7; frons smooth, with a deep longitudinal groove in upper half; stemmaticum with a distinct furrow from triangle centre to occipital carina; eyes with short sparse setae; length of eye in dorsal view 0.9 × temple; occiput concave; face transversely rugose; temple smooth; clypeus convex, slightly punctate, smooth; width of clypeus 2.3 × its medial height; tentorial pits large, distance between pits 2.4 × distance from pit to eye; malar suture absent, length of malar space 2.0 × basal width of mandible, mandible with long setae. Mesosoma. Length of mesosoma 1.5 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some long striae, reaching anterior edge of mesopleuron; notauli complete, wide and crenulate; mesoscutal lobes rather convex, densely pubescence; scutellar sulcus with distinct median carina and two weakly developed carinae; scutellum reticulate, its lateral carina lamelliform; propodeal tubercles large (Fig. 8H and 8K), obtuse apically, lateral apical angle slightly less than 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 3.5 × its maximum width; length of vein r 1.1 × width of pterostigma; vein r arising slightly from apical of pterostigma; 1-CU1:2-CU1 = 14:17. Hind wing: 1r-m:1- M = 9:17; M+CU about subequal to 1-M; 2-1A present and short, branched posteriorly. Legs. Hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.0, 11.25 and 8.75 × their width, respectively; fore tarsal claws with blackish bristles. Metasoma. Length of first tergite 1.5 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.5; remainder of tergites smooth; length of ovipositor sheath 0.16 × length of fore wing. Colour. Black; head (dorsal and frontal) black, palpi, labrum, tegulae, fore and middle legs yellow; scape, clypeus reddish-brown; several basal flagellomeres yellowish-brown, gradually lighten towards apex, but several apical flagellomeres dark brown; mandible yellow, apex reddish-brown; parastigma and base of pterostigma white; side of pronotum reddish-brown, remainder black; edges of mesoscutum reddish-brown, remainder of mesoscutum and scutellum black; mesopleuron, propodeum and first metasomal tergite black; second tergite and sternum yellow, remainder of metasomal tergites and sternum, ovipositor and sheath, veins and pterostigma brown; hind leg yellowish-brown; all basitarsi apically brown; wing membrane slightly infuscate. Male. Unknown. Biology. Unknown. Distribution. China (Liaoning). Etymology. The specific name refers to the long apical flagellomere is derived from “ longus ” (Latin for long) and “ apex ” (Latin for top). Notes. The new species is similar to B. (H.) robustus Haeselbarth, 1973 because the scutellum is coarsely rugose, the malar suture absent, the propodeal tubercle rather large, length of eye in dorsal view 0.9 × temple, and fore claw with blackish bristles, but differs by having malar space 2.0 × longer than basal width of mandible (1.7 × in B. robustus), length of first and second flagellomeres 4.8 and 3.0 × their width, respectively (4.1 and 2.6 ×), dorsal carinae of first tergite distinct in basal 0.5 (in basal 0.3), and hind leg yellowish-brown (femur, tibia and tarsus of hind leg darkened apically, base of hind tibia yellowish).

Blacus (Hysterobolus) chabarovi Belokobylskij, 1995 (Figs. 19–20) Hysterobolus chabarovi Belokobylskij, 1995: 458; Lee et al., 2020: 32. Material examined. 1 ♀, China, Chongqing, Jinfo Mt. (29º3′N, 107º7′E), 19.VIII.2012, Huang Xinlei, No. 201209624; 2 ♀♀, China, Ningxia, Longde, Liupanshan Nature Park, Sutai Tree Farm, 24. VI.2008, No. 200903090, 200903097; 1 ♀, Ningxia, Jingyuan, Liupanshan Nature Reserve, Hongxia Tree Farm, 1.VII.2008, Liu Jingxian, No. 200902112 (ZJUH). Distribution. China (Chongqing, Ningxia); Korea; Russia. New record for China. Remarks. The characters are largely consistent as described by Belokobylskij (1995), but the fore tarsal claws have weakly developed bristles., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on page 476, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330, {"references":["Belokobylskij, S. A. (1995) New and rare species of the genus Blacus (Hymenoptera: Braconidae) from the Russian Far East. European Journal of Entomology, 92 (2), 449 - 467.","Lee, H. - R., Belokobylskij, S. A., Ku, D. - S. & Byun, B. - K. (2020) Eight newly recorded species of the genus Blacus (Hymenoptera: Braconidae: Blacinae) in Korea. Animal Systematics Evolution and Diversity, 36 (1), 31 - 34."]}

Blacus (Hysterobolus) flaviceps sp. nov. (Figs. 3-4) Material examined. Holotype: ♀, China, Yunnan, Baoshan, Gaoligongshan Nature Park (24º50′N, 98º46′E), 10.V.2009, Wang Manman, No. 200904546 (ZJUH). Description. Female. Length of body 2.6 mm, length of fore wing 2.6 mm. Body setosity. Body with long pubescence, entire head with long setae, pubescence on mesonotum sparser than on remainder of mesosoma. Head. Antennomeres 20; length of first flagellomere 1.7 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.4, 2.9, 1.75 and 3.3 × their width, respectively; length of maxillary palp subequal to height of head; OOL: diameter of posterior ocellus: POL = 11:4:3; frons smooth, with a shallow longitudinal groove in upper half; stemmaticum smooth; eyes with short sparse setae; length of eye in dorsal view 0.8 × temple; occiput concave; face transversely rugose; temple smooth; clypeus convex, slightly punctate; width of clypeus 2.5 × its medial height; tentorial pits large, distance between pits twice distance from pit to eye; malar suture absent, length of malar space 2.8 × basal width of mandible. Mesosoma. Length of mesosoma 1.4 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some long striae, reaching anterior edge of mesopleuron; notauli complete, wide and crenulate; mesoscutal lobes rather convex, densely pubescence; scutellar sulcus with median carina; scutellum reticulate, its lateral carina lamelliform; propodeal tubercles small, obtuse apically, lateral apical angle about 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 3.75 × its maximum width; length of vein r 1.0 × width of pterostigma; vein r arising almost from middle of pterostigma; 1-CU1:2-CU1 = 6:7. Hind wing: 1r-m:1-M = 1:2; M+CU slightly short than 1-M; 2-1A present and short, branched down. Legs. Hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.8, 11.4 and 10.0 × their width, respectively; fore tarsal claws missing. Metasoma. Length of first tergite 1.8 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.5; second tergite smooth; length of ovipositor sheath 0.17 × length of fore wing. Colour. Dark reddish-brown; head (dorsal and anterior), scape, palpi, tegulae, hind leg (except tibia) and ovipositor yellow; antenna yellow, gradually darkened towards apex; stemmaticum, pterostigma (basally paler), parastigma, veins, ovipositor sheath brown; edges of mesoscutum yellow, remainder of mesoscutum and scutellum reddish-brown, reddish-brown area on middle lobes remain separated from each other; dorsal side of pronotum yellow, remainder of pronotum, mesopleuron, propodeum and metasomal first tergite reddish-brown; second metasomal tergite and sternum yellow, remainder of metasomal tergites blackish; fore and middle legs light yellow; all basitarsi dark brown; wing membrane slightly infuscate. Male. Unknown. Biology. Unknown. Distribution. China (Yunnan). Etymology. The specific name refers to the yellow head and is derived from “ flavus ” (Latin for yellow) and “ caput ” (Latin for head). Notes. The new species differs from other species by having a yellow head, 20 antennomeres, length of basitarsus of hind leg 10.0 × its width, and length of vein r of fore wing almost equal to width of pterostigma., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 476-477, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330

Blacus (Hysterobolus) yunnanensis sp. nov. (Figs. 11–12) Material examined. Holotype: ♀, China, Yunnan, Ruili (24º1′N, 97º51′E), 29. IV.1981, He Junhua, NO. 812372 (ZJUH). Description. Female. Length of body 2.6 mm, length of fore wing 2.5 mm. Body setosity. Body with long setae, mesopleuron and side of pronotum glabrous, propodeum denser setose than head and mesonotum. Head. Antenna with at least 18 antennomeres (apex missing); length of first flagellomere 1.8 × second flagellomere; length of first and second flagellomere 4.9 and 2.7 × their width, respectively; length of maxillary palp 1.1 ×height of head; OOL:diameter of posterior ocellus:POL=20:8:11;frons with sparse setae and a deep longitudinal groove in upper second; eyes with short setose, sparse; length of eye in dorsal view 1.2 × temple; occiput slightly concave; face almost entirely smooth; temple smooth; clypeus convex, slightly punctate, smooth; width of clypeus 2.5 × its medial height; tentorial pits large, distance between pits 2.7 × distance from pit to eye; malar suture present, length of malar space 2.0 × basal width of mandible, mandible with long setae. Mesosoma. Length of mesosoma 1.35 × its height; side of pronotum largely smooth dorsally; precoxal sulcus narrow, with a few short striae, far from the anterior edge of mesopleuron (Fig. 18G); notauli complete, wide and crenulate; mesoscutal lobes rather convex, with long and soft pubescence; scutellar sulcus with median carina; scutellum (superficially) rugulose, its lateral carina lamelliform, distinctly protruding apically; propodeal tubercles small, acute apically, lateral apical angle slightly more than 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 3.75 × its maximum width; length of vein r 1.2 × width of pterostigma; vein r arising from apical of pterostigma; 1-CU1:2-CU1 = 12:13. Hind wing: 1r-m:1-M = 20:27; M+CU almost subequal to 1-M; 2-1A absent. Legs. Hind coxa slightly rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.9, 11.5 and 7.1 × their width, respectively, fore tarsal claws with blackish bristles. Metasoma. Length of first tergite 1.6 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.7; remainder of tergites smooth; length of ovipositor sheath 0.14 × length of fore wing. Colour. Reddish-brown; dorsal head black; palpi, tegulae, parastigma, fore and middle legs yellow; medial flagellomeres lighter in colour; mandible yellow, apex red; veins, pterostigma, ovipositor sheath, and metasomal tergite (except first) brown; ovipositor, hind legs coxa, femur and telotarsus yellowish-brown, tibia brown; all apical basitarsus (basal yellow) and claws brown; wing membrane slightly infuscate. Male. Unknown. Biology. Unknown. Distribution. China (Yunnan) Etymology. The specific name refers to the locality of the holotype. Notes. The new species is similar to B. (H.) nixoni Haeselbarth, 1973 because the malar suture is present, the scutellum (superficially) rugulose, the propodeal tubercle comparatively small, length of first tergite 1.6 × its apical width, flagellomeres long setose, and face almost entirely smooth, but differs by having the eighth flagellomere 2.2 × longer than wide (about 1.7 × in B. nixoni), length of malar space of 2.0 × basal width of mandible (1.5 ×), dorsal carinae distinct in basal 0.7 (0.9 ×), precoxal sulcus with less striae (with more striae), and body reddish-brown (black)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 495-497, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330, {"references":["Belokobylskij, S. A. (1995) New and rare species of the genus Blacus (Hymenoptera: Braconidae) from the Russian Far East. European Journal of Entomology, 92 (2), 449 - 467."]}

The subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Blacus) is newly recorded from China. Nine new species are proposed, B. (H.) baishanzuensis sp. nov., B. (H.) flaviceps sp. nov., B. (H.) laculaticornis sp. nov., B. (H.) longiapicalis sp. nov., B. (H.) melanoapicalis sp. nov., B. (H.) niger sp. nov., B. (H.) variegatus sp. nov., B. (H.) xui sp. nov., B. (H.) yunnanensis sp. nov., and one species, B. (H.) chabarovi Belokobylskij, 1995, is newly recorded from China. An updated key to the currently known species of the subgenus Hysterobolus in China is provided.

Blacus (Hysterobolus) melanoapicalis sp. nov. (Figs. 9–10) Material examined. Holotype: ♀, China, Zhejiang, Tianmu Mtn, Xianren Peak (30º21′N, 119º25′E), 27.VII.2011, Song Shengnan, NO. 201102181 (ZJUH). Paratypes: 49 ♀♀, China, Fujian, Wuyi Mtn, Guadun, 1.VII.1985, Huang Donghong, No. 20004198, 20003902, 20003903; 2.VII.1985, Huang Donghong, No. 20003901, 20003902, 20003903, 20003906, 20003907; 25.VII.1985, Huang Donghong, No. 20004200; Fujian, Wuyi Mtn, 20.VII.1985, Lin Naiquan, No. 968113, 968114, 968122, 968125, 968127, 968130, 968146, 968154, 968155, 968157; Fujian, Wuyi Mtn, Erliping, 2. VI.1985, Tang Yuqing, No. 20004172; Fujian, Huanggang Mtn, VI.1980, Huang Juchang, No. 20003957, 20003958, 20003959, 20004127; Zhao Jingwei, No. 20003968, 20003969, 20003971, 20003973; Zhao Xiufu, No. 20004126, 20004128; IX.1981, Huang Juchang, No. 20003960; 9. V.1982, Xu Jianfei, No. 20003780; 28.VII.1984, Huang Juchang, No. 20004129; 6.VII.1985, Chen Xinjin, No. 20004134; Liu Minghui, No. 20004132; 13.VII.1985, Huang Donghong, No. 20004096; 14.VII.1985, Huang Donghong, No. 20004097, 20004098; Tang Yuqing, No. 20004107; Zheng Geng, No. 20004102, 20004103; 1.VIII.1985, Huang Donghong, No. 20003775, 20003960, 20004092; 30.VII.1985, Chen Xinjin, No. 20004113, 20004115, 20004116, 20004118; Liu Minghui, No. 20004123; 3 ♀♀, Fujian, Fuzhou, 22.IX.1984, Wang Jiadong, No. 20004150; 8.X.1984, Wang Jiadong, No. 20004155; Jinshan Park, 16.VII.1984, Liu Changming, No. 20004217; 1 ♀, China, Zhejiang, Tianmu Mtn, 8. VI.1992, Lin Wei, No. 922874; 9 ♀♀, Zhejiang, Tianmu Mtn, Xianren Peak, 2-4. VI.1990, Lou Yonggen, No. 900905; 29.VII.1998, Zhao Mingshui, No. 993644; 30.VII.1998, Zhao Mingshui, No. 993531; 18.VIII.1999, Ma Yun, No. 997608; 25-29.VII.2011, Liu Zhen, No. 201102647; 26.VII.2011, Song Shengnan, No. 201101970; 27.VII.2011, Song Shengnan, No. 201102181; Liu Zhen, No. 201101425, 201101431 (ZJUH). Description. Female. Length of body 2.8 mm, length of fore wing 2.75 mm. Body setosity. Body with long setae, setosity on mesopleuron and propodeum denser. Head. Antennomeres 19; length of first flagellomere 1.4 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.0, 2.8, 1.45 and 2.8 × their width, respectively; length of maxillary palp 1.2 × height of head; OOL: diameter of posterior ocellus: POL = 11:4:7; frons smooth, with a shallow longitudinal groove in upper second; eyes with short setose, sparse; length of eye in dorsal view 1.2 × temple; occiput distinctly concave; face transversely rugose; temple smooth; clypeus convex, smooth, slightly punctate; width of clypeus 2.3 × its medial height; tentorial pits large, distance between pits 1.9 × distance from pit to eye; malar suture absent, length of malar space 2.75 × basal width of mandible, mandible with long setae. Mesosoma. Length of mesosoma 1.4 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some long striae, reaching anterior edge of mesopleuron; notauli complete, wide and crenulate, with some short transversely rugae in anterior third; mesoscutal lobes rather convex; scutellar sulcus with median carina only; scutellum reticulate, its lateral carina lamelliform; propodeal tubercles small, obtuse apically, lateral apical angle almost 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 5.0 × its maximum width; length of vein r 1.5 × width of pterostigma; vein r arising almost from middle of pterostigma; 1-CU1:2-CU1 = 15:16. Hind wing: 1r-m:1- M = 5:7; M+CU about subequal to 1-M; 2-1A present and short, branched posteriorly. Legs. Fore leg tibial spur large and long; hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.7, 9.5 and 8.0 × their width, respectively; fore tarsal claws with blackish bristles. Metasoma. Length of first tergite 1.75 × its apical width, widened apically, rugose; dorsal carinae distinct in basal 0.4; remainder of tergites smooth; length of ovipositor sheath 0.15 × length of fore wing. Colour. Reddish-brown; stemmaticum, propodeum and metasomal first tergite dark reddish-brown; palpi, labrum, mandible, tegulae, all legs yellow; clypeus yellowish-red; scapus, 5 apical flagellomeres dark brown, remainder of flagellomeres brownish; parastigma and basal of pterostigma white; side of pronotum yellowish-red, remainder reddish-brown; most mesonotum and scutellum yellowish-red, consistent with dorsal head, mesoscutum companied with reddish-brown, the reddish-brown area on middle lobe separate from each other; metasomal second tergite yellowish-brown, remainder of metasomal tergites and sternum, ovipositor and sheath, veins and pterostigma brown; all basitarsi apically brown; wing membrane slightly infuscate. Variation. Length of first tergite 1.7–1.8 × its apical width; penultimate flagellomere 1.4–1.7 × their width; length of eye in dorsal view 1.0–1.2 × temple; length of fore wing 2.4–2.8 mm; length of ovipositor sheath 0.15– 0.18 × length of fore wing; dorsally head yellowish-red to reddish-brown, similar to mesonotum, propodeum and metasomal first tergite reddish-brown to blackish, remainder of metasomal tergites and sternum brownish to dark brown; 5 (or at most 6) apical flagellomeres dark brown. Male. Unknown. Biology. Unknown. Distribution. China (Zhejiang, Fujian) Etymology. The specific name refers to the dark apical flagellomeres and is derived from “ melanos ” (Greek for black) and “ apex ” (Latin for top). Notes. The new species is similar to B. (H.) fuscitibialis van Achterberg, 1988 because the scutellum is coarsely rugose, the malar suture absent, the first metasomal tergite widened apically, and the face transversely rugose, but differs by having the precoxal sulcus with long striae (only with a few medium-sized striae in B. fuscitibialis), the hind leg brownish (hind tibia infuscated medially), fore claw with blackish bristles (brownish), length of malar space 2.75 × basal width of mandible (2.0 ×), dorsal carinae distinct in basal 0.4 of tergite (basal 0.7), and length of ovipositor sheath 0.15 × length of fore wing (0.18 ×).

Key to Chinese species of subgenus Hysterobolus Viereck 1 Vein M+CU of hind wing distinctly shorter than vein 1-M; penultimate flagellomere quadrate, about as long as wide (Fig. 6C); length of malar space 3.0 × to basal width of mandible; first metasomal tergite about 2.1 × longer than its apical width (Fig. 6F)......................................................................... B. ( H.) laculaticornis sp. nov. - Vein M+CU of hind wing about subequal to vein 1-M; penultimate flagellomere distinctly longer than wide (Fig. 8D); length of malar space at most 2.8 × basal width of mandible; first tergite usually 1.5–1.8 × longer than wide apically (Figs 2H, 4H, 8E, 10J, 12F, 16F, 18E), except in B. variegatus (Fig.14E)........................................................ 2 2 Length of vein r of fore wing almost equal to width of pterostigma; antennomeres 20; length of hind basitarsus 10.0 × its width; head yellow....................................................................... B. ( H.) flaviceps sp. nov. - Length of vein r of fore wing 1.1–1.5 × width of pterostigma; antennomeres less than 20; length of hind basitarsus 7.1–8.8 × its width; colour of head variable........................................................................... 3 3 Malar suture present................................................................................... 4 - Malar suture absent.................................................................................... 5 4 Vein 2-CU1 of fore wing 1.2 × longer than vein 1-CU1 (Fig. 18F); face largely smooth; precoxal sulcus narrow, striae very short (Fig. 18G)..................................................................... B. ( H.) yunnanensis sp. nov. - vein 2-CU1 of fore wing 2.2 × longer than vein 1-CU1(Fig. 20I); face transversely rugose; precoxal sulcus wide, striae long (Fig. 20F)............................................................ B. ( H.) chabarovi Belokobylskij, 1995 5 Hind tibia yellow or reddish-brown....................................................................... 6 - Hind tibia dark brown or blackish-brown.................................................................. 8 6 First metasomal tergite 1.5 × longer than its apical width (Fig. 8E); length of pterostigma 3.5 × its maximum width; length of terminal flagellomere 3.0 × its width; body generally black............................. B. ( H.) longiapicalis sp. nov. - First tergite 1.7–2.1 × longer than its apical width; length of pterostigma 5.0 × its maximum width; length of terminal flagellomere 2.8 × its width; body generally reddish-brown............................................................... 7 7 Length of first flagellomere 1.7 × second flagellomere; scutellum smooth medially; antennomeres 18; occiput slightly concave (Fig. 14A); hind tibia reddish-brown; 2–3 apical flagellomeres dark brown; first metasomal tergite about 2.1 × longer than its apical width (Fig. 14E)............................................................ B. ( H.) variegatus sp. nov. - Length of first flagellomere 1.4 × second flagellomere; scutellum rugose medially; antennomeres 19; occiput distinctly concave (Fig. 10A); hind tibia yellow; 5–6 apical flagellomeres dark brown; first tergite 1.7–1.8 × longer than its apical width (Fig. 10J)........................................................................ B. (H.) melanoapicalis sp. nov. 8 Fore claw of ♀ with blackish bristles (Fig. 12I); face weakly rugose (Fig. 12B); striae of precoxal sulcus medium-sized; vein 2-1A of hind wing absent (Fig. 12H)...................................................... B. ( H.) niger sp. nov. - Fore claw of ♀ with brownish bristles (Figs 2J, 16I); face coarsely rugose (Figs 2B, 16B); striae of precoxal sulcus long; vein 2-1A of hind wing present (Figs 2M, 18F).................................................................. 9 9 Terminal flagellomere 2.7–2.9 × longer than wide; first discal cell of fore wing acute (Fig. 16H); setose part of ovipositor sheath slightly longer, 0.21 × length of fore wing; mesoscutum largely dark brown to black; hind femur dark brown to blackish-brown............................................................................... B. ( H.) xui sp. nov. - Terminal flagellomere about 2.2 × longer than wide; first discal cell of fore wing truncate (Fig. 2L); setose part of ovipositor sheath slightly shorter, 0.18 × length of fore wing; mesoscutum yellowish-brown; hind femur yellow............................................................................................. B. ( H.) baishanzuensis sp. nov., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on page 473, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330, {"references":["Belokobylskij, S. A. (1995) New and rare species of the genus Blacus (Hymenoptera: Braconidae) from the Russian Far East. European Journal of Entomology, 92 (2), 449 - 467."]}

Blacus (Hysterobolus) baishanzuensis sp. nov. (Figs. 1–2) Material examined. Holotype: ♀, China, Zhejiang, Qingyuan, Baishanzu, 18.VII.1994, Wu Hong, No. 946835 (ZJUH). Description. Female. Length of body 2.3 mm, length of fore wing 2.35 mm. Body setosity. Body with dense pubescence, pubescence on propodeum denser than on remainder. Head. Antennomeres 19; length of first flagellomere 1.6 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.75, 3.0, 1.3 and 2.2 × their width, respectively; length of maxillary palp 0.9 × height of head; OOL: diameter of posterior ocellus: POL = 11:3:5; frons smooth, with a very shallow longitudinal groove in upper half; eyes with short sparse setae; length of eye in dorsal view 1.0 × temple; occiput slightly concave; face transversely rugose; temple smooth; clypeus convex, slightly punctate, smooth; width of clypeus 3.0 × its medial height; tentorial pits large, distance between pits 2.3 × distance from pit to eye; malar suture absent, length of malar space 2.0 × basal width of mandible, mandible with setae. Mesosoma. Length of mesosoma 1.4 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some long striae, reaching the anterior edge of mesopleuron; notauli complete, wide and crenulate; mesoscutal lobes rather convex; scutellar sulcus with distinct median carina and four weakly developed carinae; scutellum convex, reticulate, its lateral carina lamelliform, distinctly protruding apically; propodeal tubercles small, the left acute apically, the right obtuse apically, lateral apical angle slightly less than 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 4.7 × its maximum width; length of vein r 1.3 × width of pterostigma; vein r arising slightly from apical of pterostigma; 1-CU1:2-CU1 = 11:15. Hind wing: 1r-m:1- M = 17:23; M+CU about subequal to 1-M; 2-1A present and short, branched posteriorly. Legs. Hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.0, 9.2 and 7.7 × their width, respectively; fore tarsal claws with brownish bristles. Metasoma. Length of first tergite 1.8 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.5; remainder of tergites smooth; length of ovipositor sheath 0.18 × length of fore wing. Colour. Brown; scape, head (dorsal and frontal) dark brown, palpi, labrum, mandible, tegulae, all legs (except hind coxa and tibia, all basitarsi) yellow; flagellomeres dark brown, but medial flagellomeres yellow; parastigma and base of pterostigma white; side of pronotum and mesopleuron reddish-brown; mesoscutum yellowish-brown; propodeum and first metasomal tergite black; remainder of metasoma, ovipositor and sheath, veins and pterostigma brown; hind tibia apically dark brown; all basitarsi dark brown; wing membrane slightly infuscate. Male. Unknown. Biology. Unknown. Distribution. China (Zhejiang). Etymology. The specific name refers to the name of the collection site. Notes. The new species is similar to B. (H.) robustus Haeselbarth, 1973 because the scutellum is coarsely rugose, the malar suture absent, length of eye in dorsal view as long as temple, length of the penultimate flagellomere 1.3 × its width, and the hind tibia darkened apically, but differs by having a more setose body (sparser setose in B. robustus), the temples more narrowed behind eyes in dorsal view and also in anterior view, the notauli completely crenulate (at least partly simple), the fore claw with brownish bristles (blackish), length of the malar space 2.0 × basal width of mandible (1.7 ×), length of the first tergite 1.8 × its apical width (1.4–1.5 ×), and dorsal carinae of first tergite distinct in basal 0.5 (in basal 0.3).

Gupta, Ankita, Achterberg, Cornelis Van, Pattar, Rohit (2023): First report of the genus Clinocentrus Haliday (Braconidae, Rogadinae) from India, with the description of a new species. Zootaxa 5293 (1): 196-200, DOI: 10.11646/zootaxa.5293.1.11, URL: http://dx.doi.org/10.11646/zootaxa.5293.1.11

Clinocentrus karbi Gupta & van Achterberg sp. nov. (Figs 1–2) Type material. Holotype. Female on card; India: Assam: Karbi Anglong district; sweep net; 27.ii.2016; coll. Ankita Gupta; code—NIM/ NBAIR /Hym/Brac/Clin/270216-H (NIM). Paratypes- two females on card; same data as holotype; code NIM/ NBAIR /Hym/Brac/Clin/270216 P1 and P2 (NIM). Key characters: ratio of vein r/3-SR = 0.9; second submarginal cell subquadrate; length of third antennomere nearly as long as fourth; length of third, fourth and penultimate 2.2, 2.5, 2.6 × their width, respectively; ovipositor nearly as long as metasoma; ratio of length of ovipositor/mid tibia 1.8. Holotype. Body length 4.5 mm; antenna 32 antennomeres; fore wing 4.3 mm. Body with eyes, ocellar region, prothorax (lateral), mesopleuron (anterior half), propodeum and dorsal metasoma black or dark brownish black; legs yellow except apical one third of femur, three quarters of tibia apically and tarsi brownish; pterostigma mostly dark brown, pale yellow basally, veins dark brown. Head.—Thirty two antennomeres (Fig. 1B), setose, third and fourth antennomeres nearly of equal length, length of third, fourth and penultimate antennomeres 2.2, 2.5, 2.6 × their width, respectively; antenna 1.5 × longer than fore wing; length of eye in dorsal view 2.1 × as long as temple; temple narrowed posteriorly; occipital carina complete, angular/ concave in dorsal view; OOL:OD:POL = 43:32:27; frons flat, smooth (Fig. 2A); temple and vertex smooth (Fig. 2B); face medially with slight depression, few fine transverse rugae laterally; clypeus slightly convex, punctate; length of malar space 0.85 × as long as basal width of mandibles, 0.2 × of eye length in lateral view. Mesosoma. Length of mesosoma 1.8 × its height; pronotum crenulate, dorsally and ventrally rugose (Fig. 2D); mesopleuron (Fig. 2C) anterior one third strongly crenulated and remaining part smooth and glabrous in posterior two third; mesoscutum smooth; notauli narrow, crenulate; scutellar sulcus wide, with one carina and few weak rugae; scutellum nearly smooth with few shallow punctures; propodeum irregularly reticulate-rugose, with median carina basally and areola obsolescent (Fig. 2F). Fore wing (Fig. 2G): 1-SR+M slightly curved, SR1 straight. Veins (relative length) r:3-SR: 2-SR: SR1 = 27:29:29.5:106; r-m = 13.2; 1-CU1:2-CU1 = 12:51.5. Vein m-cu far antefurcal and curved; second submarginal cell subquadrate. Hind wing vein 2-SC+R longer than wide (11.2); cu-a: 1-M: M+CU = 19.5: 60: 58; r/3-SR = 0.9. Legs. Hind coxa almost smooth; length of femur, tibia, and basitarsus of hind leg 5.0, 10.3 and 9.2 × their width, respectively; length of hind tarsus 1.1 × as long as hind tibia; length of longer hind tibial spurs 0.3 × as long as hind basitarsus. Metasoma. Length of first tergite 1.1 × its apical width, its surface longitudinally rugose (Fig. 2H), dorsal carina united at basal one third, connected to a distinct median carina; dorsope large; second tergite distinctly longitudinally rugose; third tergite weakly longitudinally rugose with transverse short rugae; length of second tergite 1.2 × as long as third tergite and third tergite narrowed posteriorly; fourth and its following tergites smooth and largely retracted (Fig. 2E); length of ovipositor sheath 0.5 × as long as fore wing and slightly shorter than metasoma. Etymology. Named after the Karbi tribe, which is the main tribe in the Karbi Anglong and West Karbi Anglong districts of Assam. Comments. The Indian species does not fit in the key to the Chinese species by Chen & He (1997). The new species shows similarity with C. cephalus, C. hubeiensis, C. nigricans and C. xinjiangensis in having vein r of fore wing 0.9 × as long as vein 3-SR. However, it can be clearly differentiated using the following characters: length of ovipositor sheath 0.7 × length of metasoma in C. xinjiangensis (vs nearly as long in C. karbi); length of third, fourth and penultimate segments 4.2, 3.2 and 2.4 × their width in C. cephalus; 3.8, 3.0 and 2.0 × in C. hubeiensis; and 4.0, 3.0, 2.5 in C. nigricans, respectively (vs 2.2, 2.5, 2.6 × in C. karbi). Clinocentrus foveatus is similar to the new species because of the robust basal antennal segments and the similar shape of the metasoma. Comparing with the images of C. foveatus (Fig. 3; coll.mfn-berlin.de/u/49d382) kindly supplied by Stefanie Krause (Museum für Naturkunde, Berlin) the following differences in fore wing venation and propodeal sculpture are observed. The areolate portion of the propodeum is minimally developed to almost smooth at basal one third with presence of median carina in C. foveatus (vs propodeum completely irregularly reticulate-rugose in C. karbi); veins 2-M and 3-SR of fore wing straight in C. foveatus (vs clearly convex or curved in C. karbi); mesopleuron with oblique row of crenulae in C. foveatus (vs smooth in middle and apical region, without any oblique row of crenulae in C. karbi). Clinicentrus compositus is easy to separate from the new species because of the elongated first metasomal tergite (about 2.0 × longer than its apical width in C. compositus (vs 1.1 × in C. karbi) and the inter-ocellar distance slightly greater than diameter of an ocellus (vs slightly smaller than diameter of one ocellus in C. karbi)., Published as part of Gupta, Ankita, Achterberg, Cornelis Van & Pattar, Rohit, 2023, First report of the genus Clinocentrus Haliday (Braconidae, Rogadinae) from India, with the description of a new species, pp. 196-200 in Zootaxa 5293 (1) on pages 196-200, DOI: 10.11646/zootaxa.5293.1.11, http://zenodo.org/record/7959509, {"references":["Chen, X. - X. & He, J. - H. (1997) Revision of the subfamily Rogadinae (Hymenoptera: Braconidae) from China. Zoologische Verhandelingen, Leiden, 308, 1 - 187."]}

The species of the genus Phaenocarpa Foerster, 1863 (Braconidae: Alysiinae) from South Korea are revised, and the genus is recorded for the first time from South Korea. Three species, Phaenocarpa artotemporalissp. nov., P. brachyurasp. nov. and P. lobatasp. nov., are new to Science, and two species, P. masha Belokobylskij, 1998 and P. fidelis Fischer, 1970, are newly recognized in South Korea. They are described and illustrated herein with a provision of the identification key to the Korean species. In addition, the DNA barcode region of the mitochondrial cytochrome c oxidase subunit I (COI) has been analyzed for the six species including P. ruficeps for genetic comparison.

Leiophron crassivena Gupta & van Achterberg sp. nov. Figs 3–5 Type material. Holotype. ♀; India, Chhattisgarh, Mainpat, 22°48′33.146′′ N, 83°17′38.919′′ E, yellow pan trap, 25– 30.iii.2022; leg Kriti A. Minz. code— NBAIR / Brac / Euph / Leio /25322H (NIM). Paratype, one female and one male, same data as holotype, code— NBAIR /Brac/Euph/Leio/25322P1 and NBAIR /Brac/Euph/Leio/25322P2 (NIM). Holotype. Body length 3.2 mm; fore wing length 2.3 mm; exserted part of ovipositor 0.15 mm. Colour. Body mainly brownish (Figs 3 & 4); antennomeres yellowish but 3–4 apical antennomeres dark brown; legs in general yellowish with comparatively paler fore and mid legs; fore wings prominently infuscated, fore wing with pterostigma dark brown in apical half and remainder pale yellowish; first tergite and majority of the second tergite yellowish brown, following tergites black. Head. Antenna with 15 antennomeres; F1 3.8 × as long as wide and 1.1 × F2; penultimate antennomere 1.4 × as long as wide; head rather square (Fig. 5A), ca. 1.4 × as wide as long in dorsal view, 1.3× as wide as long in frontal view; temple smooth; eye length 1.6 × temple in dorsal view (Fig. 5B); POL:OOL: OD = 14.3: 12.4: 4.5; POL 1.1 × OOL; malar suture present, malar space subequal to 0.7 × basal width of mandible; occipital carina ventrally present; face smooth and setose, clypeus smooth and intertentorial line 3.2 × tentorio-ocular line; vertex smooth around ocelli, with shallow punctures near temple and occiput. Mesosoma. Mesosoma 1.8 × as long as wide in dorsal view, notauli entirely absent, median and lateral lobes of mesoscutum smooth (Fig. 5C). Propodeum irregularly reticulate rugose with no median carina. Fore wing 3.3 × as long as wide (Fig. 5E); pterostigma 2.8 × as long as wide; 1- R 1 0.35–0.37 × length of pterostigma; 1-M thickened; 1-CU1, 2-CU1 present; r+3-SR+SR1 well developed; 1-SR+M weakly developed; basal cell glabrous and subbasal cell largely so. Hind femur, hind tibia and hind basitarsus 4.9, 8.0, 8.9 × as long as their width, respectively. Metasoma. Metasoma 2.5 × its width in dorsal view and 2.7 × its height in lateral view; first tergite longitudinally striate (Fig. 5D), 3.9 × longer than its apical width; spiracles situated at middle of tergite and ventral one third of T 1 closed in middle; T 2 smooth, 0.4 × length of metasoma, remaining tergites smooth. Variation. The male paratype (Fig. 4) is very similar to the holotype but differs in having 16 antennomeres. Etymology. Named after the thickened vein 1-M of the fore wing: “crassus” is Latin for thickened and “vena” is Latin for vein. Remarks. This species comes close to L. hayati by having the length of the malar space and the basal width of the mandible subequal, the face smooth, the colour pattern similar and the notauli entirely absent (Shamim et al. 2009). However, it can be differentiated with the following set of characters: length of metasoma 2.6 × its height (4.2 × its height in L. hayati); hind femur 4.9 × as long as wide (3.0 × as long as wide); T1 3.9 × longer than its apical width (3.3 × longer than its apical width) and T2 0.4 × as long as metasoma (more than 0.5 × as long as metasoma)., Published as part of Gupta, Ankita, Achterberg, Cornelis Van, Pattar, Rohit & Minz, Kriti Arpana, 2022, On the occurrence of a Palaearctic species of the genus Aphaereta Foerster (Braconidae, Alysiinae) and description of a new species of the genus Leiophron Nees (Braconidae, Euphorinae) from central India, pp. 455-462 in Zootaxa 5209 (4) on pages 458-460, DOI: 10.11646/zootaxa.5209.4.4, http://zenodo.org/record/7333850, {"references":["Shamim, M., Ahmad, Z. & Samiuddin, A. (2009) Taxonomic studies of Indian Leiophron subgen. Leiophron (Hymenoptera: Braconidae: Euphorinae), with the description of seven new species. Folia Heyrovskyana, Series A, 17, 1, 5 - 22."]}

Aphaereta vondelparkensis van Achterberg et al. 2020 Figs 1 & 2 Female, length of body 3.3 mm, of fore wing 3.4 mm. Colour. Black or dark brown (Fig. 1A); scapus yellow and pedicellus of antenna brown; palpi, tegulum, coxae, trochanters, trochantelli pale yellowish; and remainder of legs (but telotarsi infuscate) brownish-yellow; mandibles (margins darkened) and first tergite of metasoma reddish brown; wing membrane slightly infuscate. Propleuron and pronotal side ventrally orange (Fig. 1B). Head. Antenna with 24 antennomeres, length of third, fourth and penultimate antennomeres 3.7, 6.1 and 2.5 × their maximum width, respectively; OOL:diameter of posterior ocellus:POL = 5.0:1.9:2.0; frons glabrous and smooth (Fig. 2A), nearly flat; length of eye in dorsal view 2.4 × temple; eyes glabrous except for a few setae; temples slightly curved behind eyes; median groove of vertex present (Fig. 2B); head 1.5–1.7 × wider than high medially, smooth; clypeus sparsely finely punctate, rather convex medially, long setose; mandible partially rugulose, its medial length 1.6 × maximum width, second tooth longer than lateral teeth (Fig. 1B). Mesosoma. Mesoscutum glabrous, smooth, strongly shiny and with complete lateral carina (Fig. 2D); scutellar sulcus transverse and deep with one median carina; scutellum slightly convex; metanotum with two diverging sub median carinae; surface of propodeum mainly rugulose, with two sub medio-longitudinal carinae forming an areola in apical half (Fig. 2F). Wings. (Fig. 2E) Fore wing: 2-SR oblique; r:3-SR:SR1(relative) = 11.5:40.6:115.2; r about 1.3 × as long as width of pterostigma, pterostigma narrow elliptical; SR1 straight and ending nearly at apex of wing; 2-SR:3-SR: r-m (relative) = 23.8:40.6:13.1; r-m and 2-SR (apical third) interstitial. Hind wing: narrow apically; no trace of cu-a. Ratio of m-cu: 2-SR 37.7: 23.8. Second submarginal cell 2.2 × as long as its maximum width. Legs. Length of femur about 5.0 × as long as its maximum width. Ratio of ovipositor: hind femur and first tergite 13.9: 75.6: 47.8. Metasoma. Length of first tergite about 0.9 × as long as apical width, its surface distinctly convex medially and rugose, longitudinal sub median carinae distinct in basal 0.5 and tergite widened behind spiracles; dorsope rather large (Fig. 2G). Length of setose part of ovipositor sheath (Fig. 2H) 0.4 × fore wing length, sub equal to metasoma, 1.7 × hind femur length and 1.1 × hind tibia length, with very long setae. Material examined. 4♀; India, Chhattisgarh, Kanker, 20°14′48.416′′N, 81°30′28.873′′E, yellow pan trap, 8– 14.ii.2022; leg Kriti A. Minz. code— NBAIR / Brac / Aly / Aphae /08222A–D (NIM). Remarks. The species has a wide distribution as it is encountered in inhabited areas. So far known from Europe and the East Palaearctic region (pers. comm. CvA). The Indian specimens resemble A. vondelparkensis in almost all the characters and also possess the pale pronotum (ventrally) and propleuron, however, slight variations are observed which are shown in Table 1., Published as part of Gupta, Ankita, Achterberg, Cornelis Van, Pattar, Rohit & Minz, Kriti Arpana, 2022, On the occurrence of a Palaearctic species of the genus Aphaereta Foerster (Braconidae, Alysiinae) and description of a new species of the genus Leiophron Nees (Braconidae, Euphorinae) from central India, pp. 455-462 in Zootaxa 5209 (4) on pages 456-457, DOI: 10.11646/zootaxa.5209.4.4, http://zenodo.org/record/7333850, {"references":["van Achterberg, K., Schilthuizen, M., van der Meer, M., Delval, R., Dias, C., Hoynck, M, Koster, H., Maarschall, R., Peeters, N., Venema, P., Zaremba, R., Beltrami, C., Nieuwenhuis, F., de Rop, N., Njunjic, I. & Koene, J. M. (2020) A new parasitoid wasp, Aphaereta vondelparkensis sp. n. (Braconidae, Alysiinae), from a city park in the centre of Amsterdam. Biodiversity Data Journal, 8, e 49017. https: // doi. org / 10.3897 / BDJ. 8. e 49017"]}