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Homo sapiens was present in northern Asia by around 40,000 years ago, having replaced archaic populations across Eurasia after episodes of earlier population expansions and interbreeding1–4. Cultural adaptations of the last Neanderthals, the Denisovans and the incoming populations of H. sapiens into Asia remain unknown1,5–7. Here we describe Xiamabei, a well-preserved, approximately 40,000-year-old archaeological site in northern China, which includes the earliest known ochre-processing feature in east Asia, a distinctive miniaturized lithic assemblage with bladelet-like tools bearing traces of hafting, and a bone tool. The cultural assembly of traits at Xiamabei is unique for Eastern Asia and does not correspond with those found at other archaeological site assemblages inhabited by archaic populations or those generally associated with the expansion of H. sapiens, such as the Initial Upper Palaeolithic8–10. The record of northern Asia supports a process of technological innovations and cultural diversification emerging in a period of hominin hybridization and admixture2,3,6,11. Evidence for ochre processing Novel miniaturized lithics showing hafting Implications for cultural adaptations Methods

Late Pleistocene witnessed significant changes in lithic technological behaviors, mobility patterns and subsistence strategies in northeastern Eurasia, which are also correlated with paleoclimatic and paleoenvironmental changes. Integrating artifact analysis and paleoenvironmental evidence, Taoshan, a newly excavated archaeological site, provides a crucial case study for evaluating human adaptation in Northeast China. Taoshan site was excavated in 2013–2014, uncovering three prehistoric layers and abundant remains. Here we present the results of a systematic techno-typological analysis of the Late Pleistocene assemblages of the site, and place special emphasis on diachronic cultural developments and underlying dynamics.

North-east China occupies an important geographic position for understanding the process of Neolithisation in East Asia. Although archaeologists have long debated the trajectory of change in this region, a lack of intensive survey and excavation has precluded convincing interpretations. This article presents research on the newly excavated sites of Huayang and Taoshan in the southern Lesser Khingan Mountains, with a particular focus on the lithic assemblages. Comparative and environmental analyses demonstrate the largely uniform trajectory of lithic technologies across north-east China and close correspondence with Late Glacial palaeoclimatic and palaeoenvironmental changes.

The study of lithic raw material plays an important role in developing archeologists’ understanding of the different adaptive strategies and behaviors of prehistoric people. In this paper, we present new evidence from the Huayang site that reveals lithic raw material procurement and exploitation strategies dating to around 14 ka cal. BP. The Huayang site is located in the southern portion of the Lesser Khingan Mountains in Northeast China, a key region for examining migration, diffusion, and interaction among hunter-gatherers in East and Northeast Asia. Our results indicate that diversified lithic raw materials were found and procured from local riverbeds and reduced at the site. Each raw material was reduced with distinctive knapping strategies and, as a result, there is a strong discrete spatial pattern for each of the different raw material types. Intra- and inter-site comparative analyses suggest a uniform trend towards localization and diversification of lithic raw materials in the southern Lesser Khingan Mountains, Northeast China during the Late Glacial, which seems to be associated with the paleoclimatic amelioration and the demographic expansion during this period. Introduction - The Huayang site Materials and methods - Raw material identification and geological survey - Lithic raw material economy, spatial distribution, and refitting Results - Raw material types - Raw material sources - Procurement patterns and reduction sequences of raw materials - Spatial distribution and raw material exploitation Discussion and conclusions - Lithic raw material economy of Huayang - Lithic resource localization and diversification as adaptive strategies

The geographic and ecological background behind the development and spread of microblade technologies in Asia is a topic of considerable research interest. Microblade technologies are geographically widespread, and present in southern Siberia, the Russian Far East, Mongolia, northern China, the Korean Peninsula and the Japanese archipelago. Here we examine microblade sites of Northeast China which date to from ~28,000 years ago to the end of the Pleistocene. Though microblade assemblages in Northeast China are found to share a number of technological traits, regional divergences are identifiable on account of raw material differences. Technological changes through time correspond with climatic and environmental shifts during Marine Isotope Stage 2 (MIS 2). Microblade technology has its root in southern Siberia on the basis of early age ranges, and thereafter, these assemblages diffused widely, both southward and eastward into China. Microblade industries subsequently underwent a standardization process in Northeast China, leading to the formation of pressure flaking microblade technology on typical wedge-shaped cores of the Northern Microblade Industry (NMI). The NMI appears to have then diffused relatively rapidly across northern and eastern Asia, perhaps representing population movements and cultural interactions.

Knowledge about the Early and Middle Pleistocene hominin record of China is steadily increasing owing to the on-going implementation of palaeoanthropological surveys and excavations. Yet, relatively little attention has been paid to the effects of climate variability on hominin populations in China and its influence on archaeological site distributions and hominin behaviours. The Mid-Pleistocene climate transition (MPT), dating to between ∼1.2–0.7 Ma in Eastern Asia, is characterized by significant changes in the length and intensity of glacial-interglacial cycles and in monsoon intensity and terrestrial conditions, which have been implicated in influencing the occupation history of basins within China. Here, the MPT is examined relative to archaeological site distributions across China to determine potential long-term geographic and behavioural effects on hominin populations before, during and after this critical period. Changes in the geographic distribution of hominins are demonstrated across the MPT, with significant shifts in the number of sites in high and low latitudes, likely as a response to changing ecosystems. Technological innovations, including the development of Large Cutting Tools, occur in the MPT and afterwards, possibly a response to the formation of open habitats in South China. Geographic and behavioural shifts in the hominin record challenge traditional views about the long-term, conservative nature of the biological and cultural evolution of hominins in Eastern Asia, and instead demonstrate dynamic responses of populations to ecosystem changes across the Early and Middle Pleistocene.

Levallois approaches are one of the best known variants of prepared-core technologies, and are an important hallmark of stone technologies developed around 300,000 years ago in Africa and west Eurasia1,2. Existing archaeological evidence suggests that the stone technology of east Asian hominins lacked a Levallois component during the late Middle Pleistocene epoch and it is not until the Late Pleistocene (around 40,000–30,000 years ago) that this technology spread into east Asia in association with a dispersal of modern humans. Here we present evidence of Levallois technology from the lithic assemblage of the Guanyindong Cave site in southwest China, dated to approximately 170,000–80,000 years ago. To our knowledge, this is the earliest evidence of Levallois technology in east Asia. Our findings thus challenge the existing model of the origin and spread of Levallois technologies in east Asia and its links to a Late Pleistocene dispersal of modern humans. Levallois stone-tool technology found at the Guanyindong Cave site in southwest China was dated to approximately 170,000–80,000 years ago, which is much earlier than previously thought.

Donggutuo (DGT) is one of the richest archaeological localities in the Nihewan Basin of North China, thereby providing key information about the technological behaviours of early hominins in eastern Asia. Although DGT has been subject of multiple excavations and technological studies over the past several decades, few detailed studies on the lithic assemblages have been published. Here we summarize and describe the DGT lithic assemblages, examining stone tool reduction methods and technological skills. DGT dates to ca. 1.1 Ma, close to the onset of the mid-Pleistocene climate transition (MPT), indicating that occupations at DGT coincided with increased environmental instability. During this time interval, the DGT knappers began to apply innovative flaking methods, using free hand hard hammer percussion (FHHP) to manufacture pre-determined core shapes, small flakes and finely retouched tools, while occasionally using the bipolar technique, in contrast to the earlier and nearby Nihewan site of Xiaochangliang (XCL). Evidence for some degree of planning and predetermination in lithic reduction at DGT parallels technological developments in African Oldowan sites, suggesting that innovations in early industries may be situational, sometimes corresponding with adaptations to changes in environments and local conditions.

Xiaochangliang (XCL), located in the Nihewan Basin of North China, is a key archaeological locality for understanding the behavioural evolution of early humans. XCL dates to ca. 1.36 Ma, making it one of the earliest sites in Northeast Asia. Although XCL represents the first excavation of an Early Pleistocene site in the Nihewan Basin, identified and excavated in the 1970's, the lithic assemblages have never been published in full detail. Here we describe the lithic assemblages from XCL, providing information on stone tool reduction techniques and the influence of raw materials on artefact manufacture. The XCL hominins used both bipolar and freehand reduction techniques to manufacture small flakes, some of which show retouch. Bipolar reduction methods at XCL were used more frequently than previously recognized. Comparison of XCL with other Early Pleistocene sites in the Nihewan Basin indicates the variable use of bipolar and freehand reduction methods, thereby indicating a flexible approach in the utilization of raw materials. The stone tools from XCL and the Nihewan sites are classifiable as Mode I lithic assemblages, readily distinguished from bifacial industries manufactured by hominins in Eastern Asia by ca. 800 ka.