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It is widely acknowledged that opaque orthographies place additional demands on learning, often requiring many years to fully acquire. It is less widely recognized, however, that such opacity may offer certain benefits in the context of reading. For example, heterographic homophones such as ⟨knight⟩ and ⟨night⟩ (words that sound the same but which are spelled differently) impose additional costs in learning but reduce ambiguity in reading. Here, we consider the possibility that—left to evolve freely—writing systems will sometimes choose to forego some simplicity for the sake of informativeness when there is functional pressure to do so. We investigate this hypothesis by simulating the evolution of orthography as it is transmitted from one generation to the next, both with and without a communicative pressure for ambiguity avoidance. In addition, we consider two mechanisms by which informative heterography might be selected for: differentiation, in which new spellings are created to differentiate meaning (e.g., ⟨lite⟩ vs. ⟨light⟩), and conservation, in which heterography arises as a byproduct of sound change (e.g., ⟨meat⟩ vs. ⟨meet⟩). Under pressure from learning alone, orthographic systems become transparent, but when combined with communicative pressure, they tend to favor some additional informativeness. Nevertheless, our findings also suggest that, in the long term, simpler, transparent spellings may be preferred in the absence of top-down explicit teaching. [ABSTRACT FROM AUTHOR]

Language maps signals onto meanings through the use of two distinct types of structure. First, the space of meanings is discretized into categories that are shared by all users of the language. Second, the signals employed by the language are compositional: The meaning of the whole is a function of its parts and the way in which those parts are combined. In three iterated learning experiments using a vast, continuous, open-ended meaning space, we explore the conditions under which both structured categories and structured signals emerge ex nihilo. While previous experiments have been limited to either categorical structure in meanings or compositional structure in signals, these experiments demonstrate that when the meaning space lacks clear preexisting boundaries, more subtle morphological structure that lacks straightforward compositionality-as found in natural languages-may evolve as a solution to joint pressures from learning and communication. [ABSTRACT FROM AUTHOR]